270 lines
32 KiB
XML
270 lines
32 KiB
XML
<document id="3F849218DE778CEDCB70B148F2456D00" ID-DOI="10.1186/s13358-022-00261-3" ID-ISSN="1664-2384" ID-Zenodo-Dep="11999827" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1718653084494" checkinUser="felipe" docAuthor="Schwarzhans, Werner, Klots, Oleksandr, Ryabokon, Tamara & Kovalchuk, Oleksandr" docDate="2022" docId="039D7D5BFE4EFFABF872FE30FE4EFA72" docLanguage="en" docName="SwissJPalaeontol.141.18.pdf" docOrigin="Swiss Journal of Palaeontology (18) 141 (1)" docSource="http://dx.doi.org/10.1186/s13358-022-00261-3" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Gobius bratishkoi Schwarzhans & Klots & Ryabokon & Kovalchuk 2022, n. sp." docType="treatment" docVersion="3" lastPageNumber="10" masterDocId="FFA40523FE49FFA2FB5FFFDBFFBAFFF1" masterDocTitle="A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths" masterLastPageNumber="35" masterPageNumber="1" pageNumber="8" updateTime="1722278998376" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8581A27D272248F065C9460F445AB877">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<mods:roleTerm id="A08325568F1EAEDED7F595AB0309710E">Author</mods:roleTerm>
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<mods:namePart id="9F7D33C5C814A65FAA1768A5AF294C56">Schwarzhans, Werner</mods:namePart>
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<mods:affiliation id="1B43DA33DAC15EB642C55617DDA2F327">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:namePart id="03089F3F032B55067A07BDE329178E5F">Klots, Oleksandr</mods:namePart>
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<mods:roleTerm id="D3148C2659C288339E0697CE2F001452">Author</mods:roleTerm>
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<mods:namePart id="DF78386AFCB7C33D1AF8FA4B31CFA72E">Ryabokon, Tamara</mods:namePart>
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<mods:name id="B90DF930E04BF10F76B257D53684D759" type="personal">
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<mods:roleTerm id="45373784845344E0D8689E9EADB3FE2B">Author</mods:roleTerm>
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<mods:namePart id="1573AC316361D6DEDF1D77303EC9D307">Kovalchuk, Oleksandr</mods:namePart>
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<mods:typeOfResource id="45952F0310CEBC0176B6DD278B0A779C">text</mods:typeOfResource>
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<mods:title id="78B5C3804EBA30A073ED7168C5AE588B">Swiss Journal of Palaeontology</mods:title>
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<mods:part id="CDD9AB853C5C00F36EDA950BF62442F9">
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<mods:date id="97ED8AB42E8641BD5713C03EA7A43BF5">2022</mods:date>
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<mods:title id="75264D977887E36AA7494F451995E421">18</mods:title>
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<mods:number id="B5D6BA568820D9A328B0D08BF7458538">2022-11-17</mods:number>
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<mods:detail id="FAEBB269033708C883E54F20EAF1DD11" type="volume">
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<mods:number id="492CA0F65C66784BB4EB061FA2C19195">141</mods:number>
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<mods:identifier id="A9717E09DCF41C4D315DF692818BBB1E" type="ISSN">1664-2384</mods:identifier>
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<mods:identifier id="BC3D06DF8692831630D96D0AC664780F" type="Zenodo-Dep">11999827</mods:identifier>
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<treatment id="039D7D5BFE4EFFABF872FE30FE4EFA72" LSID="urn:lsid:plazi:treatment:039D7D5BFE4EFFABF872FE30FE4EFA72" httpUri="http://treatment.plazi.org/id/039D7D5BFE4EFFABF872FE30FE4EFA72" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
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<subSubSection id="C32E9FC6FE4EFFA5F872FE30FB86FDF2" box="[813,1084,491,515]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8B8BCC4DFE4EFFA5F872FE30FB86FDF2" blockId="7.[813,1084,491,515]" box="[813,1084,491,515]" pageId="7" pageNumber="8">
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<heading id="D0C37B21FE4EFFA5F872FE30FB86FDF2" bold="true" box="[813,1084,491,515]" fontSize="10" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="B940105FFE4EFFA5F872FE30FB86FDF2" bold="true" box="[813,1084,491,515]" pageId="7" pageNumber="8">
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<taxonomicName id="4C34B7CEFE4EFFA5F872FE30FC40FDF2" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[813,1018,491,515]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="bratishkoi" status="sp. nov.">
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<emphasis id="B940105FFE4EFFA5F872FE30FC40FDF2" bold="true" box="[813,1018,491,515]" italics="true" pageId="7" pageNumber="8">Gobius bratishkoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273AD24FE4EFFA5FF5FFE30FB86FDF2" box="[1024,1084,491,515]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E9FC6FE4EFFABF872FDF0FD2FFDB2" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
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<paragraph id="8B8BCC4DFE4EFFA5F872FDF0FC16FDB2" blockId="7.[813,940,555,579]" box="[813,940,555,579]" pageId="7" pageNumber="8">
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<figureCitation id="130FD0C8FE4EFFA5F872FDF0FC16FDB2" box="[813,940,555,579]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: a–e Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, b–e paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,c–e (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).g–k Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, g–i, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="7" pageNumber="8">Figure 4a–e</figureCitation>
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</paragraph>
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<paragraph id="8B8BCC4DFE4EFFA5F872FDB0FBC7FD52" blockId="7.[813,1436,619,675]" pageId="7" pageNumber="8">
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<materialsCitation id="3B5CC610FE4EFFA5F872FDB0FBC7FD52" collectionCode="NMNHU-P" country="Ukraine" location="Kozatskyi Yar" municipality="Badenian" pageId="7" pageNumber="8" specimenCode="PI 2555" specimenCount="1" typeStatus="holotype">
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<emphasis id="B940105FFE4EFFA5F872FDB0FC34FD72" box="[813,910,619,643]" italics="true" pageId="7" pageNumber="8">
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<typeStatus id="548F72EFFE4EFFA5F872FDB0FC34FD72" box="[813,910,619,643]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
<collectionCode id="ED255488FE4EFFA5F8C3FDB7FBA6FD72" box="[924,1052,620,643]" pageId="7" pageNumber="8">NMNHU-P</collectionCode>
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<specimenCode id="DB926436FE4EFFA5FF75FDB7FB3CFD72" box="[1066,1158,619,643]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="8">PI 2555</specimenCode>
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,
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<location id="8EEB9A96FE4EFFA5FFC7FDB0FA8FFD72" LSID="urn:lsid:plazi:treatment:039D7D5BFE4EFFABF872FE30FE4EFA72:8EEB9A96FE4EFFA5FFC7FDB0FA8FFD72" box="[1176,1333,619,643]" country="Ukraine" municipality="Badenian" name="Kozatskyi Yar" pageId="7" pageNumber="8">Kozatskyi Yar</location>
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, western
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<collectingCountry id="F3238CDDFE4EFFA5F872FD50FC3DFD52" box="[813,903,651,675]" name="Ukraine" pageId="7" pageNumber="8">Ukraine</collectingCountry>
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, late
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<collectingMunicipality id="6BEF5637FE4EFFA5F8E1FD50FB9CFD52" box="[958,1062,651,675]" pageId="7" pageNumber="8">Badenian</collectingMunicipality>
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,
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<figureCitation id="130FD0C8FE4EFFA5FF6FFD50FBC3FD52" box="[1072,1145,651,675]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: a–e Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, b–e paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,c–e (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).g–k Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, g–i, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="7" pageNumber="8">Fig. 4a</figureCitation>
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.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8B8BCC4DFE4EFFA5F872FD17FA2AFCF2" blockId="7.[813,1436,715,771]" pageId="7" pageNumber="8">
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<materialsCitation id="3B5CC610FE4EFFA5F872FD17FC0EFCF2" collectionCode="NMB" country="Ukraine" location="Kozatskyi Yar" municipality="Badenian" pageId="7" pageNumber="8" specimenCode="P1211" specimenCount="5" typeStatus="paratype">
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<emphasis id="B940105FFE4EFFA5F872FD17FC22FD12" box="[813,920,716,739]" italics="true" pageId="7" pageNumber="8">
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<typeStatus id="548F72EFFE4EFFA5F872FD17FC22FD12" box="[813,920,716,739]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
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</emphasis>
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7 otoliths:
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<specimenCount id="9D3207C4FE4EFFA5FF4AFD10FB21FD12" box="[1045,1179,715,739]" count="5" pageId="7" pageNumber="8" type="generic">5 specimens</specimenCount>
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same data as holotype,
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<collectionCode id="ED255488FE4EFFA5F872FD37FCD3FCF2" box="[813,873,748,771]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="7" pageNumber="8">NMB</collectionCode>
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<specimenCode id="DB926436FE4EFFA5F830FD30FC0EFCF2" box="[879,948,747,771]" collectionCode="NMB" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="7" pageNumber="8">P1211</specimenCode>
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</materialsCitation>
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;
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<materialsCitation id="3B5CC610FE4EFFA5F8E1FD30FA36FCF2" box="[958,1420,747,771]" collectionCode="NMNHU-P" country="Ukraine" pageId="7" pageNumber="8" specimenCode="PI 2556" specimenCount="2" typeStatus="paratype">
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<specimenCount id="9D3207C4FE4EFFA5F8E1FD30FBFDFCF2" box="[958,1095,747,771]" count="2" pageId="7" pageNumber="8" type="generic">2 specimens</specimenCount>
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, Mlyntsi,
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<collectionCode id="ED255488FE4EFFA5FFEEFD37FA8BFCF2" box="[1201,1329,748,771]" pageId="7" pageNumber="8">NMNHU-P</collectionCode>
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<specimenCode id="DB926436FE4EFFA5FE68FD37FA36FCF2" box="[1335,1420,747,771]" collectionCode="PI" country="Russia" name="Paleontological Institute" pageId="7" pageNumber="8">PI 2556</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8BCC4DFE4EFFA5F872FCF0FA27FC72" blockId="7.[813,1437,811,899]" pageId="7" pageNumber="8">
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<emphasis id="B940105FFE4EFFA5F872FCF0FC20FCB2" box="[813,922,811,835]" italics="true" pageId="7" pageNumber="8">Etymology</emphasis>
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Named after Andriy Bratishko, Tallahassee, Florida (formerly
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<collectingRegion id="49F002AFFE4EFFA5F8E1FC90FBA0FC92" box="[958,1050,843,867]" country="Ukraine" name="Luhans'k" pageId="7" pageNumber="8">Luhansk</collectingRegion>
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,
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<collectingCountry id="F3238CDDFE4EFFA5FF7AFC90FBC6FC92" box="[1061,1148,843,867]" name="Ukraine" pageId="7" pageNumber="8">Ukraine</collectingCountry>
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), in recognition of his contributions to the fossil fish otoliths of the Eastern Paratethys.
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</paragraph>
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<paragraph id="8B8BCC4DFE4EFFA5F872FC77FBA4FB52" blockId="7.[813,1437,938,1187]" pageId="7" pageNumber="8">
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<emphasis id="B940105FFE4EFFA5F872FC77FC2DFC32" box="[813,919,940,963]" italics="true" pageId="7" pageNumber="8">Diagnosis</emphasis>
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OL:OH = 1.2–1.3. Anterior rim nearly vertical; posterior rim with prominent, rounded postdorsal projection. Dorsal rim with distinct, broad predorsal concavity and narrow postdorsal angle positioned above cauda. Sulcus narrow, relatively short (OL:SuL = 2.1–2.35), inclined at 10–18°. Ostial lobe low; subcaudal iugum small, below anterior part of cauda. Inner face flat; outer face markedly convex.
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</paragraph>
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<paragraph id="8B8BCC4DFE4EFFA5F872FB17FB11F972" blockId="7.[813,1437,1227,1827]" pageId="7" pageNumber="8">
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<emphasis id="B940105FFE4EFFA5F872FB17FC10FB12" box="[813,938,1228,1251]" italics="true" pageId="7" pageNumber="8">Description</emphasis>
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Moderately slender, massive gobiid otoliths reaching about
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<quantity id="4CCC61A8FE4EFFA5F8BFFB30FB82FAF2" box="[992,1080,1259,1283]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="8" unit="mm" value="2.2">2.2 mm</quantity>
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in length (
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<typeStatus id="548F72EFFE4EFFA5FFE3FB30FAA7FAF2" box="[1212,1309,1259,1283]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
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||
<quantity id="4CCC61A8FE4EFFA5FE76FB30FA2BFAF2" box="[1321,1425,1259,1283]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="7" pageNumber="8" unit="mm" value="2.15">2.15 mm</quantity>
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). OH:OT = 2.2–2.6, decreasing with size. Anterior rim nearly vertical with only minor indentation above level of ostial tip. Preventral angle nearly orthogonal, predorsal angle distinct, positioned high on anterior rim. Ventral rim nearly straight to slightly bent. Dorsal rim with broad concavity across entire predorsal section followed by narrow postdorsal angle positioned above cauda and in turn followed by shorter concavity or flat, inclined section toward distinctly expanded postdorsal projection with rounded tip and not bent outward. Posterior rim below postdorsal projection nearly vertical, with orthogonal postventral angle. All rims smooth.
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</paragraph>
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<paragraph id="8B8BCC4DFE4EFFABF81DF957FCBDFEF2" blockId="7.[813,1437,1227,1827]" lastBlockId="9.[151,775,235,580]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">Inner face slightly convex, nearly flat, with narrow, somewhat deepened, centrally positioned, distinctly inclined sulcus. SuL:SuH = 2.5–2.9. Sulcus with very low ostial lobe and rounded anterior and posterior tips, inclined at 10–18°. Subcaudal iugum small, often indistinct or indiscernible after even slight erosion</paragraph>
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<caption id="DF4B9CC5FE41FFAAFBF7F935FE32F8A3" ID-DOI="http://doi.org/10.5281/zenodo.11999881" ID-Zenodo-Dep="11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="8" pageNumber="9" startId="8.[168,199,1774,1794]" targetBox="[152,1438,237,1886]" targetPageId="8" targetType="figure">
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<paragraph id="8B8BCC4DFE41FFAAFBF7F935FE32F8A3" blockId="8.[168,1352,1774,1874]" pageId="8" pageNumber="9">
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||
<emphasis id="B940105FFE41FFAAFBF7F935FF61F8F3" bold="true" box="[168,219,1774,1794]" pageId="8" pageNumber="9">Fig. 4</emphasis>
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Otoliths of
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<taxonomicName id="4C34B7CEFE41FFAAFA1CF935FE1CF8F3" box="[323,422,1774,1794]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B940105FFE41FFAAFA1CF935FEC6F8F3" box="[323,380,1774,1794]" italics="true" pageId="8" pageNumber="9">Gobius</emphasis>
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spp.
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</taxonomicName>
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||
:
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<emphasis id="B940105FFE41FFAAFAF3F935FE76F8F3" bold="true" box="[428,460,1774,1794]" pageId="8" pageNumber="9">a–e</emphasis>
|
||
<taxonomicName id="4C34B7CEFE41FFAAFA8EF935FDE7F8F3" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[465,605,1774,1794]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="bratishkoi" status="sp. nov.">
|
||
<emphasis id="B940105FFE41FFAAFA8EF935FDE7F8F3" box="[465,605,1774,1794]" italics="true" pageId="8" pageNumber="9">Gobius bratishkoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273AD24FE41FFAAF93DF935FD2AF8F3" box="[610,656,1774,1794]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B940105FFE41FFAAF9CAF935FD1AF8F3" bold="true" box="[661,672,1774,1794]" pageId="8" pageNumber="9">a</emphasis>
|
||
holotype, NMNHU-P PI 2555, Kozatskyi Yar,
|
||
<emphasis id="B940105FFE41FFAAFF50F935FB8BF8F3" bold="true" box="[1039,1073,1774,1794]" pageId="8" pageNumber="9">b–e</emphasis>
|
||
paratypes,
|
||
<emphasis id="B940105FFE41FFAAFFCDF935FB24F8F3" bold="true" box="[1170,1182,1774,1794]" pageId="8" pageNumber="9">b</emphasis>
|
||
(reversed) Mlyntsi, NMNHU-P PI 2556,
|
||
<emphasis id="B940105FFE41FFAAFA15F8D2FED0F8EC" bold="true" box="[330,362,1801,1821]" pageId="8" pageNumber="9">c–e</emphasis>
|
||
(reversed) Kozatskyi Yar, NMB P1211.
|
||
<emphasis id="B940105FFE41FFAAF9FBF8D2FD11F8EC" bold="true" box="[676,683,1801,1821]" pageId="8" pageNumber="9">f</emphasis>
|
||
<taxonomicName id="4C34B7CEFE41FFAAF9F0F8D2FBA3F8EC" authority="Schwarzhans, 2014" authorityName="Schwarzhans" authorityYear="2014" box="[687,1049,1801,1821]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="reichenbacherae">
|
||
<emphasis id="B940105FFE41FFAAF9F0F8D2FCC9F8EC" box="[687,883,1801,1821]" italics="true" pageId="8" pageNumber="9">Gobius reichenbacherae</emphasis>
|
||
<bibRefCitation id="EFA5B1BCFE41FFAAF827F8D2FBA3F8EC" author="Schwarzhans, W." box="[888,1049,1801,1821]" pageId="8" pageNumber="9" pagination="35 - 69" refId="ref23301" refString="Schwarzhans, W. (2014). Otoliths from the middle Miocene (Serravallian) of the Karaman Basin, Turkey. Cainozoic Research, 14 (1), 35 - 69." type="journal article" year="2014">Schwarzhans,2014</bibRefCitation>
|
||
</taxonomicName>
|
||
, Shydlivshchyna, NMNHU-P PI 2557 (reversed).
|
||
<emphasis id="B940105FFE41FFAAFA5BF8F8FE9CF8C6" bold="true" box="[260,294,1827,1847]" pageId="8" pageNumber="9">g–k</emphasis>
|
||
<taxonomicName id="4C34B7CEFE41FFAAFA74F8F8FE00F8C6" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[299,442,1827,1847]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="ukrainicus" status="sp. nov.">
|
||
<emphasis id="B940105FFE41FFAAFA74F8F8FE00F8C6" box="[299,442,1827,1847]" italics="true" pageId="8" pageNumber="9">Gobius ukrainicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273AD24FE41FFAAFAE0F8F8FE57F8C6" box="[447,493,1827,1847]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B940105FFE41FFAAFAADF8F8FE42F8C6" bold="true" box="[498,504,1827,1847]" pageId="8" pageNumber="9">j</emphasis>
|
||
holotype, Shydlivshchyna, NMNHU-P PI 2558,
|
||
<emphasis id="B940105FFE41FFAAF8DFF8F8FC26F8C6" bold="true" box="[896,924,1827,1847]" pageId="8" pageNumber="9">g–i</emphasis>
|
||
,
|
||
<emphasis id="B940105FFE41FFAAF8FAF8F8FC0AF8C6" bold="true" box="[933,944,1827,1847]" pageId="8" pageNumber="9">k</emphasis>
|
||
paratypes,
|
||
<emphasis id="B940105FFE41FFAAFF4FF8F8FBA6F8C6" bold="true" box="[1040,1052,1827,1847]" pageId="8" pageNumber="9">g</emphasis>
|
||
,
|
||
<emphasis id="B940105FFE41FFAAFF7AF8F8FB90F8C6" bold="true" box="[1061,1066,1827,1847]" pageId="8" pageNumber="9">i</emphasis>
|
||
,
|
||
<emphasis id="B940105FFE41FFAAFF6DF8F8FB87F8C6" bold="true" box="[1074,1085,1827,1847]" pageId="8" pageNumber="9">k</emphasis>
|
||
NMNHU-P PI 2560,
|
||
<emphasis id="B940105FFE41FFAAFFBBF8F8FB4AF8C6" bold="true" box="[1252,1264,1827,1847]" pageId="8" pageNumber="9">h</emphasis>
|
||
(reversed) Mlyntsi, NMNHU-P PI 2559
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8BCC4DFE40FFABFBC8FED0FD2FFDB2" blockId="9.[151,775,235,580]" pageId="9" pageNumber="10">
|
||
(
|
||
<figureCitation id="130FD0C8FE40FFABFBC0FED0FEB0FED2" box="[159,266,267,291]" captionStart="Fig" captionStartId="8.[168,199,1774,1794]" captionTargetBox="[152,1438,237,1886]" captionTargetId="figure-14@8.[167,1422,253,1761]" captionTargetPageId="8" captionText="Fig. 4 Otoliths of Gobius spp.: a–e Gobius bratishkoi n. sp., a holotype, NMNHU-P PI 2555, Kozatskyi Yar, b–e paratypes, b (reversed) Mlyntsi, NMNHU-P PI 2556,c–e (reversed) Kozatskyi Yar, NMB P1211. f Gobius reichenbacherae Schwarzhans,2014, Shydlivshchyna, NMNHU-P PI 2557 (reversed).g–k Gobius ukrainicus n. sp., j holotype, Shydlivshchyna, NMNHU-P PI 2558, g–i, k paratypes, g, i, k NMNHU-P PI 2560, h (reversed) Mlyntsi, NMNHU-P PI 2559" figureDoi="http://doi.org/10.5281/zenodo.11999881" httpUri="https://zenodo.org/record/11999881/files/figure.png" pageId="9" pageNumber="10">Fig. 4c, d</figureCitation>
|
||
), positioned below sulcus indentation marking ostial–caudal joint. Ventral furrow broad, deep, distinct, reaching from anterior to posterior tips of sulcus, more regularly curved than ventral rim of otolith. Ventral field between ventral furrow and sulcus slightly bulged. Dorsal field with long, narrow dorsal depression, ventrally marked by distinct crista superior toward sulcus and dorsally open to dorsal rim of otolith. Outer face convex, distinctly more convex than inner face and very regularly shaped, smooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E9FC6FE40FFABFBC8FDB7FE4EFA72" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="8B8BCC4DFE40FFABFBC8FDB7FE4EFA72" blockId="9.[151,776,619,1412]" pageId="9" pageNumber="10">
|
||
<emphasis id="B940105FFE40FFABFBC8FDB7FE62FD72" box="[151,472,619,643]" italics="true" pageId="9" pageNumber="10">
|
||
Discussion
|
||
<taxonomicName id="4C34B7CEFE40FFABFA4AFDB0FE62FD72" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[277,472,619,643]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bratishkoi">Gobius bratishkoi</taxonomicName>
|
||
</emphasis>
|
||
is easily recognized by its characteristic dorsal rim with the two concavities, the nearly vertical anterior rim, and the distinct postdorsal projection with rounded tip that is not bent outward. Further distinguishing characteristics include the outer face being more convex than the inner face and the very low ostial lobe of the sulcus.
|
||
<taxonomicName id="4C34B7CEFE40FFABFA8CFCF0FD29FCB2" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[467,659,811,835]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bratishkoi">
|
||
<emphasis id="B940105FFE40FFABFA8CFCF0FD29FCB2" box="[467,659,811,835]" italics="true" pageId="9" pageNumber="10">Gobius bratishkoi</emphasis>
|
||
</taxonomicName>
|
||
resembles the coeval
|
||
<taxonomicName id="4C34B7CEFE40FFABFA4FFC97FDE0FC92" authority="Schwarzhans, 2014" authorityName="Schwarzhans" authorityYear="2014" box="[272,602,843,867]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mustus">
|
||
<emphasis id="B940105FFE40FFABFA4FFC97FEC5FC92" box="[272,383,844,867]" italics="true" pageId="9" pageNumber="10">G. mustus</emphasis>
|
||
<bibRefCitation id="EFA5B1BCFE40FFABFAD7FC90FDE0FC92" author="Schwarzhans, W." box="[392,602,843,867]" pageId="9" pageNumber="10" pagination="35 - 69" refId="ref23301" refString="Schwarzhans, W. (2014). Otoliths from the middle Miocene (Serravallian) of the Karaman Basin, Turkey. Cainozoic Research, 14 (1), 35 - 69." type="journal article" year="2014">Schwarzhans, 2014</bibRefCitation>
|
||
</taxonomicName>
|
||
but clearly differs in shape of the dorsal rim, the near vertical anterior rim, the postdorsal projection, the strongly reduced ostial lobe, and the very small subcaudal iugum. However,
|
||
<taxonomicName id="4C34B7CEFE40FFABF9B1FC77FEB9FC12" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="bratishkoi">
|
||
<emphasis id="B940105FFE40FFABF9B1FC77FEB9FC12" italics="true" pageId="9" pageNumber="10">G. bratishkoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C34B7CEFE40FFABFA4DFC17FE3AFC12" authorityName="Schwarzhans" authorityYear="2014" box="[274,384,972,995]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mustus">
|
||
<emphasis id="B940105FFE40FFABFA4DFC17FE3AFC12" box="[274,384,972,995]" italics="true" pageId="9" pageNumber="10">G. mustus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C34B7CEFE40FFABFAE0FC17FDE8FC12" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[447,594,971,995]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ukrainicus" status="n. sp.">
|
||
<emphasis id="B940105FFE40FFABFAE0FC17FDE8FC12" box="[447,594,971,995]" italics="true" pageId="9" pageNumber="10">G. ukrainicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273AD24FE40FFABF905FC17FD2FFC12" box="[602,661,972,995]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
described below represent a distinct species group, which cannot be linked to any extant
|
||
<taxonomicName id="4C34B7CEFE40FFABFA99FBD0FDD2FBD2" box="[454,616,1035,1059]" pageId="9" pageNumber="10">
|
||
<emphasis id="B940105FFE40FFABFA99FBD0FDA9FBD2" box="[454,531,1035,1059]" italics="true" pageId="9" pageNumber="10">Gobius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and which we term the
|
||
<taxonomicName id="4C34B7CEFE40FFABFBA0FBF0FE1BFBB2" authorityName="Schwarzhans" authorityYear="2014" box="[255,417,1067,1091]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mustus">
|
||
<emphasis id="B940105FFE40FFABFBA0FBF0FE1BFBB2" box="[255,417,1067,1091]" italics="true" pageId="9" pageNumber="10">Gobius mustus</emphasis>
|
||
</taxonomicName>
|
||
complex. Among extant
|
||
<taxonomicName id="4C34B7CEFE40FFABF9E5FBF0FCBDFBB2" authorityName="Linnaeus" authorityYear="1758" box="[698,775,1067,1091]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940105FFE40FFABF9E5FBF0FCBDFBB2" box="[698,775,1067,1091]" italics="true" pageId="9" pageNumber="10">Gobius</emphasis>
|
||
</taxonomicName>
|
||
otoliths,
|
||
<taxonomicName id="4C34B7CEFE40FFABFBA4FB97FE46FB92" authority="Risso, 1810" authorityName="Risso" authorityYear="1810" box="[251,508,1099,1123]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="auratus">
|
||
<emphasis id="B940105FFE40FFABFBA4FB97FEC9FB92" box="[251,371,1100,1123]" italics="true" pageId="9" pageNumber="10">G. auratus</emphasis>
|
||
Risso, 1810
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C34B7CEFE40FFABF954FB97FCB8FB92" authority="Sarato, 1889" authorityName="Sarato" authorityYear="1889" box="[523,770,1099,1123]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="fallax">
|
||
<emphasis id="B940105FFE40FFABF954FB97FDD7FB92" box="[523,621,1099,1123]" italics="true" pageId="9" pageNumber="10">G. fallax</emphasis>
|
||
Sarato, 1889
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C34B7CEFE40FFABFB95FBB7FDAAFB72" authority="Vinciguerra, 1883" authorityName="Vinciguerra" authorityYear="1883" box="[202,528,1131,1155]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="vittatus">
|
||
<emphasis id="B940105FFE40FFABFB95FBB7FEFBFB72" box="[202,321,1132,1155]" italics="true" pageId="9" pageNumber="10">G. vittatus</emphasis>
|
||
Vinciguerra, 1883
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFA5B1BCFE40FFABF90FFBB0FCBAFB72" author="Gut, C. & Sanda, R. & Moritz, T. & Reichenbacher, B." box="[592,768,1131,1155]" pageId="9" pageNumber="10" pagination="282 - 323" refId="ref20633" refString="Gut, C., VukiC, J., Sanda, R., Moritz, T., & Reichenbacher, B. (2020). Identification of past and present gobies: Distinguishing Gobius and Pomatoschistus (Teleostei: Gobioidei) species using characters of otoliths, meristics and body morphometry. Contributions to Zoology, 89, 282 - 323." type="journal article" year="2020">Gut et al., 2020</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA5B1BCFE40FFABFBC8FB50FE00FB52" author="Schwarzhans, W. & Brzobohaty, R. & Radwanska, U." box="[151,442,1163,1187]" pageId="9" pageNumber="10" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., & Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
|
||
for figures) share the closest resemblance in terms of their compressed shapes, their inner faces being nearly flat, and their outer faces more convex. However, the degree of thickness of the outer face and the small size of the sulcus distinguish the
|
||
<taxonomicName id="4C34B7CEFE40FFABF98EFAD0FEB4FAB2" authorityName="Schwarzhans" authorityYear="2014" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="mustus">
|
||
<emphasis id="B940105FFE40FFABF98EFAD0FEB4FAB2" italics="true" pageId="9" pageNumber="10">Gobius mustus</emphasis>
|
||
</taxonomicName>
|
||
complex from those extant species and indicate that it may represent an extinct clade that is possibly separate from the genus
|
||
<taxonomicName id="4C34B7CEFE40FFABFAFDFAB0FE55FA72" authorityName="Linnaeus" authorityYear="1758" box="[418,495,1387,1411]" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940105FFE40FFABFAFDFAB0FE55FA72" box="[418,495,1387,1411]" italics="true" pageId="9" pageNumber="10">Gobius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |