853 lines
71 KiB
XML
853 lines
71 KiB
XML
<document id="DBA3ABE1D0BE0F70C5B77148AEA25A2F" ID-DOI="10.3897/zookeys.1208.117056" ID-ZooBank="ECCF2630-D984-45A8-BF19-DA219F26D495" ID-publisher-id="117056" URI-arpha="FE1DA5D8-62B9-5B21-BF52-9AF3C782709F" URI-zoobank="http://zoobank.org/ECCF2630-D984-45A8-BF19-DA219F26D495" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1722869495237" checkinUser="pensoft" docAuthor="Nahok, Benchawan, Chanlabut, Utain, Tumpeesuwan, Sakboworn & Tumpeesuwan, Chanidaporn" docDate="2024" docId="03FA6E1C6C5454AAA4E3908EC0E7EBAD" docLanguage="en" docName="ZooKeys 1208: 347-362" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/117056/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Landouria bella Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan 2024, sp. nov." docType="treatment" docUuid="DDCCEAB2-071E-4EB0-83D4-077CF7EE0232" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="362" masterDocId="FE1DA5D862B95B21BF529AF3C782709F" masterDocTitle="Two new species of Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Thailand, with a key to Thai species" masterLastPageNumber="362" masterPageNumber="347" pageNumber="347" updateTime="1722873878227" updateUser="admin">
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<mods:title id="7DE291A9F6AA701FEC4E523B7FAE31E5">Two new species of Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Thailand, with a key to Thai species</mods:title>
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<mods:name id="0DE9A54053C5201954851B2CBF6AADA3" type="personal">
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<mods:roleTerm id="75286B5ED2A466A8AD094AF505482831">Author</mods:roleTerm>
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<mods:namePart id="6507726823ABE61223D55319752C0D8C">Nahok, Benchawan</mods:namePart>
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<mods:nameIdentifier id="C9F4E7252FDD2908CF4BA7BACF77F762" type="ORCID">https://orcid.org/0009-0009-1788-6788</mods:nameIdentifier>
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<mods:affiliation id="FC51DBBCD54F6A000D9BD443310411E3">Department of General Science, Faculty of Education, Chaiyaphum Rajabhat University, Muang District, Chaiyaphum, 36000, Thailand</mods:affiliation>
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<mods:name id="1321D67BFABAD9877EDA5B10E026E496" type="personal">
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<mods:roleTerm id="232F184F168825431248A6D9F8FE5CBE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B609B79F3E95CE498EDA9FB043267D5D">Chanlabut, Utain</mods:namePart>
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<mods:nameIdentifier id="57163FF10EF7A2738652587A41FDE596" type="ORCID">0000-0001-8275-7434</mods:nameIdentifier>
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<mods:affiliation id="576AF56C613F0A16A1B474E6B60576C4">Department of General Science, Faculty of Education, Chaiyaphum Rajabhat University, Muang District, Chaiyaphum, 36000, Thailand</mods:affiliation>
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<mods:name id="E5DDE3D674D3E945EA0ABAC95AF092AD" type="personal">
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<mods:roleTerm id="79EC6CD62425E4546D071BAD089B174A">Author</mods:roleTerm>
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<mods:namePart id="9C9F546BB6798BA354928DFB0375388E">Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:nameIdentifier id="0D2A6AD994BC65B0989A3A70AE7A521E" type="ORCID">0000-0001-5900-7862</mods:nameIdentifier>
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<mods:affiliation id="78546724091708F70662C81E70162184">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150, Thailand</mods:affiliation>
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<mods:namePart id="85DD0E166D4358B6C91E51279FDA42AA">Tumpeesuwan, Chanidaporn</mods:namePart>
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<mods:affiliation id="D266D1DA808B4F0B3068B57A6A3D449A">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham, 44150, Thailand</mods:affiliation>
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<mods:typeOfResource id="BC4E72DE49542A7AF3F2286B724E124D">text</mods:typeOfResource>
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<mods:title id="EE20B40E07B4ADC4C3019E7F55193E8C">ZooKeys</mods:title>
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<mods:part id="79AE1930CD1DBF7E2D05765957B995FF">
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<mods:date id="BC1818404EB5D7D32C425B93CE4D4111">2024</mods:date>
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<mods:number id="111BF4B9B4B1104C6DF97B1538DA2864">2024-08-05</mods:number>
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<mods:number id="4C130D4D5785CEF7B779608FE9748DEE">1208</mods:number>
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<mods:classification id="B416C4D80940168CC2FCBF556B0ED8D7">journal article</mods:classification>
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<mods:identifier id="B14D749CADDA5794C2C1B26A8BACC934" type="ZooBank">ECCF2630-D984-45A8-BF19-DA219F26D495</mods:identifier>
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<treatment id="03FA6E1C6C5454AAA4E3908EC0E7EBAD" ID-arpha="03FA6E1C-6C54-54AA-A4E3-908EC0E7EBAD" ID-zoobank="https://zoobank.org/DDCCEAB2-071E-4EB0-83D4-077CF7EE0232" LSID="urn:lsid:zoobank.org:act:DDCCEAB2-071E-4EB0-83D4-077CF7EE0232" LSID-ZBK="urn:lsid:zoobank.org:act:DDCCEAB2-071E-4EB0-83D4-077CF7EE0232" httpUri="http://treatment.plazi.org/id/03FA6E1C6C5454AAA4E3908EC0E7EBAD">
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<subSubSection id="CCF608073BE8415225217DC6E10A710A" type="nomenclature">
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<paragraph id="2D6139E96C23C3F322A7B26BC1653208">
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<taxonomicName id="97553E32BD5E87337D5E497AD5078854" authority="Nahok & S. Tumpeesuwan" authorityName="Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="bella" status="sp. nov.">
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<emphasis id="1A4CC9CBBAE46440F709957CBE2C869C" italics="true">Landouria bella</emphasis>
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||
Nahok & S. Tumpeesuwan
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||
</taxonomicName>
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||
<taxonomicNameLabel id="7D03EB57892CE99E1ED68B3072262D84" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="11C4B8D0DB4A7E88760A590F641F3A3B" type="description">
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<paragraph id="AB27A344B17424F0990AA61129F420BF">
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<figureCitation id="82BD69CD8754870BF876186E6657281C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Landouria bella sp. nov. A holotype (NHMSU - 00056) B paratype (ZCPRU - 0042) C, D empty shell from Khao Nang Panthurat Forest Park, Phetchaburi Province (NHMSU - 00058)." figureDoi="10.3897/zookeys.1208.117056.figure2" httpUri="https://binary.pensoft.net/fig/1108222">Figs 2</figureCitation>
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||
,
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<figureCitation id="1D44C5D69A7DB045BF4795D3B0B4CB18" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Genital anatomy of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole genitalia B – D internal wall sculpture of B penis (p) C flagellum (fl) D vagina (va)." figureDoi="10.3897/zookeys.1208.117056.figure3" httpUri="https://binary.pensoft.net/fig/1108223">3</figureCitation>
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||
,
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<figureCitation id="C2B85AB372B1F907B652CDB4EEC5068A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Radula morphology of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole radula B close-up view of central tooth and lateral teeth C right side of central tooth and lateral teeth D right side of marginal teeth. Numbers indicate order of lateral and marginal teeth. Abbreviation: C = central tooth, L = lateral teeth, M = marginal teeth." figureDoi="10.3897/zookeys.1208.117056.figure4" httpUri="https://binary.pensoft.net/fig/1108224">4</figureCitation>
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||
,
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||
<tableCitation id="B59C8A07B222F9B39FA1CC4C89E13F8F" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shells, genital systems and radulae of Thai Landouria species. Reference: 1 = Tumpeesuwan and Tumpeesuwan (2019); 2 = Nahok et al. (2021); 3 = this study; 4 = Schileyko (2011) (question mark = no data)." httpUri="http://table.plazi.org/id/2F840F1BAB846FE53F97C0112811F9B7" tableUuid="2F840F1BAB846FE53F97C0112811F9B7">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="07E80F6C32A6364DC57E310DB593A7DB" type="reference_group">
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<paragraph id="C2BFF5D4F6F5998849CA61419231F9B4">
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||
<treatmentCitationGroup id="C09C5A42FC05CF318E72F63C772D1B80">
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||
<taxonomicName id="E41B05C34BEF1C27ABBDF516A3496287" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="29A94E2542693A658F26E5246DC08D47" italics="true">Landouria</emphasis>
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||
<comment id="478E5CC8972438609BBB6EA23106E9BD">
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||
sp. 16 —
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||
<treatmentCitation id="31108E9958FADE0D46EB084C2BB4D6D8" author="Nahok B" page="61" year="2020">
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||
<bibRefCitation id="96D3F520FE94B42B57402D11672B6F5A" author="Nahok B" firstAuthor="Nahok" page="61" refId="B15" refString="Nahok B (2020) Systematics and Taxonomy of Land Snails Genera Aegista, Thaitropis, Landouria, and Pseudobuliminus in Thailand. Unpublished PhD Thesis, Department of Biology, Faculty of Science, Mahasarakham University, Maha Sarakham." year="2020">Nahok 2020: 61</bibRefCitation>
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||
</treatmentCitation>
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||
; fig. 24 G, I, table 3
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</comment>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="51AC30D7E9063260B2153EF72CD10B67">
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<treatmentCitationGroup id="A2AE507BBE854583F27B9544D92A1534">
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<treatmentCitation id="5A4620341813F8A62564839E595AD1D4">
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<taxonomicName id="6B2F38C0C6AE0BCE5CE6D82F0854D5B9" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="undetermined">
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||
<emphasis id="681D891DB6457EDF188AED6366CF856D" italics="true">Landouria</emphasis>
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||
sp.
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||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="53D8C0806F645CB17D33D533E1F45945" DOI="10.14456/lsej.2022.20" author="Chanlabut and Nahok" firstAuthor="Chanlabut" issue="1" journalOrPublisher="Life Sciences and Environment Journal" pagination="259-270" refId="B2" refString="Chanlabut U, Nahok B (2022) Species diversity and abundance of land snails in Khao Bin limestone hills, Ratchaburi Province. Life Sciences and Environment Journal 23 (1): 259–270. https://doi.org/10.14456/lsej.2022.20" title="Species diversity and abundance of land snails in Khao Bin limestone hills, Ratchaburi Province." volume="23" year="2022">Chanlabut and Nahok 2022</bibRefCitation>
|
||
</treatmentCitation>
|
||
: figs 2 D, 3 D, table 1
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="SECID0EGJAE" type="type locality">
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||
<paragraph id="C394A3D464A31E40F2FAFC4895F1B4A4">
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||
<heading id="D4285345C75238F25F83AAE457EC9A87" reason="title">Type locality.</heading>
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||
</paragraph>
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||
<paragraph id="CEC7657A7035E985296C11C0A5EBC40F">
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||
<collectingCountry id="022E02092CA398323077E718D84320BE" name="Thailand">Thailand</collectingCountry>
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,
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<collectingRegion id="1F2F16BE00C08B4762094953284052FA" country="Thailand" name="Ratchaburi">Ratchaburi Province</collectingRegion>
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, Mueang
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<collectingRegion id="28B99B9DE979677316AAE070CAFD095F" country="Thailand" name="Ratchaburi">Ratchaburi District</collectingRegion>
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, Khao Bin Cave,
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||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EPJAE" specific-use="{"type":"Point","coordinates":[99.665714,13.593767]}">
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<geoCoordinate id="85523E0987B56BDB8008CCE5FBB8838E" degrees="13" direction="north" minutes="35" orientation="latitude" precision="1" seconds="37.56" value="13.593766">13 ° 35 ' 37.56 " N</geoCoordinate>
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,
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<geoCoordinate id="BCCF4EBCE73FF2D8D293045F5B23105F" degrees="99" direction="east" minutes="39" orientation="longitude" precision="1" seconds="56.57" value="99.66572">99 ° 39 ' 56.57 " E</geoCoordinate>
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</named-content>
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, Isolated limestone hills, alt.
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<specimenCount id="F685186684AD01B016C059837B88DB5D" count="116" type="generic">
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<quantity id="66D3F83A707BACEDC89CF1B264F44778" metricMagnitude="2" metricUnit="m" metricValue="1.16" unit="m" value="116.0">116 m</quantity>
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.
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</specimenCount>
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<date id="11C3E1416BEA54C16FA26E096E874E2C" value="2018-07-07">7 July 2018</date>
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||
. Benchawan Nahok and Utain Chanlabut leg.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="SECID0EUJAE" type="Type specimens">
|
||
<paragraph id="C677A2A8A7AB43A331FF517F7250A2BB">
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||
<heading id="52F36658813E1F84932D387F4E442437" reason="title">Type specimens.</heading>
|
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</paragraph>
|
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<paragraph id="B4BC2F20BBE7094053606838A6AB000A">
|
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<materialsCitation id="120E446A77B53C4F002CEF9E615BACF7" collectionCode="NHMSU, UW" specimenCode="NHMSU-00056" specimenCount="1" typeStatus="holotype">
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<emphasis id="656FEF0D0878FC7D6F4024B4F9F23280" bold="true" italics="true">
|
||
<typeStatus id="5A838B0C3B36537F9CD32E36D753109A" type="holotype">Holotype</typeStatus>
|
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</emphasis>
|
||
: •
|
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<specimenCode id="1571604E19210BC2ED5DE1FB1A2D0788">
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<named-content id="8F00C46DBC09F8DE6F36523D77D8D0DB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahasarakham-university" xlink_title="Natural History Museum, Mahasarakham University">NHMSU</named-content>
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-00056
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</specimenCode>
|
||
(Fig.
|
||
<figureCitation id="E5D97CE9EBE7662D77A5B921129C9C2E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Landouria bella sp. nov. A holotype (NHMSU - 00056) B paratype (ZCPRU - 0042) C, D empty shell from Khao Nang Panthurat Forest Park, Phetchaburi Province (NHMSU - 00058)." figureDoi="10.3897/zookeys.1208.117056.figure2" httpUri="https://binary.pensoft.net/fig/1108222">2 A</figureCitation>
|
||
),
|
||
<specimenCount id="3BA9B25AAE7E0213C60D8B5FCB2F76BB">1 shell</specimenCount>
|
||
,
|
||
<abbrev id="ABBRID0EHKAE" xlink_title="shell height">SH</abbrev>
|
||
= 8.0 mm,
|
||
<abbrev id="ABBRID0ELKAE" xlink_title="shell width">SW</abbrev>
|
||
=
|
||
<quantity id="F5F3CC88A417410EC0A470C44E10A124" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" unit="mm" value="11.1">11.1 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0EPKAE" xlink_title="aperture height">AH</abbrev>
|
||
=
|
||
<quantity id="C63B9CAC86350A1772EA11C472EDA31D" metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0ETKAE" xlink_title="aperture width">AW</abbrev>
|
||
=
|
||
<quantity id="B304430A59A835B2385B5DA42084F429" metricMagnitude="-3" metricUnit="m" metricValue="4.3" unit="mm" value="4.3">4.3 mm</quantity>
|
||
,
|
||
<abbrev id="ABBRID0EXKAE" xlink_title="umbilicus width">UW</abbrev>
|
||
=
|
||
<quantity id="F1FAA0DEDB43EE0CC6E42A36CDCCEBE8" metricMagnitude="-3" metricUnit="m" metricValue="3.3" unit="mm" value="3.3">3.3 mm</quantity>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="92C3FEB29A6F7C198AF2E0D307551331" collectionCode="NHMSU" location="Figs" specimenCode="NHMSU-00057" specimenCount="27" typeStatus="paratype">
|
||
<emphasis id="CD1A29B802F42ACF7F0E1CC2B8978574" bold="true" italics="true">
|
||
<typeStatus id="DDFA20D467683DA9730F64436C9AA292" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
: •
|
||
<specimenCode id="703C81F504476E9438D9C947D396C8AE">
|
||
<named-content id="B89D55775E4974A4AFB9FE9F74B7C6AB" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahasarakham-university" xlink_title="Natural History Museum, Mahasarakham University">NHMSU</named-content>
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||
-00057
|
||
</specimenCode>
|
||
(Figs
|
||
<figureCitation id="55A1A89BF250AFE02456FAA57DE523E2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Landouria bella sp. nov. A holotype (NHMSU - 00056) B paratype (ZCPRU - 0042) C, D empty shell from Khao Nang Panthurat Forest Park, Phetchaburi Province (NHMSU - 00058)." figureDoi="10.3897/zookeys.1208.117056.figure2" httpUri="https://binary.pensoft.net/fig/1108222">2 B</figureCitation>
|
||
,
|
||
<figureCitation id="DC42B1C0295098FABACC5D230FD84BB5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Genital anatomy of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole genitalia B – D internal wall sculpture of B penis (p) C flagellum (fl) D vagina (va)." figureDoi="10.3897/zookeys.1208.117056.figure3" httpUri="https://binary.pensoft.net/fig/1108223">3</figureCitation>
|
||
,
|
||
<figureCitation id="BF6EF68D39AE7F510359B89B38A07D3E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Radula morphology of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole radula B close-up view of central tooth and lateral teeth C right side of central tooth and lateral teeth D right side of marginal teeth. Numbers indicate order of lateral and marginal teeth. Abbreviation: C = central tooth, L = lateral teeth, M = marginal teeth." figureDoi="10.3897/zookeys.1208.117056.figure4" httpUri="https://binary.pensoft.net/fig/1108224">4</figureCitation>
|
||
)
|
||
<specimenCount id="18F6EC50CB1149505BFDBEC4D90A5C04">27 shells</specimenCount>
|
||
</materialsCitation>
|
||
, •
|
||
<materialsCitation id="3CDCC5BA53A0AB55DE168817E2B7FF13" collectionCode="ZMCRU" specimenCode="ZMCRU-0001" specimenCount="11">
|
||
<specimenCode id="6EE52B33C7B31FFBF6CDB5D812E381E8">
|
||
<abbrev id="ABBRID0EOLAE" xlink_title="oological Research Collection of Muban Chombueng Rajabhat University">ZMCRU</abbrev>
|
||
-0001
|
||
</specimenCode>
|
||
<specimenCount id="B592BF3B76DFD8884BC9D1C114F1B058">11 shells</specimenCount>
|
||
</materialsCitation>
|
||
, •
|
||
<materialsCitation id="7970A9A78705E39D63040A70964E93DB" collectionCode="ZCPRU" specimenCode="ZCPRU-0041" specimenCount="65">
|
||
<specimenCode id="A0B3053AE2D2486F326AF852CC5E9ED7">
|
||
<abbrev id="ABBRID0ESLAE" xlink_title="Zoological Research Collection of Chaiyaphum Rajabhat University">ZCPRU</abbrev>
|
||
-0041
|
||
</specimenCode>
|
||
<specimenCount id="780F3A33397F247AE7C0DF59BABB1203">65 shells</specimenCount>
|
||
</materialsCitation>
|
||
, •
|
||
<materialsCitation id="5127158D8A48E75F8C95D67903CD9486" collectingDate="2020-06-27" collectionCode="ZCPRU" specimenCode="ZCPRU-0042" specimenCount="5" typeStatus="paratype">
|
||
<specimenCode id="D18EF5BFEE16D01A1135E16F471DB7CF">
|
||
<abbrev id="ABBRID0EWLAE" xlink_title="Zoological Research Collection of Chaiyaphum Rajabhat University">ZCPRU</abbrev>
|
||
-0042
|
||
</specimenCode>
|
||
<specimenCount id="D3C2E503C25B7C286AF192B21E71CE6F">5 living specimens</specimenCount>
|
||
preserved in ethanol, same leg. and locality as holotype,
|
||
<date id="A7185170BB525B03EE3B8174779FB4C5" value="2020-06-27">
|
||
<collectingDate id="10A4CBA35435551FBE679A54DF0A0064" value="2020-06-27">27 June 2020</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="0334E19BB4C85204F7AD6BD64AD98AEA" bold="true" italics="true">
|
||
<collectorName id="E284C4FE48427E658E6E5CB2A9AA5EFC">Other</collectorName>
|
||
locality
|
||
</emphasis>
|
||
: •
|
||
<materialsCitation id="9FA7D209307CCAA2833D3AA4B15F7593" collectionCode="NHMSU" location="Fig." specimenCode="NHMSU-00058" specimenCount="4" typeStatus="paratype">
|
||
<specimenCode id="9C5946B485530990EA39BAAE1C44A84D">
|
||
<named-content id="6CD919AEC6E70B6FB6A4C342212259AF" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahasarakham-university" xlink_title="Natural History Museum, Mahasarakham University">NHMSU</named-content>
|
||
-00058
|
||
</specimenCode>
|
||
(Fig.
|
||
<figureCitation id="0E2BB5C8FB08DCB1C22A239739C739C8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Landouria bella sp. nov. A holotype (NHMSU - 00056) B paratype (ZCPRU - 0042) C, D empty shell from Khao Nang Panthurat Forest Park, Phetchaburi Province (NHMSU - 00058)." figureDoi="10.3897/zookeys.1208.117056.figure2" httpUri="https://binary.pensoft.net/fig/1108222">2 C, D</figureCitation>
|
||
)
|
||
<specimenCount id="D74D342BA68EEEB670FCEEB4FBF09D5A">4 shells</specimenCount>
|
||
</materialsCitation>
|
||
, •
|
||
<materialsCitation id="2399DA25B3CEC22B838C81EC91F7AC86" collectedFrom="Isolated limestone hills" collectingDate="2017-08-13" collectingDateMax="2018-07-07" collectingDateMin="2017-08-13" collectionCode="ZCPRU" collectorName="Benchawan Nahok & Utain Chanlabut" country="Thailand" elevation="42" latitude="12.8389635" location="Khao Nang Panthurat Forest Park" longLatPrecision="1" longitude="99.951904" municipality="Cha-am District" specimenCode="ZCPRU-0043" specimenCount="49" stateProvince="Phetchaburi Province" typeStatus="paratype">
|
||
<specimenCode id="0CCE3A6D75C524257367AFFAE87C5702">
|
||
<abbrev id="ABBRID0EFMAE" xlink_title="Zoological Research Collection of Chaiyaphum Rajabhat University">ZCPRU</abbrev>
|
||
-0043
|
||
</specimenCode>
|
||
<specimenCount id="97D0204F6D1C6736DAA57F43537EAF45">7 shells</specimenCount>
|
||
.
|
||
<collectingCountry id="0883D542D88B176DDD80B2187D9525A2" name="Thailand">Thailand</collectingCountry>
|
||
,
|
||
<collectingRegion id="9966456ED08887B1D1C8CC1105F3A3F3" country="Thailand" name="Phetchaburi">Phetchaburi Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6EE329BCFE309E9ED36E9470B12B249E">Cha-am District</collectingMunicipality>
|
||
,
|
||
<location id="5F3845CCA1EFBDBB56F199B2950AAD92" LSID="urn:lsid:plazi:treatment:03FA6E1C6C5454AAA4E3908EC0E7EBAD:5F3845CCA1EFBDBB56F199B2950AAD92" country="Thailand" latitude="12.8389635" longLatPrecision="1" longitude="99.951904" municipality="Cha-am District" name="Khao Nang Panthurat Forest Park" stateProvince="Phetchaburi Province">Khao Nang Panthurat Forest Park</location>
|
||
,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMMAE" specific-use="{"type":"Point","coordinates":[99.951911,12.838964]}">
|
||
<geoCoordinate id="911FF1D51583BDC5753C0561DA342382" degrees="12" direction="north" minutes="50" orientation="latitude" precision="1" seconds="20.27" value="12.8389635">12 ° 50 ' 20.27 " N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5B60B0A7CE90ED2FC7CD16C4FE2BDF96" degrees="99" direction="east" minutes="57" orientation="longitude" precision="1" seconds="6.88" value="99.951904">99 ° 57 ' 6.88 " E</geoCoordinate>
|
||
</named-content>
|
||
,
|
||
<collectedFrom id="E1279065C14EEFF3ED71DE9E48B5664E">Isolated limestone hills</collectedFrom>
|
||
, alt.
|
||
<specimenCount id="1A4751D314092EADA0013798E5952B33" count="42" type="generic">
|
||
<quantity id="882A35B9BD727EAC67F730DEF344763C" metricMagnitude="1" metricUnit="m" metricValue="4.2" unit="m" value="42.0">42 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
<date id="064999821FF4B337D03A28F285CE77E2" value="2017-08-13">
|
||
<collectingDate id="DB288C4B3A81DAB1AD78A791C43E43F7" value="2017-08-13">13 Aug. 2017</collectingDate>
|
||
</date>
|
||
and
|
||
<date id="7EA5C697DB205E4B581B45B1E6B30525" value="2018-07-07">
|
||
<collectingDate id="5C50571551EECEDD50E297031565E943" value="2018-07-07">7 July 2018</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="0752EDB9B979B95D02490844403B56AB">Benchawan Nahok</collectorName>
|
||
and
|
||
<collectorName id="BC1DB22DE294D424C0055811BDC01362">Utain Chanlabut</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<caption id="AC6CFD2E453FCE2954FFD516A9B55FE8" ID-DOI="10.3897/zookeys.1208.117056.figure2" ID-arpha="BD554122-7BAC-57A4-B6A1-F58B052B6129" httpUri="https://binary.pensoft.net/fig/1108222" startId="F2">
|
||
<paragraph id="9561B6BC5484F7FAE27238AE6FBDEF45">
|
||
<label id="E2DADE4E8C8C2AA7A33E1F51B1842363">Figure 2.</label>
|
||
</paragraph>
|
||
<paragraph id="771EE832B0993C85F397B80BA0812790">
|
||
<taxonomicName id="94F3ED2E9EEB5DCEE8B94591419EF32D" authorityName="Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="bella" status="sp. nov.">
|
||
<emphasis id="F7E21B8E82DFDA501E862AEECA09044E" italics="true">Landouria bella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="DB88518A5E7A95FD7A1AFE2E74802EEB" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="1B4145B8D12CD5360E1D292CF7C6E161" bold="true">A</emphasis>
|
||
holotype (
|
||
<specimenCode id="2AB33A8A1879C6BD44FB44F05F5B8006">
|
||
<named-content id="3530FBEE15571FBA6BE46BCADAD7E536" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahasarakham-university" xlink_title="Natural History Museum, Mahasarakham University">NHMSU</named-content>
|
||
-00056
|
||
</specimenCode>
|
||
)
|
||
<emphasis id="E9CC10BF591FDA97465C24F55B3F5D39" bold="true">B</emphasis>
|
||
paratype (
|
||
<specimenCode id="CADDB48AE5CECAC79B6D9CBC6776B6C8">
|
||
<abbrev id="ABBRID0EMNAE" xlink_title="Zoological Research Collection of Chaiyaphum Rajabhat University">ZCPRU</abbrev>
|
||
-0042
|
||
</specimenCode>
|
||
)
|
||
<emphasis id="FE1532966BED8F356761F01EB7CAFE3E" bold="true">C, D</emphasis>
|
||
empty shell from Khao Nang Panthurat Forest Park, Phetchaburi Province (
|
||
<specimenCode id="26D1A7E74AFD35C0240B4BD1B585C20C">
|
||
<named-content id="198B769204EF9440711F6B76B324DD6F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/mahasarakham-university" xlink_title="Natural History Museum, Mahasarakham University">NHMSU</named-content>
|
||
-00058
|
||
</specimenCode>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2NAE" type="diagnosis">
|
||
<paragraph id="F891D97E037413132F93C205F1F54381">
|
||
<heading id="201A37D892F60B8B0B05FADB91489E9F" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="E4C178A93D756F481271D5548A74DA35">Shell small, depressed-globose. Body whorl rounded; aperture slightly oval, oblique, with thick, slightly reflected lip. Thin, small lamella present on inner columellar side of aperture. Flagellum large, cylindrical, with bent tip; penis short, small, cylindrical; vagina and free oviduct short, swollen. Basal part of gametolytic sac swollen.</paragraph>
|
||
<caption id="2F840F1BAB846FE53F97C0112811F9B7" ID-Table-UUID="2F840F1BAB846FE53F97C0112811F9B7" httpUri="http://table.plazi.org/id/2F840F1BAB846FE53F97C0112811F9B7" startId="T1">
|
||
<paragraph id="0E15E9EF8F9C35421470A73F04134F1E">
|
||
<label id="165D60CB426676A3A0E5E9E7DAE943B5">Table 1.</label>
|
||
</paragraph>
|
||
<paragraph id="762AA19C6CAF7F0E5A3E36AEC1A47696">
|
||
Comparison of shells, genital systems and radulae of Thai
|
||
<taxonomicName id="DDAD936596FCEB73DBF2C7B766E15A03" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="80ED04227782F4CB1B592535A1B28091" italics="true">Landouria</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Reference: 1 =
|
||
<bibRefCitation id="871D2A35E1C925009335BEBEB77CF9E5" DOI="10.26107/rbz-2019-0021" author="Tumpeesuwan C & Tumpeesuwan S" firstAuthor="Tumpeesuwan" journalOrPublisher="The Raffles Bulletin of Zoology" pagination="298-305" refId="B36" refString="Tumpeesuwan C, Tumpeesuwan S (2019) First verified record of the genus Landouria Godwin- Austen, 1918 from Thailand (Gastropoda: Stylommatophora: Camaenidae) with description of a new species. The Raffles Bulletin of Zoology 67: 298–305. https://doi.org/10.26107/rbz-2019-0021" title="First verified record of the genus Landouria Godwin- Austen, 1918 from Thailand (Gastropoda: Stylommatophora: Camaenidae) with description of a new species." volume="67" year="2019">Tumpeesuwan and Tumpeesuwan (2019)</bibRefCitation>
|
||
; 2 =
|
||
<bibRefCitation id="B757E7463DF057753C8FDA832FD29F8B" DOI="10.5852/ejt.2021.767.1495" author="Nahok B & Tumpeesuwan S & Tumpeesuwan C" etAl="et al." firstAuthor="Nahok" journalOrPublisher="European Journal of Taxonomy" pagination="142-166" refId="B16" refString="Nahok B, Tumpeesuwan S, Tumpeesuwan C (2021) Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902). European Journal of Taxonomy 767: 142–166. https://doi.org/10.5852/ejt.2021.767.1495" title="Five new species of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from northeastern Thailand, with note on genitalia and radula morphology of Landouria diplogramma (Möllendorff, 1902)." volume="767" year="2021">Nahok et al. (2021)</bibRefCitation>
|
||
; 3 = this study; 4 =
|
||
<bibRefCitation id="CFCE3B430951FB4C744DD7DDFDB64D67" author="Schileyko AA" firstAuthor="Schileyko" journalOrPublisher="Ruthenica" pagination="1-68" refId="B31" refString="Schileyko AA (2011) Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). Ruthenica 21: 1–68." title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="21" year="2011">Schileyko (2011)</bibRefCitation>
|
||
(question mark = no data).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="0CE788F29C9F9E815AE4B81798882981">
|
||
<table id="TID0EVMBG" rules="all">
|
||
<tbody id="BF087DD866FC59562CB4367C9E454A09">
|
||
<tr id="7E27916F5DCC9618322FFC3A63D35C29">
|
||
<th id="1584A3F702BA2954559BC2A1D353D236" colspan="1" rowspan="1">Characters</th>
|
||
<emphasis id="BDF9D65394BC625130CC64EEDDEFB34B" italics="true">
|
||
<th id="9C401371F8F57FF0A420C2C1113D5B23" colspan="1" rowspan="1">
|
||
<taxonomicName id="B7DB765237CD27ED6AA10005B95BB726" authorityName="Tumpeesuwan & Tumpeesuwan" authorityYear="2019" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="strobiloides">L. strobiloides</taxonomicName>
|
||
</th>
|
||
<th id="5FDC78B13C5C83C2B1BC0ACDE568EAC6" colspan="1" rowspan="1">
|
||
<taxonomicName id="6961082C54E87E1CAE640F6A00008C96" authorityName="Nahok, S. Tumpeesuwan & C. Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="circinata">L. circinata</taxonomicName>
|
||
</th>
|
||
<th id="3C85A4B9B4A2B9BC81130C4129ABA093" colspan="1" rowspan="1">
|
||
<taxonomicName id="DC09940301DCE03943B076E43A0E5B01" authorityName="Nahok, S. Tumpeesuwan & C. Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="tuberculata">L. tuberculata</taxonomicName>
|
||
</th>
|
||
<th id="30870500125AC6598532871EA8551F22" colspan="1" rowspan="1">
|
||
<taxonomicName id="2E3B1D55F85ACCB087D825845145F6C5" authorityName="Nahok, S. Tumpeesuwan & C. Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="trochomorphoides">L. trochomorphoides</taxonomicName>
|
||
</th>
|
||
<th id="166E12936E7F10CD22E76B998D9CF7CC" colspan="1" rowspan="1">
|
||
<taxonomicName id="1B4D1B1A6CD4E74B961C9882EB103B6F" authorityName="Nahok, S. Tumpeesuwan & C. Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="chloritoides">L. chloritoides</taxonomicName>
|
||
</th>
|
||
<th id="4A999B6042261476F9DC2F35A806E3B4" colspan="1" rowspan="1">
|
||
<taxonomicName id="7A30A566B30C8C8BA65B366E67287DF0" authorityName="Nahok, S. Tumpeesuwan & C. Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="elegans">L. elegans</taxonomicName>
|
||
</th>
|
||
<th id="E493647F6E170BF738B41489DE22E8E6" colspan="1" rowspan="1">
|
||
<taxonomicName id="D82F6F7D3D2FB089A75B46A301BA435C" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="diplogramma">L. diplogramma</taxonomicName>
|
||
</th>
|
||
<th id="55A73B6BB65F218D0EC8D171FB84233C" colspan="1" rowspan="1">
|
||
<taxonomicName id="B11A4D4AEF4E93C7D1726E455978A854" authorityName="Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="bella" status="sp. nov.">L. bella</taxonomicName>
|
||
<taxonomicNameLabel id="13CDEE4EA0A4A3404361345DA36E4A2E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
</emphasis>
|
||
<th id="B0784230E62CAD7B2EC6406A1CF84009" colspan="1" rowspan="1">
|
||
<taxonomicName id="A5DA364B7772E696C841394F77C3B839" authorityName="Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="monodon" status="sp. nov.">
|
||
<emphasis id="334F9123CDB4498F7F7B1B7C0B239B81" italics="true">L. monodon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E80D5AF833903FC28D497FF0783EAA1E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</th>
|
||
<taxonomicName id="44D72DA3F2A13A077022009AC27CC804" authority="Shell" authorityName="Shell" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="ptychostyla">
|
||
<th id="FB08333705CC1C949100783FF956CCA9" colspan="1" rowspan="1">
|
||
<emphasis id="F37E73C10E53180141ED86738CFCA84B" italics="true">L. ptychostyla</emphasis>
|
||
</th>
|
||
<tr id="827E6EC8DC073A2A19F6144AE3BA2D62">
|
||
<td id="AF730A691402A550A8E2BF9E0429F3D4" colspan="11" rowspan="1">
|
||
<emphasis id="D82585FC5F0C83ECA3AFA6A14AA5517B" bold="true">Shell</emphasis>
|
||
:
|
||
</td>
|
||
</tr>
|
||
</taxonomicName>
|
||
</tr>
|
||
<tr id="518AA4164F9272A742DF8B01600B58D7">
|
||
<td id="80A87AF3C6F953E7B07D87079BB8A676" colspan="1" rowspan="1">Shell shape</td>
|
||
<td id="579B906AAA511C1748D9BC6C7EA025D6" colspan="1" rowspan="1">Conical-lenticular</td>
|
||
<td id="985D3E98A8EEFC6E61BE8912203DD36A" colspan="1" rowspan="1">Conical-lenticular</td>
|
||
<td id="01C17299A0756828E023F20F6B1143FD" colspan="1" rowspan="1">Slightly convex</td>
|
||
<td id="CDF078F07663CC91889495DC68CA9053" colspan="1" rowspan="1">Subconvex</td>
|
||
<td id="D8EAC6660E8639498308913AFCB2B072" colspan="1" rowspan="1">Depressed</td>
|
||
<td id="BB5300E8C7D234A8C560B81E7920AE5C" colspan="1" rowspan="1">Conical</td>
|
||
<td id="96CD5C8FC69C661AC0518EA63A21574A" colspan="1" rowspan="1">Sub-globose</td>
|
||
<td id="27AD40FB8871E030B479E129F74013BE" colspan="1" rowspan="1">Globose</td>
|
||
<td id="3B073C2E00B8093A128541A494CA9102" colspan="1" rowspan="1">Depressed- conical</td>
|
||
<td id="FB5D8F2D270B801DE349CAB476C17816" colspan="1" rowspan="1">Depressed-conical</td>
|
||
</tr>
|
||
<tr id="C76DB67D759FFD5C04A51A5640618242">
|
||
<td id="F3B803E6E9AA1D56A0DC27D7DFFD9039" colspan="1" rowspan="1">Number of whorls</td>
|
||
<td id="EA78D10B45C7E3125AC8DBBC4447010F" colspan="1" rowspan="1">5 ½</td>
|
||
<td id="10B0CDB2BA20B4E8CDD9CCEE7AF67D18" colspan="1" rowspan="1">6</td>
|
||
<td id="797AE4B27A4B6B27A38DFEDC0E062628" colspan="1" rowspan="1">6 ½</td>
|
||
<td id="CE7E87E11184F2DC4596B99E14921C89" colspan="1" rowspan="1">6</td>
|
||
<td id="7E5ACD55526D850B7D614392D349BB5C" colspan="1" rowspan="1">6 ½</td>
|
||
<td id="9254ED4DF691D28A13656D2BE54C49A8" colspan="1" rowspan="1">6</td>
|
||
<td id="0BF77E44640473EB536BA818DE6BA07B" colspan="1" rowspan="1">5 ½</td>
|
||
<td id="81EB8E283FC8D4D8CE21A0A379C33C83" colspan="1" rowspan="1">6</td>
|
||
<td id="B9973F8F7FB7AC9AD7B288F8CDD72BC4" colspan="1" rowspan="1">5 ¾</td>
|
||
<td id="1764F5D735EFE727F532C44CF9C4241D" colspan="1" rowspan="1">6</td>
|
||
</tr>
|
||
<tr id="DA15671EDA88C305B44372CD9B2868B2">
|
||
<td id="F185445617EBEC082DA41FFBA0F0657A" colspan="1" rowspan="1">Peripheral band</td>
|
||
<td id="13F28E3D3FEC4D299BDC8A7787B0999E" colspan="1" rowspan="1">Dark brown</td>
|
||
<td id="8F08899B6A40683B115D0F857878AD20" colspan="1" rowspan="1">Pale brown</td>
|
||
<td id="9F9BC8D2695FBC4D95CAF4DA469ADE99" colspan="1" rowspan="1">Absent</td>
|
||
<td id="845240A4C965527DE905824B06B1792B" colspan="1" rowspan="1">Brown</td>
|
||
<td id="4E962DA5DD8CAD672A54B1D4A80C59AD" colspan="1" rowspan="1">Absent</td>
|
||
<td id="ACED96B73BB6452F3DE245834A7CA862" colspan="1" rowspan="1">Dark brown</td>
|
||
<td id="86828C011ADB3D1B92976DE74335B415" colspan="1" rowspan="1">Pale brown</td>
|
||
<td id="18C910817C7C2A030A12FF406DD736E5" colspan="1" rowspan="1">Absent</td>
|
||
<td id="55135026F1707E635EE678F300CD1F1C" colspan="1" rowspan="1">Absent</td>
|
||
<td id="F3F9CAA5BD14208ACBF54DFB4E5879B6" colspan="1" rowspan="1">Absent</td>
|
||
</tr>
|
||
<tr id="5CDFD7621234F85CBA0D3979B2A6798A">
|
||
<td id="09EFC3501198418AD7B14B59F73F0116" colspan="1" rowspan="1">Periphery keel</td>
|
||
<td id="867822438F0F9E6A6E4CBE8693ECF06A" colspan="1" rowspan="1">Sharp</td>
|
||
<td id="BFEC9233193BB168B36EFF3CE1BED7C8" colspan="1" rowspan="1">Sharp</td>
|
||
<td id="E6F64DD6BBF32622DB47888A177E7530" colspan="1" rowspan="1">Moderate</td>
|
||
<td id="25C5782840AE7378ECD7EDD923A98B25" colspan="1" rowspan="1">Sharp</td>
|
||
<td id="28B731CD43004A13FCF6C711CE01A395" colspan="1" rowspan="1">Weak</td>
|
||
<td id="02FA47C8E0B5386383E99E9DA27C90FC" colspan="1" rowspan="1">Sharp</td>
|
||
<td id="830B10407332565F5736F2B6A4529252" colspan="1" rowspan="1">Very weak</td>
|
||
<td id="DB851084E8408B8B690106F7EA142765" colspan="1" rowspan="1">Absent</td>
|
||
<td id="7552DDE60C47C17509469F8C247FBB3A" colspan="1" rowspan="1">Slightly</td>
|
||
<td id="5CAF0A2382C504D4C248A28F2F3F57C8" colspan="1" rowspan="1">Moderate</td>
|
||
</tr>
|
||
<tr id="865003E40A0396482333BBBF0A259865">
|
||
<td id="16845723AD868D53CCDD54ADC458CB40" colspan="1" rowspan="1">Scale sculpture</td>
|
||
<td id="9DA4A451FC197E3E45E79F4AB8A997DA" colspan="1" rowspan="1">Absent</td>
|
||
<td id="2BE42B8315A1A76ED9520B2E31CF4A47" colspan="1" rowspan="1">Absent</td>
|
||
<td id="FB34C7B6E611C178885C00C02B297ACD" colspan="1" rowspan="1">Tubercle</td>
|
||
<td id="EA84DD5E1E3448EF309805034319B1D2" colspan="1" rowspan="1">Scaly</td>
|
||
<td id="C22F25C0A6D0E543E4C7240FE162DEF5" colspan="1" rowspan="1">Absent</td>
|
||
<td id="028AAAE0280C6D4FE5C5AD99C80E9156" colspan="1" rowspan="1">Absent</td>
|
||
<td id="8DDA90B7756185259C8CC04DF1C4D450" colspan="1" rowspan="1">Absent</td>
|
||
<td id="C1BAEF635BBE4B1598128700D42142C4" colspan="1" rowspan="1">Absent</td>
|
||
<td id="575A09A215A92C2E8D6F8F190089E1C4" colspan="1" rowspan="1">Absent</td>
|
||
<td id="FE0B6FF072ADF284CDA671FF7C108D6A" colspan="1" rowspan="1">Absent</td>
|
||
</tr>
|
||
<tr id="6BB3C4F0FC4CBFFA392B7F7B977C53A4">
|
||
<td id="159EFC6795C4AF7960D21AD249316BF5" colspan="1" rowspan="1">
|
||
Lamella
|
||
<abbrev id="ABBRID0EQZAE" xlink_title="atrium">at</abbrev>
|
||
inner aperture
|
||
</td>
|
||
<td id="DEB45DC722C073F9DAFF355666F8781E" colspan="1" rowspan="1">Absent</td>
|
||
<td id="E79ECC57B0B2D4D85E455595589F8D31" colspan="1" rowspan="1">Absent</td>
|
||
<td id="3CC188D4ECB73C5B3CECC8E3A058AED0" colspan="1" rowspan="1">Absent</td>
|
||
<td id="D1AB4618DAE5CBFFE04A43CFFF8BD6D9" colspan="1" rowspan="1">Absent</td>
|
||
<td id="3675963D62BD4A2C05A043561BB538BC" colspan="1" rowspan="1">Absent</td>
|
||
<td id="7EF43DF722895C9D0CDAC60E86F42DC9" colspan="1" rowspan="1">Absent</td>
|
||
<td id="67267422CA0A4F2FA0F388486F4967F8" colspan="1" rowspan="1">Absent</td>
|
||
<td id="C1EAEDF530742FCBE65AA69B5D519075" colspan="1" rowspan="1">Present (thin)</td>
|
||
<td id="0B8F8667D27B89CBD7437B6506630579" colspan="1" rowspan="1">Present (sharp)</td>
|
||
<td id="218C09DBFDFABEB44F027C7D3E463B6B" colspan="1" rowspan="1">Present (thin)</td>
|
||
</tr>
|
||
<tr id="58605511BDE7A96CCE8DDB8ED3BC904A">
|
||
<td id="CEA8A0AE54A77CB91B2C8B8A24FC3985" colspan="11" rowspan="1">
|
||
<emphasis id="820C6266A2AAED4ABB3C689E5DD29C86" bold="true">Genital system</emphasis>
|
||
:
|
||
</td>
|
||
</tr>
|
||
<tr id="5BC11388C9D2D176CDBFB63618091D5F">
|
||
<td id="1220E24B93576D3514649BF986C49A2C" colspan="1" rowspan="1">Penis</td>
|
||
<td id="9F26F163EC3242F6A54FCE3186966E0D" colspan="1" rowspan="1">Short, slightly stout</td>
|
||
<td id="E6DC4060CD275D77C6704138D38A4B03" colspan="1" rowspan="1">Long cylindrical</td>
|
||
<td id="113A4BA429BBAEFC1D9228657DBCF2A6" colspan="1" rowspan="1">Swollen</td>
|
||
<td id="98ED983DBA5D9FDA4EE887303E1F1C50" colspan="1" rowspan="1">Long slender</td>
|
||
<td id="D10294D3092FFCD224B1456C6A517E23" colspan="1" rowspan="1">Very large and stout</td>
|
||
<td id="155FF47C9D5538BDCC7A74014C3ABE3D" colspan="1" rowspan="1">Long cylindrical</td>
|
||
<td id="65B5505E4705CE5179336785D3C12F68" colspan="1" rowspan="1">Swollen</td>
|
||
<td id="BFB9DB58772F9E973E47EF162B37D368" colspan="1" rowspan="1">Small cylindrical</td>
|
||
<td id="868BD850C580BD3921EAACAC2DFF3F9D" colspan="1" rowspan="1">Long cylindrical</td>
|
||
<td id="CF7821A696D2745B1EB8A3E097AD1EE9" colspan="1" rowspan="1">Long fusiform</td>
|
||
</tr>
|
||
<tr id="32FFAE97B64FAD2D5D76152F5482C37C">
|
||
<td id="53B944677FE506824ED8CAEFC682F30C" colspan="1" rowspan="1">Verge</td>
|
||
<td id="B743D3DD45CF3FFBB702CB93AB9E47E8" colspan="1" rowspan="1">?</td>
|
||
<td id="B28B0B30C749DD19B356B8A2358BC724" colspan="1" rowspan="1">Present</td>
|
||
<td id="82E39C3EDA1E1A892857ED2B6379EAF8" colspan="1" rowspan="1">Present</td>
|
||
<td id="AF44E0BD37527CD9A9A190789FDA1469" colspan="1" rowspan="1">Absent</td>
|
||
<td id="280B8BD0C70C58355ED1B10A6AF8FAD1" colspan="1" rowspan="1">Present</td>
|
||
<td id="619C2C0EEF1136530CB15939E712335C" colspan="1" rowspan="1">Absent</td>
|
||
<td id="9D75EE528DCBABF0433C416DFF2D0E29" colspan="1" rowspan="1">Present</td>
|
||
<td id="B08AC166151E75B7E3BDE45D45AFD110" colspan="1" rowspan="1">Present</td>
|
||
<td id="3B7307C3C7932EB3BE20DFF53D2640E9" colspan="1" rowspan="1">Present</td>
|
||
<td id="CC6CD2DB2E2C11D9856B350D9CB57CF9" colspan="1" rowspan="1">Absent</td>
|
||
</tr>
|
||
<tr id="27C22F03D3E85A3B9531F95F448F8ED9">
|
||
<td id="2BF0563780D6D8A611B9351B4FB41F06" colspan="1" rowspan="1">Flagellum</td>
|
||
<td id="D32B3BFF6EAAFC40014C0BEADFF97BB7" colspan="1" rowspan="1">Strobilus-shaped</td>
|
||
<td id="EB3B5F8BBD6FF13F6717BDDCC40CF493" colspan="1" rowspan="1">Circinate-shaped</td>
|
||
<td id="C04CE38E8AFA07B4246F0CB6D7F8B395" colspan="1" rowspan="1">Protrusion</td>
|
||
<td id="C3FE09997202AEC621508187A7D77051" colspan="1" rowspan="1">Slender towards tip</td>
|
||
<td id="D1166FE695CEB98BEA408377D4B632E6" colspan="1" rowspan="1">Narrow ovate</td>
|
||
<td id="FD21133BBD11540E5B1A73A9D51348BA" colspan="1" rowspan="1">Water drop-shaped</td>
|
||
<td id="55B261F5B9BCEBF1B41743EF351990D8" colspan="1" rowspan="1">Short protrusion</td>
|
||
<td id="5FF22F3983FF49055968774305109D25" colspan="1" rowspan="1">Cylindrical, bent tip</td>
|
||
<td id="781A95CC1C168805D3674A19EA656B80" colspan="1" rowspan="1">Cylindrical, bent tip</td>
|
||
<td id="FF252BD2C36214A76B3C35ED22C708A1" colspan="1" rowspan="1">Shot, hook- shape</td>
|
||
</tr>
|
||
<tr id="0FB9EBEC7B394B136C56DBB7EC10BA37">
|
||
<td id="4B50A043DA34722903B098A3F6DB8A21" colspan="1" rowspan="1">Basal part of gametolytic sac</td>
|
||
<td id="E067C7017917697CAF7BB65FC937E9C9" colspan="1" rowspan="1">Swollen</td>
|
||
<td id="BB9C9624770394913FDA62AA60C5E342" colspan="1" rowspan="1">Thick and stout</td>
|
||
<td id="6B7BB28C1E627323B5CA0D112C01C4BA" colspan="1" rowspan="1">Thin</td>
|
||
<td id="B62E840F6886EE11B185C4DF633AB654" colspan="1" rowspan="1">Thin</td>
|
||
<td id="5CD3F26E05E550B6A78757DB780AD5EA" colspan="1" rowspan="1">Thick</td>
|
||
<td id="928B66D81CA5B0966F954D1C7F269327" colspan="1" rowspan="1">Stout</td>
|
||
<td id="CA794955B96F73A85027DEEABD75C9AE" colspan="1" rowspan="1">Thick</td>
|
||
<td id="5F50DC0985D8355FBEE52EDED47F0534" colspan="1" rowspan="1">Swollen</td>
|
||
<td id="F21B44DDF626EFE5970A2CA8744A9214" colspan="1" rowspan="1">Swollen</td>
|
||
<td id="C3E0E3A882B18B899415032CBDFBCE54" colspan="1" rowspan="1">Swollen</td>
|
||
</tr>
|
||
<tr id="C1E46590ED684E52AA222459E4E59D69">
|
||
<td id="9A6EBDB93544FAFB92D600AF7CB6A8E9" colspan="11" rowspan="1">
|
||
<emphasis id="9CAABF19000C1FD607116740495052B4" bold="true">Radula</emphasis>
|
||
:
|
||
</td>
|
||
</tr>
|
||
<tr id="FD19723A8A0D532F7EB24B5ABCF74D76">
|
||
<td id="DA3BB5047CECB8D94F562C5BA9541A83" colspan="1" rowspan="1">Number of rows</td>
|
||
<td id="B79F7BEBBA3B2D10CB3FF31D4C666687" colspan="1" rowspan="1">100</td>
|
||
<td id="41BD801D7617B3E69C6872E1F54CC171" colspan="1" rowspan="1">93–95</td>
|
||
<td id="F8DF3C148297A83BAEFAC5D2F8FE83E8" colspan="1" rowspan="1">126–130</td>
|
||
<td id="619B2624DA67E45CB458711C38BE0B10" colspan="1" rowspan="1">107–111</td>
|
||
<td id="15ED23D75ABDF07BFE30AA08A886FF04" colspan="1" rowspan="1">125–129</td>
|
||
<td id="31E3B77F8E58FB73C0FC1494AA3486FF" colspan="1" rowspan="1">110–114</td>
|
||
<td id="A8F2137B31422057EC2A2DC4A94631E1" colspan="1" rowspan="1">121–125</td>
|
||
<td id="D9E72B8B8B8BE9453E320CBE43893466" colspan="1" rowspan="1">103–107</td>
|
||
<td id="85AA1FF93BB3B5D67BFD1C5DC8541076" colspan="1" rowspan="1">136–140</td>
|
||
<td id="102469D0DD65D5EE83C4015C3A5BE900" colspan="1" rowspan="1">?</td>
|
||
</tr>
|
||
<tr id="5EAAEADB057BE5E35E484F4CE25711EF">
|
||
<td id="965828BB6D739840A3153C57EFD5AFEF" colspan="1" rowspan="1">Number of teeth in each row</td>
|
||
<td id="F34AED15D0A675FF6CDB2ECC512C8F9A" colspan="1" rowspan="1">3–53</td>
|
||
<td id="1ED007FCE4DE4CF29BA1E4E7E59ECD99" colspan="1" rowspan="1">55–63</td>
|
||
<td id="24EE72CFAC852CB7C647C10137568853" colspan="1" rowspan="1">79–87</td>
|
||
<td id="0375D450AA612282BF20C99DB5BE3DEE" colspan="1" rowspan="1">65–73</td>
|
||
<td id="C4CF8895E5983D3CE41C7ADD0B74C4F1" colspan="1" rowspan="1">75–83</td>
|
||
<td id="48269010C59A5955D78BD8279F199523" colspan="1" rowspan="1">69–77</td>
|
||
<td id="A19FD48D8DEC63612BC140729436A815" colspan="1" rowspan="1">61–69</td>
|
||
<td id="574E6DC8BFCD082D6114BBD5AB8A5CFC" colspan="1" rowspan="1">63–71</td>
|
||
<td id="820ACB044180D29CFC5E7BB5DB3E744A" colspan="1" rowspan="1">63–71</td>
|
||
<td id="B062D54BA0F273B3A10667EFB86397FC" colspan="1" rowspan="1">?</td>
|
||
</tr>
|
||
<tr id="D0247532A7CFBCECE7C5A1D035A81B1B">
|
||
<td id="1DEEC983910AF23D94B857E2138BE4CC" colspan="1" rowspan="1">- Lateral teeth</td>
|
||
<td id="D4E21DDE2F2712951DFF2B76ADDB5553" colspan="1" rowspan="1">8–10</td>
|
||
<td id="747EB25AE7A118E26A60C2E70FDC0F5F" colspan="1" rowspan="1">8–10</td>
|
||
<td id="9F2F528745DC9A5A52E2EDE6F665955B" colspan="1" rowspan="1">11–13</td>
|
||
<td id="BBAC169FB47B882A1556D07FD8CBC43E" colspan="1" rowspan="1">8–10</td>
|
||
<td id="9D6F751C26BE311F937669EB3AC2041B" colspan="1" rowspan="1">11–13</td>
|
||
<td id="33A287A149074DFEDAB6FA7056092C21" colspan="1" rowspan="1">12–14</td>
|
||
<td id="C2EEEDFAF1943D6A44CD0E190B5A5450" colspan="1" rowspan="1">7–9</td>
|
||
<td id="3ECCBD78361F9453A73651316A007159" colspan="1" rowspan="1">6–8</td>
|
||
<td id="F3B925E1CEBBBF341D00989EE68E880C" colspan="1" rowspan="1">12–14</td>
|
||
<td id="0D173871B481BA50F5BB9B384C36A1AC" colspan="1" rowspan="1">?</td>
|
||
</tr>
|
||
<tr id="5D883337B31C5EFEBD4AFF3DF7F2BEBB">
|
||
<td id="40D7A9F1CCBC04277965899DE94E8CD0" colspan="1" rowspan="1">- Marginal teeth</td>
|
||
<td id="D4A31F639B011668729869A9E8E96BD2" colspan="1" rowspan="1">15–17</td>
|
||
<td id="87BAD0D56AD81528934F22CAD14FCBE5" colspan="1" rowspan="1">19–21</td>
|
||
<td id="03D321D00512500306E6C7614BFEE756" colspan="1" rowspan="1">28–30</td>
|
||
<td id="8553A930DDD2BB67A88D8795F67F55B4" colspan="1" rowspan="1">24–26</td>
|
||
<td id="95D17A9FE99BFDF438EFFA4EE28E02EA" colspan="1" rowspan="1">26–28</td>
|
||
<td id="ACF01DF0E2134173A2578609F68A6EAC" colspan="1" rowspan="1">22–24</td>
|
||
<td id="A98ED387B641DC7D1AE72188F664A05B" colspan="1" rowspan="1">23–25</td>
|
||
<td id="BEFFC903D232C6EE3FFB13995A9DDD44" colspan="1" rowspan="1">22–24</td>
|
||
<td id="F784C94A6DE04608CF18001819AA986A" colspan="1" rowspan="1">19–21</td>
|
||
<td id="9B5C346F08F01A0ABD5FF2A16DA1AB60" colspan="1" rowspan="1">?</td>
|
||
</tr>
|
||
<tr id="931ACBB35FC2828D9694A8547C52DF71">
|
||
<td id="5FBE1027A942ACE62E98B075131E080E" colspan="1" rowspan="1">Central tooth</td>
|
||
<td id="2FFADAFD9D63774EC27F26097DAFE841" colspan="1" rowspan="1">Elongate-lanceolate</td>
|
||
<td id="0EDF5D2D4B5064B4CCA3FF06C94929C1" colspan="1" rowspan="1">Tongue-shaped</td>
|
||
<td id="EED7D01FF00AC95DFCB48B41E3EE6307" colspan="1" rowspan="1">Slender-lanceolate</td>
|
||
<td id="C8E540D9C323921119DBACA2926F9849" colspan="1" rowspan="1">Short-lanceolate</td>
|
||
<td id="31217FC6BEF7169DF62FE2675B38EFDF" colspan="1" rowspan="1">Lanceolate</td>
|
||
<td id="1A9079DC604F05A87A9F7A91450F8552" colspan="1" rowspan="1">Tongue-shaped</td>
|
||
<td id="ABD8888A4DA1BF67FA0DF3BE606E83FE" colspan="1" rowspan="1">Triangular</td>
|
||
<td id="B0833A667ACFF4954F90D6BE688004AF" colspan="1" rowspan="1">Lanceolat</td>
|
||
<td id="36CC9E17496F0ADF0E85DA0F022810C1" colspan="1" rowspan="1">Triangular</td>
|
||
<td id="694A46F4939DDA303E546FAEAB7D4F4F" colspan="1" rowspan="1">?</td>
|
||
</tr>
|
||
<tr id="EAA072B9B8FBCAF7F4FAD7FBB54072D1">
|
||
<td id="8AD957DC63A21098B95225E0C8FF921A" colspan="1" rowspan="1">References</td>
|
||
<td id="E6A84DDCE562B2B9311FFC1434EDF2B3" colspan="1" rowspan="1">1</td>
|
||
<td id="804F281614D6EAE5185C7DF3DFA8F58A" colspan="1" rowspan="1">2</td>
|
||
<td id="2134A607B72B86EC95A496F9082556BE" colspan="1" rowspan="1">2</td>
|
||
<td id="EB2B24191F8245BA57EB30955C2C1FEB" colspan="1" rowspan="1">2</td>
|
||
<td id="1D7C0CA360CEB9A1D3E5A0AF75BCCF31" colspan="1" rowspan="1">2</td>
|
||
<td id="6C47DC851AFE9BCF094C6DC0CDF5579E" colspan="1" rowspan="1">2</td>
|
||
<td id="AD8C4A92B01A374E4379631526308669" colspan="1" rowspan="1">2</td>
|
||
<td id="272755AA35109180205116121FBA61C2" colspan="1" rowspan="1">3</td>
|
||
<td id="7613E52FF94C0B8982B17DFA801E8243" colspan="1" rowspan="1">3</td>
|
||
<td id="61977A1DEF81E7175C3172A8A92AB730" colspan="1" rowspan="1">4</td>
|
||
</tr>
|
||
</tbody>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E1FAG" type="description">
|
||
<paragraph id="10715AE875BABFD74CE509AF98BD3658">
|
||
<heading id="723531FE5CA4ACDAC8F7986764B5B16C" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="49758CD8F92C0446597A6C73B5DB85D8">
|
||
(empty shells = 115, living specimens = 5).
|
||
<emphasis id="E7874BCD0158C6B39F5D5B6FB3908844" bold="true" italics="true">Shell</emphasis>
|
||
(Fig.
|
||
<figureCitation id="0991E4EDFA8CC7C877869C5B52A928DD" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Landouria bella sp. nov. A holotype (NHMSU - 00056) B paratype (ZCPRU - 0042) C, D empty shell from Khao Nang Panthurat Forest Park, Phetchaburi Province (NHMSU - 00058)." figureDoi="10.3897/zookeys.1208.117056.figure2" httpUri="https://binary.pensoft.net/fig/1108222">2</figureCitation>
|
||
) dextral, small, depressed-globose; body whorl large, well rounded. Whorls six; suture deep; apex obtuse; spire much elevated. Protoconch smooth; teleoconch with fine, irregular wrinkles, without scaly processes, last whorl with indistinct, incised spiral lines; umbilical side of last whorl with spiral lines more distinct. Umbilicus deep, very wide. Shell dimensions of specimens from type locality (
|
||
<emphasis id="FBD37DA202797E0C533A7E17DE834842" italics="true">N</emphasis>
|
||
= 104; mean values in parentheses): shell height
|
||
<quantity id="102C72EBBE5DA77AA5F55C432F6015C2" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="9.4" metricValueMin="5.7" unit="mm" value="7.55" valueMax="9.4" valueMin="5.7">5.7–9.4 mm</quantity>
|
||
(6.69 ±
|
||
<quantity id="7096046A82BDB211780B2663691CE941" metricMagnitude="-4" metricUnit="m" metricValue="7.1" unit="mm" value="0.71">0.71 mm</quantity>
|
||
), shell width
|
||
<quantity id="9979A3C415512EEA515672823BDC1529" metricMagnitude="-2" metricUnit="m" metricValue="1.035" metricValueMax="1.22" metricValueMin="0.85" unit="mm" value="10.35" valueMax="12.2" valueMin="8.5">8.5–12.2 mm</quantity>
|
||
(10.54 ±
|
||
<quantity id="4A75A3FD84E0090650D1ED32FB1BF5FB" metricMagnitude="-4" metricUnit="m" metricValue="6.5" unit="mm" value="0.65">0.65 mm</quantity>
|
||
), aperture height
|
||
<quantity id="134D76E85F7CEFCCA7BD42D756516185" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="5.3" metricValueMin="3.2" unit="mm" value="4.25" valueMax="5.3" valueMin="3.2">3.2–5.3 mm</quantity>
|
||
(4.47 ±
|
||
<quantity id="3DE24D51E11D9F76ACD16355D4AC9825" metricMagnitude="-4" metricUnit="m" metricValue="3.8" unit="mm" value="0.38">0.38 mm</quantity>
|
||
), aperture width
|
||
<quantity id="88BBD17F49F5E203E6CBBC67617DE2F9" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.1" metricValueMin="2.9" unit="mm" value="4.0" valueMax="5.1" valueMin="2.9">2.9–5.1 mm</quantity>
|
||
(4.08 ±
|
||
<quantity id="82BB9541FBBFEE34E03773EF71AA8C9B" metricMagnitude="-4" metricUnit="m" metricValue="3.4" unit="mm" value="0.34">0.34 mm</quantity>
|
||
), and umbilicus width
|
||
<quantity id="7F9D007167DB1309C040833E12DCCEC2" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="5.4" metricValueMin="2.7" unit="mm" value="4.05" valueMax="5.4" valueMin="2.7">2.7–5.4 mm</quantity>
|
||
(3.60 ±
|
||
<quantity id="81E980C31FFB96947DCBEF5BD4CDF371" metricMagnitude="-4" metricUnit="m" metricValue="3.3" unit="mm" value="0.33">0.33 mm</quantity>
|
||
). Shell dimensions of specimens from Khao Nang Panthurat (
|
||
<emphasis id="10D627362793BFC81EA2DA1461633032" italics="true">N</emphasis>
|
||
= 11): shell height
|
||
<quantity id="CB8EFEC11D5B9BB365AD210776C648D1" metricMagnitude="-3" metricUnit="m" metricValue="6.449999999999999" metricValueMax="7.09" metricValueMin="5.81" unit="mm" value="6.449999999999999" valueMax="7.09" valueMin="5.81">5.81–7.09 mm</quantity>
|
||
(6.45 ±
|
||
<quantity id="725274BF074EFDAFCF76A72FFAAF7C01" metricMagnitude="-4" metricUnit="m" metricValue="6.4" unit="mm" value="0.64">0.64 mm</quantity>
|
||
), shell width
|
||
<quantity id="10834593D377DDDC6B38284566D7FFB8" metricMagnitude="-2" metricUnit="m" metricValue="1.023" metricValueMax="1.087" metricValueMin="0.959" unit="mm" value="10.23" valueMax="10.87" valueMin="9.59">9.59–10.87 mm</quantity>
|
||
(10.23 ±
|
||
<quantity id="15E31452388F08D9A00CC64D9CBE49DD" metricMagnitude="-4" metricUnit="m" metricValue="6.4" unit="mm" value="0.64">0.64 mm</quantity>
|
||
), aperture height
|
||
<quantity id="F16563A562936099D091ABE01CA3A750" metricMagnitude="-3" metricUnit="m" metricValue="4.43" metricValueMax="4.73" metricValueMin="4.13" unit="mm" value="4.43" valueMax="4.73" valueMin="4.13">4.13–4.73 mm</quantity>
|
||
(4.43 ±
|
||
<quantity id="D1265FBA74733A17E0E7F2F61181C6F8" metricMagnitude="-4" metricUnit="m" metricValue="3.0" unit="mm" value="0.3">0.30 mm</quantity>
|
||
), aperture width
|
||
<quantity id="3A4B53C147E9F5F978211A3662510764" metricMagnitude="-3" metricUnit="m" metricValue="4.01" metricValueMax="4.38" metricValueMin="3.64" unit="mm" value="4.01" valueMax="4.38" valueMin="3.64">3.64–4.38 mm</quantity>
|
||
(4.01 ±
|
||
<quantity id="597C93DFBBCBEB0E21934353B98BF019" metricMagnitude="-4" metricUnit="m" metricValue="3.7" unit="mm" value="0.37">0.37 mm</quantity>
|
||
), and umbilicus width
|
||
<quantity id="8F13784F7763115D3A35B2B1E876DBE0" metricMagnitude="-3" metricUnit="m" metricValue="3.54" metricValueMax="3.83" metricValueMin="3.25" unit="mm" value="3.54" valueMax="3.83" valueMin="3.25">3.25–3.83 mm</quantity>
|
||
(3.54 ±
|
||
<quantity id="02A1449AF3746187A0D4A579C5109A6A" metricMagnitude="-4" metricUnit="m" metricValue="2.9" unit="mm" value="0.29">0.29 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="20C33ACFDE3278275DB0ECF1C56253FC">
|
||
<emphasis id="A46853241B0CA32D2266C8E433112DE1" bold="true" italics="true">Genital system</emphasis>
|
||
(
|
||
<emphasis id="E62E1ECF4357AEFCAEDF6462F5C51CD2" italics="true">N</emphasis>
|
||
= 3) (Fig.
|
||
<figureCitation id="291F01FC2DAFA5FCB0B90FFFE4A24359" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Genital anatomy of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole genitalia B – D internal wall sculpture of B penis (p) C flagellum (fl) D vagina (va)." figureDoi="10.3897/zookeys.1208.117056.figure3" httpUri="https://binary.pensoft.net/fig/1108223">3</figureCitation>
|
||
). Atrium short. Penis small, cylindrical, shorter than flagellum, internally with four smooth, longitudinal pilasters, which gradually become corrugated
|
||
<abbrev id="ABBRID0EYGAG" xlink_title="atrium">at</abbrev>
|
||
distal part close to verge. Verge short, swollen (Fig.
|
||
<figureCitation id="FC385C3C1EC30806B0B3A1A66ABE65E8" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Genital anatomy of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole genitalia B – D internal wall sculpture of B penis (p) C flagellum (fl) D vagina (va)." figureDoi="10.3897/zookeys.1208.117056.figure3" httpUri="https://binary.pensoft.net/fig/1108223">3 B</figureCitation>
|
||
). Penial retractor muscle present. Proximal part of epiphallus (
|
||
<abbrev id="ABBRID0EAHAG" xlink_title="proximal part of epiphallus">ep 1</abbrev>
|
||
) approximately equal to length of penis and vagina combined, but thicker and larger than penis. Distal part of epiphallus (
|
||
<abbrev id="ABBRID0EEHAG" xlink_title="distal part of epiphallus">ep 2</abbrev>
|
||
) extremely short. Flagellum apically joins epiphallus, more than twice as long as penis, regularly long-cylindrical, bent
|
||
<abbrev id="ABBRID0EIHAG" xlink_title="atrium">at</abbrev>
|
||
distal end, internally with three smooth, longitudinal pilasters (Fig.
|
||
<figureCitation id="E8D166D2F49B3941CFC012419170C71E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Genital anatomy of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole genitalia B – D internal wall sculpture of B penis (p) C flagellum (fl) D vagina (va)." figureDoi="10.3897/zookeys.1208.117056.figure3" httpUri="https://binary.pensoft.net/fig/1108223">3 C</figureCitation>
|
||
). Vas deferens a thin, cylindrical tube, laterally entering epiphallus. Vagina short, distally globularly dilated, internally with five thick, longitudinal pilasters (Fig.
|
||
<figureCitation id="BAFBD8DFB6A514B05BC172EC9949DF91" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Genital anatomy of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole genitalia B – D internal wall sculpture of B penis (p) C flagellum (fl) D vagina (va)." figureDoi="10.3897/zookeys.1208.117056.figure3" httpUri="https://binary.pensoft.net/fig/1108223">3 D</figureCitation>
|
||
). Free oviduct shorter than vagina. Proximal part of gametolytic sac very stout, swollen, with a long, narrow, cylindrical tube, and with
|
||
<abbrev id="ABBRID0EUHAG" xlink_title="atrium">at</abbrev>
|
||
its distal end sac-like, small, swollen, and spherical. Prostate gland very long; uterus long, swollen.
|
||
</paragraph>
|
||
<caption id="D9233816096F1CCA6798F4891EB71EF3" ID-DOI="10.3897/zookeys.1208.117056.figure3" ID-arpha="7910B41C-878E-5CD8-827D-83440BFD71B0" httpUri="https://binary.pensoft.net/fig/1108223" startId="F3">
|
||
<paragraph id="1B944E4801A6828169120A1F346D1BFC">
|
||
<label id="CCB27B2D27069D579A87021D75483DEE">Figure 3.</label>
|
||
</paragraph>
|
||
<paragraph id="7EE981A7FB0C03BD014EBB03613E87D5">
|
||
Genital anatomy of
|
||
<taxonomicName id="9A62BCCA18FAE617E7B15F03E5C44D2E" authorityName="Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="bella" status="sp. nov.">
|
||
<emphasis id="490EB0A49D60831A0063239B72C11D8C" italics="true">Landouria bella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5FAC101ED89904269C73FBECC147C57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (
|
||
<specimenCode id="6C1A455167AB8D6972559DBE7E21DABF">
|
||
<abbrev id="ABBRID0ELIAG" xlink_title="Zoological Research Collection of Chaiyaphum Rajabhat University">ZCPRU</abbrev>
|
||
- 0042
|
||
</specimenCode>
|
||
)
|
||
<emphasis id="CD7AD53DE7BD6972F096CBB53E0BF63E" bold="true">A</emphasis>
|
||
whole genitalia
|
||
<emphasis id="DDF552A110649D05DE35866F1A392733" bold="true">B – D</emphasis>
|
||
internal wall sculpture of
|
||
<emphasis id="5DB6034F97F3A6403930B9E837B7A5CF" bold="true">B</emphasis>
|
||
penis (p)
|
||
<emphasis id="36E6C3F92C6FEB90C7C8C87B8C7EC16E" bold="true">C</emphasis>
|
||
flagellum (
|
||
<abbrev id="ABBRID0EXIAG" xlink_title="flagellum">fl</abbrev>
|
||
)
|
||
<emphasis id="BDA67006760200E04E166943192B67D8" bold="true">D</emphasis>
|
||
vagina (
|
||
<abbrev id="ABBRID0E4IAG" xlink_title="vagina">va</abbrev>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="640D9231483534A138189C5FC0E76D68">
|
||
<emphasis id="7B6A89261337C3DE9EF82C7D515E541E" bold="true" italics="true">Radula</emphasis>
|
||
(
|
||
<emphasis id="E5D0156777115D58DE5A4EB68F0AFC35" italics="true">N</emphasis>
|
||
= 3) Comprises 103–107 transverse rows with 63–71 teeth per row; radular formula: (22–24) + (6–8) + 1 + (6–8) + (22–24). Central tooth usually symmetric, unicuspid, lanceolate (Fig.
|
||
<figureCitation id="A5A77EC0FD8E14EDDAB8A5FEE35DA0CF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Radula morphology of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole radula B close-up view of central tooth and lateral teeth C right side of central tooth and lateral teeth D right side of marginal teeth. Numbers indicate order of lateral and marginal teeth. Abbreviation: C = central tooth, L = lateral teeth, M = marginal teeth." figureDoi="10.3897/zookeys.1208.117056.figure4" httpUri="https://binary.pensoft.net/fig/1108224">4 B</figureCitation>
|
||
). Lateral teeth quite like central tooth, but oblique, larger, and longer. Teeth on both sides begin to transform into indistinct bicuspid marginal teeth with tiny ectocone
|
||
<abbrev id="ABBRID0EYJAG" xlink_title="atrium">at</abbrev>
|
||
tooth number 6–8 (Fig.
|
||
<figureCitation id="CFA6986C37E56A71AF3334CE585F5119" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Radula morphology of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole radula B close-up view of central tooth and lateral teeth C right side of central tooth and lateral teeth D right side of marginal teeth. Numbers indicate order of lateral and marginal teeth. Abbreviation: C = central tooth, L = lateral teeth, M = marginal teeth." figureDoi="10.3897/zookeys.1208.117056.figure4" httpUri="https://binary.pensoft.net/fig/1108224">4 C</figureCitation>
|
||
) and gradually change from bicuspid to tricuspid; endocone small; mesocone large, lanceolate; ectocone triangular, with 2–4 tiny cusps (Fig.
|
||
<figureCitation id="89CAA0B2F012F8590E2BFC7FC50827B6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Radula morphology of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole radula B close-up view of central tooth and lateral teeth C right side of central tooth and lateral teeth D right side of marginal teeth. Numbers indicate order of lateral and marginal teeth. Abbreviation: C = central tooth, L = lateral teeth, M = marginal teeth." figureDoi="10.3897/zookeys.1208.117056.figure4" httpUri="https://binary.pensoft.net/fig/1108224">4 D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="A9B313877B234AE8F63C626BDEEE544D" ID-DOI="10.3897/zookeys.1208.117056.figure4" ID-arpha="559CA5AB-8B2D-53A4-B342-243C3A9327DC" httpUri="https://binary.pensoft.net/fig/1108224" startId="F4">
|
||
<paragraph id="33FEB64769B624BE0FE14580B939BA1E">
|
||
<label id="1D296D906FDBB58582CCEA2CECF864F9">Figure 4.</label>
|
||
</paragraph>
|
||
<paragraph id="12354F78FC4959E3DCFAE14D4CFC97EA">
|
||
Radula morphology of
|
||
<taxonomicName id="C82E4002544A17A4ABF4E4306F403DF8" authorityName="Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="bella" status="sp. nov.">
|
||
<emphasis id="968A286AED10BC71B9F6DE152D9F14CB" italics="true">Landouria bella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="190033FB6B2269B79F55F70114994455" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (
|
||
<specimenCode id="CC19CB0B1869DC4920A2DF18093DCB1C">
|
||
<abbrev id="ABBRID0E4KAG" xlink_title="Zoological Research Collection of Chaiyaphum Rajabhat University">ZCPRU</abbrev>
|
||
- 0042
|
||
</specimenCode>
|
||
)
|
||
<emphasis id="C609D061EBBAD1842C77ABDA8983FB95" bold="true">A</emphasis>
|
||
whole radula
|
||
<emphasis id="F75A3442086C1E6D2FA6778B2B7C2AE5" bold="true">B</emphasis>
|
||
close-up view of central tooth and lateral teeth
|
||
<emphasis id="3BBAA26DEF5C18DB7CA409FB8D5872CF" bold="true">C</emphasis>
|
||
right side of central tooth and lateral teeth
|
||
<emphasis id="225B580862D263FC37AE579AF4B788A3" bold="true">D</emphasis>
|
||
right side of marginal teeth. Numbers indicate order of lateral and marginal teeth. Abbreviation: C = central tooth, L = lateral teeth, M = marginal teeth.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EOLAG" type="etymology">
|
||
<paragraph id="7159F3E147A110A191ADA57DAF1D0021">
|
||
<heading id="580873524626CC87FE06F2EB04E04963" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="ED4A5BA70D8ABF38534E10CD5B386862">
|
||
Specific epithet
|
||
<taxonomicName id="84123AFB4EF89A83A2D3D17A4FC0FC0C" authorityName="Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="bella">
|
||
“
|
||
<emphasis id="9F1A485DA26B64F16AB6F32C37E72765" italics="true">bella</emphasis>
|
||
</taxonomicName>
|
||
” is derived from the Latin word “ bellus ”, which means lovely fine, pretty, and beauty. This name refers to the lovely shell of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E5LAG" type="habitat">
|
||
<paragraph id="B3A197FFA1D595D33C5A5673F6007C94">
|
||
<heading id="8CAEE7CF75A13E07610359ADA73704C1" reason="title">Habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="0AA15EEEE1B620467B21257316FC714B">
|
||
This species inhabits isolated limestone hills, where it lives on rocks and the ground in a natural forest
|
||
<abbrev id="ABBRID0EEMAG" xlink_title="atrium">at</abbrev>
|
||
the foot of limestone cliffs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EIMAG" type="distribution">
|
||
<paragraph id="F9DDDE3262A913C2E25D8F96823F5E3A">
|
||
<heading id="1E22C128FC546484BE90FFB44C23725A" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="5BC67DF813D57953B08051603ACECB04">
|
||
<taxonomicName id="E437520CFBAF5EA2611CEFC42B02B8B5" authorityName="Nahok & Chanlabut & Tumpeesuwan & Tumpeesuwan" authorityYear="2024" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="bella" status="sp. nov.">
|
||
<emphasis id="EEDD940AEC2BAB3DD4DA6DE97541B34C" italics="true">Landouria bella</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CC1C071CDEDA4B91ACB2204181101439" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is currently known only from Khao Bin Cave,
|
||
<collectingRegion id="09776520CE20704B35B9E27418477D04" country="Thailand" name="Ratchaburi">Ratchaburi Province</collectingRegion>
|
||
, and Khao Nang Panthurat Forest Park,
|
||
<collectingRegion id="2FEBF44A7FFF37C63601904FE31C971C" country="Thailand" name="Phetchaburi">Phetchaburi Province</collectingRegion>
|
||
, in western
|
||
<collectingCountry id="0E363F42DBE5D442C3FF6605E75CAD8E" name="Thailand">Thailand</collectingCountry>
|
||
(Fig.
|
||
<figureCitation id="1511177200C52352DD67A3702894E8EC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographic distribution of verified Thai Landouria species and showing the type localities of L. bella sp. nov. and L. monodon sp. nov." figureDoi="10.3897/zookeys.1208.117056.figure1" httpUri="https://binary.pensoft.net/fig/1108221">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E4MAG" type="remarks">
|
||
<paragraph id="26CFDA8560D781CB9842F6BC377CB114">
|
||
<heading id="A29EAB3FF7E503684F8221F80FAD6B25" reason="title">Remarks.</heading>
|
||
</paragraph>
|
||
<paragraph id="8A694B8B7F80D40A66446FB1162BAE39">
|
||
The new species is distinguished from other species of
|
||
<taxonomicName id="D529A356334A28503FBE15AB1D27379E" authorityName="Godwin-Austen" authorityYear="1918" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
|
||
<emphasis id="160EFA4BC7C5E755DFB30446EC5C6D54" italics="true">Landouria</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="8BF5A0CF8FF8689FF27DF004B54021F5" name="Thailand">Thailand</collectingCountry>
|
||
by its depressed-globose shell, the presence of a thin, small columellar lamella on the inner side of the aperture, and the absence of an angular and peripheral keel (Fig.
|
||
<figureCitation id="521FEE213188F92A5E84088FADC61C6B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Landouria bella sp. nov. A holotype (NHMSU - 00056) B paratype (ZCPRU - 0042) C, D empty shell from Khao Nang Panthurat Forest Park, Phetchaburi Province (NHMSU - 00058)." figureDoi="10.3897/zookeys.1208.117056.figure2" httpUri="https://binary.pensoft.net/fig/1108222">2</figureCitation>
|
||
). Its genital system is distinguished by its flagellum, which is regularly cylindrical, with a bent tip. The internal structure of the penis has a rounded swollen verge and resembles that of
|
||
<taxonomicName id="EEBA6661D3339FF75F03230BE02BC57C" authorityName="Nahok, S. Tumpeesuwan & C. Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="circinata">
|
||
<emphasis id="E093E887930DD199265706B70BE741A9" italics="true">L. circinata</emphasis>
|
||
</taxonomicName>
|
||
, as well as
|
||
<taxonomicName id="BA6A844D612EBE7F910990BEF6B97143" authorityName="Nahok, S. Tumpeesuwan & C. Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="tuberculata">
|
||
<emphasis id="93EDE84A506BC0EC3829F64296A1ABAE" italics="true">L. tuberculata</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="CD0F33F9A7C8EC9AE26DD447A1709F44" country="Thailand" name="Loei">Loei</collectingRegion>
|
||
,
|
||
<taxonomicName id="B1050FCB8097533A7814DBA0B3136145" authorityName="Nahok, S. Tumpeesuwan & C. Tumpeesuwan" authorityYear="2021" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="chloritoides">
|
||
<emphasis id="A302A93F928A47E19F2160534ACE3BD0" italics="true">L. chloritoides</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="64D187B957E0C5B18657F52B67BBB824" country="Thailand" name="Khon Kaen">Khon Kaen</collectingRegion>
|
||
, and
|
||
<taxonomicName id="E233886C88D5AB0E2489B051DEBC112C" baseAuthorityName="Mollendorff" baseAuthorityYear="1902" class="Gastropoda" family="Camaenidae" genus="Landouria" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="diplogramma">
|
||
<emphasis id="A3BFBB541DB0C9119543B0A6F4A87B43" italics="true">L. diplogramma</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="82588C51B70398650B2F94EAA73B332B" country="Thailand" name="Nakhon Ratchasima">Nakhon Ratchasima</collectingRegion>
|
||
, northeastern
|
||
<collectingCountry id="AD0E5428CCB06BE4FD8A59C6C0B8037E" name="Thailand">Thailand</collectingCountry>
|
||
, but the new species differs in having the proximal part of the penis with thin longitudinal pilasters and the distal part which becomes gradually corrugated (Fig.
|
||
<figureCitation id="CAB78DE5374058C499842CA6ACE957B4" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Genital anatomy of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole genitalia B – D internal wall sculpture of B penis (p) C flagellum (fl) D vagina (va)." figureDoi="10.3897/zookeys.1208.117056.figure3" httpUri="https://binary.pensoft.net/fig/1108223">3 B</figureCitation>
|
||
). Most radular characteristics are quite similar to those of other species, except for the central tooth, which is lanceolate (Fig.
|
||
<figureCitation id="2972B728F5118C49EB2A9E2E635550FA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Radula morphology of Landouria bella sp. nov., paratype (ZCPRU - 0042) A whole radula B close-up view of central tooth and lateral teeth C right side of central tooth and lateral teeth D right side of marginal teeth. Numbers indicate order of lateral and marginal teeth. Abbreviation: C = central tooth, L = lateral teeth, M = marginal teeth." figureDoi="10.3897/zookeys.1208.117056.figure4" httpUri="https://binary.pensoft.net/fig/1108224">4 B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |