314 lines
31 KiB
XML
314 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="A18048685394159FF2742C27204F3356" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole moramanaensis Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="B9BFD481-97AC-43D0-BABA-9C3BD42C67E6" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161777021" LSID="urn:lsid:zoobank.org:act:B9BFD481-97AC-43D0-BABA-9C3BD42C67E6" httpUri="http://treatment.plazi.org/id/A18048685394159FF2742C27204F3356" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/B9BFD481-97AC-43D0-BABA-9C3BD42C67E6" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole moramanaensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="moramanaensis" status="sp. nov.">Pheidole moramanaensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">Figs 53A-F</figureCitation>
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85L</figureCitation>
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<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87P</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toamasina;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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Ambatovy, 14.3 km 57° Moramanga; -18.85083, 48.32; alt. 1075 m; 23 Mar 2004; B.L. Fisher et al. leg.; BLF10624, CASENT0050486 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •2w., 2s.; same data as for holotype; CASENT0923172, CASENT0051295, CASENT0051296, CASENT0872186 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Fianarantsoa</emphasis>
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: •2w., 1s.; 9.0 km NE Ivohibe;
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<geoCoordinate degrees="22.42667" direction="south" orientation="latitude" precision="1" value="-22.42667">-22.42667</geoCoordinate>
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,
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<geoCoordinate degrees="46.93833" direction="east" orientation="longitude" precision="1" value="46.93833">46.93833</geoCoordinate>
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; alt. 900 m; 12 Nov 1997; B.L. Fisher et al. leg.; CASENT0198400, CASENT0198899 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mahajanga</emphasis>
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: •2w., 1s., 1q.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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Marotandrano, Marotandrano 48.3 km S Mandritsara;
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<geoCoordinate degrees="16.28322" direction="south" orientation="latitude" precision="1" value="-16.28322">-16.28322</geoCoordinate>
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,
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<geoCoordinate degrees="48.81443" direction="east" orientation="longitude" precision="1" value="48.81443">48.81443</geoCoordinate>
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; alt. 865 m; 7 Dec 2007; B.L. Fisher et al. leg.; CASENT0134236, CASENT0134237 (CASC). -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: •1w., 1q.; Bevolota 17.1 km N Andasibe;
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<geoCoordinate degrees="18.77071" direction="south" orientation="latitude" precision="1" value="-18.77071">-18.77071</geoCoordinate>
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,
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<geoCoordinate degrees="48.43164" direction="east" orientation="longitude" precision="1" value="48.43164">48.43164</geoCoordinate>
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; alt. 995 m; 12 Dec 2007; B.L. Fisher et al. leg.; CASENT0135181 (CASC). •3w., 2s., 1q.; Corridor Forestier Analamay-Mantadia, Ambatoharanana;
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<geoCoordinate degrees="18.80388" direction="south" orientation="latitude" precision="1" value="-18.80388">-18.80388</geoCoordinate>
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,
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<geoCoordinate degrees="48.40506" direction="east" orientation="longitude" precision="1" value="48.40506">48.40506</geoCoordinate>
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; alt. 1013 m; 12 Dec 2012; B.L. Fisher et al. leg.; CASENT0300854, CASENT0300855, CASENT0301765 (CASC). •1w., 1s.; Corridor Forestier Analamay-Mantadia, Ambatoharanana;
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<geoCoordinate degrees="18.80424" direction="south" orientation="latitude" precision="1" value="-18.80424">-18.80424</geoCoordinate>
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,
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<geoCoordinate degrees="48.40081" direction="east" orientation="longitude" precision="1" value="48.40081">48.40081</geoCoordinate>
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; alt. 968 m; 12 Dec 2012; B.L. Fisher et al. leg.; CASENT0301827 (CASC). •1w., 1s.; Corridor Forestier Analamay-Mantadia, Tsaravoniana;
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<geoCoordinate degrees="18.76124" direction="south" orientation="latitude" precision="1" value="-18.76124">-18.76124</geoCoordinate>
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,
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<geoCoordinate degrees="48.42134" direction="east" orientation="longitude" precision="1" value="48.42134">48.42134</geoCoordinate>
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; alt. 939 m; 2 Dec 2012; B.L. Fisher et al. leg.; CASENT0297007 (CASC). •2w., 1s., 1q.;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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Ambatovy, 14.3 km 57° Moramanga; -18.85083, 48.32; alt. 1075 m; 18 Dec 2004; B.L. Fisher et al. leg.; CASENT0110600, CASENT0110601 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Head in full face view rectangular, slightly widening posteriorly; sides of the head with moderately dense, long, erect pilosity; frons with thin, moderately dense, longitudinal rugae, interspaces smooth or with indistinct but dense rugulae; low, thick and triangular, with rounded apex; outer hypostomal teeth higher than inner hypostomal teeth, thick, with rounded tips, triangular inner and outer teeth closely spaced and connected by indistinct concavity; propodeal spines moderately long, with base slightly wider than top; first gastral tergite finely shagreened.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Whole head foveolate, fading on frons and genae; thin, sparse, longitudinal to irregular rugae overlie foveolate sculpture; promesonotum low, short, flat, or slightly convex, with steep posterior declivity; mesosoma foveolate, katepisternum and mesonotum smooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 9): HL: 1.61-1.79 (1.69); HW: 1.49-1.68 (1.57); SL: 0.62-0. 7 (0.67); EL: 0.17-0.2 (0.19); WL: 1.15-1.26 (1.2); PSL: 0.23-0.27 (0.26); MTL: 0.62-0.74 (0.68); PNW: 0.71-0.8 (0.75); PTW: 0.2-0.25 (0.22); PPW: 0.58-0.77 (0.7); CI: 91.5-95.2 (93.5); SI: 40.1-45.6 (42.7); PSLI: 14.0-16.2 (15.4); PPI: 28.4-34.0 (31.0); PNI: 45.7-49.5 (47.9); MTI: 41.7-45.1 (43.4).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view rectangular, slightly widening posteriorly, anterior of eyes relatively straight, posterior of eyes convex (Fig.
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming shallow transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Sides of head with moderately dense, long, erect pilosity; whole head with moderately long, dense, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae. Occipital lobes shiny, with thick, sparse, irregular rugae, interspaces smooth to finely rugulose; frons with thin, moderately dense, longitudinal rugae, interspaces smooth or with indistinct but dense rugulae; lateral sides of head with thin, dense, longitudinal rugulae, area between rugulae with sparse foveolae; malar area with dense and thin longitudinal rugulae; genae with fine, thin, dense rugulae. Clypeus shiny and smooth, with thin, longitudinal rugulae on the lateral sides; median notch present, wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, reaching the midlength of head; pilosity decumbent to erect (Fig.
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53B, D</figureCitation>
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). Inner hypostomal teeth distinct, low, thick and triangular, with rounded apex; outer hypostomal teeth higher than inner hypostomal teeth, thick, with rounded tips, triangular; inner and outer teeth closely spaced and connected by indistinct concavity (Fig.
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<figureCitation captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85L</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and relatively low, posterior mesonotum steep, with small, tubercle-like projections; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, thin, with base slightly wider than top, apex rounded; humeral area with small and flat tubercles (Fig.
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53D</figureCitation>
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). Surface shiny, with fine and sparse foveolae; mesonotum and katepisternum smooth, lateral sides of propodeum with few thick, longitudinal rugae; surface with additional indistinct and sparse rugulae on promesonotum. Pilosity moderately sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny, finely shagreened; peduncle short, with small, rounded, horizontal lobes on its basal part; node moderately high and narrow, with convex apex, in rear view node straight; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Shiny and finely shagreened; short and rounded; in dorsal view sides with relatively long, acute, horn-like projections; pilosity long, moderately dense, and erect (Fig.
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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First gastral tergite shiny and finely shagreened; pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Unicolourous, brown to dark brown; legs yellow (Fig.
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<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53D, F</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1" start="Figure 53" startId="F53">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 53.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole moramanaensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="moramanaensis">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole moramanaensis</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), profile (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
), and dorsal view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
) of paratype minor worker (CASENT0923172) and full-face view (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), profile (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
), and dorsal view (
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|
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
) of holotype major worker (CASENT0050486).
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 0.55-0.63 (0.59); HW: 0.51-0.57 (0.53); SL: 0.57-0.62 (0.59); EL: 0.1-0.13 (0.11); WL: 0.67-0.79 (0.74); PSL: 0.1-0.15 (0.11); MTL: 0.43-0.49 (0.46); PNW: 0.36-0.4 (0.38); PTW: 0.06-0.1 (0.08); PPW: 0.11-0.17 (0.08); CI: 87.2-93.8 (90.2); SI: 107.8-114.0 (111.4); PSLI: 16.2-23.5 (19.3); PPI: 52.1-61.8 (56.1); PNI: 68.8-71.8 (70.6); MTI: 83.0-93.3 (87.4).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig.
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|
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53A</figureCitation>
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|
). Pilosity moderately dense, long, erect. Whole head foveolate, fading on frons and genae; thin, sparse, longitudinal to irregular rugae overlie foveolate sculpture. Clypeus with median longitudinal carina present; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity erect (Fig.
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|
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53A, C</figureCitation>
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).
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|
<emphasis bold="true" pageId="0" pageNumber="1">
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|
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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|
.
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</emphasis>
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In lateral view, promesonotum low, short, flat or slightly convex, with steep posterior declivity; promesonotal groove absent; metanotal groove absent or indistinct; propodeal spines very short, triangular, apex acute (Fig.
|
|
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53C</figureCitation>
|
|
). Sculpture foveolate; katepisternum and mesosoma smooth. Pilosity sparse, long, and erect (Fig.
|
|
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
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|
</emphasis>
|
|
Peduncle moderately short and thin; node low, triangular, and small; with few long, erect setae (Fig.
|
|
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Short, low, and slightly convex; with few short, erect setae (Fig.
|
|
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
With sparse, erect pilosity (Fig.
|
|
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Unicolourous, yellow to yellowish brown (Fig.
|
|
<figureCitation captionStart="Figure 53" captionStartId="F53" captionText="Figure 53. Pheidole moramanaensis sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923172) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0050486)." figureDoi="10.3897/zookeys.905.39592.figure53" httpUri="https://binary.pensoft.net/fig/373100" pageId="0" pageNumber="1">53C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">The species was collected between 865-1075 m in elevation, in transition humid forest, rainforest, and montane rainforest. Nests were located in rotten logs, rotted twigs on the ground, and dead bamboo above the ground.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="comments">
|
|
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole moramanaensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="moramanaensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole moramanaensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is most similar to
|
|
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName lsidName="P. mantadia" pageId="0" pageNumber="1" rank="species" species="mantadia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. mantadia</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole moramanaensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="moramanaensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole moramanaensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by presence of fine foveolate sculpture on frons and mesosoma, low and thick inner hypostomal teeth which are wider than high, and thick outer hypostomal teeth; form
|
|
<taxonomicName lsidName="P. mantadia" pageId="0" pageNumber="1" rank="species" species="mantadia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. mantadia</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by denser and longer pilosity on head sides, and higher and narrower outer hypostomal teeth.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole moramanaensis" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="moramanaensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole moramanaensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. can be distinguished from
|
|
<taxonomicName lsidName="P. curvistriata" pageId="0" pageNumber="1" rank="species" species="curvistriata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. curvistriata</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by frons and malar area with thin, longitudinal to irregular rugae, and vertex with thin irregular and never arcuate rugae, absence of irregular rugae on mesosoma, and smooth mesonotum; from
|
|
<taxonomicName lsidName="P. mantadia" pageId="0" pageNumber="1" rank="species" species="mantadia">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. mantadia</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by frons and malar area with thin but distinct, irregular rugae and smooth clypeus.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |