253 lines
30 KiB
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253 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.949.51269" ID-GBIF-Dataset="e4d2c425-4596-487c-8c4f-b09330b48203" ID-PMC="PMC7381721" ID-Pensoft-Pub="1313-2970-949-1" ID-Pensoft-UUID="303D64593B3B550DBB7CE01AA547409D" ID-PubMed="32765176" ID-ZooBank="93BFA448BB7343CDBC4EA86090DA63D5" ModsDocID="1313-2970-949-1" checkinTime="1594832489974" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="2DA61EF8ABA45A0BB374D03E1E8A4F18" docLanguage="en" docName="ZooKeys 949: 1-185" docOrigin="ZooKeys 949" docSource="http://dx.doi.org/10.3897/zookeys.949.51269" docTitle="Pheidole analavelona Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="DC4E9F6D-96E6-42F5-8076-02D067506117" docUuidSource="ZooBank" docVersion="4" id="303D64593B3B550DBB7CE01AA547409D" lastPageNumber="1" masterDocId="303D64593B3B550DBB7CE01AA547409D" masterDocTitle="Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar" masterLastPageNumber="185" masterPageNumber="1" pageNumber="1" updateTime="1668169097828" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-0811-2309</mods:nameIdentifier>
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<mods:nameIdentifier type="email">sdsalata@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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<mods:affiliation>Department of Entomology, California Academy of Sciences, San Francisco, CA 94118, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>949</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>185</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.949.51269</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.949.51269</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-949-1</mods:identifier>
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<mods:identifier type="ZooBank">93BFA448BB7343CDBC4EA86090DA63D5</mods:identifier>
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<mods:identifier type="Pensoft-UUID">303D64593B3B550DBB7CE01AA547409D</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="165090193" LSID="urn:lsid:zoobank.org:act:DC4E9F6D-96E6-42F5-8076-02D067506117" httpUri="http://treatment.plazi.org/id/2DA61EF8ABA45A0BB374D03E1E8A4F18" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="2DA61EF8-ABA4-5A0B-B374-D03E1E8A4F18" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole analavelona" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="analavelona" status="sp. nov.">Pheidole analavelona</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">Figs 21A-F</figureCitation>
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">, 63C</figureCitation>
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<figureCitation captionStart="Figure 65" captionStartId="F65" captionText="Figure 65. Distribution. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T)." figureDoi="10.3897/zookeys.949.51269.figure65" httpUri="https://binary.pensoft.net/fig/432234" pageId="0" pageNumber="1">, 65C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype.</emphasis>
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Madagascar. • 1 major worker; Toliara;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken originalValue="Classée">Classee</normalizedToken>
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<normalizedToken originalValue="d’Analavelona">d'Analavelona</normalizedToken>
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, 33.2 km 344°NNW Mahaboboka;
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<geoCoordinate degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
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,
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<geoCoordinate degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
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; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07970; CASENT0491849, top specimen on the pin (CASC).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes.</emphasis>
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• 9w., 6s., 2m.; same data as for holotype; CASENT0491847, CASENT0491846, CASENT0872248, CASENT0491850, CASENT0491845, CASENT0491844, CASENT0491848 (CASC, MHNG, PBZT).
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</paragraph>
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<caption doi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1" start="Figure 21" startId="F21">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 21.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole analavelona" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="analavelona">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole analavelona</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0491846) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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) of holotype major worker (CASENT0491849).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" pageId="0" pageNumber="1">Toliara</emphasis>
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: • 3w.;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken originalValue="Classée">Classee</normalizedToken>
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<normalizedToken originalValue="d’Analavelona">d'Analavelona</normalizedToken>
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, 29.2 km 343°NNW Mahaboboka; -22.675, 44.19; alt. 1100 m; 18 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07833 (CASC). • 3w., 3s.; ibid.; BLF07835 (CASC). • 3w.; ibid.; BLF07844 (CASC). • 3w., 3s.; ibid.; BLF07857 (CASC). •3w.; ibid.; BLF07887 (CASC). • 3s.; ibid.; under stone; BLF07853 (CASC). • 9w., 3s.;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken originalValue="Classée">Classee</normalizedToken>
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<normalizedToken originalValue="d’Analavelona">d'Analavelona</normalizedToken>
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, 33.2 km 344°NNW Mahaboboka;
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<geoCoordinate degrees="22.64333" direction="south" orientation="latitude" precision="1" value="-22.64333">-22.64333</geoCoordinate>
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,
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<geoCoordinate degrees="44.17167" direction="east" orientation="longitude" precision="1" value="44.17167">44.17167</geoCoordinate>
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; alt. 1300 m; 22 Feb 2003; Fisher et al. leg.; montane rainforest, ex rotten log; BLF07953 (CASC). • 1w., 1s.; ibid.; BLF07966 (CASC). • 3., 3s.; ibid.; BLF07971 (CASC). • 6w., 1s., 1q., 6m.; ibid.; BLF07989 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Moderately large species.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Head in full-face view oval, not widening posteriorly, with anterior and posterior sides convex, in lateral view sub-oval, ventral and dorsal faces convex, occipital cleft moderately deep; sides of the head with dense, very long, suberect to erect pilosity; frons with thick, irregular rugae, moderately dense on lateral sides and sparser on medial part; interspaces shiny with distinct and sparse rugoreticulae; occipital lobes, area posterolateral from eyes never smooth; scape, when laid back, exceeding the midlength of head by one-fifth of its length; inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed inward; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth not closely spaced and not connected by concavity; mesosoma with dense rugofoveolae; pronotum, mesonotum, katepisternum, and anepisternum with additional thick to thin, irregular to transverse rugae; body dark brown.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Head foveolate; medial frons with foveolae sparser and with moderately thick longitudinal and interrupted rugae; lateral sides of frons and vertex with additional moderately thick rugae; area posterolateral from eyes with reduced sculpture and smooth notches; scape, when laid back, surpassing the posterior head margin by two-fifths of its length; promesonotum moderately high and short; promesonotal groove absent; propodeal spines minute and triangular; propodeum predominantly smooth with indistinct rugofoveolae on lateral sides; mesonotum, anepisternum, katepisternum, and propodeum with sparse and indistinct rugofoveolae; body dark orange.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 0.97-1.02 (1.0); HW: 0.95-1.0 (0.98); SL: 0.64-0.67 (0.66); EL: 0.11-0.13 (0.12); WL: 0.88-0.95 (0.92); PSL: 0.13-0.15 (0.14); MTL: 0.57-0.6 (0.58); PNW: 0.44-0.48 (0.46); PTW: 0.13-0.17 (0.14); PPW: 0.26-0.29 (0.28); CI: 101.0-104.4 (102.0); SI: 65.3-69.3 (67.1); PSLI: 13.1-15.4 (14.1); PPI: 44.8-57.2 (52.2); PNI: 46.2-48.8 (47.4); MTI: 57.0-61.8 (59.3).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
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In full-face view oval, not widening posteriorly, with anterior and posterior sides convex (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21B</figureCitation>
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). In lateral view sub-oval; ventral and dorsal faces convex; inner hypostomal teeth visible. Sides of the head with dense, very long, suberect to erect pilosity; whole head with dense, long, decumbent to erect pilosity. Frons with thick, irregular rugae, moderately dense on lateral sides and sparser on medial part; interspaces shiny with distinct and sparse rugoreticulae. Occipital lobes with relatively dense, thick, and irregular rugae; interspaces rugoreticulate. Gena with relatively dense, thick, and longitudinal rugae and rugoreticulate interspaces. Area posterolateral from eyes with moderately dense, thick, and irregular rugae; interspaces shiny with distinct and sparse rugoreticulae. Centre of clypeus smooth and shiny, lateral sides with indistinct rugulae; median notch present, moderately wide, and shallow; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, exceeding the midlength of head by one-fifth of its length; pilosity subdecumbent to erect (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21B, D</figureCitation>
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). Inner hypostomal teeth distinct, large, closely spaced, triangular, with rounded apex directed inward; outer hypostomal teeth lobe-like, wider than inner hypostomal teeth and approximately the same height, apex directed outward; inner and outer hypostomal teeth not closely spaced and not connected by concavity (Fig.
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<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Major worker, hypostomal teeth. Pheidole alina sp. nov. (A). Pheidole ambohimanga sp. nov. (B). Pheidole analavelona sp. nov. (C). Pheidole andohahela sp. nov. (D). Pheidole anomala sp. nov. (E). Pheidole anosyenne sp. nov. (F). Pheidole antranohofa sp. nov. (G). Pheidole beanka sp. nov. (H). Pheidole befotaka sp. nov. (I). Pheidole dasos sp. nov. (J). Pheidole flavominuta sp. nov. (K). Pheidole gracilis sp. nov. (L). Pheidole haboka sp. nov. (M). Pheidole havoana sp. nov. (N). Pheidole hazo sp. nov. (O). Pheidole itremo sp. nov. (P). Pheidole joffreville sp. nov. (Q). Pheidole kely sp. nov. (R). Pheidole lavasoa sp. nov. (S). Pheidole litigiosa Forel (T). Pheidole mahamavo sp. nov. (U). Pheidole mainty sp. nov. (V). Pheidole mamiratra sp. nov. (W). Pheidole manantenina sp. nov. (X)." figureDoi="10.3897/zookeys.949.51269.figure63" httpUri="https://binary.pensoft.net/fig/432232" pageId="0" pageNumber="1">63C</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
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In lateral view, promesonotum short, angular, and moderately low, posterior mesonotum moderately steep, mesonotal process indistinct, tubercle-like; promesonotal groove absent; metanotal groove indistinct; propodeal spines moderately short, with wide base and acute apex; humeral area laterally weakly produced (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21D</figureCitation>
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). Surface shiny with dense rugofoveolae; pronotum, mesonotum, katepisternum, and anepisternum with additional thick to thin, irregular to transverse rugae. Pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Petiole.</emphasis>
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Shiny with fine foveolae; node smooth to finely foveolate, low, triangular, with rounded and thin apex, in rear view node dorsoventrally straight to slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postpetiole.</emphasis>
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Shiny and foveolate; dorsum with reduced sculpture and smooth notch; in dorsal view oval, lateral margins medially with two dentate projections; pilosity long, moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
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Shiny and smooth; pilosity dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21D, F</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
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Dark brown; legs and antennae yellow (Fig.
|
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 10): HL: 0.55-0.59 (0.57); HW: 0.46-0.48 (0.47); SL: 0.59-0.63 (0.62); EL: 0.09-0.11 (0.1); WL: 0.69-0.73 (0.71); PSL: 0.06-0.09 (0.08); MTL: 0.46-0.49 (0.47); PNW: 0.32-0.35 (0.33); PTW: 0.07-0.1 (0.08); PPW: 0.12-0.14 (0.13); CI: 116.6-124.7 (120.7); SI: 125.9-133.5 (130.5); PSLI: 10.6-15.1 (14.0); PPI: 55.0-81.5 (66.0); PNI: 67.4-72.0 (69.8); MTI: 97.3-102.3 (99.4).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head.</emphasis>
|
||
Cephalic margin indistinctly concave or straight (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21A</figureCitation>
|
||
). Pilosity relatively dense, long, decumbent to suberect. Sculpture foveolate; medial frons with foveolae sparser and with moderately thick, longitudinal, and interrupted rugae; lateral sides of frons and vertex with additional moderately thick rugae; area posterolateral from eyes with reduced sculpture and smooth notches; antennal sockets with few thick, curved outward rugae and foveolate interspaces. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity dense, subdecumbent to erect (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21A, C</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Mesosoma.</emphasis>
|
||
In lateral view, promesonotum moderately high and short, arched; promesonotal groove absent; metanotal groove distinct; propodeal spines minute, triangular (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21C</figureCitation>
|
||
). Sculpture shiny; propodeum predominantly smooth with indistinct rugofoveolae on lateral sides; mesonotum, anepisternum, katepisternum, and propodeum with sparse and indistinct rugofoveolae. Pilosity moderately dense, long, and erect (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gaster.</emphasis>
|
||
With sparse, erect pilosity (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colour.</emphasis>
|
||
Dark orange, gaster slightly darker (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Pheidole analavelona sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0491846) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0491849)." figureDoi="10.3897/zookeys.949.51269.figure21" httpUri="https://binary.pensoft.net/fig/432189" pageId="0" pageNumber="1">21C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The species was collected between 1100-1300 m in elevation, in montane rainforest. Nests were located in rotten logs and under stones.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole analavelona" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="analavelona">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole analavelona</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is a member of the group of species characterised by major workers with head in full-face view oval and not widening posteriorly and medial part of frons with thick, moderately sparse, and irregular rugae or medial frons with moderately dense, thin, longitudinal anteriorly to irregular posteriorly, and interrupted rugae and very shallow occipital cleft. Minor workers of this group have short and moderately high promesonotum and dark body colouration ranging from orange to brown. The group consists of three species:
|
||
<taxonomicName lsidName="P. vadum" pageId="0" pageNumber="1" rank="species" species="vadum">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. vadum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName lsidName="P. analavelona" pageId="0" pageNumber="1" rank="species" species="analavelona">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. analavelona</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., and
|
||
<taxonomicName lsidName="P. ambohimanga" pageId="0" pageNumber="1" rank="species" species="ambohimanga">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. ambohimanga</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole analavelona" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="analavelona">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole analavelona</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is known from
|
||
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
|
||
<normalizedToken originalValue="Classée">Classee</normalizedToken>
|
||
<normalizedToken originalValue="d’Analavelona">d'Analavelona</normalizedToken>
|
||
in Toliara and its distribution does not overlap with that of the two remaining members of the group known from the vicinity of Antananarivo. Morphologically both minor and major workers of
|
||
<taxonomicName lsidName="P. analavelona" pageId="0" pageNumber="1" rank="species" species="analavelona">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. analavelona</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. are most similar to
|
||
<taxonomicName lsidName="P. ambohimanga" pageId="0" pageNumber="1" rank="species" species="ambohimanga">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. ambohimanga</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Majors of
|
||
<taxonomicName lsidName="P. analavelona" pageId="0" pageNumber="1" rank="species" species="analavelona">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. analavelona</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished based on distinctly rugoreticulate interspaces on head while minor workers have head sculpture foveolate with foveolae sparser on frons and area posterolateral from eyes, and sparse but never absent rugofoveolae on mesonotum, anepisternum, katepisternum, and propodeum. Minors of
|
||
<taxonomicName lsidName="P. analavelona" pageId="0" pageNumber="1" rank="species" species="analavelona">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. analavelona</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be easily separated from other members of the
|
||
<taxonomicName lsidName="sikorae" pageId="0" pageNumber="1" rank="species" species="sikorae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">sikorae</emphasis>
|
||
</taxonomicName>
|
||
group based on short and moderately high promesonotum and additional, moderately thick to thick, longitudinal, and interrupted rugae on head.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |