93 lines
13 KiB
XML
93 lines
13 KiB
XML
<document id="BEE75C53653C2DC37A406CFE2AAFED09" ID-DOI="10.5281/zenodo.157458" ID-GBIF-Dataset="913d472f-21c0-4891-a893-561570d80b58" ID-ISSN="11755326" ID-Zenodo-Dep="157458" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461021562626" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2004" docId="039D87C49C72FFCC3E3CFDA2FC8CFCFF" docLanguage="en" docName="zt00559.pdf" docOrigin="Zootaxa 559" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Nucleotops Perkins and Balfour-Browne 1994" docType="treatment" docVersion="8" lastPageNumber="4" masterDocId="FFA4FFBC9C70FFCF3F34FFF3FF88FFAC" masterDocTitle="A revision of the South African endemic humicolous beetle genus Nucleotops Perkins and BalfourBrowne (Coleoptera: Hydraenidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1698570255036" updateUser="plazi">
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<mods:title id="D841A77B61FA6830E95E58E26422784E">A revision of the South African endemic humicolous beetle genus Nucleotops Perkins and BalfourBrowne (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="BF4E82EB3F08496C9527461D3FE2EB70">Perkins, Philip D.</mods:namePart>
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<mods:date id="CB7C5D28E6303D27062BFBCFAC4FC999">2004</mods:date>
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<mods:number id="9E7CBE1A71CB058E1DCF13F511657C2F">559</mods:number>
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<treatment id="039D87C49C72FFCC3E3CFDA2FC8CFCFF" ID-DOI="http://doi.org/10.5281/zenodo.6271911" ID-GBIF-Taxon="119603393" ID-Zenodo-Dep="6271911" LSID="urn:lsid:plazi:treatment:039D87C49C72FFCC3E3CFDA2FC8CFCFF" httpUri="http://treatment.plazi.org/id/039D87C49C72FFCC3E3CFDA2FC8CFCFF" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<subSubSection id="C32E65599C72FFCD3E3CFDA2FCCFFDC7" box="[264,839,593,619]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8B8B36D29C72FFCD3E3CFDA2FCCFFDC7" blockId="2.[264,839,593,619]" box="[264,839,593,619]" pageId="2" pageNumber="3">
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<taxonomicName id="4C344D519C72FFCD3E3CFDA2FCCFFDC7" ID-CoL="8VZSJ" authority="Perkins and Balfour-Browne, 1994" authorityName="Perkins and Balfour-Browne" authorityYear="1994" box="[264,839,593,619]" class="Insecta" family="Hydraenidae" genus="Nucleotops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B940EAC09C72FFCD3E3CFDA2FCCFFDC7" bold="true" box="[264,839,593,619]" pageId="2" pageNumber="3">
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<emphasis id="B940EAC09C72FFCD3E3CFDA2FE07FDC7" bold="true" box="[264,399,593,619]" italics="true" pageId="2" pageNumber="3">Nucleotops</emphasis>
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<bibRefCitation id="EFA54B239C72FFCD3EA3FDA2FCCFFDC7" author="Perkins" box="[407,839,593,619]" pageId="2" pageNumber="8" refString="Perkins, P. D. & Balfour-Browne, J. (1994) Contribution to the taxonomy of aquatic and humicolous beetles of the family Hydraenidae in southern Africa. Fieldiana: Zoology (New Series), 77, 1 - 159." type="journal article" year="1994">Perkins and BalfourBrowne, 1994</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C32E65599C72FFCD3E3CFD6CFDBAFD70" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="8B8B36D29C72FFCD3E3CFD6CFDBAFD70" blockId="2.[264,1323,670,732]" pageId="2" pageNumber="3">
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<treatmentCitationGroup id="AB2411FC9C72FFCD3E3CFD6CFDBAFD70" pageId="2" pageNumber="3">
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<taxonomicName id="4C344D519C72FFCD3E3CFD6CFCAAFD19" ID-CoL="8VZSJ" authority="Perkins and Balfour-Browne, 1994: 13" authorityName="Perkins and Balfour-Browne" authorityPageNumber="13" authorityYear="1994" box="[264,802,670,694]" class="Insecta" family="Hydraenidae" genus="Nucleotops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
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<emphasis id="B940EAC09C72FFCD3E3CFD6CFE08FD1A" box="[264,384,671,694]" italics="true" pageId="2" pageNumber="3">Nucleotops</emphasis>
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<treatmentCitation id="0A9510C39C72FFCD3EB2FD6DFCAAFD19" author="Perkins" box="[390,802,670,693]" page="13" pageId="2" pageNumber="3" year="1994">
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<bibRefCitation id="EFA54B239C72FFCD3EB2FD6DFD75FD19" author="Perkins" box="[390,765,670,693]" pageId="2" pageNumber="8" refString="Perkins, P. D. & Balfour-Browne, J. (1994) Contribution to the taxonomy of aquatic and humicolous beetles of the family Hydraenidae in southern Africa. Fieldiana: Zoology (New Series), 77, 1 - 159." type="journal article" year="1994">Perkins and BalfourBrowne, 1994</bibRefCitation>
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: 13
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</taxonomicName>
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(
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<typeStatus id="548F88709C72FFCD3C1BFD6DFCD6FD19" box="[815,862,670,693]" pageId="2" pageNumber="3">type</typeStatus>
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species
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<taxonomicName id="4C344D519C72FFCD3C8CFD6CFDA2FD70" ID-CoL="482P8" authority="Perkins and Balfour-Browne, 1994" authorityName="Perkins and Balfour-Browne" authorityYear="1994" class="Insecta" family="Hydraenidae" genus="Nucleotops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nimbaceps">
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<emphasis id="B940EAC09C72FFCD3C8CFD6CFB20FD1A" box="[952,1192,671,694]" italics="true" pageId="2" pageNumber="3">Nucleotops nimbaceps</emphasis>
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<bibRefCitation id="EFA54B239C72FFCD3B9AFD6DFDA2FD70" author="Perkins" pageId="2" pageNumber="8" refString="Perkins, P. D. & Balfour-Browne, J. (1994) Contribution to the taxonomy of aquatic and humicolous beetles of the family Hydraenidae in southern Africa. Fieldiana: Zoology (New Series), 77, 1 - 159." type="journal article" year="1994">Perkins and BalfourBrowne, 1994</bibRefCitation>
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)
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</paragraph>
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<subSubSection id="C32E65599C72FFCD3E3CFCE1FDC1FC0D" pageId="2" pageNumber="3" type="diagnosis">
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<paragraph id="8B8B36D29C72FFCD3E3CFCE1FDC1FC0D" blockId="2.[264,1324,786,1895]" pageId="2" pageNumber="3">
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<emphasis id="B940EAC09C72FFCD3E3CFCE1FE00FC80" bold="true" box="[264,392,786,812]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
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Recognized by the costate elytra (intervals 2, 4 and 6), the admedian longitudinal reliefs of the pronotum, the lateral fringe of setae on the mentum, the eleven articles of the antenna, the lack of hydrofuge pubescence on the venter, the short maxillary palpi and legs, and the gibbose frons.
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</paragraph>
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<subSubSection id="C32E65599C72FFCC3E0CFC5EFC8CFCFF" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
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<paragraph id="8B8B36D29C72FFCD3E0CFC5EFB97FB50" blockId="2.[264,1324,786,1895]" pageId="2" pageNumber="3">
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<emphasis id="B940EAC09C72FFCD3E0CFC5EFE47FC6B" bold="true" box="[312,463,941,967]" pageId="2" pageNumber="3">Description.</emphasis>
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Body size ca.
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to
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<quantity id="4CCC9B379C72FFCD3C28FC5EFC01FC6B" box="[796,905,941,967]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="3" unit="mm" value="1.9">1.90 mm</quantity>
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, form oblong, transversely convex. Antennomeres eleven (6 + 5); third subequal length of first two; pubescent club articles tightly articulated. Eyes small. Frons tumid to gibbose. Margin of head angulate at frontoclypeal suture. Frontoclypeal suture bisinuate. Anterior margin of clypeus straight. Labrum set at angle to clypeus, margin with setal fringe. Maxillary palpus short, shorter than antennae, palpomeres 2, 3 and 4 length ratios ca. 2:1:3, palpomere 3 subglobose, palpomere 4 penicillate. Mentum length slightly less than basal width; lateral border with row of arcuate setae; anterior margin arcuate. Genae rounded, separated by median longitudinal depression; a distinct lateral sulcus for reception of antenna.
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</paragraph>
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<paragraph id="8B8B36D29C72FFCD3E0CFAFAFEF6F90A" blockId="2.[264,1324,786,1895]" pageId="2" pageNumber="3">Pronotum with well developed lateral depressions, each with anterior and posterior pit; disc with submedian longitudinal costa on each side, smaller elevation lateral to each costa, on posterior 0.5 of disc; anterior margin with hyaline border in middle 0.33; lateral margins converging to base posterior to lateral depression; posterior margin slightly arcuate. Prosternum with procoxae contiguous; prosternum anterior to coxae with one carina on each side and a median carina; antennal cavity very small; a low carina separating pubescent sternal area from nonpubescent inflexed margin of pronotum, latter with two foveae; coxal cavities open behind. Mesosternum at base of intercoxal process with ridge in form of inverted "U"; mesosternum also with short basomedian carina and two low carinae on each side. Metasternum with sparse pubescence, a well developed median fovea near base.</paragraph>
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<paragraph id="8B8B36D29C72FFCC3E0CF940FD26FEF0" blockId="2.[264,1324,786,1895]" lastBlockId="3.[264,1324,284,851]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">Elytra with ten rows of punctures. Intervals 2, 4 and 6 costate, at least in part. Explanate margin thick, continuing virtually to apices. Abdomen lacking hydrofuge pubescence. First ventrite with apicomedian fovea between metacoxae, on each side an oblique carina from margin of fovea to posterior margin of first ventrite. Intersegmental impressions deep. Intercoxal sternite small, triangular. Legs of moderate build, rather short, each femur slightly excavate to receive tibia; without apparent sexual dimorphism; tarsi 555; claws simple; metatrochanter small.</paragraph>
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<paragraph id="8B8B36D29C73FFCC3E0CFE9AFC8CFCFF" blockId="3.[264,1324,284,851]" pageId="3" pageNumber="4">
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Antennal pocket (terminology as in
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<bibRefCitation id="EFA54B239C73FFCC3DD8FE9AFC19FE2F" author="Perkins" box="[748,913,361,387]" pageId="3" pageNumber="8" refString="Perkins, P. D. (1997) Life on the effective bubble: exocrine secretion delivery systems (ESDS) and the evolution and classification of beetles in the family Hydraenidae (Insecta: Coleoptera). Annals of Carnegie Museum, 66 (2), 89 - 207." type="journal article" year="1997">Perkins, 1997</bibRefCitation>
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) hypomeral, very shallow depression on anterior face of hypomeron, with spiniform setae contiguous with pubescence of prosternum, but not with that of postcoxal projection. The hypomeral carina welldeveloped; over middle onethird carina contiguous with the notosternal suture, hence hydrofuge pubescence of antennal pocket and that of postcoxal projection are not contiguous; few (twoten) exocrine pores present along part of carina bordering antennal pocket. Wethypomeron wide, varying from smooth to strongly microreticulate, with random exocrine pores varying in density and placement according to species. No organized clusters of pores, or secretion sulcus on hypomeron. Postocular antennal pocket formed by shallowly concave area behind eye, and by contiguous, small concavity on the gena; pocket bordered by distinct postocular carina; setae in postocular part of antennal pocket spiniform. Prosternal antennal cleaner, a cluster of spiniform setae, located on anterior margin of prosternum, adjacent to antennal pocket.
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</paragraph>
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