1930 lines
465 KiB
XML
1930 lines
465 KiB
XML
<document id="57C42B84AF0B0A5DE395BA41724BBFAC" ID-CLB-Dataset="20710" ID-DOI="10.11646/zootaxa.5099.1.3" ID-GBIF-Dataset="fea6b915-1809-41f9-b4f6-0bceffa50a8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6036906" ID-ZooBank="3BD6F566-9B77-48C6-8538-DD3E9BCA08BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1644487694507" checkinUser="plazi" docAuthor="Rucci, Kevin A., Neuhaus, Birger & Bulnes, Verónica N." docDate="2022" docId="03C1B533FFE776347F8DFC0DFC679672" docLanguage="en" docName="zootaxa.5099.1.3.pdf" docOrigin="Zootaxa 5099 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Echinoderes goku Rucci & Neuhaus & Bulnes 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="82" masterDocId="FFF8CD4BFFE476257F1AFA1FFF849167" masterDocTitle="A new species of Echinoderes (Kinorhyncha: Cyclorhagida: Echinoderidae) from the Argentinean continental shelf with notes on its postembryonic development and on subcuticular morphological characters unreported for Kinorhyncha" masterLastPageNumber="90" masterPageNumber="65" pageNumber="68" updateTime="1699308980906" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="803B5CDB8EDB24E3575B0B0A38BC1FDF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="560B815E9EAC97D15C819B0C34B7FDB7">A new species of Echinoderes (Kinorhyncha: Cyclorhagida: Echinoderidae) from the Argentinean continental shelf with notes on its postembryonic development and on subcuticular morphological characters unreported for Kinorhyncha</mods:title>
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<mods:name id="8A039A74B2D6159C041DF820A787791D" type="personal">
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<mods:roleTerm id="10357F4723B8D7F994ACB1AF16DD0B77">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A1FB2ECD470AD367968A513F193173A0">Rucci, Kevin A.</mods:namePart>
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<mods:affiliation id="6B70FA4FF3F90CFA02B08F94F99E9B00">Departamento de Biología, Bioquímica y Farmacia, Universidad Nacional del Sur, San Juan 670, B 8000 ICN Bahía Blanca, Buenos Aires, Argentina, & current address: Laboratorio de Arbovirus - Instituto de Virología “ Dr. J. M. Vanella ” (InViV), Universidad Nacional de Córdoba, Enfermera Gordillo Gómez s / n, 5000 Córdoba, Córdoba, Argentina.</mods:affiliation>
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</mods:name>
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<mods:name id="B90E21F224B73D654AA164F911ACD4D8" type="personal">
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<mods:roleTerm id="9BAE1F551D1F408DDCADEB39BB505B2B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EE4BEDC09340ABE350ED4F36BA58A632">Neuhaus, Birger</mods:namePart>
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<mods:affiliation id="C44B7A9D8EB044C08EB89709A6795F74">Museum für Naturkunde Berlin-Leibniz Institute for Evolution and Biodiversity Science, Invalidenstr. 43, D- 10115 Berlin, Germany. INBIOSUR, CONICET, San Juan 671, B 8000 ICN Bahía Blanca, Buenos Aires, Argentina.</mods:affiliation>
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<mods:name id="D39E94D621D1AE0C1C1337BBB5F7160E" type="personal">
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<mods:roleTerm id="4243DD5BCDAB7525898D1885A70C5B50">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="07D8BE60328FA7230430BD590A07CBEE">Bulnes, Verónica N.</mods:namePart>
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<mods:title id="177CEEEE2A673FBD884D09DD8589BCFA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="45A1EC5AFFAF00BF4FC6EE9450381B53">
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<mods:date id="2D157F4BD6D81FF84DDF661EE7BDAB60">2022</mods:date>
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<mods:number id="B62B1C232FD2ED6022733B5A65993F3C">2022-02-10</mods:number>
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<mods:detail id="12BABEE7CFD02F7CF7D14D405725CF27" type="volume">
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<mods:number id="D00DDA007359686F725D2E0A9D5EE4E2">5099</mods:number>
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<treatment id="03C1B533FFE776347F8DFC0DFC679672" ID-DOI="http://doi.org/10.5281/zenodo.6309107" ID-GBIF-Taxon="193331530" ID-Zenodo-Dep="6309107" LSID="urn:lsid:plazi:treatment:03C1B533FFE776347F8DFC0DFC679672" httpUri="http://treatment.plazi.org/id/03C1B533FFE776347F8DFC0DFC679672" lastPageId="17" lastPageNumber="82" pageId="3" pageNumber="68">
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<subSubSection id="C37257AEFFE776267F8DFC0DFB079736" pageId="3" pageNumber="68" type="nomenclature">
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<paragraph id="8BD70425FFE776267F8DFC0DFE53974A" blockId="3.[151,1155,1554,1617]" box="[151,471,1554,1581]" pageId="3" pageNumber="68">
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<heading id="D09FB349FFE776267F8DFC0DFE53974A" bold="true" box="[151,471,1554,1581]" fontSize="11" level="1" pageId="3" pageNumber="68" reason="1">
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<emphasis id="B91CD837FFE776267F8DFC0DFE53974A" bold="true" box="[151,471,1554,1581]" pageId="3" pageNumber="68">
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<taxonomicName id="4C687FA6FFE776267F8DFC0DFEF6974A" ID-CoL="8YJD6" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[151,370,1554,1581]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="3" pageNumber="68" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
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<emphasis id="B91CD837FFE776267F8DFC0DFEF6974A" bold="true" box="[151,370,1554,1581]" italics="true" pageId="3" pageNumber="68">Echinoderes goku</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22F654CFFE776267E60FC0CFE53974A" box="[378,471,1555,1581]" pageId="3" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD70425FFE776267F8DFC29FB079736" blockId="3.[151,1155,1554,1617]" box="[151,1155,1590,1617]" pageId="3" pageNumber="68">
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http://zoobank.org/
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urn:lsid:zoobank.org:pub:
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<uuid id="FFCE3EF0FFE776267D90FC29FB079736" box="[650,1155,1590,1617]" pageId="3" pageNumber="68">93EF677E-1755-41C7-8906-952614E71018</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37257AEFFE776267F8DFC61FD2396F2" pageId="3" pageNumber="68" type="diagnosis">
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<paragraph id="8BD70425FFE776267F8DFC61FD2396F2" blockId="3.[151,1437,1662,2013]" pageId="3" pageNumber="68">
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<emphasis id="B91CD837FFE776267F8DFC61FE9697FF" bold="true" box="[151,274,1662,1688]" pageId="3" pageNumber="68">Diagnosis.</emphasis>
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<taxonomicName id="4C687FA6FFE776267E06FC60FE2397FE" authorityName="Claparede" authorityYear="1863" box="[284,423,1663,1689]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="3" pageNumber="68" phylum="Kinorhyncha" rank="genus">
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<emphasis id="B91CD837FFE776267E06FC60FE2397FE" box="[284,423,1663,1689]" italics="true" pageId="3" pageNumber="68">Echinoderes</emphasis>
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</taxonomicName>
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with middorsal spine on segments 4, 6, and 8, and spine in lateroventral positions on segments 6–9 and 11 (lateral terminal spines). Tube present in subdorsal position on segment 2, sublateral to midlateral position on segments 7 and 9, lateral accessory position on segment 8, lateroventral position on segment 5 and ventrolateral position on segment 2. Type-2 gland cell outlets, female-specific papillae or subcuticular funnel-like structures not present. Male with a sublateral tube on segment 10 with a basal cylindrical reinforcement structure and three pairs of penile spines on segment 11. Female with sublateral fringe on surface connected with a subcuticular conical structure surrounded basally by a cylindrical reinforcement structure in segment 10 and with a lateral terminal accessory spine on segment 11.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37257AEFFE776267FDDFD80FBD296BA" pageId="3" pageNumber="68" type="etymology">
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<paragraph id="8BD70425FFE776267FDDFD80FBD296BA" blockId="3.[151,1437,1662,2013]" pageId="3" pageNumber="68">
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<emphasis id="B91CD837FFE776267FDDFD80FECA96DE" bold="true" box="[199,334,1951,1977]" pageId="3" pageNumber="68">Etymology.</emphasis>
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The specific epithet is a noun in apposition and alludes to the adoptive name of the Saiyan Son Gokū, the major character from the Dragon Ball manga, authored by Akira Toriyama.
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</paragraph>
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</subSubSection>
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||
<caption id="DF1754ADFFE076217F8DFC59FD439618" ID-DOI="http://doi.org/10.5281/zenodo.6036917" ID-Zenodo-Dep="6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="4" pageNumber="69" startId="4.[151,250,1606,1630]" targetBox="[151,1436,198,1582]" targetPageId="4">
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<paragraph id="8BD70425FFE076217F8DFC59FD439618" blockId="4.[151,1437,1606,1919]" pageId="4" pageNumber="69">
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<emphasis id="B91CD837FFE076217F8DFC59FE899739" bold="true" box="[151,269,1606,1630]" pageId="4" pageNumber="69">FIGURE 2</emphasis>
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. Line art illustrations of
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<taxonomicName id="4C687FA6FFE076217D1DFC59FD3F9739" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[519,699,1606,1630]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="4" pageNumber="69" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
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<emphasis id="B91CD837FFE076217D1DFC59FD3F9739" box="[519,699,1606,1630]" italics="true" pageId="4" pageNumber="69">Echinoderes goku</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B91CD837FFE076217DD8FC59FC969739" bold="true" box="[706,786,1606,1630]" pageId="4" pageNumber="69">
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<taxonomicNameLabel id="A22F654CFFE076217DD8FC59FC969739" box="[706,786,1606,1630]" pageId="4" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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,
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<emphasis id="B91CD837FFE076217C05FC59FCB59739" bold="true" box="[799,817,1606,1630]" pageId="4" pageNumber="69">A</emphasis>
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,
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<emphasis id="B91CD837FFE076217C27FC59FCCA9739" bold="true" box="[829,846,1606,1630]" pageId="4" pageNumber="69">B</emphasis>
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, male holotype (MACN-In 43755),
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<emphasis id="B91CD837FFE076217BA7FC59FB4B9739" bold="true" box="[1213,1231,1606,1630]" pageId="4" pageNumber="69">C</emphasis>
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,
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<emphasis id="B91CD837FFE076217BC1FC59FB699739" bold="true" box="[1243,1261,1606,1630]" pageId="4" pageNumber="69">D</emphasis>
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, female paratype (MACN-In 43756b).
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<emphasis id="B91CD837FFE076217E6CFC75FE0C97E5" bold="true" box="[374,392,1642,1666]" pageId="4" pageNumber="69">A</emphasis>
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. Dorsal view.
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<emphasis id="B91CD837FFE076217D38FC75FDB797E5" bold="true" box="[546,563,1642,1666]" pageId="4" pageNumber="69">B</emphasis>
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. Ventral view.
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<emphasis id="B91CD837FFE076217DC8FC75FD6097E5" bold="true" box="[722,740,1642,1666]" pageId="4" pageNumber="69">C</emphasis>
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,
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<emphasis id="B91CD837FFE076217DEEFC75FC8297E5" bold="true" box="[756,774,1642,1666]" pageId="4" pageNumber="69">D</emphasis>
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. Segments 10 and 11, dorsal (
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<emphasis id="B91CD837FFE076217B5DFC75FBDD97E5" bold="true" box="[1095,1113,1642,1666]" pageId="4" pageNumber="69">C</emphasis>
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) and ventral view (
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<emphasis id="B91CD837FFE076217A33FC75FABF97E5" bold="true" box="[1321,1339,1642,1666]" pageId="4" pageNumber="69">D</emphasis>
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). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm.
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</paragraph>
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</caption>
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<caption id="DF1754ADFFE176207F8DFC91FEA096C5" ID-DOI="http://doi.org/10.5281/zenodo.6036925" ID-Zenodo-Dep="6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="5" pageNumber="70" startId="5.[151,250,1678,1702]" targetBox="[153,1436,182,1654]" targetPageId="5">
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<paragraph id="8BD70425FFE176207F8DFC91FEA096C5" blockId="5.[151,1437,1678,1955]" pageId="5" pageNumber="70">
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<emphasis id="B91CD837FFE176207F8DFC91FE9097C1" bold="true" box="[151,276,1678,1702]" pageId="5" pageNumber="70">FIGURE 3.</emphasis>
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<taxonomicName id="4C687FA6FFE176207E07FC91FE5797C1" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[285,467,1678,1702]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="5" pageNumber="70" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
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<emphasis id="B91CD837FFE176207E07FC91FE5797C1" box="[285,467,1678,1702]" italics="true" pageId="5" pageNumber="70">Echinoderes goku</emphasis>
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</taxonomicName>
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||
<emphasis id="B91CD837FFE176207EC1FC90FDA997C0" bold="true" box="[475,557,1679,1703]" pageId="5" pageNumber="70">
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||
<taxonomicNameLabel id="A22F654CFFE176207EC1FC90FDA997C0" box="[475,557,1679,1703]" pageId="5" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, male paratype ZMB 12334 (
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<emphasis id="B91CD837FFE176207C79FC91FCF197C1" bold="true" box="[867,885,1678,1702]" pageId="5" pageNumber="70">A</emphasis>
|
||
,
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||
<emphasis id="B91CD837FFE176207C99FC91FC1197C1" bold="true" box="[899,917,1678,1702]" pageId="5" pageNumber="70">C</emphasis>
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), juvenile non-type ZMB 12347 (
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<emphasis id="B91CD837FFE176207BECFC90FA8397C0" bold="true" box="[1270,1287,1679,1703]" pageId="5" pageNumber="70">B</emphasis>
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,
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<emphasis id="B91CD837FFE176207A0CFC90FAAC97C0" bold="true" box="[1302,1320,1679,1703]" pageId="5" pageNumber="70">D</emphasis>
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), and male holotype MACN-In 43755 (
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<emphasis id="B91CD837FFE176207EA3FCACFE6097AC" bold="true" box="[441,484,1715,1739]" pageId="5" pageNumber="70">E–J</emphasis>
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), in ventral (
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<emphasis id="B91CD837FFE176207D77FCADFDFB97AD" bold="true" box="[621,639,1714,1738]" pageId="5" pageNumber="70">A</emphasis>
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), lateral (
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<emphasis id="B91CD837FFE176207DFCFCACFD7397AC" bold="true" box="[742,759,1715,1739]" pageId="5" pageNumber="70">B</emphasis>
|
||
,
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<emphasis id="B91CD837FFE176207C1FFCACFC9397AC" bold="true" box="[773,791,1715,1739]" pageId="5" pageNumber="70">D</emphasis>
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), and dorsal view (
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<emphasis id="B91CD837FFE176207CF9FCADFC7197AD" bold="true" box="[995,1013,1714,1738]" pageId="5" pageNumber="70">C</emphasis>
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,
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<emphasis id="B91CD837FFE176207B1EFCACFBAA97AC" bold="true" box="[1028,1070,1715,1739]" pageId="5" pageNumber="70">E–J</emphasis>
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), DIC.
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<emphasis id="B91CD837FFE176207B65FCADFB3497AC" bold="true" box="[1151,1200,1714,1739]" pageId="5" pageNumber="70">A–D</emphasis>
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. Habitus of specimens infected with fungus before (
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<emphasis id="B91CD837FFE176207ED6FCC9FE5A9789" bold="true" box="[460,478,1750,1774]" pageId="5" pageNumber="70">A</emphasis>
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,
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<emphasis id="B91CD837FFE176207EF5FCC8FD849788" bold="true" box="[495,512,1751,1775]" pageId="5" pageNumber="70">B</emphasis>
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) and after treatment with formaldehyde for two weeks (
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<emphasis id="B91CD837FFE176207B40FCC9FBE89789" bold="true" box="[1114,1132,1750,1774]" pageId="5" pageNumber="70">C</emphasis>
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,
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<emphasis id="B91CD837FFE176207B66FCC8FB0A9788" bold="true" box="[1148,1166,1751,1775]" pageId="5" pageNumber="70">D</emphasis>
|
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).
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<emphasis id="B91CD837FFE176207BB2FCC8FB3D9788" bold="true" box="[1192,1209,1751,1775]" pageId="5" pageNumber="70">E</emphasis>
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. Segments 1–5.
|
||
<emphasis id="B91CD837FFE176207A76FCC8FA129788" bold="true" box="[1388,1430,1751,1775]" pageId="5" pageNumber="70">F, J</emphasis>
|
||
. Segments 6–9 at different focal levels.
|
||
<emphasis id="B91CD837FFE176207D3AFCE5FDD59675" bold="true" box="[544,593,1786,1810]" pageId="5" pageNumber="70">G–I.</emphasis>
|
||
Segments 10 (
|
||
<emphasis id="B91CD837FFE176207DFDFCE5FD7F9675" bold="true" box="[743,763,1786,1810]" pageId="5" pageNumber="70">G</emphasis>
|
||
), 10 and 11 (
|
||
<emphasis id="B91CD837FFE176207C9AFCE4FC109674" bold="true" box="[896,916,1787,1811]" pageId="5" pageNumber="70">H</emphasis>
|
||
), and 11 (
|
||
<emphasis id="B91CD837FFE176207CE0FCE4FB809674" bold="true" box="[1018,1028,1787,1811]" pageId="5" pageNumber="70">I</emphasis>
|
||
) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in
|
||
<emphasis id="B91CD837FFE176207CBCFD78FC3C9618" bold="true" box="[934,952,1895,1919]" pageId="5" pageNumber="70">D</emphasis>
|
||
100 μm, valid for
|
||
<emphasis id="B91CD837FFE176207B61FD79FB289618" bold="true" box="[1147,1196,1894,1919]" pageId="5" pageNumber="70">A–D</emphasis>
|
||
. Scale bar in
|
||
<emphasis id="B91CD837FFE176207A26FD78FACF9618" bold="true" box="[1340,1355,1895,1919]" pageId="5" pageNumber="70">F</emphasis>
|
||
50 µm, valid for
|
||
<emphasis id="B91CD837FFE176207FEEFD94FE9B96C4" bold="true" box="[244,287,1931,1955]" pageId="5" pageNumber="70">E–J</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1754ADFFE276237F8DFC71FE51968A" ID-DOI="http://doi.org/10.5281/zenodo.6036933" ID-Zenodo-Dep="6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="6" pageNumber="71" startId="6.[151,250,1646,1670]" targetBox="[166,1421,181,1622]" targetPageId="6">
|
||
<paragraph id="8BD70425FFE276237F8DFC71FE51968A" blockId="6.[151,1437,1646,2030]" pageId="6" pageNumber="71">
|
||
<emphasis id="B91CD837FFE276237F8DFC71FE9297E1" bold="true" box="[151,278,1646,1670]" pageId="6" pageNumber="71">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE276237E05FC71FE5297E1" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[287,470,1646,1670]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="6" pageNumber="71" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFE276237E05FC71FE5297E1" box="[287,470,1646,1670]" italics="true" pageId="6" pageNumber="71">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CD837FFE276237EC5FC71FDB697E1" bold="true" box="[479,562,1646,1670]" pageId="6" pageNumber="71">
|
||
<taxonomicNameLabel id="A22F654CFFE276237EC5FC71FDB697E1" box="[479,562,1646,1670]" pageId="6" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype MACN-In 43755 (
|
||
<emphasis id="B91CD837FFE276237CBBFC71FC5697E1" bold="true" box="[929,978,1646,1670]" pageId="6" pageNumber="71">A–C</emphasis>
|
||
), male paratype ZMB 12318 (
|
||
<emphasis id="B91CD837FFE276237A0FFC71FAA397E1" bold="true" box="[1301,1319,1646,1670]" pageId="6" pageNumber="71">D</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237A2CFC71FAE397E1" bold="true" box="[1334,1383,1646,1670]" pageId="6" pageNumber="71">F–H</emphasis>
|
||
) and female paratypes ZMB 12337 (
|
||
<emphasis id="B91CD837FFE276237EF9FC8DFE7097CD" bold="true" box="[483,500,1682,1706]" pageId="6" pageNumber="71">E</emphasis>
|
||
), ZMB 12265 (
|
||
<emphasis id="B91CD837FFE276237D84FC8DFD2C97CD" bold="true" box="[670,680,1682,1706]" pageId="6" pageNumber="71">I</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237DA2FC8DFD4897CD" bold="true" box="[696,716,1682,1706]" pageId="6" pageNumber="71">K</emphasis>
|
||
), and MACN-In 43756b (
|
||
<emphasis id="B91CD837FFE276237CFEFC8DFC7597CD" bold="true" box="[996,1009,1682,1706]" pageId="6" pageNumber="71">J</emphasis>
|
||
), ventral view (
|
||
<emphasis id="B91CD837FFE276237B87FC8DFB4997CD" bold="true" box="[1181,1229,1682,1706]" pageId="6" pageNumber="71">A–E</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237BC7FC8DFB6397CD" bold="true" box="[1245,1255,1682,1706]" pageId="6" pageNumber="71">I</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237BEDFC8DFA8F97CD" bold="true" box="[1271,1291,1682,1706]" pageId="6" pageNumber="71">K</emphasis>
|
||
), dorsal view (
|
||
<emphasis id="B91CD837FFE276237FBAFCA9FF5497A9" bold="true" box="[160,208,1718,1742]" pageId="6" pageNumber="71">F–H</emphasis>
|
||
), and optical section (
|
||
<emphasis id="B91CD837FFE276237EAEFCA9FE4597A9" bold="true" box="[436,449,1718,1742]" pageId="6" pageNumber="71">J</emphasis>
|
||
), extended focus image from four images (
|
||
<emphasis id="B91CD837FFE276237C67FCA9FC0B97A9" bold="true" box="[893,911,1718,1742]" pageId="6" pageNumber="71">D</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237C87FCA9FC3597A9" bold="true" box="[925,945,1718,1742]" pageId="6" pageNumber="71">K</emphasis>
|
||
), DIC.
|
||
<emphasis id="B91CD837FFE276237CE5FCA9FB9C97A9" bold="true" box="[1023,1048,1718,1742]" pageId="6" pageNumber="71">A.</emphasis>
|
||
Segments 1–6.
|
||
<emphasis id="B91CD837FFE276237BA6FCA9FB4997A9" bold="true" box="[1212,1229,1718,1742]" pageId="6" pageNumber="71">B</emphasis>
|
||
. Segments 7–10.
|
||
<emphasis id="B91CD837FFE276237A9EFCA9FA1897A9" bold="true" box="[1412,1436,1718,1742]" pageId="6" pageNumber="71">C.</emphasis>
|
||
Segments 9–11.
|
||
<emphasis id="B91CD837FFE276237E25FCC5FEDC9795" bold="true" box="[319,344,1754,1778]" pageId="6" pageNumber="71">D.</emphasis>
|
||
Segments 1–4.
|
||
<emphasis id="B91CD837FFE276237EE1FCC5FD979795" bold="true" box="[507,531,1754,1778]" pageId="6" pageNumber="71">E.</emphasis>
|
||
Segment 10.
|
||
<emphasis id="B91CD837FFE276237D85FCC5FD549795" bold="true" box="[671,720,1754,1778]" pageId="6" pageNumber="71">F–H</emphasis>
|
||
. Segments 10 and 11 at different focal levels.
|
||
<emphasis id="B91CD837FFE276237BB3FCC5FB379795" bold="true" box="[1193,1203,1754,1778]" pageId="6" pageNumber="71">I</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237BDAFCC5FB579795" bold="true" box="[1216,1235,1754,1778]" pageId="6" pageNumber="71">J.</emphasis>
|
||
Segments 8
|
||
<emphasis id="B91CD837FFE276237A4AFCC5FAD99795" bold="true" box="[1360,1373,1754,1778]" pageId="6" pageNumber="71">–</emphasis>
|
||
11.
|
||
<emphasis id="B91CD837FFE276237A98FCC5FA189795" bold="true" box="[1410,1436,1754,1778]" pageId="6" pageNumber="71">K.</emphasis>
|
||
Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in
|
||
<emphasis id="B91CD837FFE276237ED5FDADFE6496AD" bold="true" box="[463,480,1970,1994]" pageId="6" pageNumber="71">B</emphasis>
|
||
50 μm, valid for
|
||
<emphasis id="B91CD837FFE276237D8BFDADFD4696AD" bold="true" box="[657,706,1970,1994]" pageId="6" pageNumber="71">A–D</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237DD5FDADFD5D96AD" bold="true" box="[719,729,1970,1994]" pageId="6" pageNumber="71">I</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237DFFFDADFD7696AD" bold="true" box="[741,754,1970,1994]" pageId="6" pageNumber="71">J</emphasis>
|
||
. Scale bar in
|
||
<emphasis id="B91CD837FFE276237C67FDADFC1596AD" bold="true" box="[893,913,1970,1994]" pageId="6" pageNumber="71">G</emphasis>
|
||
50 µm, valid for
|
||
<emphasis id="B91CD837FFE276237B58FDADFBD596AD" bold="true" box="[1090,1105,1970,1994]" pageId="6" pageNumber="71">F</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237B44FDADFBF696AD" bold="true" box="[1118,1138,1970,1994]" pageId="6" pageNumber="71">G</emphasis>
|
||
. Scale bar in
|
||
<emphasis id="B91CD837FFE276237BE7FDADFA9596AD" bold="true" box="[1277,1297,1970,1994]" pageId="6" pageNumber="71">H</emphasis>
|
||
20 µm. Scale bar in
|
||
<emphasis id="B91CD837FFE276237FC2FDC9FF689689" bold="true" box="[216,236,2006,2030]" pageId="6" pageNumber="71">K</emphasis>
|
||
30 µm, valid for
|
||
<emphasis id="B91CD837FFE276237E87FDC9FE2A9689" bold="true" box="[413,430,2006,2030]" pageId="6" pageNumber="71">E</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE276237EA1FDC9FE4B9689" bold="true" box="[443,463,2006,2030]" pageId="6" pageNumber="71">K</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1754ADFFE376227F8DFC25FBB996B9" ID-DOI="http://doi.org/10.5281/zenodo.6036938" ID-Zenodo-Dep="6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="7" pageNumber="72" startId="7.[151,250,1594,1618]" targetBox="[151,1355,151,1575]" targetPageId="7">
|
||
<paragraph id="8BD70425FFE376227F8DFC25FBB996B9" blockId="7.[151,1437,1594,2014]" pageId="7" pageNumber="72">
|
||
<emphasis id="B91CD837FFE376227F8DFC25FE929735" bold="true" box="[151,278,1594,1618]" pageId="7" pageNumber="72">FIGURE 5.</emphasis>
|
||
Female specimens of species of
|
||
<taxonomicName id="4C687FA6FFE376227D6EFC25FD769735" authorityName="Claparede" authorityYear="1863" box="[628,754,1594,1618]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="72" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="B91CD837FFE376227D6EFC25FD769735" box="[628,754,1594,1618]" italics="true" pageId="7" pageNumber="72">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
, dorsal view (
|
||
<emphasis id="B91CD837FFE376227C91FC25FC199735" bold="true" box="[907,925,1594,1618]" pageId="7" pageNumber="72">A</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE376227CB7FC25FBBD9735" bold="true" box="[941,1081,1594,1618]" pageId="7" pageNumber="72">B, H, I, L–N</emphasis>
|
||
), lateral view (
|
||
<emphasis id="B91CD837FFE376227BC6FC25FA969735" bold="true" box="[1244,1298,1594,1618]" pageId="7" pageNumber="72">C, K</emphasis>
|
||
), and optical section (
|
||
<emphasis id="B91CD837FFE376227FF7FC41FEA49711" bold="true" box="[237,288,1630,1654]" pageId="7" pageNumber="72">D–G</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE376227E31FC41FEBC9711" bold="true" box="[299,312,1630,1654]" pageId="7" pageNumber="72">J</emphasis>
|
||
), DIC.
|
||
<emphasis id="B91CD837FFE376227E99FC41FE3E9711" bold="true" box="[387,442,1630,1654]" pageId="7" pageNumber="72">A–E.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE376227EA5FC41FDF79711" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[447,627,1630,1654]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="72" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFE376227EA5FC41FDF79711" box="[447,627,1630,1654]" italics="true" pageId="7" pageNumber="72">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CD837FFE376227D62FC41FD5B9711" bold="true" box="[632,735,1630,1654]" pageId="7" pageNumber="72">
|
||
<taxonomicNameLabel id="A22F654CFFE376227D62FC41FD4C9711" box="[632,712,1630,1654]" pageId="7" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE376227DF1FC41FC869711" bold="true" box="[747,770,1630,1654]" pageId="7" pageNumber="72">B.</emphasis>
|
||
paratype ZMB 12335, segments 1–5 (
|
||
<emphasis id="B91CD837FFE376227B99FC41FB119711" bold="true" box="[1155,1173,1630,1654]" pageId="7" pageNumber="72">A</emphasis>
|
||
) and 6–9 (
|
||
<emphasis id="B91CD837FFE376227A1BFC41FA969711" bold="true" box="[1281,1298,1630,1654]" pageId="7" pageNumber="72">B</emphasis>
|
||
).
|
||
<emphasis id="B91CD837FFE376227A3CFC41FABA9711" bold="true" box="[1318,1342,1630,1654]" pageId="7" pageNumber="72">C.</emphasis>
|
||
Paratype ZMB 12344, segments 2–7.
|
||
<emphasis id="B91CD837FFE376227EA1FC9DFE7697FD" bold="true" box="[443,498,1666,1690]" pageId="7" pageNumber="72">D, E.</emphasis>
|
||
Paratypes ZMB 12342 (
|
||
<emphasis id="B91CD837FFE376227DF5FC9DFC8597FD" bold="true" box="[751,769,1666,1690]" pageId="7" pageNumber="72">D</emphasis>
|
||
) and ZMB 12264 (
|
||
<emphasis id="B91CD837FFE376227CD0FC9DFC5F97FD" bold="true" box="[970,987,1666,1690]" pageId="7" pageNumber="72">E</emphasis>
|
||
), segments 10 and 11 (
|
||
<emphasis id="B91CD837FFE376227BD0FC9DFB5897FD" bold="true" box="[1226,1244,1666,1690]" pageId="7" pageNumber="72">D</emphasis>
|
||
) and 10 (
|
||
<emphasis id="B91CD837FFE376227A5BFC9DFAD697FD" bold="true" box="[1345,1362,1666,1690]" pageId="7" pageNumber="72">E</emphasis>
|
||
).
|
||
<emphasis id="B91CD837FFE376227A7DFC9DFA1297FD" bold="true" box="[1383,1430,1666,1690]" pageId="7" pageNumber="72">F–N</emphasis>
|
||
. Segments 10 and 11.
|
||
<emphasis id="B91CD837FFE376227E6FFCB9FE0097D9" bold="true" box="[373,388,1702,1726]" pageId="7" pageNumber="72">F</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE376227E89FCB9FE2997D9" bold="true" box="[403,429,1702,1726]" pageId="7" pageNumber="72">G.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE376227EAEFCB9FD5897DA" authority=", ZMB" authorityName="ZMB" box="[436,732,1702,1726]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="72" phylum="Kinorhyncha" rank="species" species="capitatus">
|
||
<emphasis id="B91CD837FFE376227EAEFCB9FD1397D9" box="[436,663,1702,1726]" italics="true" pageId="7" pageNumber="72">Echinoderes capitatus</emphasis>
|
||
, ZMB
|
||
</taxonomicName>
|
||
11913 (
|
||
<emphasis id="B91CD837FFE376227C28FCB9FCC597D9" bold="true" box="[818,833,1702,1726]" pageId="7" pageNumber="72">F</emphasis>
|
||
) and 11921 (
|
||
<emphasis id="B91CD837FFE376227CD6FCB9FC6497D9" bold="true" box="[972,992,1702,1726]" pageId="7" pageNumber="72">G</emphasis>
|
||
).
|
||
<emphasis id="B91CD837FFE376227CECFCB9FB8E97D9" bold="true" box="[1014,1034,1702,1726]" pageId="7" pageNumber="72">H</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE376227B02FCB9FBAC97D9" bold="true" box="[1048,1064,1702,1726]" pageId="7" pageNumber="72">I.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE376227B2AFCB9FAA997D9" box="[1072,1325,1702,1726]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="72" phylum="Kinorhyncha" rank="species" species="meteorensis">
|
||
<emphasis id="B91CD837FFE376227B2AFCB9FAA997D9" box="[1072,1325,1702,1726]" italics="true" pageId="7" pageNumber="72">Echinoderes meteorensis</emphasis>
|
||
</taxonomicName>
|
||
, paratypes ZMB 11571b (
|
||
<emphasis id="B91CD837FFE376227E34FCD5FEC69785" bold="true" box="[302,322,1738,1762]" pageId="7" pageNumber="72">H</emphasis>
|
||
) and ZMB 11572b (
|
||
<emphasis id="B91CD837FFE376227D08FCD5FD989785" bold="true" box="[530,540,1738,1762]" pageId="7" pageNumber="72">I</emphasis>
|
||
).
|
||
<emphasis id="B91CD837FFE376227D2BFCD5FDC09785" bold="true" box="[561,580,1738,1762]" pageId="7" pageNumber="72">J.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE376227D51FCD5FCC39785" box="[587,839,1738,1762]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="72" phylum="Kinorhyncha" rank="species" species="unispinosus">
|
||
<emphasis id="B91CD837FFE376227D51FCD5FCC39785" box="[587,839,1738,1762]" italics="true" pageId="7" pageNumber="72">Echinoderes unispinosus</emphasis>
|
||
</taxonomicName>
|
||
, paratype ZMB 11589b.
|
||
<emphasis id="B91CD837FFE376227B5FFCD5FBDD9785" bold="true" box="[1093,1113,1738,1762]" pageId="7" pageNumber="72">K</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE376227B7CFCD5FBF99785" bold="true" box="[1126,1149,1738,1762]" pageId="7" pageNumber="72">L.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE376227B99FCD5FAB39785" box="[1155,1335,1738,1762]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="72" phylum="Kinorhyncha" rank="species" species="apex">
|
||
<emphasis id="B91CD837FFE376227B99FCD5FAB39785" box="[1155,1335,1738,1762]" italics="true" pageId="7" pageNumber="72">Echinoderes apex</emphasis>
|
||
</taxonomicName>
|
||
, holotype ZMB 11561, paratype ZMB 11562a.
|
||
<emphasis id="B91CD837FFE376227D0EFCF1FDB79661" bold="true" box="[532,563,1774,1798]" pageId="7" pageNumber="72">M.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE376227D23FCF1FCF39662" authority=", ZMB" authorityName="ZMB" box="[569,887,1774,1798]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="72" phylum="Kinorhyncha" rank="species" species="ferrugineus">
|
||
<emphasis id="B91CD837FFE376227D23FCF1FCB79661" box="[569,819,1774,1798]" italics="true" pageId="7" pageNumber="72">Echinoderes ferrugineus</emphasis>
|
||
, ZMB
|
||
</taxonomicName>
|
||
11929.
|
||
<emphasis id="B91CD837FFE376227CD0FCF1FC679661" bold="true" box="[970,995,1774,1798]" pageId="7" pageNumber="72">N.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE376227CF0FCF1FB329661" authorityName="Lang" authorityYear="1949" box="[1002,1206,1774,1798]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="7" pageNumber="72" phylum="Kinorhyncha" rank="species" species="pilosus">
|
||
<emphasis id="B91CD837FFE376227CF0FCF1FB329661" box="[1002,1206,1774,1798]" italics="true" pageId="7" pageNumber="72">Echinoderes pilosus</emphasis>
|
||
</taxonomicName>
|
||
, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in
|
||
<emphasis id="B91CD837FFE376227FC2FDD9FF6D96B9" bold="true" box="[216,233,1990,2014]" pageId="7" pageNumber="72">B</emphasis>
|
||
50 μm, valid for
|
||
<emphasis id="B91CD837FFE376227E80FDD9FE4F96B9" bold="true" box="[410,459,1990,2014]" pageId="7" pageNumber="72">A–C</emphasis>
|
||
. Scale bar in
|
||
<emphasis id="B91CD837FFE376227D4FFDD9FDE396B9" bold="true" box="[597,615,1990,2014]" pageId="7" pageNumber="72">D</emphasis>
|
||
20 µm, valid for
|
||
<emphasis id="B91CD837FFE376227C00FDD9FCD596B9" bold="true" box="[794,849,1990,2014]" pageId="7" pageNumber="72">D–M</emphasis>
|
||
. Scale bar in
|
||
<emphasis id="B91CD837FFE376227CC1FDD9FC6996B9" bold="true" box="[987,1005,1990,2014]" pageId="7" pageNumber="72">N</emphasis>
|
||
20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1754ADFFEC762D7F8DFEEEFE629485" ID-DOI="http://doi.org/10.5281/zenodo.6036942" ID-Zenodo-Dep="6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="8" pageNumber="73" startId="8.[151,250,1265,1289]" targetBox="[151,1436,181,1241]" targetPageId="8">
|
||
<paragraph id="8BD70425FFEC762D7F8DFEEEFE629485" blockId="8.[151,1437,1265,1506]" pageId="8" pageNumber="73">
|
||
<emphasis id="B91CD837FFEC762D7F8DFEEEFE97946E" bold="true" box="[151,275,1265,1289]" pageId="8" pageNumber="73">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C687FA6FFEC762D7E00FEEEFE54946E" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[282,464,1265,1289]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="8" pageNumber="73" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFEC762D7E00FEEEFE54946E" box="[282,464,1265,1289]" italics="true" pageId="8" pageNumber="73">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22F654CFFEC762D7ECCFEEEFDAD946E" box="[470,553,1265,1289]" pageId="8" pageNumber="73" rank="species">
|
||
<emphasis id="B91CD837FFEC762D7ECCFEEEFDA6946E" bold="true" box="[470,546,1265,1289]" pageId="8" pageNumber="73">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, female, ZMB 12271 (
|
||
<emphasis id="B91CD837FFEC762D7C0EFEEEFCA2946E" bold="true" box="[788,806,1265,1289]" pageId="8" pageNumber="73">A</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFEC762D7C2EFEEEFCC2946E" bold="true" box="[820,838,1265,1289]" pageId="8" pageNumber="73">C</emphasis>
|
||
) and ZMB 12272 (
|
||
<emphasis id="B91CD837FFEC762D7B16FEEEFB99946E" bold="true" box="[1036,1053,1265,1289]" pageId="8" pageNumber="73">B</emphasis>
|
||
), in laterodorsal (
|
||
<emphasis id="B91CD837FFEC762D7BCEFEEEFB62946E" bold="true" box="[1236,1254,1265,1289]" pageId="8" pageNumber="73">A</emphasis>
|
||
), ventral (
|
||
<emphasis id="B91CD837FFEC762D7A4BFEEEFAE6946E" bold="true" box="[1361,1378,1265,1289]" pageId="8" pageNumber="73">B</emphasis>
|
||
), and dorsal-right (
|
||
<emphasis id="B91CD837FFEC762D7E07FF0AFEAB944A" bold="true" box="[285,303,1301,1325]" pageId="8" pageNumber="73">C</emphasis>
|
||
) view, SEM.
|
||
<emphasis id="B91CD837FFEC762D7EA4FF0AFE53944A" bold="true" box="[446,471,1301,1325]" pageId="8" pageNumber="73">A.</emphasis>
|
||
Habitus.
|
||
<emphasis id="B91CD837FFEC762D7D21FF0AFDD5944A" bold="true" box="[571,593,1301,1325]" pageId="8" pageNumber="73">B.</emphasis>
|
||
Mouth cone. White arrow points to bifurcated tip of posterior spinose process.
|
||
<emphasis id="B91CD837FFEC762D7A9EFF0AFA18944A" bold="true" box="[1412,1436,1301,1325]" pageId="8" pageNumber="73">C.</emphasis>
|
||
Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (
|
||
<emphasis id="B91CD837FFEC762D7C8CFFBAFC2C94DA" bold="true" box="[918,936,1445,1469]" pageId="8" pageNumber="73">A</emphasis>
|
||
) or introvert ring number (
|
||
<emphasis id="B91CD837FFEC762D7BA0FFBAFB4894DA" bold="true" box="[1210,1228,1445,1469]" pageId="8" pageNumber="73">C</emphasis>
|
||
). Scale bar in
|
||
<emphasis id="B91CD837FFEC762D7A44FFBAFAF494DA" bold="true" box="[1374,1392,1445,1469]" pageId="8" pageNumber="73">A</emphasis>
|
||
100 μm. Scale bar in
|
||
<emphasis id="B91CD837FFEC762D7E59FFD6FED09486" bold="true" box="[323,340,1481,1505]" pageId="8" pageNumber="73">B</emphasis>
|
||
and
|
||
<emphasis id="B91CD837FFEC762D7E9FFFD6FE139486" bold="true" box="[389,407,1481,1505]" pageId="8" pageNumber="73">C</emphasis>
|
||
10 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37257AEFFEC762D7FDDFC16FDB59690" pageId="8" pageNumber="73" type="materials_examined">
|
||
<paragraph id="8BD70425FFEC762D7FDDFC16FABA96C8" blockId="8.[151,1437,1545,2040]" pageId="8" pageNumber="73">
|
||
<emphasis id="B91CD837FFEC762D7FDDFC16FE2A9744" bold="true" box="[199,430,1545,1571]" pageId="8" pageNumber="73">Material examined.</emphasis>
|
||
<materialsCitation id="3B000E78FFEC762D7EA8FC16FE4597E8" ID-GBIF-Occurrence="3464450303" collectingDate="2019-03-31" collectionCode="IADO, CONICET" collectorName="Male & Agustin G. Menechella" country="Argentina" county="Cobb" elevation="40" location="Motorsailer" municipality="Argentinean" pageId="8" pageNumber="73" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54D3BA87FFEC762D7EA8FC16FD9B9744" box="[434,543,1545,1571]" pageId="8" pageNumber="73" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<collectorName id="269D61F3FFEC762D7D3DFC16FDE59744" box="[551,609,1545,1571]" pageId="8" pageNumber="73">Male</collectorName>
|
||
(MACN-In 43755), collected on
|
||
<date id="FFD622E5FFEC762D7CD6FC16FBFD9744" box="[972,1145,1545,1571]" pageId="8" pageNumber="73" value="2019-03-31">
|
||
<collectingDate id="EF92DB0DFFEC762D7CD6FC16FBFD9744" box="[972,1145,1545,1571]" pageId="8" pageNumber="73" value="2019-03-31">March 31, 2019</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="269D61F3FFEC762D7B87FC16FA199744" box="[1181,1437,1545,1571]" pageId="8" pageNumber="73">Agustín G. Menechella</collectorName>
|
||
(
|
||
<collectionCode id="ED799CE0FFEC762D7F85FC32FF6D9720" box="[159,233,1581,1607]" pageId="8" pageNumber="73">IADO</collectionCode>
|
||
,
|
||
<collectionCode id="ED799CE0FFEC762D7FEDFC32FEF59720" box="[247,369,1581,1607]" pageId="8" pageNumber="73">CONICET</collectionCode>
|
||
,
|
||
<collectingCountry id="F37F44B5FFEC762D7E67FC32FE75972F" box="[381,497,1581,1608]" name="Argentina" pageId="8" pageNumber="73">Argentina</collectingCountry>
|
||
), at station E29 during the campaign aboard the
|
||
<location id="8EB752FEFFEC762D7B3FFC32FB2C9720" LSID="urn:lsid:plazi:treatment:03C1B533FFE776347F8DFC0DFC679672:8EB752FEFFEC762D7B3FFC32FB2C9720" box="[1061,1192,1581,1607]" country="Argentina" county="Cobb" municipality="Argentinean" name="Motorsailer" pageId="8" pageNumber="73">Motorsailer</location>
|
||
“Bernardo Houssay” through the
|
||
<collectingMunicipality id="6BB39E5FFFEC762D7E06FC4EFE22970C" box="[284,422,1617,1643]" pageId="8" pageNumber="73">Argentinean</collectingMunicipality>
|
||
continental shelf (
|
||
<figureCitation id="135318A0FFEC762D7D6FFC4EFD32970C" box="[629,694,1617,1643]" captionStart="FIGURE 1" captionStartId="2.[151,250,949,973]" captionTargetBox="[354,1232,202,919]" captionTargetId="figure-20@2.[352,1236,198,922]" captionTargetPageId="2" captionText="FIGURE 1. Map of the sampling station at the Argentinean continental shelf (Station E29)." figureDoi="http://doi.org/10.5281/zenodo.6036912" httpUri="https://zenodo.org/record/6036912/files/figure.png" pageId="8" pageNumber="73">Fig. 1</figureCitation>
|
||
), at
|
||
<elevation id="0045E316FFEC762D7DF3FC4EFCE1970C" box="[745,869,1617,1643]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="73" unit="m" value="40.0">
|
||
<quantity id="4C90A9C0FFEC762D7DF3FC4EFCA5970C" box="[745,801,1617,1643]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="73" unit="m" value="40.0">40 m</quantity>
|
||
depth
|
||
</elevation>
|
||
using a dredge; mounted as glycerol-paraffin slide on
|
||
<collectingCounty id="62B67CA9FFEC762D7FA6FC6AFF7D97E8" box="[188,249,1653,1679]" pageId="8" pageNumber="73">Cobb</collectingCounty>
|
||
aluminum frame
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B000E78FFEC762D7ED4FC6AFABF96C8" ID-GBIF-Occurrence="3464450304" accessionNumber="ZMB 12262, ZMB 12263, ZMB 12270, ZMB 12350, ZMB 12271" collectingDate="2019-03-31" collectionCode="ZMB" collectorName="Male & Agustin G. Menechella" country="Argentina" county="Cobb" elevation="40" location="Museum" municipality="Museo Argentino de Ciencias Naturales" pageId="8" pageNumber="73" specimenCode="12264, 12265, 12314-12331, 12334-12344, 12266, 12332, 12333, ZMB 12267-12269, 12345- 12347, 12351, 12272, 12348, 12349" specimenCount="38" specimenCount-adult="1" specimenCount-female="14" specimenCount-juvenile="1" specimenCount-male="22" stateProvince="Berlin" typeStatus="paratype">
|
||
<typeStatus id="54D3BA87FFEC762D7ED4FC6AFDBF97E8" box="[462,571,1653,1679]" pageId="8" pageNumber="73" type="paratype">Paratypes</typeStatus>
|
||
mounted as glycerol-paraffin slides on
|
||
<collectingCounty id="62B67CA9FFEC762D7B19FC6AFBC497E8" box="[1027,1088,1653,1679]" pageId="8" pageNumber="73">Cobb</collectingCounty>
|
||
aluminum frames: 1) material deposited at the
|
||
<collectingMunicipality id="6BB39E5FFFEC762D7E4CFC86FC9A97D4" box="[342,798,1689,1715]" pageId="8" pageNumber="73">Museo Argentino de Ciencias Naturales</collectingMunicipality>
|
||
“Bernardino Rivadavia”, Buenos Aires,
|
||
<collectingCountry id="F37F44B5FFEC762D7BE9FC86FAE397D4" box="[1267,1383,1689,1715]" name="Argentina" pageId="8" pageNumber="73">Argentina</collectingCountry>
|
||
:
|
||
<specimenCount id="9D6ECFACFFEC762D7A6EFC86FF4997B0" pageId="8" pageNumber="73" type="male">one male</specimenCount>
|
||
and
|
||
<specimenCount id="9D6ECFACFFEC762D7E1DFCA2FE1597B0" box="[263,401,1725,1751]" pageId="8" pageNumber="73" type="female">two females</specimenCount>
|
||
(catalogue numbers MACN-In 43756a –43756c); 2) material deposited at the
|
||
<location id="8EB752FEFFEC762D7A0BFCA2FAF797B0" LSID="urn:lsid:plazi:treatment:03C1B533FFE776347F8DFC0DFC679672:8EB752FEFFEC762D7A0BFCA2FAF797B0" box="[1297,1395,1725,1751]" country="Argentina" county="Cobb" municipality="Museo Argentino de Ciencias Naturales" name="Museum" pageId="8" pageNumber="73" stateProvince="Berlin">Museum</location>
|
||
für
|
||
<location id="8EB752FEFFEC762D7F8DFCFEFE9F979C" LSID="urn:lsid:plazi:treatment:03C1B533FFE776347F8DFC0DFC679672:8EB752FEFFEC762D7F8DFCFEFE9F979C" box="[151,283,1761,1787]" country="Argentina" county="Cobb" municipality="Museo Argentino de Ciencias Naturales" name="Naturkunde" pageId="8" pageNumber="73" stateProvince="Berlin">Naturkunde</location>
|
||
<collectingRegion id="49ACCAC7FFEC762D7E3BFCFEFEEE979C" box="[289,362,1761,1787]" country="Germany" name="Berlin" pageId="8" pageNumber="73">Berlin</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37F44B5FFEC762D7E6EFCFEFE59979C" box="[372,477,1761,1787]" name="Germany" pageId="8" pageNumber="73">Germany</collectingCountry>
|
||
, collection “Vermes”, “Generalkatalog freilebende Würmer”:
|
||
<specimenCount id="9D6ECFACFFEC762D7B96FCFEFA81979C" box="[1164,1285,1761,1787]" pageId="8" pageNumber="73" type="female">12 females</specimenCount>
|
||
and
|
||
<specimenCount id="9D6ECFACFFEC762D7A23FCFEFA18979C" box="[1337,1436,1761,1787]" pageId="8" pageNumber="73" type="male">19 males</specimenCount>
|
||
(catalogue numbers
|
||
<collectionCode id="ED799CE0FFEC762D7E6DFD1AFE309678" box="[375,436,1797,1823]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">ZMB</collectionCode>
|
||
12262,
|
||
<specimenCode id="DBCEAC5EFFEC762D7D16FD1AFDD19678" box="[524,597,1797,1823]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12264</specimenCode>
|
||
,
|
||
<specimenCode id="DBCEAC5EFFEC762D7D45FD1AFD2C9647" box="[607,680,1797,1824]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12265</specimenCode>
|
||
,
|
||
<specimenCode id="DBCEAC5EFFEC762D7DABFD1AFCCB9678" box="[689,847,1797,1823]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12314–12331</specimenCode>
|
||
,
|
||
<specimenCode id="DBCEAC5EFFEC762D7C42FD1AFC739678" box="[856,1015,1797,1823]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12334–12344</specimenCode>
|
||
). Non-types, deposited at Museum für Naturkunde
|
||
<collectingRegion id="49ACCAC7FFEC762D7E3BFD36FEEF9624" box="[289,363,1833,1859]" country="Germany" name="Berlin" pageId="8" pageNumber="73">Berlin</collectingRegion>
|
||
:
|
||
<specimenCount id="9D6ECFACFFEC762D7E6FFD36FE6E9624" box="[373,490,1833,1859]" pageId="8" pageNumber="73" type="male">four males</specimenCount>
|
||
(
|
||
<collectionCode id="ED799CE0FFEC762D7EE2FD36FDB29624" box="[504,566,1833,1859]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">ZMB</collectionCode>
|
||
12263,
|
||
<specimenCode id="DBCEAC5EFFEC762D7D8AFD36FD5D9624" box="[656,729,1833,1859]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12266</specimenCode>
|
||
,
|
||
<specimenCode id="DBCEAC5EFFEC762D7DF8FD36FCAF9624" box="[738,811,1833,1859]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12332</specimenCode>
|
||
,
|
||
<specimenCode id="DBCEAC5EFFEC762D7C2FFD36FCF89624" box="[821,892,1833,1859]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12333</specimenCode>
|
||
) and
|
||
<specimenCount id="9D6ECFACFFEC762D7CA3FD36FBCB9624" box="[953,1103,1833,1859]" pageId="8" pageNumber="73" type="juvenile">five juveniles</specimenCount>
|
||
(
|
||
<specimenCode id="DBCEAC5EFFEC762D7B47FD36FABA9624" box="[1117,1342,1833,1859]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">ZMB 12267–12269</specimenCode>
|
||
,
|
||
<specimenCode id="DBCEAC5EFFEC762D7A52FD36FF5A9600" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12345– 12347</specimenCode>
|
||
) mounted as glycerol-paraffin slides on Cobb aluminum frames and
|
||
<specimenCount id="9D6ECFACFFEC762D7CF1FD52FBE49600" box="[1003,1120,1869,1895]" pageId="8" pageNumber="73" type="adult">two adults</specimenCount>
|
||
of undetermined sex (
|
||
<collectionCode id="ED799CE0FFEC762D7A44FD52FA189600" box="[1374,1436,1869,1895]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">ZMB</collectionCode>
|
||
12270,
|
||
<specimenCode id="DBCEAC5EFFEC762D7FF0FD6EFEB096EB" box="[234,308,1905,1932]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12351</specimenCode>
|
||
),
|
||
<specimenCount id="9D6ECFACFFEC762D7E5CFD6EFE2F96EC" box="[326,427,1905,1931]" pageId="8" pageNumber="73" type="male">one male</specimenCount>
|
||
(
|
||
<collectionCode id="ED799CE0FFEC762D7EA3FD6EFE7396EC" box="[441,503,1905,1931]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">ZMB</collectionCode>
|
||
12350), and
|
||
<specimenCount id="9D6ECFACFFEC762D7D9DFD6EFC9796EC" box="[647,787,1905,1931]" pageId="8" pageNumber="73" type="female">four females</specimenCount>
|
||
(
|
||
<collectionCode id="ED799CE0FFEC762D7C3AFD6EFCDA96EC" box="[800,862,1905,1931]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">ZMB</collectionCode>
|
||
12271,
|
||
<specimenCode id="DBCEAC5EFFEC762D7CADFD6EFB8496EC" box="[951,1024,1905,1931]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12272</specimenCode>
|
||
,
|
||
<specimenCode id="DBCEAC5EFFEC762D7B10FD6EFBD796EC" box="[1034,1107,1905,1931]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12348</specimenCode>
|
||
,
|
||
<specimenCode id="DBCEAC5EFFEC762D7B47FD6EFB2196EC" box="[1117,1189,1905,1931]" collectionCode="ZMB" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="8" pageNumber="73" type="Museum">12349</specimenCode>
|
||
) mounted for SEM on two stubs. All
|
||
<typeStatus id="54D3BA87FFEC762D7E23FD8AFE2096C8" box="[313,420,1941,1967]" pageId="8" pageNumber="73" type="paratype">paratypes</typeStatus>
|
||
and non-types collected at same locality and date by same person as the
|
||
<typeStatus id="54D3BA87FFEC762D7BCFFD8AFABF96C8" box="[1237,1339,1941,1967]" pageId="8" pageNumber="73" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFEC762D7FDDFDA6FDB59690" blockId="8.[151,1437,1545,2040]" pageId="8" pageNumber="73">
|
||
<materialsCitation id="3B000E78FFEC762D7FDDFDA6FDA99690" ID-GBIF-Occurrence="3464450302" county="Monte Hermoso" elevation="40" latitude="-39.111668" location="South West Atlantic Ocean" longLatPrecision="116" longitude="-61.441666" municipality="Argentinian" pageId="8" pageNumber="73" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B91CD837FFEC762D7FDDFDA6FEE296B4" bold="true" box="[199,358,1977,2003]" pageId="8" pageNumber="73">
|
||
<typeStatus id="54D3BA87FFEC762D7FDDFDA6FE8696B4" box="[199,258,1977,2003]" pageId="8" pageNumber="73">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EB752FEFFEC762D7E74FDA6FD2496B4" LSID="urn:lsid:plazi:treatment:03C1B533FFE776347F8DFC0DFC679672:8EB752FEFFEC762D7E74FDA6FD2496B4" box="[366,672,1977,2003]" county="Monte Hermoso" latitude="-39.111668" longLatPrecision="116" longitude="-61.441666" municipality="Argentinian" name="South West Atlantic Ocean" pageId="8" pageNumber="73">South West Atlantic Ocean</location>
|
||
,
|
||
<collectingMunicipality id="6BB39E5FFFEC762D7DB1FDA6FCB496B4" box="[683,816,1977,2003]" pageId="8" pageNumber="73">Argentinian</collectingMunicipality>
|
||
continental shelf, off Pehuen-Có and
|
||
<collectingCounty id="62B67CA9FFEC762D7BC4FDA6FA1D96B4" box="[1246,1433,1977,2003]" pageId="8" pageNumber="73">Monte Hermoso</collectingCounty>
|
||
,
|
||
<geoCoordinate id="EE5C62E2FFEC762D7F8DFDC2FE8F9690" box="[151,267,2013,2039]" degrees="39" direction="south" minutes="06.7" orientation="latitude" pageId="8" pageNumber="73" precision="92" value="-39.111668">39°06.7’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5C62E2FFEC762D7E0CFDC2FE24969F" box="[278,416,2013,2040]" degrees="061" direction="west" minutes="26.5" orientation="longitude" pageId="8" pageNumber="73" precision="92" value="-61.441666">061°26.5’W</geoCoordinate>
|
||
,
|
||
<elevation id="0045E316FFEC762D7EB6FDC2FDA99690" box="[428,557,2013,2039]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="73" unit="m" value="40.0">
|
||
<quantity id="4C90A9C0FFEC762D7EB6FDC2FE619690" box="[428,485,2013,2039]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="73" unit="m" value="40.0">40 m</quantity>
|
||
depth
|
||
</elevation>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37257AEFFED76347FDDFA89FC679672" lastPageId="17" lastPageNumber="82" pageId="9" pageNumber="74" type="description">
|
||
<paragraph id="8BD70425FFED762C7FDDFA89FD72919E" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="74">
|
||
<emphasis id="B91CD837FFED762C7FDDFA89FED591D7" bold="true" box="[199,337,150,176]" pageId="9" pageNumber="74">Description</emphasis>
|
||
. Adults with head, neck, and 11 trunk segments (
|
||
<figureCitation id="135318A0FFED762C7C66FA88FC7791D6" box="[892,1011,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7B1AFA88FBBA91D6" box="[1024,1086,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">3A,C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7B50FA88FBF491D5" box="[1098,1136,151,178]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">6A</figureCitation>
|
||
). For a complete overview of measures and dimensions see
|
||
<tableCitation id="C6EA319EFFED762C7D19FAA4FDDD91B2" box="[515,601,187,213]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="9" pageNumber="74" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Table 2</tableCitation>
|
||
. Distribution of cuticular structures, i.e., sensory spots, gland cells, conical structures, spines and tubes, is summarised in
|
||
<tableCitation id="C6EA319EFFED762C7D86FAC0FD75919E" box="[668,753,223,249]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="9" pageNumber="74" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">Table 3</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFED762C7FDDFB1CFE659396" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="74">
|
||
<emphasis id="B91CD837FFED762C7FDDFB1CFE8A907A" bold="true" box="[199,270,259,285]" pageId="9" pageNumber="74">Head.</emphasis>
|
||
Head consisting of a retractable mouth cone and an introvert (
|
||
<figureCitation id="135318A0FFED762C7CD1FB1CFBC6907A" box="[971,1090,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">Figs 3A,C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7B57FB1CFB119079" box="[1101,1173,259,286]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">6A–C</figureCitation>
|
||
). Mouth cone equipped with 9 outer oral styles, each with two articulating elements and neighbouring styles slightly alternating in length (
|
||
<figureCitation id="135318A0FFED762C7F85FB54FF7A9002" box="[159,254,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">Figs 4K</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7E11FB54FEAB9001" box="[267,303,331,358]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">6B</figureCitation>
|
||
). Each outer oral style possessing at its base a posterior central fringe of spinose processes and one anterior lateral spinose process on each side (
|
||
<figureCitation id="135318A0FFED762C7D86FB70FD7290ED" box="[668,758,367,394]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">Fig. 6B</figureCitation>
|
||
). Posterior spinose processes may be split apically into two processes (
|
||
<figureCitation id="135318A0FFED762C7E08FB8CFEE890C9" box="[274,364,403,430]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">Fig. 6B</figureCitation>
|
||
). Although two out of
|
||
<specimenCount id="9D6ECFACFFED762C7D71FB8CFC9B90CA" box="[619,799,403,429]" pageId="9" pageNumber="74" type="generic">three specimens</specimenCount>
|
||
mounted for SEM appeared with the introvert extended, the information on scalid numbers and arrangement is scarce because of particles partially masking characters. Both odd and even-numbered sectors showing seemingly 6 spinoscalid rings (
|
||
<figureCitation id="135318A0FFED762C7B1EFBC4FBD99091" box="[1028,1117,475,502]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">Fig. 6C</figureCitation>
|
||
). All spinoscalids composed of a broader sheath-like base with a terminal fringe of spinose processes and an articulating terminal element ending bluntly in the primary spinoscalids and spinose in rings 02–06 (
|
||
<figureCitation id="135318A0FFED762C7CD3F83CFBA09359" box="[969,1060,547,574]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">Fig. 6C</figureCitation>
|
||
). Terminal element covered with short cuticular hairs. Each primary spinoscalid revealing apically on its base two prominent fringes of spinose processes (
|
||
<figureCitation id="135318A0FFED762C7E08F874FEEF93E1" box="[274,363,619,646]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">Fig. 6C</figureCitation>
|
||
). One prominent apical process also found on the basis of ring 03 spinoscalids but not on those of the following rings. Spinoscalid of ring 06 of each second sector appearing to be much smaller than that of its neighbouring sector. Six short and hairy trichoscalids with trichoscalid plates being present, opposite to the neck placids 2, 6, 8, 10, 12 and 16.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFED762C7FDDF8E4FBB092AE" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="74">
|
||
<emphasis id="B91CD837FFED762C7FDDF8E4FE8F9272" bold="true" box="[199,267,763,789]" pageId="9" pageNumber="74">Neck.</emphasis>
|
||
Neck equipped with 16 trapezoidal placids (
|
||
<figureCitation id="135318A0FFED762C7C08F8E4FC0F9272" box="[786,907,763,789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7C80F8E4FC3F9272" box="[922,955,763,789]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7CD0F8E4FC6B9272" box="[970,1007,763,789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4D</figureCitation>
|
||
), wider posteriorly and with rounded edges anteriorly in SEM of at least
|
||
<specimenCount id="9D6ECFACFFED762C7D2AF900FD4F925E" box="[560,715,799,825]" pageId="9" pageNumber="74" type="generic">one specimen</specimenCount>
|
||
revealing placids at full length (ZMB 12270; not shown here). In light microscopy, anterior end of each placid of almost all specimens looking as if bent partly backwards (
|
||
<figureCitation id="135318A0FFED762C7A70F95CFF5192E6" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7FFBF978FE8692E6" box="[225,258,871,897]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7E14F978FEB792E6" box="[270,307,871,897]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7E24F979FEE692E6" box="[318,354,870,897]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A</figureCitation>
|
||
), possibly because of the thin cuticle revealed by most specimens (see below). Midventral placid wider than the remaining ones. All placids articulating with the first trunk segment. Trichoscalid plates present on paradorsal (8 and 10), laterodorsal (6 and 12) and ventromedial (2 and 16) placids.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFED762C7FDDF9CCFA849736" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="74">
|
||
<emphasis id="B91CD837FFED762C7FDDF9CCFE9E928A" bold="true" box="[199,282,979,1005]" pageId="9" pageNumber="74">Trunk.</emphasis>
|
||
Segments 1 and 2 formed by a closed cuticular ring (
|
||
<figureCitation id="135318A0FFED762C7C69F9CCFC6E928A" box="[883,1002,979,1005]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7CEDF9CCFBB3928A" box="[1015,1079,979,1005]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4A,D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7B58F9CCFBE7928A" box="[1090,1123,979,1005]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8C</figureCitation>
|
||
) and the remaining ones by one tergal and two sternal plates (
|
||
<figureCitation id="135318A0FFED762C7D0DF9E8FD0B9576" box="[535,655,1015,1041]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7D87F9E8FD609576" box="[669,740,1015,1041]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">3E,F,J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7DE8F9E8FCC19576" box="[754,837,1015,1041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4A–E,I</figureCitation>
|
||
,
|
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<figureCitation id="135318A0FFED762C7C49F9E9FC1C9576" box="[851,920,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A–C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7CBFF9E8FC4E9576" box="[933,970,1015,1041]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">6A</figureCitation>
|
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,
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<figureCitation id="135318A0FFED762C7CCCF9E8FC629576" box="[982,998,1015,1041]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">7</figureCitation>
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<figureCitation id="135318A0FFED762C7CEEF9E8FBD69576" box="[1012,1106,1015,1041]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8B,D–F</figureCitation>
|
||
). Midsternal and tergosternal junctions well-developed, however, the latter being less conspicuous and sometime hard to see under the light microscope. Pachycycli present anteriorly on segments 1–11 (
|
||
<figureCitation id="135318A0FFED762C7C7BFE20FC61953E" box="[865,997,1087,1113]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">Figs 3E,F,J</figureCitation>
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,
|
||
<figureCitation id="135318A0FFED762C7CEEFE20FBDE953E" box="[1012,1114,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4A–E,I,J</figureCitation>
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<figureCitation id="135318A0FFED762C7B72FE21FB34953E" box="[1128,1200,1086,1113]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A–C</figureCitation>
|
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). Cuticular hairs not abundant and filiform (
|
||
<figureCitation id="135318A0FFED762C7E82FE7CFD9C951A" box="[408,536,1123,1149]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">Figs 3E,F,J</figureCitation>
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,
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<figureCitation id="135318A0FFED762C7D39FE7DFDEC951A" box="[547,616,1122,1149]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A–C</figureCitation>
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,
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<figureCitation id="135318A0FFED762C7D69FE7CFD1C951A" box="[627,664,1123,1149]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">6A</figureCitation>
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<figureCitation id="135318A0FFED762C7DB9FE7CFD63951A" box="[675,743,1123,1149]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">7A–E</figureCitation>
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,
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<figureCitation id="135318A0FFED762C7DE8FE7CFCB0951A" box="[754,820,1123,1149]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8B–F</figureCitation>
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). The bracteate hairs on segments 2–10 arranged in one or two irregular wavy rows (
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<figureCitation id="135318A0FFED762C7EC4FE98FDBA95C6" box="[478,574,1159,1185]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">Figs 6A</figureCitation>
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,
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<figureCitation id="135318A0FFED762C7D50FE98FD0A95C6" box="[586,654,1159,1185]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">7A–E</figureCitation>
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,
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<figureCitation id="135318A0FFED762C7D81FE98FD5995C6" box="[667,733,1159,1185]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8C–F</figureCitation>
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). Hairs of anterior row shorter than those of the posterior one (
|
||
<figureCitation id="135318A0FFED762C7F85FEB4FF7995A2" box="[159,253,1195,1221]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">Figs 6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7E12FEB4FEC895A2" box="[264,332,1195,1221]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">7A–E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7E42FEB4FE1E95A2" box="[344,410,1195,1221]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8C–F</figureCitation>
|
||
). Well-developed patch of short, stronger cuticular hairs in three rows present paraventrally and centrally on segments 3–6 (
|
||
<figureCitation id="135318A0FFED762C7D1DFED0FDE3958E" box="[519,615,1231,1257]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">Figs 4D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7D6FFED0FD32958E" box="[629,694,1231,1257]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8C–E</figureCitation>
|
||
) and, less developed, also on segment 7 (
|
||
<figureCitation id="135318A0FFED762C7B80FED0FB70958E" box="[1178,1268,1231,1257]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">Fig. 8E</figureCitation>
|
||
). Segments 4– 10 with a ventrolateral to paraventral patch of thin cuticular hairs organised in a single or sometimes in two or three rows (
|
||
<figureCitation id="135318A0FFED762C7E04FF08FE1C9456" box="[286,408,1303,1329]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">Fig. 8D–F</figureCitation>
|
||
). Most adult specimens having moulted recently recognizable by differing extent of fusion of button-like structures in cuticular flap at posterior end of each segment (comp.
|
||
<figureCitation id="135318A0FFED762C7B50FF24FB529432" box="[1098,1238,1339,1365]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">Figs 4D,F–I</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7BFFFF25FA8D9432" box="[1253,1289,1338,1365]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A</figureCitation>
|
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,
|
||
<figureCitation id="135318A0FFED762C7A0DFF24FA149432" box="[1303,1424,1339,1365]" captionStart="FIGURE 9" captionStartId="12.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-20@12.[151,1436,181,1531]" captionTargetPageId="12" captionText="FIGURE 9. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, ventral (B–E), optical section (A, F), and dorsal side (G, H). A. Habitus. B. Segments 1–4. C, D. Segments 5–8 at different focal levels showing aspects of the juvenile (C) and adult (D) cuticle. E. Segments 8–11. F. Segments 9–11. G. Segments 1–4. H. Segments 4–7.Abbreviations: bs, button-like structures; fe, refers to female character; juv, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in A 100 µm. Scale bar in F 50 µm, valid for B–H." figureDoi="http://doi.org/10.5281/zenodo.6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="9" pageNumber="74">9B,D,F–H</figureCitation>
|
||
). Primary pectinate fringe well-developed in segments 2–11 and formed by teeth usually of even length and width within a segment (
|
||
<figureCitation id="135318A0FFED762C7E75FF9CFE7694FA" box="[367,498,1411,1437]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">Figs 3E,F,J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7D1BFF9CFD9F94FA" box="[513,539,1411,1437]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4I</figureCitation>
|
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,
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<figureCitation id="135318A0FFED762C7D33FF9DFDE294FA" box="[553,614,1410,1437]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5B,C</figureCitation>
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,
|
||
<figureCitation id="135318A0FFED762C7D6FFF9CFD0194FA" box="[629,645,1411,1437]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">7</figureCitation>
|
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,
|
||
<figureCitation id="135318A0FFED762C7D8EFF9CFD5394FA" box="[660,727,1411,1437]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8B–F</figureCitation>
|
||
). Fringe appearing shorter on anterior three segments than in remaining segments and almost lacking dorsally on segment 10 (
|
||
<figureCitation id="135318A0FFED762C7C90FFB8FC7B94A6" box="[906,1023,1447,1473]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">Fig. 7D,E</figureCitation>
|
||
), narrower ventrally on segment 10 than on remaining segments (
|
||
<figureCitation id="135318A0FFED762C7EF3FFD4FDC49482" box="[489,576,1483,1509]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">Fig. 8F</figureCitation>
|
||
). Secondary pectinate fringe present as a single row of very short cuticular processes dorsally and ventromedially (
|
||
<figureCitation id="135318A0FFED762C7D48FFF0FD29976E" box="[594,685,1519,1545]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">Figs 7C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7DA3FFF0FD5E976E" box="[697,730,1519,1545]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8F</figureCitation>
|
||
). Type-1 sensory spots (
|
||
<figureCitation id="135318A0FFED762C7CF2FFF0FBB5976E" box="[1000,1073,1519,1545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">
|
||
Figs
|
||
<geoCoordinate id="EE5C62E2FFED762C7B3BFFF0FBB5976E" box="[1057,1073,1519,1545]" degrees="2, 3" direction="east" orientation="longitude" pageId="9" pageNumber="74" precision="5555" value="2.3">2</geoCoordinate>
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</figureCitation>
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,
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||
<figureCitation id="135318A0FFED762C7B27FFF0FBF3976E" box="[1085,1143,1519,1545]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">3E–J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7B99FFF0FB7C976E" box="[1155,1272,1519,1545]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4A–D,F–I</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7A19FFF1FACC976E" box="[1283,1352,1518,1545]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A–C</figureCitation>
|
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,
|
||
<figureCitation id="135318A0FFED762C7A4EFFF0FA1C976E" box="[1364,1432,1519,1545]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">7A–E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7F8DFC0CFF53974A" box="[151,215,1555,1581]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8C–F</figureCitation>
|
||
) and type-1 gland cell outlets (
|
||
<figureCitation id="135318A0FFED762C7D21FC0CFD02974A" box="[571,646,1555,1581]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">
|
||
Figs
|
||
<geoCoordinate id="EE5C62E2FFED762C7D6CFC0CFD02974A" box="[630,646,1555,1581]" degrees="2, 3" direction="east" orientation="longitude" pageId="9" pageNumber="74" precision="5555" value="2.3">2</geoCoordinate>
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</figureCitation>
|
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,
|
||
<figureCitation id="135318A0FFED762C7D89FC0CFD79974A" box="[659,765,1555,1581]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">3E,G,H,J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7C10FC0CFC13974A" box="[778,919,1555,1581]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4A,B,D–G,I</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7CBEFC0DFC64974A" box="[932,992,1554,1581]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A,B</figureCitation>
|
||
) present. Type-2 gland cell outlets and female-specific papillae or funnel-like or pear-shaped subcuticular structures absent on any segment.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFED762C7FDDFC44FBC89662" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="74">
|
||
Segment 1 (
|
||
<figureCitation id="135318A0FFED762C7E51FC44FE459712" box="[331,449,1627,1653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7ED7FC44FE6A9712" box="[461,494,1627,1653]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7EE3FC44FDBD9712" box="[505,569,1627,1653]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4A,D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7D5EFC45FDEC9712" box="[580,616,1626,1653]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7D69FC44FD1C9712" box="[627,664,1627,1653]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7DB9FC44FD4C9712" box="[675,712,1627,1653]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">7A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7DC8FC44FD729712" box="[722,758,1627,1653]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8C</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFED762C7C1AFC44FCD69712" box="[768,850,1627,1653]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="9" pageNumber="74" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">Table 3</tableCitation>
|
||
) lacking spines and tubes. Sensory spot usually with 2 but sometimes with up to 5 pores present centrally in subdorsal, midlateral and ventromedial position, flanked by long cuticular hairs. Type-1 gland cell outlet occurring centrally in middorsal and anteriorly in lateroventral position. Other hairs than those associated with sensory spots not existing on this segment. Primary pectinate fringe as row of very short teeth with a blunt tip, broader than teeth of remaining segments.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFED762C7FDDFD10FED096DE" blockId="9.[151,1437,150,1977]" pageId="9" pageNumber="74">
|
||
Segment 2 (
|
||
<figureCitation id="135318A0FFED762C7E50FD10FE44964E" box="[330,448,1807,1833]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="9" pageNumber="74">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7ED1FD10FE68964E" box="[459,492,1807,1833]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="9" pageNumber="74">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7EECFD10FDB2964E" box="[502,566,1807,1833]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="9" pageNumber="74">4A,D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7D5BFD11FDFA964E" box="[577,638,1806,1833]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="9" pageNumber="74">5A,C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7D93FD10FD2A964E" box="[649,686,1807,1833]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="9" pageNumber="74">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7DA2FD10FD59964E" box="[696,733,1807,1833]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="9" pageNumber="74">7A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFED762C7DFDFD10FC8F964E" box="[743,779,1807,1833]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="9" pageNumber="74">8C</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFED762C7C0FFD10FCF7964E" box="[789,883,1807,1833]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="9" pageNumber="74" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6EA319EFFED762C7C64FD10FC08964E" box="[894,908,1807,1833]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="9" pageNumber="74" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) with subdorsal and ventrolateral tube. Sensory spot present posteriorly in middorsal and midlateral position and centrally to posteriorly in ventromedial position. Type-1 gland cell outlet occurring anteriorly in middorsal and ventromedial position. Cuticular hairs scarce on this segment, but more evident if associated with the sensory spots. A few additional hairs emerging from cuticular perforation sites.
|
||
</paragraph>
|
||
<caption id="DF1754ADFFEE762F7F8DFCB9FDA896DC" ID-DOI="http://doi.org/10.5281/zenodo.6036947" ID-Zenodo-Dep="6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="10" pageNumber="75" startId="10.[151,250,1702,1726]" targetBox="[151,1435,181,1678]" targetPageId="10">
|
||
<paragraph id="8BD70425FFEE762F7F8DFCB9FDA896DC" blockId="10.[151,1437,1702,1979]" pageId="10" pageNumber="75">
|
||
<emphasis id="B91CD837FFEE762F7F8DFCB9FE9197D8" bold="true" box="[151,277,1702,1727]" pageId="10" pageNumber="75">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C687FA6FFEE762F7E07FCB9FE5097D9" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[285,468,1702,1726]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="10" pageNumber="75" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFEE762F7E07FCB9FE5097D9" box="[285,468,1702,1726]" italics="true" pageId="10" pageNumber="75">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CD837FFEE762F7EC7FCB8FDAB97D8" bold="true" box="[477,559,1703,1727]" pageId="10" pageNumber="75">
|
||
<taxonomicNameLabel id="A22F654CFFEE762F7EC7FCB8FDAB97D8" box="[477,559,1703,1727]" pageId="10" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female, ZMB 12271 (
|
||
<emphasis id="B91CD837FFEE762F7C3BFCB9FCD597D8" bold="true" box="[801,849,1702,1727]" pageId="10" pageNumber="75">A–E</emphasis>
|
||
) and ZMB 12349 (
|
||
<emphasis id="B91CD837FFEE762F7B04FCB8FBA997D8" bold="true" box="[1054,1069,1703,1727]" pageId="10" pageNumber="75">F</emphasis>
|
||
), right side (
|
||
<emphasis id="B91CD837FFEE762F7BACFCB9FB6397D9" bold="true" box="[1206,1255,1702,1726]" pageId="10" pageNumber="75">A–D</emphasis>
|
||
) and dorsal view
|
||
<emphasis id="B91CD837FFEE762F7F8DFCD5FF359785" bold="true" box="[151,177,1738,1762]" pageId="10" pageNumber="75">(E</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFEE762F7FA4FCD5FF539785" bold="true" box="[190,215,1738,1762]" pageId="10" pageNumber="75">F)</emphasis>
|
||
, SEM.
|
||
<emphasis id="B91CD837FFEE762F7E3CFCD5FEBB9785" bold="true" box="[294,319,1738,1762]" pageId="10" pageNumber="75">A.</emphasis>
|
||
Segments 1
|
||
<emphasis id="B91CD837FFEE762F7EA6FCD4FE4D9784" bold="true" box="[444,457,1739,1763]" pageId="10" pageNumber="75">–</emphasis>
|
||
3.
|
||
<emphasis id="B91CD837FFEE762F7EFEFCD4FE7E9784" bold="true" box="[484,506,1739,1763]" pageId="10" pageNumber="75">B.</emphasis>
|
||
Segments 4
|
||
<emphasis id="B91CD837FFEE762F7D63FCD4FD029784" bold="true" box="[633,646,1739,1763]" pageId="10" pageNumber="75">–</emphasis>
|
||
6.
|
||
<emphasis id="B91CD837FFEE762F7DBAFCD5FD3C9785" bold="true" box="[672,696,1738,1762]" pageId="10" pageNumber="75">C.</emphasis>
|
||
Segments 7
|
||
<emphasis id="B91CD837FFEE762F7C2DFCD4FCC09784" bold="true" box="[823,836,1739,1763]" pageId="10" pageNumber="75">–</emphasis>
|
||
8.
|
||
<emphasis id="B91CD837FFEE762F7C44FCD4FCF29784" bold="true" box="[862,886,1739,1763]" pageId="10" pageNumber="75">D.</emphasis>
|
||
Segments 9
|
||
<emphasis id="B91CD837FFEE762F7CEEFCD4FB859784" bold="true" box="[1012,1025,1739,1763]" pageId="10" pageNumber="75">–</emphasis>
|
||
11.
|
||
<emphasis id="B91CD837FFEE762F7B3DFCD4FBBC9784" bold="true" box="[1063,1080,1739,1763]" pageId="10" pageNumber="75">E</emphasis>
|
||
. Segments 10 and 11.
|
||
<emphasis id="B91CD837FFEE762F7A39FCD4FAB29784" bold="true" box="[1315,1334,1739,1763]" pageId="10" pageNumber="75">F.</emphasis>
|
||
Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in
|
||
<emphasis id="B91CD837FFEE762F7FC2FDBDFE8C96DD" bold="true" box="[216,264,1954,1978]" pageId="10" pageNumber="75">A–E</emphasis>
|
||
10 μm. Scale bar in
|
||
<emphasis id="B91CD837FFEE762F7EC0FDBCFE6D96DC" bold="true" box="[474,489,1955,1979]" pageId="10" pageNumber="75">F</emphasis>
|
||
5 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1754ADFFEF762E7F8DFCBBFE5796F2" ID-DOI="http://doi.org/10.5281/zenodo.6036949" ID-Zenodo-Dep="6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="11" pageNumber="76" startId="11.[151,250,1700,1724]" targetBox="[151,1436,181,1676]" targetPageId="11">
|
||
<paragraph id="8BD70425FFEF762E7F8DFCBBFE5796F2" blockId="11.[151,1436,1700,1941]" pageId="11" pageNumber="76">
|
||
<emphasis id="B91CD837FFEF762E7F8DFCBBFE9797DB" bold="true" box="[151,275,1700,1724]" pageId="11" pageNumber="76">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C687FA6FFEF762E7E03FCBBFE4A97DB" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[281,462,1700,1724]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="11" pageNumber="76" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFEF762E7E03FCBBFE4A97DB" box="[281,462,1700,1724]" italics="true" pageId="11" pageNumber="76">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CD837FFEF762E7ECFFCBBFDA197DB" bold="true" box="[469,549,1700,1724]" pageId="11" pageNumber="76">
|
||
<taxonomicNameLabel id="A22F654CFFEF762E7ECFFCBBFDA197DB" box="[469,549,1700,1724]" pageId="11" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male (ZMB 12350,
|
||
<emphasis id="B91CD837FFEF762E7DE1FCBBFC8997DB" bold="true" box="[763,781,1700,1724]" pageId="11" pageNumber="76">A</emphasis>
|
||
) and female (ZMB 12271,
|
||
<emphasis id="B91CD837FFEF762E7B39FCBBFBD497DB" bold="true" box="[1059,1104,1700,1724]" pageId="11" pageNumber="76">B–F</emphasis>
|
||
), right side (
|
||
<emphasis id="B91CD837FFEF762E7BC9FCBBFB6197DB" bold="true" box="[1235,1253,1700,1724]" pageId="11" pageNumber="76">A</emphasis>
|
||
) and ventral view (
|
||
<emphasis id="B91CD837FFEF762E7FBAFCD7FF499787" bold="true" box="[160,205,1736,1760]" pageId="11" pageNumber="76">B–F</emphasis>
|
||
), SEM.
|
||
<emphasis id="B91CD837FFEF762E7E38FCD7FEBF9787" bold="true" box="[290,315,1736,1760]" pageId="11" pageNumber="76">A.</emphasis>
|
||
Segment 11 with penile spines.
|
||
<emphasis id="B91CD837FFEF762E7D9BFCD7FD1C9787" bold="true" box="[641,664,1736,1760]" pageId="11" pageNumber="76">B.</emphasis>
|
||
Segments 10
|
||
<emphasis id="B91CD837FFEF762E7C3AFCD7FCA99787" bold="true" box="[800,813,1736,1760]" pageId="11" pageNumber="76">–</emphasis>
|
||
11.
|
||
<emphasis id="B91CD837FFEF762E7C48FCD7FCEE9787" bold="true" box="[850,874,1736,1760]" pageId="11" pageNumber="76">C.</emphasis>
|
||
Segments 1
|
||
<emphasis id="B91CD837FFEF762E7CFCFCD7FC779787" bold="true" box="[998,1011,1736,1760]" pageId="11" pageNumber="76">–</emphasis>
|
||
3.
|
||
<emphasis id="B91CD837FFEF762E7B16FCD7FBA19787" bold="true" box="[1036,1061,1736,1760]" pageId="11" pageNumber="76">D.</emphasis>
|
||
Segments 4
|
||
<emphasis id="B91CD837FFEF762E7BBAFCD7FB299787" bold="true" box="[1184,1197,1736,1760]" pageId="11" pageNumber="76">–</emphasis>
|
||
6.
|
||
<emphasis id="B91CD837FFEF762E7BDCFCD7FB599787" bold="true" box="[1222,1245,1736,1760]" pageId="11" pageNumber="76">E.</emphasis>
|
||
Segments 7 and 8.
|
||
<emphasis id="B91CD837FFEF762E7F8DFCF3FF2E9663" bold="true" box="[151,170,1772,1796]" pageId="11" pageNumber="76">F.</emphasis>
|
||
Segment 9
|
||
<emphasis id="B91CD837FFEF762E7E05FCF3FEA89663" bold="true" box="[287,300,1772,1796]" pageId="11" pageNumber="76">–</emphasis>
|
||
11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in
|
||
<emphasis id="B91CD837FFEF762E7E4CFD63FE0096F3" bold="true" box="[342,388,1916,1940]" pageId="11" pageNumber="76">A–F</emphasis>
|
||
10 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1754ADFFE876297F8DFC0CFF769640" ID-DOI="http://doi.org/10.5281/zenodo.6036953" ID-Zenodo-Dep="6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="12" pageNumber="77" startId="12.[151,250,1555,1579]" targetBox="[151,1436,181,1531]" targetPageId="12">
|
||
<paragraph id="8BD70425FFE876297F8DFC0CFF769640" blockId="12.[151,1437,1555,1831]" pageId="12" pageNumber="77">
|
||
<emphasis id="B91CD837FFE876297F8DFC0CFE97974C" bold="true" box="[151,275,1555,1579]" pageId="12" pageNumber="77">FIGURE 9.</emphasis>
|
||
<taxonomicName id="4C687FA6FFE876297E00FC0CFE4B974C" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[282,463,1555,1579]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="12" pageNumber="77" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFE876297E00FC0CFE4B974C" box="[282,463,1555,1579]" italics="true" pageId="12" pageNumber="77">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CD837FFE876297ECCFC0CFDA3974C" bold="true" box="[470,551,1555,1579]" pageId="12" pageNumber="77">
|
||
<taxonomicNameLabel id="A22F654CFFE876297ECCFC0CFDA3974C" box="[470,551,1555,1579]" pageId="12" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, last juvenile stage moulting to female, ZMB 12267, ventral (
|
||
<emphasis id="B91CD837FFE876297B84FC0CFB48974C" bold="true" box="[1182,1228,1555,1579]" pageId="12" pageNumber="77">B–E</emphasis>
|
||
), optical section (
|
||
<emphasis id="B91CD837FFE876297A9EFC0CFA12974C" bold="true" box="[1412,1430,1555,1579]" pageId="12" pageNumber="77">A</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE876297F8DFC28FF229728" bold="true" box="[151,166,1591,1615]" pageId="12" pageNumber="77">F</emphasis>
|
||
), and dorsal side (
|
||
<emphasis id="B91CD837FFE876297E7CFC28FEFE9728" bold="true" box="[358,378,1591,1615]" pageId="12" pageNumber="77">G</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE876297E9DFC28FE1F9728" bold="true" box="[391,411,1591,1615]" pageId="12" pageNumber="77">H</emphasis>
|
||
).
|
||
<emphasis id="B91CD837FFE876297EABFC28FE4E9728" bold="true" box="[433,458,1591,1615]" pageId="12" pageNumber="77">A.</emphasis>
|
||
Habitus.
|
||
<emphasis id="B91CD837FFE876297D37FC28FDC09728" bold="true" box="[557,580,1591,1615]" pageId="12" pageNumber="77">B.</emphasis>
|
||
Segments 1–4.
|
||
<emphasis id="B91CD837FFE876297DF3FC28FD7F9728" bold="true" box="[745,763,1591,1615]" pageId="12" pageNumber="77">C</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFE876297C13FC28FCA69728" bold="true" box="[777,802,1591,1615]" pageId="12" pageNumber="77">D.</emphasis>
|
||
Segments 5–8 at different focal levels showing aspects of the juvenile (
|
||
<emphasis id="B91CD837FFE876297FEDFC44FE8D9714" bold="true" box="[247,265,1627,1651]" pageId="12" pageNumber="77">C</emphasis>
|
||
) and adult (
|
||
<emphasis id="B91CD837FFE876297E98FC44FE109714" bold="true" box="[386,404,1627,1651]" pageId="12" pageNumber="77">D</emphasis>
|
||
) cuticle.
|
||
<emphasis id="B91CD837FFE876297EEBFC44FD8C9714" bold="true" box="[497,520,1627,1651]" pageId="12" pageNumber="77">E.</emphasis>
|
||
Segments 8–11.
|
||
<emphasis id="B91CD837FFE876297DA9FC44FD429714" bold="true" box="[691,710,1627,1651]" pageId="12" pageNumber="77">F.</emphasis>
|
||
Segments 9–11.
|
||
<emphasis id="B91CD837FFE876297C68FC44FC089714" bold="true" box="[882,908,1627,1651]" pageId="12" pageNumber="77">G.</emphasis>
|
||
Segments 1–4.
|
||
<emphasis id="B91CD837FFE876297B31FC44FBC29714" bold="true" box="[1067,1094,1627,1651]" pageId="12" pageNumber="77">H.</emphasis>
|
||
Segments 4–7. Abbreviations: bs, button-like structures;
|
||
<superScript id="7C1DA96DFFE876297E67FC60FE0C97EA" attach="right" box="[381,392,1663,1677]" fontSize="6" pageId="12" pageNumber="77">fe</superScript>
|
||
, refers to female character;
|
||
<superScript id="7C1DA96DFFE876297DBDFC60FD3D97EA" attach="right" box="[679,697,1663,1677]" fontSize="6" pageId="12" pageNumber="77">juv</superScript>
|
||
, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in
|
||
<emphasis id="B91CD837FFE876297B01FCF4FBA99664" bold="true" box="[1051,1069,1771,1795]" pageId="12" pageNumber="77">A</emphasis>
|
||
100 µm. Scale bar in
|
||
<emphasis id="B91CD837FFE876297A12FCF4FA939664" bold="true" box="[1288,1303,1771,1795]" pageId="12" pageNumber="77">F</emphasis>
|
||
50 µm, valid for
|
||
<emphasis id="B91CD837FFE876297FA1FD10FF699640" bold="true" box="[187,237,1807,1831]" pageId="12" pageNumber="77">B–H</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD70425FFE876297FDDFD49FDB296BB" blockId="12.[151,1436,1877,2012]" pageId="12" pageNumber="77">
|
||
Segment 3 (
|
||
<figureCitation id="135318A0FFE876297E55FD4AFE429617" box="[335,454,1877,1904]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="12" pageNumber="77">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE876297EC9FD49FE709617" box="[467,500,1878,1904]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="12" pageNumber="77">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE876297D1BFD49FDC59617" box="[513,577,1878,1904]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="12" pageNumber="77">4A,D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE876297D57FD4AFD0F9617" box="[589,651,1877,1904]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="12" pageNumber="77">5A,C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE876297D8DFD4AFD399608" box="[663,701,1877,1903]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="12" pageNumber="77">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE876297DD2FD49FD699617" box="[712,749,1878,1904]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="12" pageNumber="77">7A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE876297DE3FD49FC999617" box="[761,797,1878,1904]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="12" pageNumber="77">8C</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE876297C33FD4AFCF99617" box="[809,893,1877,1904]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="12" pageNumber="77" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">Table 3</tableCitation>
|
||
) without spines and tubes. Sensory spot present posteriorly in subdorsal and sublateral position. Type-1 gland cell outlet occurring anteriorly in middorsal and centrally in ventromedial position. Cuticular hairs more abundant than on the previous segment and organised similarly on the remaining segments.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDFA88FE62919E" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
Segment 4 (
|
||
<figureCitation id="135318A0FFE976287E51FA88FE4491D6" box="[331,448,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287ED1FA88FE6891D6" box="[459,492,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="13" pageNumber="78">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287EEDFA88FDB391D6" box="[503,567,151,177]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4A,D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D5BFA89FDFB91D6" box="[577,639,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5A,C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D90FA88FD2B91D6" box="[650,687,151,177]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287DA3FA88FD5891D6" box="[697,732,151,177]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287DFDFA88FC8F91D6" box="[743,779,151,177]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8D</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE976287C0CFA88FCF091D6" box="[790,884,151,177]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6EA319EFFE976287C65FA88FC0991D6" box="[895,909,151,177]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) with a middorsal acicular spine. Sensory spots lacking on this segment. Type-1 gland cell outlet occurring anteriorly in paradorsal and centrally in paraventral to almost ventromedial position.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDFB1CFB3290EE" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
Segment 5 (
|
||
<figureCitation id="135318A0FFE976287E4AFB1CFE4C907A" box="[336,456,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287ECFFB1CFE72907A" box="[469,502,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="13" pageNumber="78">3E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D1EFB1CFDAC907A" box="[516,552,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D2FFB1DFDF7907A" box="[565,627,258,285]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5A,C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D9AFB1CFD22907A" box="[640,678,259,285]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
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|
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<figureCitation id="135318A0FFE976287DA8FB1CFD51907A" box="[690,725,259,285]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7B</figureCitation>
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,
|
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<figureCitation id="135318A0FFE976287DF8FB1CFC83907A" box="[738,775,259,285]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8D</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE976287C0EFB1CFCF1907A" box="[788,885,259,285]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
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,
|
||
<tableCitation id="C6EA319EFFE976287C98FB1CFC14907A" box="[898,912,259,285]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) with lateroventral tube. Sensory spot present posteriorly in subdorsal, sublateral to midlateral position and almost centrally in ventromedial position. Sublateral to midlateral spot of segments 5 and 6 slightly more dorsally than spot of segment 3. Type-1 gland cell outlets occurring anteriorly in middorsal and centrally in paraventral to almost ventromedial position.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDFB8CFE62937E" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
Segment 6 (
|
||
<figureCitation id="135318A0FFE976287E49FB8CFE4990CA" box="[339,461,403,429]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2A,B</figureCitation>
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||
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|
||
<figureCitation id="135318A0FFE976287EC6FB8CFD8890CA" box="[476,524,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="13" pageNumber="78">3F,J</figureCitation>
|
||
,
|
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<figureCitation id="135318A0FFE976287D00FB8CFDB990CA" box="[538,573,403,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D56FB8DFD0C90CA" box="[588,648,402,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5B,C</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D82FB8CFD3990CA" box="[664,701,403,429]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287DD1FB8CFD6A90CA" box="[715,750,403,429]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7B</figureCitation>
|
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,
|
||
<figureCitation id="135318A0FFE976287DE7FB8CFCA590CA" box="[765,801,403,429]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8D</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE976287C2AFB8CFC1790CA" box="[816,915,403,429]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6EA319EFFE976287CB8FB8CFC3490CA" box="[930,944,403,429]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) with middorsal and lateroventral acicular spine. Sensory spot present posteriorly in paradorsal and sublateral to midlateral position and almost centrally in ventromedial position. Type-1 gland cell outlet occurring anteriorly in paradorsal and centrally in paraventral to almost ventromedial position.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDF83CFAC693E2" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
Segment 7 (
|
||
<figureCitation id="135318A0FFE976287E5DF83CFE3F935A" box="[327,443,547,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287EDEF83CFE70935A" box="[452,500,547,573]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="13" pageNumber="78">3F,J</figureCitation>
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,
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<figureCitation id="135318A0FFE976287EE7F83CFD9B935A" box="[509,543,547,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4B</figureCitation>
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<figureCitation id="135318A0FFE976287D33F83DFDE1935A" box="[553,613,546,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5B,C</figureCitation>
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<figureCitation id="135318A0FFE976287D74F83CFD17935A" box="[622,659,547,573]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
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<figureCitation id="135318A0FFE976287D86F83CFD3A935A" box="[668,702,547,573]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7C</figureCitation>
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<figureCitation id="135318A0FFE976287DD2F83CFD68935A" box="[712,748,547,573]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8D</figureCitation>
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||
;
|
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<tableCitation id="C6EA319EFFE976287DEFF83CFCD6935A" box="[757,850,547,573]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
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<tableCitation id="C6EA319EFFE976287C41F83CFCED935A" box="[859,873,547,573]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) with sublateral to midlateral tube and lateroventral acicular spine. Sensory spot present posteriorly in subdorsal position and almost centrally in ventromedial position. Type-1 gland cell outlet occurring anteriorly in middorsal and paraventral to almost ventromedial position.
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||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDF890FBB19272" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
Segment 8 (
|
||
<figureCitation id="135318A0FFE976287E48F890FE4F93CE" box="[338,459,655,681]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2A,B</figureCitation>
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<figureCitation id="135318A0FFE976287EC0F890FD8D93CE" box="[474,521,655,681]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="13" pageNumber="78">3F,J</figureCitation>
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<figureCitation id="135318A0FFE976287D02F890FDD993CE" box="[536,605,655,681]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4B,I,J</figureCitation>
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<figureCitation id="135318A0FFE976287D71F891FD0A93CE" box="[619,654,654,681]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5B</figureCitation>
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<figureCitation id="135318A0FFE976287D86F890FD4693CE" box="[668,706,655,681]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
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<figureCitation id="135318A0FFE976287DD5F890FD7693CE" box="[719,754,655,681]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7C</figureCitation>
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<figureCitation id="135318A0FFE976287C1BF890FCBA93CE" box="[769,830,655,681]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8D,E</figureCitation>
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;
|
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<tableCitation id="C6EA319EFFE976287C56F890FC2A93CE" box="[844,942,655,681]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
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<tableCitation id="C6EA319EFFE976287CA7F890FC4F93CE" box="[957,971,655,681]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) with a middorsal acicular spine, lateral accessory tube and lateroventral acicular spine. Middorsal spine of female much longer (92–98 µm) than that of male (54–79 µm) (
|
||
<tableCitation id="C6EA319EFFE976287E89F8C8FE6A9396" box="[403,494,727,753]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Table 2</tableCitation>
|
||
). Sensory spot present posteriorly in paradorsal position. Type-1 gland cell outlet occurring anteriorly in paradorsal and paraventral to almost ventromedial position.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDF900FF7D92C2" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
Segment 9 (
|
||
<figureCitation id="135318A0FFE976287E4EF900FE24925E" box="[340,416,799,825]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287EAAF900FE7E925E" box="[432,506,799,825]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="13" pageNumber="78">3F,G,J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D13F900FDED925E" box="[521,617,799,825]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4B,C,I,J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D6DF901FD4E925E" box="[631,714,798,825]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5B,F,G</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287DC3F900FD7A925E" box="[729,766,799,825]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287C17F900FCB5925E" box="[781,817,799,825]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287C5AF900FCFD925E" box="[832,889,799,825]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8E,F</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE976287C92F900FC6F925E" box="[904,1003,799,825]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6EA319EFFE976287CE0F900FB8C925E" box="[1018,1032,799,825]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) with sublateral to midlateral tube and lateroventral acicular spine. Sensory spot present posteriorly in paradorsal and centrally in laterodorsal and ventrolateral position. Type-1 gland cell outlet occurring anteriorly in paradorsal and centrally in paraventral position.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDF9B0FEFC95A2" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
Segment 10 (
|
||
<figureCitation id="135318A0FFE976287E43F9B0FE2592AE" box="[345,417,943,969]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287EB6F9B0FE7992AE" box="[428,509,943,969]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="13" pageNumber="78">3G,H,J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D12F9B0FDF292AE" box="[520,630,943,969]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4B,C,E–J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D9AF9B1FD5292AE" box="[640,726,942,969]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5B,D,E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287DFBF9B0FC8292AE" box="[737,774,943,969]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287C0AF9B0FCD592AE" box="[784,849,943,969]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7D–F</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287C41F9B0FC1192AE" box="[859,917,943,969]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8B,F</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE976287C85F9B0FC7A92AE" box="[927,1022,943,969]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6EA319EFFE976287B13F9B0FB9392AE" box="[1033,1047,943,969]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) without spines but with sublateral to midlateral straight short tube with basal cuticular cylindrical reinforcement structure below trunk cuticle in male (
|
||
<figureCitation id="135318A0FFE976287FBAF9E8FE8B9576" box="[160,271,1015,1041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">Fig. 4F,H</figureCitation>
|
||
). Female without tube but with sublateral to midlateral subcuticular, conical structure (
|
||
<figureCitation id="135318A0FFE976287BF8F9E8FADC9576" box="[1250,1368,1014,1041]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">Figs 5D,E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287A7EF9E8FA109576" box="[1380,1428,1015,1041]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="13" pageNumber="78">10C</figureCitation>
|
||
) ending in short apical cuticular duct through trunk cuticle, with basal cuticular cylindrical reinforcement structure below trunk cuticle (
|
||
<figureCitation id="135318A0FFE976287E99FE20FE72953E" box="[387,502,1086,1113]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">Fig. 5D,E</figureCitation>
|
||
), opening to outside via fringed pore (
|
||
<figureCitation id="135318A0FFE976287CB6FE20FB9F953E" box="[940,1051,1087,1113]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">Fig. 7E,F</figureCitation>
|
||
). Sensory spot present posteriorly in subdorsal and ventrolateral position. Type-1 gland cell outlet occurring anteriorly in middorsal and centrally in middorsal and paraventral position. Posterior margin of each sternal plate progressing more posteriorly midventrally than lateroventrally.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDFED0FEE69482" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
Segment 11 (
|
||
<figureCitation id="135318A0FFE976287E7AFED0FE28958E" box="[352,428,1231,1257]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287EA1FED0FE6B958E" box="[443,495,1231,1257]" captionStart="FIGURE 3" captionStartId="5.[151,250,1678,1702]" captionTargetBox="[153,1436,182,1654]" captionTargetId="figure-17@5.[151,1436,181,1654]" captionTargetPageId="5" captionText="FIGURE 3. Echinoderes goku sp. nov., male paratype ZMB 12334 (A, C), juvenile non-type ZMB 12347 (B, D), and male holotype MACN-In 43755 (E–J), in ventral (A), lateral (B, D), and dorsal view (C, E–J), DIC. A–D. Habitus of specimens infected with fungus before (A, B) and after treatment with formaldehyde for two weeks (C, D). E. Segments 1–5. F, J. Segments 6–9 at different focal levels. G–I. Segments 10 (G), 10 and 11 (H), and 11 (I) at different focal levels. Abbreviations: fu, fungus; hr, trunk cuticular hair; mds, middorsal spine; pa, pachycycli; pl, placid; ppf, primary pectinate fringe; s, segment; sdt, subdorsal tube; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number.Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in D 100 μm, valid for A–D. Scale bar in F 50 µm, valid for E–J." figureDoi="http://doi.org/10.5281/zenodo.6036925" httpUri="https://zenodo.org/record/6036925/files/figure.png" pageId="13" pageNumber="78">3H,I</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287EE5FED0FDBC958E" box="[511,568,1231,1257]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4F–J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D5CFED1FD07958E" box="[582,643,1230,1257]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5D,E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D88FED0FD33958E" box="[658,695,1231,1257]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287DDFFED0FC85958E" box="[709,769,1231,1257]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7D,E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287C0AFED0FCE1958E" box="[784,869,1231,1257]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8A,B,F</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE976287C6EFED0FC53958E" box="[884,983,1231,1257]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6EA319EFFE976287CFCFED0FC70958E" box="[998,1012,1231,1257]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) with lateral terminal spine, varying considerably in length (146–208 µm) in both females and males (
|
||
<tableCitation id="C6EA319EFFE976287C6BFEECFC4D946A" box="[881,969,1267,1293]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Table 2</tableCitation>
|
||
). Straight lateral terminal accessory spine present in females (
|
||
<figureCitation id="135318A0FFE976287E68FF08FE6D9456" box="[370,489,1303,1329]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2C,D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287EEFFF08FDA49456" box="[501,544,1303,1329]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4I,J</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D36FF09FDD59456" box="[556,593,1302,1329]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="13" pageNumber="78">5D</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D46FF08FD059456" box="[604,641,1303,1329]" captionStart="FIGURE 6" captionStartId="8.[151,250,1265,1289]" captionTargetBox="[151,1436,181,1241]" captionTargetId="figure-20@8.[151,1436,181,1241]" captionTargetPageId="8" captionText="FIGURE 6. Echinoderes goku sp. nov., female, ZMB 12271 (A, C) and ZMB 12272 (B), in laterodorsal (A), ventral (B), and dorsal-right (C) view, SEM. A. Habitus. B. Mouth cone. White arrow points to bifurcated tip of posterior spinose process. C. Two neighbouring introvert sector. Abbreviations: asp, anterior spinose process; in, introvert; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mc, mouth cone; mds, middorsal spine; oos, outer oral style; psc, primary spinoscalid; psp, posterior spinose process; s, segment; sc, spinoscalid; slt, sublateral tube; tr, trichoscalid. Digits after abbreviations indicate the corresponding segment (A) or introvert ring number (C). Scale bar in A 100 μm. Scale bar in B and C 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036942" httpUri="https://zenodo.org/record/6036942/files/figure.png" pageId="13" pageNumber="78">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287D97FF08FD4D9456" box="[653,713,1303,1329]" captionStart="FIGURE 7" captionStartId="10.[151,250,1702,1726]" captionTargetBox="[151,1435,181,1678]" captionTargetId="figure-20@10.[151,1435,181,1678]" captionTargetPageId="10" captionText="FIGURE 7. Echinoderes goku sp. nov., female, ZMB 12271 (A–E) and ZMB 12349 (F), right side (A–D) and dorsal view (E, F), SEM. A. Segments 1–3. B. Segments 4–6. C. Segments 7–8. D. Segments 9–11. E. Segments 10 and 11. F. Fringe of sublateral, subcuticular conical structure opening on segment 10. Abbreviations: co, fringe of sublateral, subcuticular conical structure opening on segment 10; gc1, type-1 gland cell outlet; hr, trunk cuticular hair; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; mds, middorsal spine; pf, primary pectinate fringe; s, segment; sdt, subdorsal tube (lost on segment 2); sf, secondary pectinate fringe; slt, sublateral to midlateral tube; te, tergal extension. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–E 10 μm. Scale bar in F 5 µm." figureDoi="http://doi.org/10.5281/zenodo.6036947" httpUri="https://zenodo.org/record/6036947/files/figure.png" pageId="13" pageNumber="78">7D,E</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287DCFFF08FD729456" box="[725,758,1303,1329]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8F</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE976287C1BFF08FCE59456" box="[769,865,1303,1329]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6EA319EFFE976287C77FF08FCFF9456" box="[877,891,1303,1329]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
) and three penile spines appearing in males, one bent, tubular, longer, in lateral accessory position, one shorter, broader, almost triangular, slightly more dorsally, and one bent, tubular, longer sublaterally (
|
||
<figureCitation id="135318A0FFE976287D61FF40FD77941E" box="[635,755,1375,1401]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1630]" captionTargetBox="[151,1436,198,1582]" captionTargetId="figure-20@4.[151,1436,198,1582]" captionTargetPageId="4" captionText="FIGURE 2. Line art illustrations of Echinoderes goku sp. nov., A, B, male holotype (MACN-In 43755), C, D, female paratype (MACN-In 43756b). A. Dorsal view. B. Ventral view. C, D. Segments 10 and 11, dorsal (C) and ventral view (D). Placids illustrated as appearing in SEM. Sublateral structures often hidden by the neighbouring body cuticle in the more or less dorsoventral mount and often only illustrated for one side. Abbreviations: co, female-specific, subcuticular conical structure; hr, cuticular hair; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mdgc, middorsal gland cell outlet; mds, middorsal spine; pa, pachycycli; pe, penile spine; pl, placid; pvp, paraventral patch of cuticular hairs; sdt, subdorsal tube; slt, sublateral to midlateral tube; te, tergal extension; vlp, ventrolateral to paraventral patch of cuticular hairs; vlt, ventrolateral tube; vmss, ventromedial sensory spot. Digits after abbreviations indicate the corresponding segment number. Scale bar: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6036917" httpUri="https://zenodo.org/record/6036917/files/figure.png" pageId="13" pageNumber="78">Figs 2A,B</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287C1BFF40FCBE941E" box="[769,826,1375,1401]" captionStart="FIGURE 4" captionStartId="6.[151,250,1646,1670]" captionTargetBox="[166,1421,181,1622]" captionTargetId="figure-20@6.[166,1421,181,1622]" captionTargetPageId="6" captionText="FIGURE 4. Echinoderes goku sp. nov., male holotype MACN-In 43755 (A–C), male paratype ZMB 12318 (D, F–H) and female paratypes ZMB 12337 (E), ZMB 12265 (I, K), and MACN-In 43756b (J), ventral view (A–E, I, K), dorsal view (F–H), and optical section (J), extended focus image from four images (D, K), DIC. A. Segments 1–6. B. Segments 7–10. C. Segments 9–11. D. Segments 1–4. E. Segment 10. F–H. Segments 10 and 11 at different focal levels. I, J. Segments 8–11. K. Mouth cone with outer oral styles. Abbreviations: bs, button-like structures; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mvpl, midventral placid; oos, outer oral style; ov, ovarium with egg; pa, pachycycli; peb, broad penile spine; pet, tubular penile spine; pvp, paraventral patch of short cuticular hairs; re, cuticular reinforcement; s, segment; slt, sublateral tube; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–D, I, J. Scale bar in G 50 µm, valid for F, G. Scale bar in H 20 µm. Scale bar in K 30 µm, valid for E, K." figureDoi="http://doi.org/10.5281/zenodo.6036933" httpUri="https://zenodo.org/record/6036933/files/figure.png" pageId="13" pageNumber="78">4F,G</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFE976287C52FF40FCE9941E" box="[840,877,1375,1401]" captionStart="FIGURE 8" captionStartId="11.[151,250,1700,1724]" captionTargetBox="[151,1436,181,1676]" captionTargetId="figure-17@11.[151,1436,181,1676]" captionTargetPageId="11" captionText="FIGURE 8. Echinoderes goku sp. nov., male (ZMB 12350, A) and female (ZMB 12271, B–F), right side (A) and ventral view (B–F), SEM. A. Segment 11 with penile spines. B. Segments 10–11. C. Segments 1–3. D. Segments 4–6. E. Segments 7 and 8. F. Segment 9–11. Abbreviations: dv, attachment site of dorsoventral muscle; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; peb, broad penile spine; pet, tubular penile spine; pf, primary pectinate fringe; pvp, paraventral patch of short cuticular hairs; s, segment; sf, secondary pectinate fringe; te, tergal extension; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. Scale bar in A–F 10 μm." figureDoi="http://doi.org/10.5281/zenodo.6036949" httpUri="https://zenodo.org/record/6036949/files/figure.png" pageId="13" pageNumber="78">8A</figureCitation>
|
||
;
|
||
<tableCitation id="C6EA319EFFE976287C60FF40FC5F941E" box="[890,987,1375,1401]" captionStart="TABLE 2" captionStartId="14.[151,239,518,544]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in µm) for holotype and paratypes of Echinoderes goku sp. nov. from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number." httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="13" pageNumber="78" tableUuid="DF1754ADFFEA762B7F8DF819FD2E93EB">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="C6EA319EFFE976287CF3FF40FC7D941E" box="[1001,1017,1375,1401]" captionStart="TABLE 3" captionStartId="16.[151,239,691,717]" captionTargetBox="[165,1982,947,1502]" captionTargetPageId="16" captionText="TABLE 3. Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in Echinoderes goku sp. nov. from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial." httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="13" pageNumber="78" tableUuid="DF1754ADFFF476357F8DF8ACFE6B92C2">3</tableCitation>
|
||
). Sensory spot present posteriorly in subdorsal position. Males often with two ventrolateral pores, unclear if belonging to a sensory spot. Two type-1 gland cell outlets occurring anteriorly and centrally behind each other in middorsal position. Tergal extensions triangular-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDFFF0FDE29712" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
<materialsCitation id="3B000E78FFE976287FDDFFF0FDE69712" ID-GBIF-Occurrence="3464450301" accessionNumber="ZMB 12323" collectionCode="ZMB" pageId="13" pageNumber="78" specimenCount="41">
|
||
Among the
|
||
<specimenCount id="9D6ECFACFFE976287E52FFF0FE5A976E" box="[328,478,1519,1545]" pageId="13" pageNumber="78" type="generic">39 specimens</specimenCount>
|
||
studied by LM and
|
||
<specimenCount id="9D6ECFACFFE976287DADFFF0FCED976E" box="[695,873,1519,1545]" pageId="13" pageNumber="78" type="generic">three specimens</specimenCount>
|
||
investigatable with SEM no variation of characters was found except that the position of sensory spots on segment 5 and 6 and of tubes on segments 7 and 9 varied slightly from more sublaterally to more midlaterally. Probably,
|
||
<specimenCount id="9D6ECFACFFE976287C57FC28FC619736" box="[845,997,1591,1617]" pageId="13" pageNumber="78" type="generic">one specimen</specimenCount>
|
||
(
|
||
<collectionCode id="ED799CE0FFE976287CE8FC28FBB49736" box="[1010,1072,1591,1617]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="13" pageNumber="78" type="Museum">ZMB</collectionCode>
|
||
12323) possessed a lateroventral spine only on the right side of segment 7
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFE976287FDDFC61FED396BA" blockId="13.[151,1437,150,2013]" pageId="13" pageNumber="78">
|
||
<emphasis id="B91CD837FFE976287FDDFC61FEF897FE" bold="true" box="[199,380,1662,1689]" pageId="13" pageNumber="78">Juvenile stages</emphasis>
|
||
. Five specimens could be identified as juvenile stages based on their thin trunk cuticle, the postmarginal spicula, a gap in the arrangement of cuticular hairs and postmarginal spicula about ventromedially to paraventrally where the dorsoventral muscles attach (
|
||
<figureCitation id="135318A0FFE976287DEBFCD8FCF89786" box="[753,892,1735,1761]" captionStart="FIGURE 9" captionStartId="12.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-20@12.[151,1436,181,1531]" captionTargetPageId="12" captionText="FIGURE 9. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, ventral (B–E), optical section (A, F), and dorsal side (G, H). A. Habitus. B. Segments 1–4. C, D. Segments 5–8 at different focal levels showing aspects of the juvenile (C) and adult (D) cuticle. E. Segments 8–11. F. Segments 9–11. G. Segments 1–4. H. Segments 4–7.Abbreviations: bs, button-like structures; fe, refers to female character; juv, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in A 100 µm. Scale bar in F 50 µm, valid for B–H." figureDoi="http://doi.org/10.5281/zenodo.6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="13" pageNumber="78">Fig. 9B,C,E</figureCitation>
|
||
), papillate sensory spots (
|
||
<figureCitation id="135318A0FFE976287BBAFCD8FA839786" box="[1184,1287,1735,1761]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="13" pageNumber="78">Fig. 10A</figureCitation>
|
||
) if present at all, the lack of recognisable gland cells, an acicular spine middorsally on segment 9 (
|
||
<figureCitation id="135318A0FFE976287B2CFCF4FB1E9662" box="[1078,1178,1771,1797]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="13" pageNumber="78">Fig. 10A</figureCitation>
|
||
) and lateral accessorily on segment 10 (
|
||
<figureCitation id="135318A0FFE976287E51FD10FE35964E" box="[331,433,1807,1833]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="13" pageNumber="78">Fig. 10C</figureCitation>
|
||
), and a well developed lateral terminal accessory spine (
|
||
<figureCitation id="135318A0FFE976287B37FD10FB32964E" box="[1069,1206,1807,1833]" captionStart="FIGURE 9" captionStartId="12.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-20@12.[151,1436,181,1531]" captionTargetPageId="12" captionText="FIGURE 9. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, ventral (B–E), optical section (A, F), and dorsal side (G, H). A. Habitus. B. Segments 1–4. C, D. Segments 5–8 at different focal levels showing aspects of the juvenile (C) and adult (D) cuticle. E. Segments 8–11. F. Segments 9–11. G. Segments 1–4. H. Segments 4–7.Abbreviations: bs, button-like structures; fe, refers to female character; juv, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in A 100 µm. Scale bar in F 50 µm, valid for B–H." figureDoi="http://doi.org/10.5281/zenodo.6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="13" pageNumber="78">Fig. 9A,E,F</figureCitation>
|
||
). It was not possible to identify different life history stages with certainty, because the specimens came out crumpled, twisted, and folded during mounting due to their thin cuticle. In all specimens, 11 trunk segments delimited by postmarginal spicula could be recognised, whereas sensory spots were difficult to trace. Four specimens seemed to represent late juvenile stages if not the last stage, and the fifth was moulting to a female, and this is subsequently described in more detail and documented.
|
||
</paragraph>
|
||
<caption id="DF1754ADFFEA762B7F8DF819FD2E93EB" ID-Table-UUID="DF1754ADFFEA762B7F8DF819FD2E93EB" httpUri="http://table.plazi.org/id/DF1754ADFFEA762B7F8DF819FD2E93EB" pageId="14" pageNumber="79" startId="14.[151,239,518,544]" targetBox="[165,2021,667,1621]" targetIsTable="true" targetPageId="14">
|
||
<paragraph id="8BD70425FFEA762B7F8DF819FD2E93EB" blockId="14.[151,2041,518,652]" pageId="14" pageNumber="79">
|
||
<emphasis id="B91CD837FFEA762B7F8DF819FE8D9347" bold="true" box="[151,265,518,544]" pageId="14" pageNumber="79">TABLE 2.</emphasis>
|
||
Measurements (in µm) for holotype and paratypes of
|
||
<taxonomicName id="4C687FA6FFEA762B7C59F819FB859347" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[835,1025,518,544]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="14" pageNumber="79" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFEA762B7C59F819FB859347" box="[835,1025,518,544]" italics="true" pageId="14" pageNumber="79">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CD837FFEA762B7B1DF819FBD89347" bold="true" box="[1031,1116,518,544]" pageId="14" pageNumber="79">
|
||
<taxonomicNameLabel id="A22F654CFFEA762B7B1DF819FBD89347" box="[1031,1116,518,544]" pageId="14" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Argentina. Abbreviations: –, character not present; ♀, female; ♂, male; lat, lateral accessory tube; ltas, lateral terminal accessory spine; lts; lateral terminal spine; lvs/t, lateroventral spine or tube; mds, middorsal spine; n. a., character not measurable; peb, broad penile spine; pet, tubular penile spine; s, segment length; sdt, subdorsal tube; slt, sublateral tube; sw, standard width; tl, trunk length; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD70425FFEA762B7FBFF884F8279732" pageId="14" pageNumber="79">
|
||
<table id="F968F685FFEA89DA7FBFF884F8619732" box="[165,2021,667,1621]" colsContinueIn="15.[151,2006,776,1459]" gridcols="11" gridrows="21" pageId="14" pageNumber="79">
|
||
<tr id="35580667FFEA89DA7FBFF884F86193BE" box="[165,2021,667,729]" gridrow="0" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFF884FE9093BE" box="[165,276,667,729]" gridcol="0" gridrow="0" pageId="14" pageNumber="79">
|
||
<emphasis id="B91CD837FFEA762B7FBFF884FE8993D3" bold="true" box="[165,269,667,692]" pageId="14" pageNumber="79">Specimen</emphasis>
|
||
</th>
|
||
<th id="76896F1BFFEA89DA7E48F884FE4E93BE" box="[338,458,667,729]" gridcol="1" gridrow="0" pageId="14" pageNumber="79">♂ holotype, MACN-In</th>
|
||
<th id="76896F1BFFEA89DA7D0FF884FD0893BE" box="[533,652,667,729]" gridcol="2" gridrow="0" pageId="14" pageNumber="79">♂ paratype, MACN-In</th>
|
||
<th id="76896F1BFFEA89DA7DCDF884FCD093BE" box="[727,852,667,729]" gridcol="3" gridrow="0" pageId="14" pageNumber="79">♂ paratype, ZMB 12262</th>
|
||
<th id="76896F1BFFEA89DA7C69F884FC6B93BE" box="[883,1007,667,729]" gridcol="4" gridrow="0" pageId="14" pageNumber="79">♂ non-type, ZMB 12263</th>
|
||
<th id="76896F1BFFEA89DA7B15F884FB0F93BE" box="[1039,1163,667,729]" gridcol="5" gridrow="0" pageId="14" pageNumber="79">♂ non-type, ZMB 12266</th>
|
||
<th id="76896F1BFFEA89DA7BB4F884FAA193BE" box="[1198,1317,667,729]" gridcol="6" gridrow="0" pageId="14" pageNumber="79">♀ paratype, MACN-In</th>
|
||
<th id="76896F1BFFEA89DA7A74F884FA6193BE" box="[1390,1509,667,729]" gridcol="7" gridrow="0" pageId="14" pageNumber="79">♀ paratype, MACN-In</th>
|
||
<th id="76896F1BFFEA89DA792BF884F92993BE" box="[1585,1709,667,729]" gridcol="8" gridrow="0" pageId="14" pageNumber="79">♀ paratype, ZMB 12264</th>
|
||
<th id="76896F1BFFEA89DA79D7F884F8CD93BE" box="[1741,1865,667,729]" gridcol="9" gridrow="0" pageId="14" pageNumber="79">♀ paratype, ZMB 12265</th>
|
||
<th id="76896F1BFFEA89DA7872F884F86193BE" box="[1896,2021,667,729]" gridcol="10" gridrow="0" pageId="14" pageNumber="79">♀ paratype, ZMB 12341</th>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFF8FCF861939A" box="[165,2021,739,765]" gridrow="1" pageId="14" pageNumber="79" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="76896F1BFFEA89DA7FBFF8FCFE90939A" box="[165,276,739,765]" gridcol="0" gridrow="1" pageId="14" pageNumber="79">
|
||
<emphasis id="B91CD837FFEA762B7FBFF8FCFE90939B" bold="true" box="[165,276,739,764]" pageId="14" pageNumber="79">Character</emphasis>
|
||
</th>
|
||
<td id="76896F1BFFEA89DA7E48F8FCFE4E939A" box="[338,458,739,765]" gridcol="1" gridrow="1" pageId="14" pageNumber="79">43755</td>
|
||
<td id="76896F1BFFEA89DA7D0FF8FCFD08939A" box="[533,652,739,765]" gridcol="2" gridrow="1" pageId="14" pageNumber="79">43756a</td>
|
||
<td id="76896F1BFFEA89DA7BB4F8FCFAA1939A" box="[1198,1317,739,765]" gridcol="6" gridrow="1" pageId="14" pageNumber="79">43756b</td>
|
||
<td id="76896F1BFFEA89DA7A74F8FCFA61939A" box="[1390,1509,739,765]" gridcol="7" gridrow="1" pageId="14" pageNumber="79">43756c</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFF913F8619242" box="[165,2021,780,805]" gridrow="2" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFF913FE909242" box="[165,276,780,805]" gridcol="0" gridrow="2" pageId="14" pageNumber="79">tl</th>
|
||
<td id="76896F1BFFEA89DA7E48F913FE4E9242" box="[338,458,780,805]" gridcol="1" gridrow="2" pageId="14" pageNumber="79">277</td>
|
||
<td id="76896F1BFFEA89DA7D0FF913FD089242" box="[533,652,780,805]" gridcol="2" gridrow="2" pageId="14" pageNumber="79">271</td>
|
||
<td id="76896F1BFFEA89DA7DCDF913FCD09242" box="[727,852,780,805]" gridcol="3" gridrow="2" pageId="14" pageNumber="79">244</td>
|
||
<td id="76896F1BFFEA89DA7C69F913FC6B9242" box="[883,1007,780,805]" gridcol="4" gridrow="2" pageId="14" pageNumber="79">286</td>
|
||
<td id="76896F1BFFEA89DA7B15F913FB0F9242" box="[1039,1163,780,805]" gridcol="5" gridrow="2" pageId="14" pageNumber="79">298</td>
|
||
<td id="76896F1BFFEA89DA7BB4F913FAA19242" box="[1198,1317,780,805]" gridcol="6" gridrow="2" pageId="14" pageNumber="79">276</td>
|
||
<td id="76896F1BFFEA89DA7A74F913FA619242" box="[1390,1509,780,805]" gridcol="7" gridrow="2" pageId="14" pageNumber="79">283</td>
|
||
<td id="76896F1BFFEA89DA792BF913F9299242" box="[1585,1709,780,805]" gridcol="8" gridrow="2" pageId="14" pageNumber="79">287</td>
|
||
<td id="76896F1BFFEA89DA79D7F913F8CD9242" box="[1741,1865,780,805]" gridcol="9" gridrow="2" pageId="14" pageNumber="79">254</td>
|
||
<td id="76896F1BFFEA89DA7872F913F8619242" box="[1896,2021,780,805]" gridcol="10" gridrow="2" pageId="14" pageNumber="79">259</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFF926F8619235" box="[165,2021,825,850]" gridrow="3" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFF926FE909235" box="[165,276,825,850]" gridcol="0" gridrow="3" pageId="14" pageNumber="79">sw7</th>
|
||
<td id="76896F1BFFEA89DA7E48F926FE4E9235" box="[338,458,825,850]" gridcol="1" gridrow="3" pageId="14" pageNumber="79">48</td>
|
||
<td id="76896F1BFFEA89DA7D0FF926FD089235" box="[533,652,825,850]" gridcol="2" gridrow="3" pageId="14" pageNumber="79">49</td>
|
||
<td id="76896F1BFFEA89DA7DCDF926FCD09235" box="[727,852,825,850]" gridcol="3" gridrow="3" pageId="14" pageNumber="79">52</td>
|
||
<td id="76896F1BFFEA89DA7C69F926FC6B9235" box="[883,1007,825,850]" gridcol="4" gridrow="3" pageId="14" pageNumber="79">53</td>
|
||
<td id="76896F1BFFEA89DA7B15F926FB0F9235" box="[1039,1163,825,850]" gridcol="5" gridrow="3" pageId="14" pageNumber="79">53</td>
|
||
<td id="76896F1BFFEA89DA7BB4F926FAA19235" box="[1198,1317,825,850]" gridcol="6" gridrow="3" pageId="14" pageNumber="79">53</td>
|
||
<td id="76896F1BFFEA89DA7A74F926FA619235" box="[1390,1509,825,850]" gridcol="7" gridrow="3" pageId="14" pageNumber="79">55</td>
|
||
<td id="76896F1BFFEA89DA792BF926F9299235" box="[1585,1709,825,850]" gridcol="8" gridrow="3" pageId="14" pageNumber="79">53</td>
|
||
<td id="76896F1BFFEA89DA79D7F926F8CD9235" box="[1741,1865,825,850]" gridcol="9" gridrow="3" pageId="14" pageNumber="79">55</td>
|
||
<td id="76896F1BFFEA89DA7872F926F8619235" box="[1896,2021,825,850]" gridcol="10" gridrow="3" pageId="14" pageNumber="79">54</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFF978F86192E7" box="[165,2021,871,896]" gridrow="4" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFF978FE9092E7" box="[165,276,871,896]" gridcol="0" gridrow="4" pageId="14" pageNumber="79">sw10</th>
|
||
<td id="76896F1BFFEA89DA7E48F978FE4E92E7" box="[338,458,871,896]" gridcol="1" gridrow="4" pageId="14" pageNumber="79">40</td>
|
||
<td id="76896F1BFFEA89DA7D0FF978FD0892E7" box="[533,652,871,896]" gridcol="2" gridrow="4" pageId="14" pageNumber="79">41</td>
|
||
<td id="76896F1BFFEA89DA7DCDF978FCD092E7" box="[727,852,871,896]" gridcol="3" gridrow="4" pageId="14" pageNumber="79">41</td>
|
||
<td id="76896F1BFFEA89DA7C69F978FC6B92E7" box="[883,1007,871,896]" gridcol="4" gridrow="4" pageId="14" pageNumber="79">47</td>
|
||
<td id="76896F1BFFEA89DA7B15F978FB0F92E7" box="[1039,1163,871,896]" gridcol="5" gridrow="4" pageId="14" pageNumber="79">47</td>
|
||
<td id="76896F1BFFEA89DA7BB4F978FAA192E7" box="[1198,1317,871,896]" gridcol="6" gridrow="4" pageId="14" pageNumber="79">43</td>
|
||
<td id="76896F1BFFEA89DA7A74F978FA6192E7" box="[1390,1509,871,896]" gridcol="7" gridrow="4" pageId="14" pageNumber="79">44</td>
|
||
<td id="76896F1BFFEA89DA792BF978F92992E7" box="[1585,1709,871,896]" gridcol="8" gridrow="4" pageId="14" pageNumber="79">49</td>
|
||
<td id="76896F1BFFEA89DA79D7F978F8CD92E7" box="[1741,1865,871,896]" gridcol="9" gridrow="4" pageId="14" pageNumber="79">51</td>
|
||
<td id="76896F1BFFEA89DA7872F978F86192E7" box="[1896,2021,871,896]" gridcol="10" gridrow="4" pageId="14" pageNumber="79">45</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFF98BF86192CA" box="[165,2021,916,941]" gridrow="5" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFF98BFE9092CA" box="[165,276,916,941]" gridcol="0" gridrow="5" pageId="14" pageNumber="79">s1</th>
|
||
<td id="76896F1BFFEA89DA7E48F98BFE4E92CA" box="[338,458,916,941]" gridcol="1" gridrow="5" pageId="14" pageNumber="79">28</td>
|
||
<td id="76896F1BFFEA89DA7D0FF98BFD0892CA" box="[533,652,916,941]" gridcol="2" gridrow="5" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA7DCDF98BFCD092CA" box="[727,852,916,941]" gridcol="3" gridrow="5" pageId="14" pageNumber="79">25</td>
|
||
<td id="76896F1BFFEA89DA7C69F98BFC6B92CA" box="[883,1007,916,941]" gridcol="4" gridrow="5" pageId="14" pageNumber="79">32</td>
|
||
<td id="76896F1BFFEA89DA7B15F98BFB0F92CA" box="[1039,1163,916,941]" gridcol="5" gridrow="5" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA7BB4F98BFAA192CA" box="[1198,1317,916,941]" gridcol="6" gridrow="5" pageId="14" pageNumber="79">31</td>
|
||
<td id="76896F1BFFEA89DA7A74F98BFA6192CA" box="[1390,1509,916,941]" gridcol="7" gridrow="5" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA792BF98BF92992CA" box="[1585,1709,916,941]" gridcol="8" gridrow="5" pageId="14" pageNumber="79">31</td>
|
||
<td id="76896F1BFFEA89DA79D7F98BF8CD92CA" box="[1741,1865,916,941]" gridcol="9" gridrow="5" pageId="14" pageNumber="79">31</td>
|
||
<td id="76896F1BFFEA89DA7872F98BF86192CA" box="[1896,2021,916,941]" gridcol="10" gridrow="5" pageId="14" pageNumber="79">27</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFF9DEF86192BD" box="[165,2021,961,986]" gridrow="6" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFF9DEFE9092BD" box="[165,276,961,986]" gridcol="0" gridrow="6" pageId="14" pageNumber="79">s2</th>
|
||
<td id="76896F1BFFEA89DA7E48F9DEFE4E92BD" box="[338,458,961,986]" gridcol="1" gridrow="6" pageId="14" pageNumber="79">25</td>
|
||
<td id="76896F1BFFEA89DA7D0FF9DEFD0892BD" box="[533,652,961,986]" gridcol="2" gridrow="6" pageId="14" pageNumber="79">26</td>
|
||
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|
||
<td id="76896F1BFFEA89DA7C69F9DEFC6B92BD" box="[883,1007,961,986]" gridcol="4" gridrow="6" pageId="14" pageNumber="79">26</td>
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||
<td id="76896F1BFFEA89DA7B15F9DEFB0F92BD" box="[1039,1163,961,986]" gridcol="5" gridrow="6" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA7BB4F9DEFAA192BD" box="[1198,1317,961,986]" gridcol="6" gridrow="6" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA7A74F9DEFA6192BD" box="[1390,1509,961,986]" gridcol="7" gridrow="6" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA792BF9DEF92992BD" box="[1585,1709,961,986]" gridcol="8" gridrow="6" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA79D7F9DEF8CD92BD" box="[1741,1865,961,986]" gridcol="9" gridrow="6" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA7872F9DEF86192BD" box="[1896,2021,961,986]" gridcol="10" gridrow="6" pageId="14" pageNumber="79">25</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFF9F0F861956F" box="[165,2021,1007,1032]" gridrow="7" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFF9F0FE90956F" box="[165,276,1007,1032]" gridcol="0" gridrow="7" pageId="14" pageNumber="79">s3</th>
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||
<td id="76896F1BFFEA89DA7E48F9F0FE4E956F" box="[338,458,1007,1032]" gridcol="1" gridrow="7" pageId="14" pageNumber="79">25</td>
|
||
<td id="76896F1BFFEA89DA7D0FF9F0FD08956F" box="[533,652,1007,1032]" gridcol="2" gridrow="7" pageId="14" pageNumber="79">24</td>
|
||
<td id="76896F1BFFEA89DA7DCDF9F0FCD0956F" box="[727,852,1007,1032]" gridcol="3" gridrow="7" pageId="14" pageNumber="79">21</td>
|
||
<td id="76896F1BFFEA89DA7C69F9F0FC6B956F" box="[883,1007,1007,1032]" gridcol="4" gridrow="7" pageId="14" pageNumber="79">21</td>
|
||
<td id="76896F1BFFEA89DA7B15F9F0FB0F956F" box="[1039,1163,1007,1032]" gridcol="5" gridrow="7" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA7BB4F9F0FAA1956F" box="[1198,1317,1007,1032]" gridcol="6" gridrow="7" pageId="14" pageNumber="79">24</td>
|
||
<td id="76896F1BFFEA89DA7A74F9F0FA61956F" box="[1390,1509,1007,1032]" gridcol="7" gridrow="7" pageId="14" pageNumber="79">25</td>
|
||
<td id="76896F1BFFEA89DA792BF9F0F929956F" box="[1585,1709,1007,1032]" gridcol="8" gridrow="7" pageId="14" pageNumber="79">25</td>
|
||
<td id="76896F1BFFEA89DA79D7F9F0F8CD956F" box="[1741,1865,1007,1032]" gridcol="9" gridrow="7" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA7872F9F0F861956F" box="[1896,2021,1007,1032]" gridcol="10" gridrow="7" pageId="14" pageNumber="79">25</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFE03F8619552" box="[165,2021,1052,1077]" gridrow="8" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFE03FE909552" box="[165,276,1052,1077]" gridcol="0" gridrow="8" pageId="14" pageNumber="79">s4</th>
|
||
<td id="76896F1BFFEA89DA7E48FE03FE4E9552" box="[338,458,1052,1077]" gridcol="1" gridrow="8" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA7D0FFE03FD089552" box="[533,652,1052,1077]" gridcol="2" gridrow="8" pageId="14" pageNumber="79">23</td>
|
||
<td id="76896F1BFFEA89DA7DCDFE03FCD09552" box="[727,852,1052,1077]" gridcol="3" gridrow="8" pageId="14" pageNumber="79">22</td>
|
||
<td id="76896F1BFFEA89DA7C69FE03FC6B9552" box="[883,1007,1052,1077]" gridcol="4" gridrow="8" pageId="14" pageNumber="79">25</td>
|
||
<td id="76896F1BFFEA89DA7B15FE03FB0F9552" box="[1039,1163,1052,1077]" gridcol="5" gridrow="8" pageId="14" pageNumber="79">broken</td>
|
||
<td id="76896F1BFFEA89DA7BB4FE03FAA19552" box="[1198,1317,1052,1077]" gridcol="6" gridrow="8" pageId="14" pageNumber="79">28</td>
|
||
<td id="76896F1BFFEA89DA7A74FE03FA619552" box="[1390,1509,1052,1077]" gridcol="7" gridrow="8" pageId="14" pageNumber="79">25</td>
|
||
<td id="76896F1BFFEA89DA792BFE03F9299552" box="[1585,1709,1052,1077]" gridcol="8" gridrow="8" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA79D7FE03F8CD9552" box="[1741,1865,1052,1077]" gridcol="9" gridrow="8" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA7872FE03F8619552" box="[1896,2021,1052,1077]" gridcol="10" gridrow="8" pageId="14" pageNumber="79">23</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFE56F8619505" box="[165,2021,1097,1122]" gridrow="9" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFE56FE909505" box="[165,276,1097,1122]" gridcol="0" gridrow="9" pageId="14" pageNumber="79">s5</th>
|
||
<td id="76896F1BFFEA89DA7E48FE56FE4E9505" box="[338,458,1097,1122]" gridcol="1" gridrow="9" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA7D0FFE56FD089505" box="[533,652,1097,1122]" gridcol="2" gridrow="9" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA7DCDFE56FCD09505" box="[727,852,1097,1122]" gridcol="3" gridrow="9" pageId="14" pageNumber="79">24</td>
|
||
<td id="76896F1BFFEA89DA7C69FE56FC6B9505" box="[883,1007,1097,1122]" gridcol="4" gridrow="9" pageId="14" pageNumber="79">24</td>
|
||
<td id="76896F1BFFEA89DA7B15FE56FB0F9505" box="[1039,1163,1097,1122]" gridcol="5" gridrow="9" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA7BB4FE56FAA19505" box="[1198,1317,1097,1122]" gridcol="6" gridrow="9" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA7A74FE56FA619505" box="[1390,1509,1097,1122]" gridcol="7" gridrow="9" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA792BFE56F9299505" box="[1585,1709,1097,1122]" gridcol="8" gridrow="9" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA79D7FE56F8CD9505" box="[1741,1865,1097,1122]" gridcol="9" gridrow="9" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA7872FE56F8619505" box="[1896,2021,1097,1122]" gridcol="10" gridrow="9" pageId="14" pageNumber="79">25</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFE68F86195F7" box="[165,2021,1143,1168]" gridrow="10" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFE68FE9095F7" box="[165,276,1143,1168]" gridcol="0" gridrow="10" pageId="14" pageNumber="79">s6</th>
|
||
<td id="76896F1BFFEA89DA7E48FE68FE4E95F7" box="[338,458,1143,1168]" gridcol="1" gridrow="10" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA7D0FFE68FD0895F7" box="[533,652,1143,1168]" gridcol="2" gridrow="10" pageId="14" pageNumber="79">24</td>
|
||
<td id="76896F1BFFEA89DA7DCDFE68FCD095F7" box="[727,852,1143,1168]" gridcol="3" gridrow="10" pageId="14" pageNumber="79">26</td>
|
||
<td id="76896F1BFFEA89DA7C69FE68FC6B95F7" box="[883,1007,1143,1168]" gridcol="4" gridrow="10" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA7B15FE68FB0F95F7" box="[1039,1163,1143,1168]" gridcol="5" gridrow="10" pageId="14" pageNumber="79">28</td>
|
||
<td id="76896F1BFFEA89DA7BB4FE68FAA195F7" box="[1198,1317,1143,1168]" gridcol="6" gridrow="10" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA7A74FE68FA6195F7" box="[1390,1509,1143,1168]" gridcol="7" gridrow="10" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA792BFE68F92995F7" box="[1585,1709,1143,1168]" gridcol="8" gridrow="10" pageId="14" pageNumber="79">28</td>
|
||
<td id="76896F1BFFEA89DA79D7FE68F8CD95F7" box="[1741,1865,1143,1168]" gridcol="9" gridrow="10" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA7872FE68F86195F7" box="[1896,2021,1143,1168]" gridcol="10" gridrow="10" pageId="14" pageNumber="79">27</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFEBBF86195DA" box="[165,2021,1188,1213]" gridrow="11" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFEBBFE9095DA" box="[165,276,1188,1213]" gridcol="0" gridrow="11" pageId="14" pageNumber="79">s7</th>
|
||
<td id="76896F1BFFEA89DA7E48FEBBFE4E95DA" box="[338,458,1188,1213]" gridcol="1" gridrow="11" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA7D0FFEBBFD0895DA" box="[533,652,1188,1213]" gridcol="2" gridrow="11" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA7DCDFEBBFCD095DA" box="[727,852,1188,1213]" gridcol="3" gridrow="11" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA7C69FEBBFC6B95DA" box="[883,1007,1188,1213]" gridcol="4" gridrow="11" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA7B15FEBBFB0F95DA" box="[1039,1163,1188,1213]" gridcol="5" gridrow="11" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA7BB4FEBBFAA195DA" box="[1198,1317,1188,1213]" gridcol="6" gridrow="11" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA7A74FEBBFA6195DA" box="[1390,1509,1188,1213]" gridcol="7" gridrow="11" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA792BFEBBF92995DA" box="[1585,1709,1188,1213]" gridcol="8" gridrow="11" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA79D7FEBBF8CD95DA" box="[1741,1865,1188,1213]" gridcol="9" gridrow="11" pageId="14" pageNumber="79">31</td>
|
||
<td id="76896F1BFFEA89DA7872FEBBF86195DA" box="[1896,2021,1188,1213]" gridcol="10" gridrow="11" pageId="14" pageNumber="79">29</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFECEF861958D" box="[165,2021,1233,1258]" gridrow="12" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFECEFE90958D" box="[165,276,1233,1258]" gridcol="0" gridrow="12" pageId="14" pageNumber="79">s8</th>
|
||
<td id="76896F1BFFEA89DA7E48FECEFE4E958D" box="[338,458,1233,1258]" gridcol="1" gridrow="12" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA7D0FFECEFD08958D" box="[533,652,1233,1258]" gridcol="2" gridrow="12" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA7DCDFECEFCD0958D" box="[727,852,1233,1258]" gridcol="3" gridrow="12" pageId="14" pageNumber="79">33</td>
|
||
<td id="76896F1BFFEA89DA7C69FECEFC6B958D" box="[883,1007,1233,1258]" gridcol="4" gridrow="12" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA7B15FECEFB0F958D" box="[1039,1163,1233,1258]" gridcol="5" gridrow="12" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA7BB4FECEFAA1958D" box="[1198,1317,1233,1258]" gridcol="6" gridrow="12" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA7A74FECEFA61958D" box="[1390,1509,1233,1258]" gridcol="7" gridrow="12" pageId="14" pageNumber="79">34</td>
|
||
<td id="76896F1BFFEA89DA792BFECEF929958D" box="[1585,1709,1233,1258]" gridcol="8" gridrow="12" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA79D7FECEF8CD958D" box="[1741,1865,1233,1258]" gridcol="9" gridrow="12" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA7872FECEF861958D" box="[1896,2021,1233,1258]" gridcol="10" gridrow="12" pageId="14" pageNumber="79">34</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFEE0F861947F" box="[165,2021,1279,1304]" gridrow="13" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFEE0FE90947F" box="[165,276,1279,1304]" gridcol="0" gridrow="13" pageId="14" pageNumber="79">s9</th>
|
||
<td id="76896F1BFFEA89DA7E48FEE0FE4E947F" box="[338,458,1279,1304]" gridcol="1" gridrow="13" pageId="14" pageNumber="79">37</td>
|
||
<td id="76896F1BFFEA89DA7D0FFEE0FD08947F" box="[533,652,1279,1304]" gridcol="2" gridrow="13" pageId="14" pageNumber="79">36</td>
|
||
<td id="76896F1BFFEA89DA7DCDFEE0FCD0947F" box="[727,852,1279,1304]" gridcol="3" gridrow="13" pageId="14" pageNumber="79">33</td>
|
||
<td id="76896F1BFFEA89DA7C69FEE0FC6B947F" box="[883,1007,1279,1304]" gridcol="4" gridrow="13" pageId="14" pageNumber="79">37</td>
|
||
<td id="76896F1BFFEA89DA7B15FEE0FB0F947F" box="[1039,1163,1279,1304]" gridcol="5" gridrow="13" pageId="14" pageNumber="79">39</td>
|
||
<td id="76896F1BFFEA89DA7BB4FEE0FAA1947F" box="[1198,1317,1279,1304]" gridcol="6" gridrow="13" pageId="14" pageNumber="79">38</td>
|
||
<td id="76896F1BFFEA89DA7A74FEE0FA61947F" box="[1390,1509,1279,1304]" gridcol="7" gridrow="13" pageId="14" pageNumber="79">39</td>
|
||
<td id="76896F1BFFEA89DA792BFEE0F929947F" box="[1585,1709,1279,1304]" gridcol="8" gridrow="13" pageId="14" pageNumber="79">41</td>
|
||
<td id="76896F1BFFEA89DA79D7FEE0F8CD947F" box="[1741,1865,1279,1304]" gridcol="9" gridrow="13" pageId="14" pageNumber="79">39</td>
|
||
<td id="76896F1BFFEA89DA7872FEE0F861947F" box="[1896,2021,1279,1304]" gridcol="10" gridrow="13" pageId="14" pageNumber="79">35</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFF33F8619422" box="[165,2021,1324,1349]" gridrow="14" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFF33FE909422" box="[165,276,1324,1349]" gridcol="0" gridrow="14" pageId="14" pageNumber="79">s10</th>
|
||
<td id="76896F1BFFEA89DA7E48FF33FE4E9422" box="[338,458,1324,1349]" gridcol="1" gridrow="14" pageId="14" pageNumber="79">32</td>
|
||
<td id="76896F1BFFEA89DA7D0FFF33FD089422" box="[533,652,1324,1349]" gridcol="2" gridrow="14" pageId="14" pageNumber="79">30</td>
|
||
<td id="76896F1BFFEA89DA7DCDFF33FCD09422" box="[727,852,1324,1349]" gridcol="3" gridrow="14" pageId="14" pageNumber="79">31</td>
|
||
<td id="76896F1BFFEA89DA7C69FF33FC6B9422" box="[883,1007,1324,1349]" gridcol="4" gridrow="14" pageId="14" pageNumber="79">36</td>
|
||
<td id="76896F1BFFEA89DA7B15FF33FB0F9422" box="[1039,1163,1324,1349]" gridcol="5" gridrow="14" pageId="14" pageNumber="79">34</td>
|
||
<td id="76896F1BFFEA89DA7BB4FF33FAA19422" box="[1198,1317,1324,1349]" gridcol="6" gridrow="14" pageId="14" pageNumber="79">34</td>
|
||
<td id="76896F1BFFEA89DA7A74FF33FA619422" box="[1390,1509,1324,1349]" gridcol="7" gridrow="14" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA792BFF33F9299422" box="[1585,1709,1324,1349]" gridcol="8" gridrow="14" pageId="14" pageNumber="79">35</td>
|
||
<td id="76896F1BFFEA89DA79D7FF33F8CD9422" box="[1741,1865,1324,1349]" gridcol="9" gridrow="14" pageId="14" pageNumber="79">38</td>
|
||
<td id="76896F1BFFEA89DA7872FF33F8619422" box="[1896,2021,1324,1349]" gridcol="10" gridrow="14" pageId="14" pageNumber="79">33</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFF46F8619415" box="[165,2021,1369,1394]" gridrow="15" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFF46FE909415" box="[165,276,1369,1394]" gridcol="0" gridrow="15" pageId="14" pageNumber="79">s11</th>
|
||
<td id="76896F1BFFEA89DA7E48FF46FE4E9415" box="[338,458,1369,1394]" gridcol="1" gridrow="15" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA7D0FFF46FD089415" box="[533,652,1369,1394]" gridcol="2" gridrow="15" pageId="14" pageNumber="79">25</td>
|
||
<td id="76896F1BFFEA89DA7DCDFF46FCD09415" box="[727,852,1369,1394]" gridcol="3" gridrow="15" pageId="14" pageNumber="79">24</td>
|
||
<td id="76896F1BFFEA89DA7C69FF46FC6B9415" box="[883,1007,1369,1394]" gridcol="4" gridrow="15" pageId="14" pageNumber="79">28</td>
|
||
<td id="76896F1BFFEA89DA7B15FF46FB0F9415" box="[1039,1163,1369,1394]" gridcol="5" gridrow="15" pageId="14" pageNumber="79">29</td>
|
||
<td id="76896F1BFFEA89DA7BB4FF46FAA19415" box="[1198,1317,1369,1394]" gridcol="6" gridrow="15" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA7A74FF46FA619415" box="[1390,1509,1369,1394]" gridcol="7" gridrow="15" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA792BFF46F9299415" box="[1585,1709,1369,1394]" gridcol="8" gridrow="15" pageId="14" pageNumber="79">27</td>
|
||
<td id="76896F1BFFEA89DA79D7FF46F8CD9415" box="[1741,1865,1369,1394]" gridcol="9" gridrow="15" pageId="14" pageNumber="79">28</td>
|
||
<td id="76896F1BFFEA89DA7872FF46F8619415" box="[1896,2021,1369,1394]" gridcol="10" gridrow="15" pageId="14" pageNumber="79">27</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFF98F86194C7" box="[165,2021,1415,1440]" gridrow="16" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFF98FE9094C7" box="[165,276,1415,1440]" gridcol="0" gridrow="16" pageId="14" pageNumber="79">mds4</th>
|
||
<td id="76896F1BFFEA89DA7E48FF98FE4E94C7" box="[338,458,1415,1440]" gridcol="1" gridrow="16" pageId="14" pageNumber="79">47</td>
|
||
<td id="76896F1BFFEA89DA7D0FFF98FD0894C7" box="[533,652,1415,1440]" gridcol="2" gridrow="16" pageId="14" pageNumber="79">37</td>
|
||
<td id="76896F1BFFEA89DA7DCDFF98FCD094C7" box="[727,852,1415,1440]" gridcol="3" gridrow="16" pageId="14" pageNumber="79">32</td>
|
||
<td id="76896F1BFFEA89DA7C69FF98FC6B94C7" box="[883,1007,1415,1440]" gridcol="4" gridrow="16" pageId="14" pageNumber="79">49</td>
|
||
<td id="76896F1BFFEA89DA7B15FF98FB0F94C7" box="[1039,1163,1415,1440]" gridcol="5" gridrow="16" pageId="14" pageNumber="79">broken</td>
|
||
<td id="76896F1BFFEA89DA7BB4FF98FAA194C7" box="[1198,1317,1415,1440]" gridcol="6" gridrow="16" pageId="14" pageNumber="79">53</td>
|
||
<td id="76896F1BFFEA89DA7A74FF98FA6194C7" box="[1390,1509,1415,1440]" gridcol="7" gridrow="16" pageId="14" pageNumber="79">42</td>
|
||
<td id="76896F1BFFEA89DA792BFF98F92994C7" box="[1585,1709,1415,1440]" gridcol="8" gridrow="16" pageId="14" pageNumber="79">50</td>
|
||
<td id="76896F1BFFEA89DA79D7FF98F8CD94C7" box="[1741,1865,1415,1440]" gridcol="9" gridrow="16" pageId="14" pageNumber="79">50</td>
|
||
<td id="76896F1BFFEA89DA7872FF98F86194C7" box="[1896,2021,1415,1440]" gridcol="10" gridrow="16" pageId="14" pageNumber="79">39</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFFABF86194AA" box="[165,2021,1460,1485]" gridrow="17" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFFABFE9094AA" box="[165,276,1460,1485]" gridcol="0" gridrow="17" pageId="14" pageNumber="79">mds6</th>
|
||
<td id="76896F1BFFEA89DA7E48FFABFE4E94AA" box="[338,458,1460,1485]" gridcol="1" gridrow="17" pageId="14" pageNumber="79">59</td>
|
||
<td id="76896F1BFFEA89DA7D0FFFABFD0894AA" box="[533,652,1460,1485]" gridcol="2" gridrow="17" pageId="14" pageNumber="79">46</td>
|
||
<td id="76896F1BFFEA89DA7DCDFFABFCD094AA" box="[727,852,1460,1485]" gridcol="3" gridrow="17" pageId="14" pageNumber="79">46</td>
|
||
<td id="76896F1BFFEA89DA7C69FFABFC6B94AA" box="[883,1007,1460,1485]" gridcol="4" gridrow="17" pageId="14" pageNumber="79">57</td>
|
||
<td id="76896F1BFFEA89DA7B15FFABFB0F94AA" box="[1039,1163,1460,1485]" gridcol="5" gridrow="17" pageId="14" pageNumber="79">68</td>
|
||
<td id="76896F1BFFEA89DA7BB4FFABFAA194AA" box="[1198,1317,1460,1485]" gridcol="6" gridrow="17" pageId="14" pageNumber="79">67</td>
|
||
<td id="76896F1BFFEA89DA7A74FFABFA6194AA" box="[1390,1509,1460,1485]" gridcol="7" gridrow="17" pageId="14" pageNumber="79">64</td>
|
||
<td id="76896F1BFFEA89DA792BFFABF92994AA" box="[1585,1709,1460,1485]" gridcol="8" gridrow="17" pageId="14" pageNumber="79">66</td>
|
||
<td id="76896F1BFFEA89DA79D7FFABF8CD94AA" box="[1741,1865,1460,1485]" gridcol="9" gridrow="17" pageId="14" pageNumber="79">75</td>
|
||
<td id="76896F1BFFEA89DA7872FFABF86194AA" box="[1896,2021,1460,1485]" gridcol="10" gridrow="17" pageId="14" pageNumber="79">68</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFFFEF861949D" box="[165,2021,1505,1530]" gridrow="18" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFFFEFE90949D" box="[165,276,1505,1530]" gridcol="0" gridrow="18" pageId="14" pageNumber="79">mds8</th>
|
||
<td id="76896F1BFFEA89DA7E48FFFEFE4E949D" box="[338,458,1505,1530]" gridcol="1" gridrow="18" pageId="14" pageNumber="79">broken</td>
|
||
<td id="76896F1BFFEA89DA7D0FFFFEFD08949D" box="[533,652,1505,1530]" gridcol="2" gridrow="18" pageId="14" pageNumber="79">61</td>
|
||
<td id="76896F1BFFEA89DA7DCDFFFEFCD0949D" box="[727,852,1505,1530]" gridcol="3" gridrow="18" pageId="14" pageNumber="79">54</td>
|
||
<td id="76896F1BFFEA89DA7C69FFFEFC6B949D" box="[883,1007,1505,1530]" gridcol="4" gridrow="18" pageId="14" pageNumber="79">69</td>
|
||
<td id="76896F1BFFEA89DA7B15FFFEFB0F949D" box="[1039,1163,1505,1530]" gridcol="5" gridrow="18" pageId="14" pageNumber="79">79</td>
|
||
<td id="76896F1BFFEA89DA7BB4FFFEFAA1949D" box="[1198,1317,1505,1530]" gridcol="6" gridrow="18" pageId="14" pageNumber="79">93</td>
|
||
<td id="76896F1BFFEA89DA7A74FFFEFA61949D" box="[1390,1509,1505,1530]" gridcol="7" gridrow="18" pageId="14" pageNumber="79">92</td>
|
||
<td id="76896F1BFFEA89DA792BFFFEF929949D" box="[1585,1709,1505,1530]" gridcol="8" gridrow="18" pageId="14" pageNumber="79">94</td>
|
||
<td id="76896F1BFFEA89DA79D7FFFEF8CD949D" box="[1741,1865,1505,1530]" gridcol="9" gridrow="18" pageId="14" pageNumber="79">98</td>
|
||
<td id="76896F1BFFEA89DA7872FFFEF861949D" box="[1896,2021,1505,1530]" gridcol="10" gridrow="18" pageId="14" pageNumber="79">broken</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFC10F861974F" box="[165,2021,1551,1576]" gridrow="19" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFC10FE90974F" box="[165,276,1551,1576]" gridcol="0" gridrow="19" pageId="14" pageNumber="79">sdt2</th>
|
||
<td id="76896F1BFFEA89DA7E48FC10FE4E974F" box="[338,458,1551,1576]" gridcol="1" gridrow="19" pageId="14" pageNumber="79">15/16</td>
|
||
<td id="76896F1BFFEA89DA7D0FFC10FD08974F" box="[533,652,1551,1576]" gridcol="2" gridrow="19" pageId="14" pageNumber="79">15/broken</td>
|
||
<td id="76896F1BFFEA89DA7DCDFC10FCD0974F" box="[727,852,1551,1576]" gridcol="3" gridrow="19" pageId="14" pageNumber="79">11/n. a.</td>
|
||
<td id="76896F1BFFEA89DA7C69FC10FC6B974F" box="[883,1007,1551,1576]" gridcol="4" gridrow="19" pageId="14" pageNumber="79">11</td>
|
||
<td id="76896F1BFFEA89DA7B15FC10FB0F974F" box="[1039,1163,1551,1576]" gridcol="5" gridrow="19" pageId="14" pageNumber="79">12/n.a.</td>
|
||
<td id="76896F1BFFEA89DA7BB4FC10FAA1974F" box="[1198,1317,1551,1576]" gridcol="6" gridrow="19" pageId="14" pageNumber="79">18/17</td>
|
||
<td id="76896F1BFFEA89DA7A74FC10FA61974F" box="[1390,1509,1551,1576]" gridcol="7" gridrow="19" pageId="14" pageNumber="79">14/10</td>
|
||
<td id="76896F1BFFEA89DA792BFC10F929974F" box="[1585,1709,1551,1576]" gridcol="8" gridrow="19" pageId="14" pageNumber="79">17/10</td>
|
||
<td id="76896F1BFFEA89DA79D7FC10F8CD974F" box="[1741,1865,1551,1576]" gridcol="9" gridrow="19" pageId="14" pageNumber="79">15</td>
|
||
<td id="76896F1BFFEA89DA7872FC10F861974F" box="[1896,2021,1551,1576]" gridcol="10" gridrow="19" pageId="14" pageNumber="79">9/13</td>
|
||
</tr>
|
||
<tr id="35580667FFEA89DA7FBFFC23F8619732" box="[165,2021,1596,1621]" gridrow="20" pageId="14" pageNumber="79">
|
||
<th id="76896F1BFFEA89DA7FBFFC23FE909732" box="[165,276,1596,1621]" gridcol="0" gridrow="20" pageId="14" pageNumber="79">vlt2</th>
|
||
<td id="76896F1BFFEA89DA7E48FC23FE4E9732" box="[338,458,1596,1621]" gridcol="1" gridrow="20" pageId="14" pageNumber="79">17</td>
|
||
<td id="76896F1BFFEA89DA7D0FFC23FD089732" box="[533,652,1596,1621]" gridcol="2" gridrow="20" pageId="14" pageNumber="79">17/broken</td>
|
||
<td id="76896F1BFFEA89DA7DCDFC23FCD09732" box="[727,852,1596,1621]" gridcol="3" gridrow="20" pageId="14" pageNumber="79">18/17</td>
|
||
<td id="76896F1BFFEA89DA7C69FC23FC6B9732" box="[883,1007,1596,1621]" gridcol="4" gridrow="20" pageId="14" pageNumber="79">n. a./13</td>
|
||
<td id="76896F1BFFEA89DA7B15FC23FB0F9732" box="[1039,1163,1596,1621]" gridcol="5" gridrow="20" pageId="14" pageNumber="79">n. a.</td>
|
||
<td id="76896F1BFFEA89DA7BB4FC23FAA19732" box="[1198,1317,1596,1621]" gridcol="6" gridrow="20" pageId="14" pageNumber="79">broken/19</td>
|
||
<td id="76896F1BFFEA89DA7A74FC23FA619732" box="[1390,1509,1596,1621]" gridcol="7" gridrow="20" pageId="14" pageNumber="79">17</td>
|
||
<td id="76896F1BFFEA89DA792BFC23F9299732" box="[1585,1709,1596,1621]" gridcol="8" gridrow="20" pageId="14" pageNumber="79">18/n.a.</td>
|
||
<td id="76896F1BFFEA89DA79D7FC23F8CD9732" box="[1741,1865,1596,1621]" gridcol="9" gridrow="20" pageId="14" pageNumber="79">17/18</td>
|
||
<td id="76896F1BFFEA89DA7872FC23F8619732" box="[1896,2021,1596,1621]" gridcol="10" gridrow="20" pageId="14" pageNumber="79">18/17</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFEA762B79ABFC75F86397E4" blockId="14.[1713,2023,1642,1667]" box="[1713,2023,1642,1667]" pageId="14" pageNumber="79">
|
||
<tableNote id="768E05ABFFEA762B79ABFC75F86397E4" box="[1713,2023,1642,1667]" pageId="14" pageNumber="79" targetBox="[165,2021,667,1621]" targetPageId="14">......continued on the next page</tableNote>
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFEB762A7F8DF8C4FE0A9393" blockId="15.[151,398,731,756]" box="[151,398,731,756]" pageId="15" pageNumber="80">
|
||
<tableNote id="768E05ABFFEB762A7F8DF8C4FE0A9393" box="[151,398,731,756]" pageId="15" pageNumber="80" targetBox="[151,2006,776,1459]" targetPageId="15">TABLE 2. (Continued)</tableNote>
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFEB762A7F8DF917F81794D4" pageId="15" pageNumber="80">
|
||
<table id="F968F685FFEB89DA7F8DF917F85294D4" box="[151,2006,776,1459]" colsContinueFrom="14.[165,2021,667,1621]" gridcols="11" gridrows="15" pageId="15" pageNumber="80">
|
||
<tr id="35580667FFEB89DA7F8DF917F8529221" box="[151,2006,776,838]" gridrow="0" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DF917FE829221" box="[151,262,776,838]" gridcol="0" gridrow="0" pageId="15" pageNumber="80">
|
||
<emphasis id="B91CD837FFEB762A7F8DF917FF7B9246" bold="true" box="[151,255,776,801]" pageId="15" pageNumber="80">Specimen</emphasis>
|
||
</th>
|
||
<th id="76896F1BFFEB89DA7E59F917FE389221" box="[323,444,776,838]" gridcol="1" gridrow="0" pageId="15" pageNumber="80">♂ holotype, MACN-In</th>
|
||
<th id="76896F1BFFEB89DA7D1CF917FDF99221" box="[518,637,776,838]" gridcol="2" gridrow="0" pageId="15" pageNumber="80">♂ paratype, MACN-In</th>
|
||
<th id="76896F1BFFEB89DA7DD3F917FCC19221" box="[713,837,776,838]" gridcol="3" gridrow="0" pageId="15" pageNumber="80">♂ paratype, ZMB 12262</th>
|
||
<th id="76896F1BFFEB89DA7C7FF917FC659221" box="[869,993,776,838]" gridcol="4" gridrow="0" pageId="15" pageNumber="80">♂ non-type, ZMB 12263</th>
|
||
<th id="76896F1BFFEB89DA7B1BF917FBF99221" box="[1025,1149,776,838]" gridcol="5" gridrow="0" pageId="15" pageNumber="80">♂ non-type, ZMB 12266</th>
|
||
<th id="76896F1BFFEB89DA7B85F917FA939221" box="[1183,1303,776,838]" gridcol="6" gridrow="0" pageId="15" pageNumber="80">♀ paratype, MACN-In</th>
|
||
<th id="76896F1BFFEB89DA7A7AF917FA539221" box="[1376,1495,776,838]" gridcol="7" gridrow="0" pageId="15" pageNumber="80">♀ paratype, MACN-In</th>
|
||
<th id="76896F1BFFEB89DA7938F917F91B9221" box="[1570,1695,776,838]" gridcol="8" gridrow="0" pageId="15" pageNumber="80">♀ paratype, ZMB 12264</th>
|
||
<th id="76896F1BFFEB89DA79A4F917F8BE9221" box="[1726,1850,776,838]" gridcol="9" gridrow="0" pageId="15" pageNumber="80">♀ paratype, ZMB 12265</th>
|
||
<th id="76896F1BFFEB89DA7840F917F8529221" box="[1882,2006,776,838]" gridcol="10" gridrow="0" pageId="15" pageNumber="80">♀ paratype, ZMB 12341</th>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DF94FF852920D" box="[151,2006,848,874]" gridrow="1" pageId="15" pageNumber="80" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="76896F1BFFEB89DA7F8DF94FFE82920D" box="[151,262,848,874]" gridcol="0" gridrow="1" pageId="15" pageNumber="80">
|
||
<emphasis id="B91CD837FFEB762A7F8DF94FFE82920E" bold="true" box="[151,262,848,873]" pageId="15" pageNumber="80">Character</emphasis>
|
||
</th>
|
||
<td id="76896F1BFFEB89DA7E59F94FFE38920D" box="[323,444,848,874]" gridcol="1" gridrow="1" pageId="15" pageNumber="80">43755</td>
|
||
<td id="76896F1BFFEB89DA7D1CF94FFDF9920D" box="[518,637,848,874]" gridcol="2" gridrow="1" pageId="15" pageNumber="80">43756a</td>
|
||
<td id="76896F1BFFEB89DA7B85F94FFA93920D" box="[1183,1303,848,874]" gridcol="6" gridrow="1" pageId="15" pageNumber="80">43756b</td>
|
||
<td id="76896F1BFFEB89DA7A7AF94FFA53920D" box="[1376,1495,848,874]" gridcol="7" gridrow="1" pageId="15" pageNumber="80">43756c</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DF965F85292F4" box="[151,2006,890,915]" gridrow="2" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DF965FE8292F4" box="[151,262,890,915]" gridcol="0" gridrow="2" pageId="15" pageNumber="80">lvt5</th>
|
||
<td id="76896F1BFFEB89DA7E59F965FE3892F4" box="[323,444,890,915]" gridcol="1" gridrow="2" pageId="15" pageNumber="80">21/20</td>
|
||
<td id="76896F1BFFEB89DA7D1CF965FDF992F4" box="[518,637,890,915]" gridcol="2" gridrow="2" pageId="15" pageNumber="80">19/broken</td>
|
||
<td id="76896F1BFFEB89DA7DD3F965FCC192F4" box="[713,837,890,915]" gridcol="3" gridrow="2" pageId="15" pageNumber="80">21</td>
|
||
<td id="76896F1BFFEB89DA7C7FF965FC6592F4" box="[869,993,890,915]" gridcol="4" gridrow="2" pageId="15" pageNumber="80">20/24</td>
|
||
<td id="76896F1BFFEB89DA7B1BF965FBF992F4" box="[1025,1149,890,915]" gridcol="5" gridrow="2" pageId="15" pageNumber="80">19/20</td>
|
||
<td id="76896F1BFFEB89DA7B85F965FA9392F4" box="[1183,1303,890,915]" gridcol="6" gridrow="2" pageId="15" pageNumber="80">22</td>
|
||
<td id="76896F1BFFEB89DA7A7AF965FA5392F4" box="[1376,1495,890,915]" gridcol="7" gridrow="2" pageId="15" pageNumber="80">20/22</td>
|
||
<td id="76896F1BFFEB89DA7938F965F91B92F4" box="[1570,1695,890,915]" gridcol="8" gridrow="2" pageId="15" pageNumber="80">21/n.a.</td>
|
||
<td id="76896F1BFFEB89DA79A4F965F8BE92F4" box="[1726,1850,890,915]" gridcol="9" gridrow="2" pageId="15" pageNumber="80">21/22</td>
|
||
<td id="76896F1BFFEB89DA7840F965F85292F4" box="[1882,2006,890,915]" gridcol="10" gridrow="2" pageId="15" pageNumber="80">19/17</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DF9B8F85292A7" box="[151,2006,935,960]" gridrow="3" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DF9B8FE8292A7" box="[151,262,935,960]" gridcol="0" gridrow="3" pageId="15" pageNumber="80">lvs6</th>
|
||
<td id="76896F1BFFEB89DA7E59F9B8FE3892A7" box="[323,444,935,960]" gridcol="1" gridrow="3" pageId="15" pageNumber="80">broken/30</td>
|
||
<td id="76896F1BFFEB89DA7D1CF9B8FDF992A7" box="[518,637,935,960]" gridcol="2" gridrow="3" pageId="15" pageNumber="80">29/26</td>
|
||
<td id="76896F1BFFEB89DA7DD3F9B8FCC192A7" box="[713,837,935,960]" gridcol="3" gridrow="3" pageId="15" pageNumber="80">26/25</td>
|
||
<td id="76896F1BFFEB89DA7C7FF9B8FC6592A7" box="[869,993,935,960]" gridcol="4" gridrow="3" pageId="15" pageNumber="80">21/29</td>
|
||
<td id="76896F1BFFEB89DA7B1BF9B8FBF992A7" box="[1025,1149,935,960]" gridcol="5" gridrow="3" pageId="15" pageNumber="80">30/broken</td>
|
||
<td id="76896F1BFFEB89DA7B85F9B8FA9392A7" box="[1183,1303,935,960]" gridcol="6" gridrow="3" pageId="15" pageNumber="80">27/30</td>
|
||
<td id="76896F1BFFEB89DA7A7AF9B8FA5392A7" box="[1376,1495,935,960]" gridcol="7" gridrow="3" pageId="15" pageNumber="80">30/broken</td>
|
||
<td id="76896F1BFFEB89DA7938F9B8F91B92A7" box="[1570,1695,935,960]" gridcol="8" gridrow="3" pageId="15" pageNumber="80">30/26</td>
|
||
<td id="76896F1BFFEB89DA79A4F9B8F8BE92A7" box="[1726,1850,935,960]" gridcol="9" gridrow="3" pageId="15" pageNumber="80">36/33</td>
|
||
<td id="76896F1BFFEB89DA7840F9B8F85292A7" box="[1882,2006,935,960]" gridcol="10" gridrow="3" pageId="15" pageNumber="80">31/broken</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DF9CAF8529289" box="[151,2006,981,1006]" gridrow="4" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DF9CAFE829289" box="[151,262,981,1006]" gridcol="0" gridrow="4" pageId="15" pageNumber="80">slt7</th>
|
||
<td id="76896F1BFFEB89DA7E59F9CAFE389289" box="[323,444,981,1006]" gridcol="1" gridrow="4" pageId="15" pageNumber="80">19</td>
|
||
<td id="76896F1BFFEB89DA7D1CF9CAFDF99289" box="[518,637,981,1006]" gridcol="2" gridrow="4" pageId="15" pageNumber="80">18</td>
|
||
<td id="76896F1BFFEB89DA7DD3F9CAFCC19289" box="[713,837,981,1006]" gridcol="3" gridrow="4" pageId="15" pageNumber="80">23/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7C7FF9CAFC659289" box="[869,993,981,1006]" gridcol="4" gridrow="4" pageId="15" pageNumber="80">n. a.</td>
|
||
<td id="76896F1BFFEB89DA7B1BF9CAFBF99289" box="[1025,1149,981,1006]" gridcol="5" gridrow="4" pageId="15" pageNumber="80">n. a./17</td>
|
||
<td id="76896F1BFFEB89DA7B85F9CAFA939289" box="[1183,1303,981,1006]" gridcol="6" gridrow="4" pageId="15" pageNumber="80">21</td>
|
||
<td id="76896F1BFFEB89DA7A7AF9CAFA539289" box="[1376,1495,981,1006]" gridcol="7" gridrow="4" pageId="15" pageNumber="80">n. a./19</td>
|
||
<td id="76896F1BFFEB89DA7938F9CAF91B9289" box="[1570,1695,981,1006]" gridcol="8" gridrow="4" pageId="15" pageNumber="80">21</td>
|
||
<td id="76896F1BFFEB89DA79A4F9CAF8BE9289" box="[1726,1850,981,1006]" gridcol="9" gridrow="4" pageId="15" pageNumber="80">21/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7840F9CAF8529289" box="[1882,2006,981,1006]" gridcol="10" gridrow="4" pageId="15" pageNumber="80">18</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFE1DF852957C" box="[151,2006,1026,1051]" gridrow="5" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFE1DFE82957C" box="[151,262,1026,1051]" gridcol="0" gridrow="5" pageId="15" pageNumber="80">lvs7</th>
|
||
<td id="76896F1BFFEB89DA7E59FE1DFE38957C" box="[323,444,1026,1051]" gridcol="1" gridrow="5" pageId="15" pageNumber="80">33/34</td>
|
||
<td id="76896F1BFFEB89DA7D1CFE1DFDF9957C" box="[518,637,1026,1051]" gridcol="2" gridrow="5" pageId="15" pageNumber="80">34/31</td>
|
||
<td id="76896F1BFFEB89DA7DD3FE1DFCC1957C" box="[713,837,1026,1051]" gridcol="3" gridrow="5" pageId="15" pageNumber="80">30/29</td>
|
||
<td id="76896F1BFFEB89DA7C7FFE1DFC65957C" box="[869,993,1026,1051]" gridcol="4" gridrow="5" pageId="15" pageNumber="80">25/29</td>
|
||
<td id="76896F1BFFEB89DA7B1BFE1DFBF9957C" box="[1025,1149,1026,1051]" gridcol="5" gridrow="5" pageId="15" pageNumber="80">37/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7B85FE1DFA93957C" box="[1183,1303,1026,1051]" gridcol="6" gridrow="5" pageId="15" pageNumber="80">37/35</td>
|
||
<td id="76896F1BFFEB89DA7A7AFE1DFA53957C" box="[1376,1495,1026,1051]" gridcol="7" gridrow="5" pageId="15" pageNumber="80">37</td>
|
||
<td id="76896F1BFFEB89DA7938FE1DF91B957C" box="[1570,1695,1026,1051]" gridcol="8" gridrow="5" pageId="15" pageNumber="80">37/34</td>
|
||
<td id="76896F1BFFEB89DA79A4FE1DF8BE957C" box="[1726,1850,1026,1051]" gridcol="9" gridrow="5" pageId="15" pageNumber="80">38</td>
|
||
<td id="76896F1BFFEB89DA7840FE1DF852957C" box="[1882,2006,1026,1051]" gridcol="10" gridrow="5" pageId="15" pageNumber="80">35/34</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFE2FF852952E" box="[151,2006,1072,1097]" gridrow="6" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFE2FFE82952E" box="[151,262,1072,1097]" gridcol="0" gridrow="6" pageId="15" pageNumber="80">lat8</th>
|
||
<td id="76896F1BFFEB89DA7E59FE2FFE38952E" box="[323,444,1072,1097]" gridcol="1" gridrow="6" pageId="15" pageNumber="80">21/25</td>
|
||
<td id="76896F1BFFEB89DA7D1CFE2FFDF9952E" box="[518,637,1072,1097]" gridcol="2" gridrow="6" pageId="15" pageNumber="80">25/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7DD3FE2FFCC1952E" box="[713,837,1072,1097]" gridcol="3" gridrow="6" pageId="15" pageNumber="80">n.a./23</td>
|
||
<td id="76896F1BFFEB89DA7C7FFE2FFC65952E" box="[869,993,1072,1097]" gridcol="4" gridrow="6" pageId="15" pageNumber="80">n.a./23</td>
|
||
<td id="76896F1BFFEB89DA7B1BFE2FFBF9952E" box="[1025,1149,1072,1097]" gridcol="5" gridrow="6" pageId="15" pageNumber="80">19/22</td>
|
||
<td id="76896F1BFFEB89DA7B85FE2FFA93952E" box="[1183,1303,1072,1097]" gridcol="6" gridrow="6" pageId="15" pageNumber="80">22/23</td>
|
||
<td id="76896F1BFFEB89DA7A7AFE2FFA53952E" box="[1376,1495,1072,1097]" gridcol="7" gridrow="6" pageId="15" pageNumber="80">21</td>
|
||
<td id="76896F1BFFEB89DA7938FE2FF91B952E" box="[1570,1695,1072,1097]" gridcol="8" gridrow="6" pageId="15" pageNumber="80">22</td>
|
||
<td id="76896F1BFFEB89DA79A4FE2FF8BE952E" box="[1726,1850,1072,1097]" gridcol="9" gridrow="6" pageId="15" pageNumber="80">15/22</td>
|
||
<td id="76896F1BFFEB89DA7840FE2FF852952E" box="[1882,2006,1072,1097]" gridcol="10" gridrow="6" pageId="15" pageNumber="80">21/20</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFE42F8529511" box="[151,2006,1117,1142]" gridrow="7" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFE42FE829511" box="[151,262,1117,1142]" gridcol="0" gridrow="7" pageId="15" pageNumber="80">lvs8</th>
|
||
<td id="76896F1BFFEB89DA7E59FE42FE389511" box="[323,444,1117,1142]" gridcol="1" gridrow="7" pageId="15" pageNumber="80">38/40</td>
|
||
<td id="76896F1BFFEB89DA7D1CFE42FDF99511" box="[518,637,1117,1142]" gridcol="2" gridrow="7" pageId="15" pageNumber="80">43/37</td>
|
||
<td id="76896F1BFFEB89DA7DD3FE42FCC19511" box="[713,837,1117,1142]" gridcol="3" gridrow="7" pageId="15" pageNumber="80">35/32</td>
|
||
<td id="76896F1BFFEB89DA7C7FFE42FC659511" box="[869,993,1117,1142]" gridcol="4" gridrow="7" pageId="15" pageNumber="80">39/44</td>
|
||
<td id="76896F1BFFEB89DA7B1BFE42FBF99511" box="[1025,1149,1117,1142]" gridcol="5" gridrow="7" pageId="15" pageNumber="80">43/51</td>
|
||
<td id="76896F1BFFEB89DA7B85FE42FA939511" box="[1183,1303,1117,1142]" gridcol="6" gridrow="7" pageId="15" pageNumber="80">50/49</td>
|
||
<td id="76896F1BFFEB89DA7A7AFE42FA539511" box="[1376,1495,1117,1142]" gridcol="7" gridrow="7" pageId="15" pageNumber="80">47/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7938FE42F91B9511" box="[1570,1695,1117,1142]" gridcol="8" gridrow="7" pageId="15" pageNumber="80">49/42</td>
|
||
<td id="76896F1BFFEB89DA79A4FE42F8BE9511" box="[1726,1850,1117,1142]" gridcol="9" gridrow="7" pageId="15" pageNumber="80">47/46</td>
|
||
<td id="76896F1BFFEB89DA7840FE42F8529511" box="[1882,2006,1117,1142]" gridcol="10" gridrow="7" pageId="15" pageNumber="80">n. a./46</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFE95F85295C4" box="[151,2006,1162,1187]" gridrow="8" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFE95FE8295C4" box="[151,262,1162,1187]" gridcol="0" gridrow="8" pageId="15" pageNumber="80">slt9</th>
|
||
<td id="76896F1BFFEB89DA7E59FE95FE3895C4" box="[323,444,1162,1187]" gridcol="1" gridrow="8" pageId="15" pageNumber="80">20</td>
|
||
<td id="76896F1BFFEB89DA7D1CFE95FDF995C4" box="[518,637,1162,1187]" gridcol="2" gridrow="8" pageId="15" pageNumber="80">18/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7DD3FE95FCC195C4" box="[713,837,1162,1187]" gridcol="3" gridrow="8" pageId="15" pageNumber="80">18/19</td>
|
||
<td id="76896F1BFFEB89DA7C7FFE95FC6595C4" box="[869,993,1162,1187]" gridcol="4" gridrow="8" pageId="15" pageNumber="80">n. a.</td>
|
||
<td id="76896F1BFFEB89DA7B1BFE95FBF995C4" box="[1025,1149,1162,1187]" gridcol="5" gridrow="8" pageId="15" pageNumber="80">n. a./15</td>
|
||
<td id="76896F1BFFEB89DA7B85FE95FA9395C4" box="[1183,1303,1162,1187]" gridcol="6" gridrow="8" pageId="15" pageNumber="80">20</td>
|
||
<td id="76896F1BFFEB89DA7A7AFE95FA5395C4" box="[1376,1495,1162,1187]" gridcol="7" gridrow="8" pageId="15" pageNumber="80">19/20</td>
|
||
<td id="76896F1BFFEB89DA7938FE95F91B95C4" box="[1570,1695,1162,1187]" gridcol="8" gridrow="8" pageId="15" pageNumber="80">22</td>
|
||
<td id="76896F1BFFEB89DA79A4FE95F8BE95C4" box="[1726,1850,1162,1187]" gridcol="9" gridrow="8" pageId="15" pageNumber="80">n. a.</td>
|
||
<td id="76896F1BFFEB89DA7840FE95F85295C4" box="[1882,2006,1162,1187]" gridcol="10" gridrow="8" pageId="15" pageNumber="80">19/n. a.</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFEA7F85295B6" box="[151,2006,1208,1233]" gridrow="9" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFEA7FE8295B6" box="[151,262,1208,1233]" gridcol="0" gridrow="9" pageId="15" pageNumber="80">lv9</th>
|
||
<td id="76896F1BFFEB89DA7E59FEA7FE3895B6" box="[323,444,1208,1233]" gridcol="1" gridrow="9" pageId="15" pageNumber="80">47/45</td>
|
||
<td id="76896F1BFFEB89DA7D1CFEA7FDF995B6" box="[518,637,1208,1233]" gridcol="2" gridrow="9" pageId="15" pageNumber="80">43/42</td>
|
||
<td id="76896F1BFFEB89DA7DD3FEA7FCC195B6" box="[713,837,1208,1233]" gridcol="3" gridrow="9" pageId="15" pageNumber="80">35</td>
|
||
<td id="76896F1BFFEB89DA7C7FFEA7FC6595B6" box="[869,993,1208,1233]" gridcol="4" gridrow="9" pageId="15" pageNumber="80">47/39</td>
|
||
<td id="76896F1BFFEB89DA7B1BFEA7FBF995B6" box="[1025,1149,1208,1233]" gridcol="5" gridrow="9" pageId="15" pageNumber="80">51</td>
|
||
<td id="76896F1BFFEB89DA7B85FEA7FA9395B6" box="[1183,1303,1208,1233]" gridcol="6" gridrow="9" pageId="15" pageNumber="80">47/49</td>
|
||
<td id="76896F1BFFEB89DA7A7AFEA7FA5395B6" box="[1376,1495,1208,1233]" gridcol="7" gridrow="9" pageId="15" pageNumber="80">broken/49</td>
|
||
<td id="76896F1BFFEB89DA7938FEA7F91B95B6" box="[1570,1695,1208,1233]" gridcol="8" gridrow="9" pageId="15" pageNumber="80">51/48</td>
|
||
<td id="76896F1BFFEB89DA79A4FEA7F8BE95B6" box="[1726,1850,1208,1233]" gridcol="9" gridrow="9" pageId="15" pageNumber="80">45/49</td>
|
||
<td id="76896F1BFFEB89DA7840FEA7F85295B6" box="[1882,2006,1208,1233]" gridcol="10" gridrow="9" pageId="15" pageNumber="80">45/50</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFEFAF8529599" box="[151,2006,1253,1278]" gridrow="10" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFEFAFE829599" box="[151,262,1253,1278]" gridcol="0" gridrow="10" pageId="15" pageNumber="80">slt10 (♂)</th>
|
||
<td id="76896F1BFFEB89DA7E59FEFAFE389599" box="[323,444,1253,1278]" gridcol="1" gridrow="10" pageId="15" pageNumber="80">23/24</td>
|
||
<td id="76896F1BFFEB89DA7D1CFEFAFDF99599" box="[518,637,1253,1278]" gridcol="2" gridrow="10" pageId="15" pageNumber="80">24/20</td>
|
||
<td id="76896F1BFFEB89DA7DD3FEFAFCC19599" box="[713,837,1253,1278]" gridcol="3" gridrow="10" pageId="15" pageNumber="80">18/20</td>
|
||
<td id="76896F1BFFEB89DA7C7FFEFAFC659599" box="[869,993,1253,1278]" gridcol="4" gridrow="10" pageId="15" pageNumber="80">21/22</td>
|
||
<td id="76896F1BFFEB89DA7B1BFEFAFBF99599" box="[1025,1149,1253,1278]" gridcol="5" gridrow="10" pageId="15" pageNumber="80">17/18</td>
|
||
<td id="76896F1BFFEB89DA7B85FEFAFA939599" box="[1183,1303,1253,1278]" gridcol="6" gridrow="10" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7A7AFEFAFA539599" box="[1376,1495,1253,1278]" gridcol="7" gridrow="10" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7938FEFAF91B9599" box="[1570,1695,1253,1278]" gridcol="8" gridrow="10" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA79A4FEFAF8BE9599" box="[1726,1850,1253,1278]" gridcol="9" gridrow="10" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7840FEFAF8529599" box="[1882,2006,1253,1278]" gridcol="10" gridrow="10" pageId="15" pageNumber="80">–</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFF0DF852944C" box="[151,2006,1298,1323]" gridrow="11" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFF0DFE82944C" box="[151,262,1298,1323]" gridcol="0" gridrow="11" pageId="15" pageNumber="80">peb (♂)</th>
|
||
<td id="76896F1BFFEB89DA7E59FF0DFE38944C" box="[323,444,1298,1323]" gridcol="1" gridrow="11" pageId="15" pageNumber="80">n. a.</td>
|
||
<td id="76896F1BFFEB89DA7D1CFF0DFDF9944C" box="[518,637,1298,1323]" gridcol="2" gridrow="11" pageId="15" pageNumber="80">16/17</td>
|
||
<td id="76896F1BFFEB89DA7DD3FF0DFCC1944C" box="[713,837,1298,1323]" gridcol="3" gridrow="11" pageId="15" pageNumber="80">15/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7C7FFF0DFC65944C" box="[869,993,1298,1323]" gridcol="4" gridrow="11" pageId="15" pageNumber="80">17/n.a.</td>
|
||
<td id="76896F1BFFEB89DA7B1BFF0DFBF9944C" box="[1025,1149,1298,1323]" gridcol="5" gridrow="11" pageId="15" pageNumber="80">15/17</td>
|
||
<td id="76896F1BFFEB89DA7B85FF0DFA93944C" box="[1183,1303,1298,1323]" gridcol="6" gridrow="11" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7A7AFF0DFA53944C" box="[1376,1495,1298,1323]" gridcol="7" gridrow="11" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7938FF0DF91B944C" box="[1570,1695,1298,1323]" gridcol="8" gridrow="11" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA79A4FF0DF8BE944C" box="[1726,1850,1298,1323]" gridcol="9" gridrow="11" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7840FF0DF852944C" box="[1882,2006,1298,1323]" gridcol="10" gridrow="11" pageId="15" pageNumber="80">–</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFF5FF852943E" box="[151,2006,1344,1369]" gridrow="12" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFF5FFE82943E" box="[151,262,1344,1369]" gridcol="0" gridrow="12" pageId="15" pageNumber="80">pet (♂)</th>
|
||
<td id="76896F1BFFEB89DA7E59FF5FFE38943E" box="[323,444,1344,1369]" gridcol="1" gridrow="12" pageId="15" pageNumber="80">25/26</td>
|
||
<td id="76896F1BFFEB89DA7D1CFF5FFDF9943E" box="[518,637,1344,1369]" gridcol="2" gridrow="12" pageId="15" pageNumber="80">25/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7DD3FF5FFCC1943E" box="[713,837,1344,1369]" gridcol="3" gridrow="12" pageId="15" pageNumber="80">25/n. a.</td>
|
||
<td id="76896F1BFFEB89DA7C7FFF5FFC65943E" box="[869,993,1344,1369]" gridcol="4" gridrow="12" pageId="15" pageNumber="80">31/n.a.</td>
|
||
<td id="76896F1BFFEB89DA7B1BFF5FFBF9943E" box="[1025,1149,1344,1369]" gridcol="5" gridrow="12" pageId="15" pageNumber="80">27/26</td>
|
||
<td id="76896F1BFFEB89DA7B85FF5FFA93943E" box="[1183,1303,1344,1369]" gridcol="6" gridrow="12" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7A7AFF5FFA53943E" box="[1376,1495,1344,1369]" gridcol="7" gridrow="12" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7938FF5FF91B943E" box="[1570,1695,1344,1369]" gridcol="8" gridrow="12" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA79A4FF5FF8BE943E" box="[1726,1850,1344,1369]" gridcol="9" gridrow="12" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7840FF5FF852943E" box="[1882,2006,1344,1369]" gridcol="10" gridrow="12" pageId="15" pageNumber="80">–</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFF72F85294E1" box="[151,2006,1389,1414]" gridrow="13" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFF72FE8294E1" box="[151,262,1389,1414]" gridcol="0" gridrow="13" pageId="15" pageNumber="80">lts</th>
|
||
<td id="76896F1BFFEB89DA7E59FF72FE3894E1" box="[323,444,1389,1414]" gridcol="1" gridrow="13" pageId="15" pageNumber="80">146/166</td>
|
||
<td id="76896F1BFFEB89DA7D1CFF72FDF994E1" box="[518,637,1389,1414]" gridcol="2" gridrow="13" pageId="15" pageNumber="80">154/broken</td>
|
||
<td id="76896F1BFFEB89DA7DD3FF72FCC194E1" box="[713,837,1389,1414]" gridcol="3" gridrow="13" pageId="15" pageNumber="80">broken/128</td>
|
||
<td id="76896F1BFFEB89DA7C7FFF72FC6594E1" box="[869,993,1389,1414]" gridcol="4" gridrow="13" pageId="15" pageNumber="80">168/169</td>
|
||
<td id="76896F1BFFEB89DA7B1BFF72FBF994E1" box="[1025,1149,1389,1414]" gridcol="5" gridrow="13" pageId="15" pageNumber="80">broken</td>
|
||
<td id="76896F1BFFEB89DA7B85FF72FA9394E1" box="[1183,1303,1389,1414]" gridcol="6" gridrow="13" pageId="15" pageNumber="80">186/broken</td>
|
||
<td id="76896F1BFFEB89DA7A7AFF72FA5394E1" box="[1376,1495,1389,1414]" gridcol="7" gridrow="13" pageId="15" pageNumber="80">148/broken</td>
|
||
<td id="76896F1BFFEB89DA7938FF72F91B94E1" box="[1570,1695,1389,1414]" gridcol="8" gridrow="13" pageId="15" pageNumber="80">broken/191</td>
|
||
<td id="76896F1BFFEB89DA79A4FF72F8BE94E1" box="[1726,1850,1389,1414]" gridcol="9" gridrow="13" pageId="15" pageNumber="80">206/208</td>
|
||
<td id="76896F1BFFEB89DA7840FF72F85294E1" box="[1882,2006,1389,1414]" gridcol="10" gridrow="13" pageId="15" pageNumber="80">181/162</td>
|
||
</tr>
|
||
<tr id="35580667FFEB89DA7F8DFF85F85294D4" box="[151,2006,1434,1459]" gridrow="14" pageId="15" pageNumber="80">
|
||
<th id="76896F1BFFEB89DA7F8DFF85FE8294D4" box="[151,262,1434,1459]" gridcol="0" gridrow="14" pageId="15" pageNumber="80">ltas (♀)</th>
|
||
<td id="76896F1BFFEB89DA7E59FF85FE3894D4" box="[323,444,1434,1459]" gridcol="1" gridrow="14" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7D1CFF85FDF994D4" box="[518,637,1434,1459]" gridcol="2" gridrow="14" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7DD3FF85FCC194D4" box="[713,837,1434,1459]" gridcol="3" gridrow="14" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7C7FFF85FC6594D4" box="[869,993,1434,1459]" gridcol="4" gridrow="14" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7B1BFF85FBF994D4" box="[1025,1149,1434,1459]" gridcol="5" gridrow="14" pageId="15" pageNumber="80">–</td>
|
||
<td id="76896F1BFFEB89DA7B85FF85FA9394D4" box="[1183,1303,1434,1459]" gridcol="6" gridrow="14" pageId="15" pageNumber="80">31/28</td>
|
||
<td id="76896F1BFFEB89DA7A7AFF85FA5394D4" box="[1376,1495,1434,1459]" gridcol="7" gridrow="14" pageId="15" pageNumber="80">29/27</td>
|
||
<td id="76896F1BFFEB89DA7938FF85F91B94D4" box="[1570,1695,1434,1459]" gridcol="8" gridrow="14" pageId="15" pageNumber="80">25/29</td>
|
||
<td id="76896F1BFFEB89DA79A4FF85F8BE94D4" box="[1726,1850,1434,1459]" gridcol="9" gridrow="14" pageId="15" pageNumber="80">31</td>
|
||
<td id="76896F1BFFEB89DA7840FF85F85294D4" box="[1882,2006,1434,1459]" gridcol="10" gridrow="14" pageId="15" pageNumber="80">26/25</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF1754ADFFF476357F8DF8ACFE6B92C2" ID-Table-UUID="DF1754ADFFF476357F8DF8ACFE6B92C2" httpUri="http://table.plazi.org/id/DF1754ADFFF476357F8DF8ACFE6B92C2" pageId="16" pageNumber="81" startId="16.[151,239,691,717]" subCaptionStartIDs="16.[1710,1865,763,789]" subCaptionStarts="Abbr" targetBox="[165,1982,947,1502]" targetIsTable="true" targetPageId="16">
|
||
<paragraph id="8BD70425FFF476357F8DF8ACFE6B92C2" blockId="16.[151,2041,691,933]" pageId="16" pageNumber="81">
|
||
<emphasis id="B91CD837FFF476357F8DF8ACFE8F93AA" bold="true" box="[151,267,691,717]" pageId="16" pageNumber="81">TABLE 3.</emphasis>
|
||
Summary of nature and location of sensory spots, gland cell outlets and spines arranged by series in
|
||
<taxonomicName id="4C687FA6FFF476357A4FF8ACF99193AA" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[1365,1557,691,717]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="16" pageNumber="81" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFF476357A4FF8ACF99193AA" box="[1365,1557,691,717]" italics="true" pageId="16" pageNumber="81">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CD837FFF476357906F8ACF9F793AA" bold="true" box="[1564,1651,691,717]" pageId="16" pageNumber="81">
|
||
<taxonomicNameLabel id="A22F654CFFF476357906F8ACF9F793AA" box="[1564,1651,691,717]" pageId="16" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Argentina. Features appearing anteriorly on a segment do not show any background, features located centrally on a segment are indicated by a grey background, and characters occurring posteriorly on a segment are marked in black. Sensory spots on segments 5 and 6 and tubes on segments 7 and 9 are located in a sublateral to almost midlateral position. Abbreviations: ♀ or ♂, female or male condition of sexually dimorphic character; ac, acicular spine; co, fringe on trunk surface with conical subcuticular structure and basal cylindrical reinforcement structure in female; gc1, type-1 gland cell outlet; la, lateral accessory; ld, laterodorsal; ltas, lateral terminal accessory spine; lts, lateral terminal spine; lv, lateroventral; md, middorsal; pd, paradorsal; pe, penile spine; pv, paraventral; sd, subdorsal; sl, sublateral; ssp1, type-1 sensory spot; tu, tube; tuc, tube with basal cylindrical reinforcement structure in male; vl, ventrolateral; vm, ventromedial.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD70425FFF476357FBFF9ACF9AC94B9" pageId="16" pageNumber="81">
|
||
<table id="F968F685FFF489DA7FBFF9ACF83A94B9" box="[165,1982,947,1502]" gridcols="12" gridrows="12" pageId="16" pageNumber="81">
|
||
<tr id="35580667FFF489DA7FBFF9ACF83A9297" box="[165,1982,947,1008]" gridrow="0" pageId="16" pageNumber="81">
|
||
<th id="76896F1BFFF489DA7FBFF9ACFE869297" box="[165,258,947,1008]" gridcol="0" gridrow="0" pageId="16" pageNumber="81">Position Segment</th>
|
||
<th id="76896F1BFFF489DA7E77F9ACFE579297" box="[365,467,947,1008]" gridcol="1" gridrow="0" pageId="16" pageNumber="81">md</th>
|
||
<th id="76896F1BFFF489DA7D2DF9ACFD129297" box="[567,662,947,1008]" gridcol="2" gridrow="0" pageId="16" pageNumber="81">pd</th>
|
||
<th id="76896F1BFFF489DA7DFFF9ACFC979297" box="[741,787,947,1008]" gridcol="3" gridrow="0" pageId="16" pageNumber="81">sd</th>
|
||
<th id="76896F1BFFF489DA7C53F9ACFCF39297" box="[841,887,947,1008]" gridcol="4" gridrow="0" pageId="16" pageNumber="81">ld</th>
|
||
<th id="76896F1BFFF489DA7CB7F9ACFC5F9297" box="[941,987,947,1008]" gridcol="5" gridrow="0" pageId="16" pageNumber="81">ml</th>
|
||
<th id="76896F1BFFF489DA7B1EF9ACFB2D9297" box="[1028,1193,947,1008]" gridcol="6" gridrow="0" pageId="16" pageNumber="81">sl</th>
|
||
<th id="76896F1BFFF489DA7A16F9ACFA319297" box="[1292,1461,947,1008]" gridcol="7" gridrow="0" pageId="16" pageNumber="81">la</th>
|
||
<th id="76896F1BFFF489DA7910F9ACF9AA9297" box="[1546,1582,947,1008]" gridcol="8" gridrow="0" pageId="16" pageNumber="81">lv</th>
|
||
<th id="76896F1BFFF489DA7973F9ACF9139297" box="[1641,1687,947,1008]" gridcol="9" gridrow="0" pageId="16" pageNumber="81">vl</th>
|
||
<th id="76896F1BFFF489DA79C0F9ACF8BD9297" box="[1754,1849,947,1008]" gridcol="10" gridrow="0" pageId="16" pageNumber="81">vm</th>
|
||
<th id="76896F1BFFF489DA7875F9ACF83A9297" box="[1903,1982,947,1008]" gridcol="11" gridrow="0" pageId="16" pageNumber="81">
|
||
<emphasis id="B91CD837FFF476357884F9ACF83D92AB" bold="true" box="[1950,1977,947,972]" pageId="16" pageNumber="81">pv</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFF9E0F83A957F" box="[165,1982,1023,1048]" gridrow="1" pageId="16" pageNumber="81" rowspan-11="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76896F1BFFF489DA7FBFF9E0FE86957F" box="[165,258,1023,1048]" gridcol="0" gridrow="1" pageId="16" pageNumber="81">1</th>
|
||
<td id="76896F1BFFF489DA7E77F9E0FE57957F" box="[365,467,1023,1048]" gridcol="1" gridrow="1" pageId="16" pageNumber="81">gc1</td>
|
||
<td id="76896F1BFFF489DA7DFFF9E0FC97957F" box="[741,787,1023,1048]" gridcol="3" gridrow="1" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7CB7F9E0FC5F957F" box="[941,987,1023,1048]" gridcol="5" gridrow="1" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7910F9E0F9AA957F" box="[1546,1582,1023,1048]" gridcol="8" gridrow="1" pageId="16" pageNumber="81">gc1</td>
|
||
<td id="76896F1BFFF489DA79C0F9E0F8BD957F" box="[1754,1849,1023,1048]" gridcol="10" gridrow="1" pageId="16" pageNumber="81">ssp1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFE32F83A952F" box="[165,1982,1069,1096]" gridrow="2" pageId="16" pageNumber="81" rowspan-11="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="76896F1BFFF489DA7FBFFE32FE86952F" box="[165,258,1069,1096]" gridcol="0" gridrow="2" pageId="16" pageNumber="81">2</th>
|
||
<td id="76896F1BFFF489DA7E77FE32FE57952F" box="[365,467,1069,1096]" gridcol="1" gridrow="2" pageId="16" pageNumber="81">gc1; ssp1</td>
|
||
<td id="76896F1BFFF489DA7DFFFE32FC97952F" box="[741,787,1069,1096]" gridcol="3" gridrow="2" pageId="16" pageNumber="81">tu</td>
|
||
<td id="76896F1BFFF489DA7CB7FE32FC5F952F" box="[941,987,1069,1096]" gridcol="5" gridrow="2" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7973FE32F913952F" box="[1641,1687,1069,1096]" gridcol="9" gridrow="2" pageId="16" pageNumber="81">tu</td>
|
||
<td id="76896F1BFFF489DA79C0FE32F8BD952F" box="[1754,1849,1069,1096]" gridcol="10" gridrow="2" pageId="16" pageNumber="81">ssp1; gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFE45F83A9514" box="[165,1982,1114,1139]" gridrow="3" pageId="16" pageNumber="81" rowspan-11="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="76896F1BFFF489DA7FBFFE45FE869514" box="[165,258,1114,1139]" gridcol="0" gridrow="3" pageId="16" pageNumber="81">3</th>
|
||
<td id="76896F1BFFF489DA7E77FE45FE579514" box="[365,467,1114,1139]" gridcol="1" gridrow="3" pageId="16" pageNumber="81">gc1</td>
|
||
<td id="76896F1BFFF489DA7DFFFE45FC979514" box="[741,787,1114,1139]" gridcol="3" gridrow="3" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7B1EFE45FB2D9514" box="[1028,1193,1114,1139]" gridcol="6" gridrow="3" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA79C0FE45F8BD9514" box="[1754,1849,1114,1139]" gridcol="10" gridrow="3" pageId="16" pageNumber="81">gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFE97F83A95C6" box="[165,1982,1160,1185]" gridrow="4" pageId="16" pageNumber="81" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="76896F1BFFF489DA7FBFFE97FE8695C6" box="[165,258,1160,1185]" gridcol="0" gridrow="4" pageId="16" pageNumber="81">4</th>
|
||
<td id="76896F1BFFF489DA7E77FE97FE5795C6" box="[365,467,1160,1185]" gridcol="1" gridrow="4" pageId="16" pageNumber="81">ac</td>
|
||
<td id="76896F1BFFF489DA7D2DFE97FD1295C6" box="[567,662,1160,1185]" gridcol="2" gridrow="4" pageId="16" pageNumber="81">gc1</td>
|
||
<td id="76896F1BFFF489DA7875FE97F83A95C6" box="[1903,1982,1160,1185]" gridcol="11" gridrow="4" pageId="16" pageNumber="81">gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFEAAF83A95A9" box="[165,1982,1205,1230]" gridrow="5" pageId="16" pageNumber="81" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76896F1BFFF489DA7FBFFEAAFE8695A9" box="[165,258,1205,1230]" gridcol="0" gridrow="5" pageId="16" pageNumber="81">5</th>
|
||
<td id="76896F1BFFF489DA7E77FEAAFE5795A9" box="[365,467,1205,1230]" gridcol="1" gridrow="5" pageId="16" pageNumber="81">gc1</td>
|
||
<td id="76896F1BFFF489DA7DFFFEAAFC9795A9" box="[741,787,1205,1230]" gridcol="3" gridrow="5" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7B1EFEAAFB2D95A9" box="[1028,1193,1205,1230]" gridcol="6" gridrow="5" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7910FEAAF9AA95A9" box="[1546,1582,1205,1230]" gridcol="8" gridrow="5" pageId="16" pageNumber="81">tu</td>
|
||
<td id="76896F1BFFF489DA79C0FEAAF8BD95A9" box="[1754,1849,1205,1230]" gridcol="10" gridrow="5" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7875FEAAF83A95A9" box="[1903,1982,1205,1230]" gridcol="11" gridrow="5" pageId="16" pageNumber="81">gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFEFDF83A959C" box="[165,1982,1250,1275]" gridrow="6" pageId="16" pageNumber="81" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76896F1BFFF489DA7FBFFEFDFE86959C" box="[165,258,1250,1275]" gridcol="0" gridrow="6" pageId="16" pageNumber="81">6</th>
|
||
<td id="76896F1BFFF489DA7E77FEFDFE57959C" box="[365,467,1250,1275]" gridcol="1" gridrow="6" pageId="16" pageNumber="81">ac</td>
|
||
<td id="76896F1BFFF489DA7D2DFEFDFD12959C" box="[567,662,1250,1275]" gridcol="2" gridrow="6" pageId="16" pageNumber="81">gc1; ssp1</td>
|
||
<td id="76896F1BFFF489DA7B1EFEFDFB2D959C" box="[1028,1193,1250,1275]" gridcol="6" gridrow="6" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7910FEFDF9AA959C" box="[1546,1582,1250,1275]" gridcol="8" gridrow="6" pageId="16" pageNumber="81">ac</td>
|
||
<td id="76896F1BFFF489DA79C0FEFDF8BD959C" box="[1754,1849,1250,1275]" gridcol="10" gridrow="6" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7875FEFDF83A959C" box="[1903,1982,1250,1275]" gridcol="11" gridrow="6" pageId="16" pageNumber="81">gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFF0FF83A944E" box="[165,1982,1296,1321]" gridrow="7" pageId="16" pageNumber="81" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="76896F1BFFF489DA7FBFFF0FFE86944E" box="[165,258,1296,1321]" gridcol="0" gridrow="7" pageId="16" pageNumber="81">7</th>
|
||
<td id="76896F1BFFF489DA7E77FF0FFE57944E" box="[365,467,1296,1321]" gridcol="1" gridrow="7" pageId="16" pageNumber="81">gc1</td>
|
||
<td id="76896F1BFFF489DA7DFFFF0FFC97944E" box="[741,787,1296,1321]" gridcol="3" gridrow="7" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7B1EFF0FFB2D944E" box="[1028,1193,1296,1321]" gridcol="6" gridrow="7" pageId="16" pageNumber="81">tu</td>
|
||
<td id="76896F1BFFF489DA7910FF0FF9AA944E" box="[1546,1582,1296,1321]" gridcol="8" gridrow="7" pageId="16" pageNumber="81">ac</td>
|
||
<td id="76896F1BFFF489DA79C0FF0FF8BD944E" box="[1754,1849,1296,1321]" gridcol="10" gridrow="7" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7875FF0FF83A944E" box="[1903,1982,1296,1321]" gridcol="11" gridrow="7" pageId="16" pageNumber="81">gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFF22F83A9431" box="[165,1982,1341,1366]" gridrow="8" pageId="16" pageNumber="81" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-9="1">
|
||
<th id="76896F1BFFF489DA7FBFFF22FE869431" box="[165,258,1341,1366]" gridcol="0" gridrow="8" pageId="16" pageNumber="81">8</th>
|
||
<td id="76896F1BFFF489DA7E77FF22FE579431" box="[365,467,1341,1366]" gridcol="1" gridrow="8" pageId="16" pageNumber="81">ac</td>
|
||
<td id="76896F1BFFF489DA7D2DFF22FD129431" box="[567,662,1341,1366]" gridcol="2" gridrow="8" pageId="16" pageNumber="81">gc1; ssp1</td>
|
||
<td id="76896F1BFFF489DA7A16FF22FA319431" box="[1292,1461,1341,1366]" gridcol="7" gridrow="8" pageId="16" pageNumber="81">tu</td>
|
||
<td id="76896F1BFFF489DA7910FF22F9AA9431" box="[1546,1582,1341,1366]" gridcol="8" gridrow="8" pageId="16" pageNumber="81">ac</td>
|
||
<td id="76896F1BFFF489DA7875FF22F83A9431" box="[1903,1982,1341,1366]" gridcol="11" gridrow="8" pageId="16" pageNumber="81">gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFF75F83A94E4" box="[165,1982,1386,1411]" gridrow="9" pageId="16" pageNumber="81" rowspan-1="1" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-7="1">
|
||
<th id="76896F1BFFF489DA7FBFFF75FE8694E4" box="[165,258,1386,1411]" gridcol="0" gridrow="9" pageId="16" pageNumber="81">9</th>
|
||
<td id="76896F1BFFF489DA7D2DFF75FD1294E4" box="[567,662,1386,1411]" gridcol="2" gridrow="9" pageId="16" pageNumber="81">gc1; ssp1</td>
|
||
<td id="76896F1BFFF489DA7C53FF75FCF394E4" box="[841,887,1386,1411]" gridcol="4" gridrow="9" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7B1EFF75FB2D94E4" box="[1028,1193,1386,1411]" gridcol="6" gridrow="9" pageId="16" pageNumber="81">tu</td>
|
||
<td id="76896F1BFFF489DA7910FF75F9AA94E4" box="[1546,1582,1386,1411]" gridcol="8" gridrow="9" pageId="16" pageNumber="81">ac</td>
|
||
<td id="76896F1BFFF489DA7973FF75F91394E4" box="[1641,1687,1386,1411]" gridcol="9" gridrow="9" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7875FF75F83A94E4" box="[1903,1982,1386,1411]" gridcol="11" gridrow="9" pageId="16" pageNumber="81">gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFF87F83A94D6" box="[165,1982,1432,1457]" gridrow="10" pageId="16" pageNumber="81" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="76896F1BFFF489DA7FBFFF87FE8694D6" box="[165,258,1432,1457]" gridcol="0" gridrow="10" pageId="16" pageNumber="81">10</th>
|
||
<td id="76896F1BFFF489DA7E77FF87FE5794D6" box="[365,467,1432,1457]" gridcol="1" gridrow="10" pageId="16" pageNumber="81">gc1; gc1</td>
|
||
<td id="76896F1BFFF489DA7DFFFF87FC9794D6" box="[741,787,1432,1457]" gridcol="3" gridrow="10" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7B1EFF87FB2D94D6" box="[1028,1193,1432,1457]" gridcol="6" gridrow="10" pageId="16" pageNumber="81">
|
||
<specimenCount id="9D6ECFACFFF476357B3EFF87FBBE94D6" box="[1060,1082,1432,1457]" pageId="16" pageNumber="81" type="male">♂</specimenCount>
|
||
: tuc;
|
||
<specimenCount id="9D6ECFACFFF476357B6BFF87FB0294D6" box="[1137,1158,1432,1457]" pageId="16" pageNumber="81" type="female">♀</specimenCount>
|
||
: co
|
||
</td>
|
||
<td id="76896F1BFFF489DA7973FF87F91394D6" box="[1641,1687,1432,1457]" gridcol="9" gridrow="10" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7875FF87F83A94D6" box="[1903,1982,1432,1457]" gridcol="11" gridrow="10" pageId="16" pageNumber="81">gc1</td>
|
||
</tr>
|
||
<tr id="35580667FFF489DA7FBFFFDAF83A94B9" box="[165,1982,1477,1502]" gridrow="11" pageId="16" pageNumber="81" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-9="1">
|
||
<th id="76896F1BFFF489DA7FBFFFDAFE8694B9" box="[165,258,1477,1502]" gridcol="0" gridrow="11" pageId="16" pageNumber="81">11</th>
|
||
<td id="76896F1BFFF489DA7E77FFDAFE5794B9" box="[365,467,1477,1502]" gridcol="1" gridrow="11" pageId="16" pageNumber="81">gc1; gc1</td>
|
||
<td id="76896F1BFFF489DA7DFFFFDAFC9794B9" box="[741,787,1477,1502]" gridcol="3" gridrow="11" pageId="16" pageNumber="81">ssp1</td>
|
||
<td id="76896F1BFFF489DA7A16FFDAFA3194B9" box="[1292,1461,1477,1502]" gridcol="7" gridrow="11" pageId="16" pageNumber="81">
|
||
<specimenCount id="9D6ECFACFFF476357A16FFDAFAA594B9" box="[1292,1313,1477,1502]" pageId="16" pageNumber="81" type="female">♀</specimenCount>
|
||
: ltas;
|
||
<specimenCount id="9D6ECFACFFF476357A47FFDAFAF794B9" box="[1373,1395,1477,1502]" pageId="16" pageNumber="81" type="male">♂</specimenCount>
|
||
: 3x pe
|
||
</td>
|
||
<td id="76896F1BFFF489DA7910FFDAF9AA94B9" box="[1546,1582,1477,1502]" gridcol="8" gridrow="11" pageId="16" pageNumber="81">lts</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFF576347FDDFA88FABC90B6" blockId="17.[151,1436,150,609]" pageId="17" pageNumber="82">
|
||
<emphasis id="B91CD837FFF576347FDDFA88FD9A91D7" bold="true" box="[199,542,150,177]" pageId="17" pageNumber="82">Last juvenile stage moulting.</emphasis>
|
||
One specimen was fixed while moulting from the last juvenile stage to the adult stage (ZMB 12267,
|
||
<figureCitation id="135318A0FFF576347E6FFAA4FE4C91B1" box="[373,456,187,214]" captionStart="FIGURE 9" captionStartId="12.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-20@12.[151,1436,181,1531]" captionTargetPageId="12" captionText="FIGURE 9. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, ventral (B–E), optical section (A, F), and dorsal side (G, H). A. Habitus. B. Segments 1–4. C, D. Segments 5–8 at different focal levels showing aspects of the juvenile (C) and adult (D) cuticle. E. Segments 8–11. F. Segments 9–11. G. Segments 1–4. H. Segments 4–7.Abbreviations: bs, button-like structures; fe, refers to female character; juv, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in A 100 µm. Scale bar in F 50 µm, valid for B–H." figureDoi="http://doi.org/10.5281/zenodo.6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="17" pageNumber="82">Fig.9A</figureCitation>
|
||
). Each of the 11 trunk segments of the juvenile specimen was covered by numerous short cuticular hairs in its central part and showed a row of postmarginal spicula posteriorly on each segment (
|
||
<figureCitation id="135318A0FFF576347A00FAC0FA1C919E" box="[1306,1432,223,249]" captionStart="FIGURE 9" captionStartId="12.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-20@12.[151,1436,181,1531]" captionTargetPageId="12" captionText="FIGURE 9. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, ventral (B–E), optical section (A, F), and dorsal side (G, H). A. Habitus. B. Segments 1–4. C, D. Segments 5–8 at different focal levels showing aspects of the juvenile (C) and adult (D) cuticle. E. Segments 8–11. F. Segments 9–11. G. Segments 1–4. H. Segments 4–7.Abbreviations: bs, button-like structures; fe, refers to female character; juv, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in A 100 µm. Scale bar in F 50 µm, valid for B–H." figureDoi="http://doi.org/10.5281/zenodo.6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="17" pageNumber="82">Figs 9B–H</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFF576347F8DFB1CFF61907A" box="[151,229,259,285]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="17" pageNumber="82">10A,B</figureCitation>
|
||
). An acicular spine occurred middorsally on segments 4, 6, 8 and 9 (
|
||
<figureCitation id="135318A0FFF576347B1DFB1CFBE3907A" box="[1031,1127,259,285]" captionStart="FIGURE 9" captionStartId="12.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-20@12.[151,1436,181,1531]" captionTargetPageId="12" captionText="FIGURE 9. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, ventral (B–E), optical section (A, F), and dorsal side (G, H). A. Habitus. B. Segments 1–4. C, D. Segments 5–8 at different focal levels showing aspects of the juvenile (C) and adult (D) cuticle. E. Segments 8–11. F. Segments 9–11. G. Segments 1–4. H. Segments 4–7.Abbreviations: bs, button-like structures; fe, refers to female character; juv, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in A 100 µm. Scale bar in F 50 µm, valid for B–H." figureDoi="http://doi.org/10.5281/zenodo.6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="17" pageNumber="82">Figs 9H</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFF576347B6CFB1CFB2D907A" box="[1142,1193,259,285]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="17" pageNumber="82">10A</figureCitation>
|
||
), in lateral accessory position on segments 10 (
|
||
<figureCitation id="135318A0FFF576347EA9FB38FD939026" box="[435,535,295,321]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="17" pageNumber="82">Fig. 10C</figureCitation>
|
||
) and 11 (= lateral terminal accessory spine), and lateroventrally on segments 6–9 and 11 (= lateral terminal spine) (
|
||
<figureCitation id="135318A0FFF576347D11FB54FDD79002" box="[523,595,331,357]" captionStart="FIGURE 9" captionStartId="12.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-20@12.[151,1436,181,1531]" captionTargetPageId="12" captionText="FIGURE 9. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, ventral (B–E), optical section (A, F), and dorsal side (G, H). A. Habitus. B. Segments 1–4. C, D. Segments 5–8 at different focal levels showing aspects of the juvenile (C) and adult (D) cuticle. E. Segments 8–11. F. Segments 9–11. G. Segments 1–4. H. Segments 4–7.Abbreviations: bs, button-like structures; fe, refers to female character; juv, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in A 100 µm. Scale bar in F 50 µm, valid for B–H." figureDoi="http://doi.org/10.5281/zenodo.6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="17" pageNumber="82">Figs 9</figureCitation>
|
||
,
|
||
<figureCitation id="135318A0FFF576347D45FB54FD179002" box="[607,659,331,357]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="17" pageNumber="82">10A</figureCitation>
|
||
). A straight tube was found sublaterally on segments 7, 9 and 10 (
|
||
<figureCitation id="135318A0FFF576347A75FB54FF4D90EE" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="17" pageNumber="82">Fig. 10B</figureCitation>
|
||
), lateral accessorily on segment 8, lateroventrally on segment 5, and ventrolaterally on segment 2. A papillate sensory spot appeared at least paradorsally on segments 6, 8 and 9 (
|
||
<figureCitation id="135318A0FFF576347C8EFB8CFC7990CA" box="[916,1021,403,429]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="17" pageNumber="82">Fig. 10A</figureCitation>
|
||
), subdorsally on segments 10 and 11 and possibly on segment 5, ventrolaterally on segments 9–11, and ventromedially on segments 1 and 5–7.
|
||
</paragraph>
|
||
<paragraph id="8BD70425FFF576347FDDFBC4FBE39306" blockId="17.[151,1436,150,609]" pageId="17" pageNumber="82">
|
||
The moulting adult possessed in the free flap of segments 1–10 the button-like structures, which are typical of recently moulted adults (
|
||
<figureCitation id="135318A0FFF576347EC2FBE0FD01937E" box="[472,645,511,537]" captionStart="FIGURE 9" captionStartId="12.[151,250,1555,1579]" captionTargetBox="[151,1436,181,1531]" captionTargetId="figure-20@12.[151,1436,181,1531]" captionTargetPageId="12" captionText="FIGURE 9. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, ventral (B–E), optical section (A, F), and dorsal side (G, H). A. Habitus. B. Segments 1–4. C, D. Segments 5–8 at different focal levels showing aspects of the juvenile (C) and adult (D) cuticle. E. Segments 8–11. F. Segments 9–11. G. Segments 1–4. H. Segments 4–7.Abbreviations: bs, button-like structures; fe, refers to female character; juv, refers to character of last juvenile stage; lat, lateral accessory tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; mv, midventral articulation of sternal plates; mvpl, midventral placid; ps, postmarginal spicula; s, segment; vlt, ventrolateral spine. Digits after abbreviations indicate the corresponding segment number. Scale bar in A 100 µm. Scale bar in F 50 µm, valid for B–H." figureDoi="http://doi.org/10.5281/zenodo.6036953" httpUri="https://zenodo.org/record/6036953/files/figure.png" pageId="17" pageNumber="82">Fig. 9B,D,G,H</figureCitation>
|
||
). The adult lacked the middorsal spine on segment 9 and the lateral accessory spine on segment 10 as well as the sublateral tube on segment 10. In the latter position, it revealed a subcuticular conical structure with a basal cuticular reinforcement structure (
|
||
<figureCitation id="135318A0FFF576347CE9F858FBDE9306" box="[1011,1114,583,609]" captionStart="FIGURE 10" captionStartId="17.[151,250,1177,1201]" captionTargetBox="[151,1435,633,1153]" captionTargetId="figure-253@17.[151,1435,633,1153]" captionTargetPageId="17" captionText="FIGURE 10. Echinoderes goku sp. nov., last juvenile stage moulting to female, ZMB 12267, dorsal view (A, B) and optical section (C). A. Segments 7–10. B, C. Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10; fe, refers to female character; juv, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in A mark papillate sensory spots. Scale bar in A 50 µm. Scale bar in B 30 µm, valid for B and C." figureDoi="http://doi.org/10.5281/zenodo.6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="17" pageNumber="82">Fig. 10C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF1754ADFFF576347F8DFE86FD309403" ID-DOI="http://doi.org/10.5281/zenodo.6036957" ID-Zenodo-Dep="6036957" httpUri="https://zenodo.org/record/6036957/files/figure.png" pageId="17" pageNumber="82" startId="17.[151,250,1177,1201]" targetBox="[151,1435,633,1153]" targetPageId="17">
|
||
<paragraph id="8BD70425FFF576347F8DFE86FD309403" blockId="17.[151,1437,1177,1381]" pageId="17" pageNumber="82">
|
||
<emphasis id="B91CD837FFF576347F8DFE86FEA595D6" bold="true" box="[151,289,1177,1201]" pageId="17" pageNumber="82">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C687FA6FFF576347E33FE86FE5B95D6" authority="Rucci & Neuhaus & Bulnes, 2022" authorityName="Rucci & Neuhaus & Bulnes" authorityYear="2022" box="[297,479,1177,1201]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="goku" status="sp. nov.">
|
||
<emphasis id="B91CD837FFF576347E33FE86FE5B95D6" box="[297,479,1177,1201]" italics="true" pageId="17" pageNumber="82">Echinoderes goku</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B91CD837FFF576347EFDFE86FDBC95D6" bold="true" box="[487,568,1177,1201]" pageId="17" pageNumber="82">
|
||
<taxonomicNameLabel id="A22F654CFFF576347EFDFE86FDBC95D6" box="[487,568,1177,1201]" pageId="17" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, last juvenile stage moulting to female, ZMB 12267, dorsal view (
|
||
<emphasis id="B91CD837FFF576347BF0FE86FB7895D6" bold="true" box="[1258,1276,1177,1201]" pageId="17" pageNumber="82">A</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFF576347A10FE86FA9F95D6" bold="true" box="[1290,1307,1177,1201]" pageId="17" pageNumber="82">B</emphasis>
|
||
) and optical section (
|
||
<emphasis id="B91CD837FFF576347FF7FEA2FF7B95B2" bold="true" box="[237,255,1213,1237]" pageId="17" pageNumber="82">C</emphasis>
|
||
).
|
||
<emphasis id="B91CD837FFF576347E0EFEA2FEA995B2" bold="true" box="[276,301,1213,1237]" pageId="17" pageNumber="82">A.</emphasis>
|
||
Segments 7–10.
|
||
<emphasis id="B91CD837FFF576347EC0FEA2FE6F95B2" bold="true" box="[474,491,1213,1237]" pageId="17" pageNumber="82">B</emphasis>
|
||
,
|
||
<emphasis id="B91CD837FFF576347EE2FEA2FD9495B2" bold="true" box="[504,528,1213,1237]" pageId="17" pageNumber="82">C.</emphasis>
|
||
Segments 10 and 11. Abbreviations: co, sublateral, subcuticular conical structure opening on segment 10;
|
||
<superScript id="7C1DA96DFFF576347E23FEFEFEC09588" attach="right" box="[313,324,1249,1263]" fontSize="6" pageId="17" pageNumber="82">fe</superScript>
|
||
, refers to female character;
|
||
<superScript id="7C1DA96DFFF576347D78FEFEFDF09588" attach="right" box="[610,628,1249,1263]" fontSize="6" pageId="17" pageNumber="82">juv</superScript>
|
||
, refers to character of last juvenile stage; las, lateral accessory spine; lts, lateral terminal spine; mds, middorsal spine; ps, postmarginal spicula; re, cuticular reinforcement; s, segment; slt, sublateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads in
|
||
<emphasis id="B91CD837FFF576347CE6FF36FB8A9426" bold="true" box="[1020,1038,1321,1345]" pageId="17" pageNumber="82">A</emphasis>
|
||
mark papillate sensory spots. Scale bar in
|
||
<emphasis id="B91CD837FFF576347FABFF52FF479402" bold="true" box="[177,195,1357,1381]" pageId="17" pageNumber="82">A</emphasis>
|
||
50 µm. Scale bar in
|
||
<emphasis id="B91CD837FFF576347E8CFF52FE239402" bold="true" box="[406,423,1357,1381]" pageId="17" pageNumber="82">B</emphasis>
|
||
30 µm, valid for
|
||
<emphasis id="B91CD837FFF576347D43FF52FDEE9402" bold="true" box="[601,618,1357,1381]" pageId="17" pageNumber="82">B</emphasis>
|
||
and
|
||
<emphasis id="B91CD837FFF576347D81FF52FD299402" bold="true" box="[667,685,1357,1381]" pageId="17" pageNumber="82">C</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD70425FFF576347FDDFF8CFC679672" blockId="17.[151,1437,1427,1814]" pageId="17" pageNumber="82">
|
||
<emphasis id="B91CD837FFF576347FDDFF8CFE3794CA" bold="true" box="[199,435,1427,1453]" pageId="17" pageNumber="82">Additional material</emphasis>
|
||
. A reinvestigation of sublateral to midlateral subcuticular characters in segment 10 of 14 species of
|
||
<taxonomicName id="4C687FA6FFF576347E15FFA8FE3594B6" authorityName="Carus" authorityYear="1885" box="[271,433,1463,1489]" class="Cyclorhagida" family="Echinoderidae" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="family">Echinoderidae</taxonomicName>
|
||
revealed the following results: (1) Female and male
|
||
<taxonomicName id="4C687FA6FFF576347B13FFA8FA9B94B6" box="[1033,1311,1463,1489]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="unispinosus">
|
||
<emphasis id="B91CD837FFF576347B13FFA8FA9B94B6" box="[1033,1311,1463,1489]" italics="true" pageId="17" pageNumber="82">Echinoderes unispinosus</emphasis>
|
||
</taxonomicName>
|
||
unveiled a subcuticular conical structure (
|
||
<figureCitation id="135318A0FFF576347EE9FFC4FDC09491" box="[499,580,1499,1526]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="17" pageNumber="82">Fig. 5J</figureCitation>
|
||
). (2) Female and male
|
||
<taxonomicName id="4C687FA6FFF576347C51FFC3FC749492" box="[843,1008,1500,1525]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="meteorensis">
|
||
<emphasis id="B91CD837FFF576347C51FFC3FC749492" box="[843,1008,1500,1525]" italics="true" pageId="17" pageNumber="82">E. meteorensis</emphasis>
|
||
</taxonomicName>
|
||
had a subcuticular very small conical structure plus a cylindrical reinforcement structure at the base of the laterodorsal tube (
|
||
<figureCitation id="135318A0FFF576347B6BFFE0FB58977D" box="[1137,1244,1535,1562]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="17" pageNumber="82">Fig. 5H,I</figureCitation>
|
||
). (3) Female and male
|
||
<taxonomicName id="4C687FA6FFF576347FCEFC3BFEA3975A" box="[212,295,1572,1597]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="apex">
|
||
<emphasis id="B91CD837FFF576347FCEFC3BFEA3975A" box="[212,295,1572,1597]" italics="true" pageId="17" pageNumber="82">E. apex</emphasis>
|
||
</taxonomicName>
|
||
possessed a subcuticular spherical structure at the base of the laterodorsal tube (
|
||
<figureCitation id="135318A0FFF576347B85FC3CFA949759" box="[1183,1296,1571,1598]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="17" pageNumber="82">Fig. 5K,L</figureCitation>
|
||
). (4) Female and male
|
||
<taxonomicName id="4C687FA6FFF576347E12FC57FE0B9706" box="[264,399,1608,1633]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="capitatus">
|
||
<emphasis id="B91CD837FFF576347E12FC57FE0B9706" box="[264,399,1608,1633]" italics="true" pageId="17" pageNumber="82">E. capitatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C687FA6FFF576347ED3FC57FDEF9706" box="[457,619,1607,1633]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="ferrugineus">
|
||
<emphasis id="B91CD837FFF576347ED3FC57FDEF9706" box="[457,619,1607,1633]" italics="true" pageId="17" pageNumber="82">E. ferrugineus</emphasis>
|
||
</taxonomicName>
|
||
expressed a subcuticular cylindrical reinforcement structure at the base of the laterodorsal tube (
|
||
<taxonomicName id="4C687FA6FFF576347EB5FC73FDB397E2" box="[431,567,1644,1669]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="capitatus">
|
||
<emphasis id="B91CD837FFF576347EB5FC73FDB397E2" box="[431,567,1644,1669]" italics="true" pageId="17" pageNumber="82">E. capitatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="135318A0FFF576347D5CFC74FD3697E1" box="[582,690,1643,1670]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="17" pageNumber="82">Fig. 5F,G</figureCitation>
|
||
) or a tube was missing (
|
||
<taxonomicName id="4C687FA6FFF576347CD6FC73FBE997E2" box="[972,1133,1643,1669]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="ferrugineus">
|
||
<emphasis id="B91CD837FFF576347CD6FC73FBE997E2" box="[972,1133,1643,1669]" italics="true" pageId="17" pageNumber="82">E. ferrugineus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="135318A0FFF576347B66FC74FB5897E1" box="[1148,1244,1643,1670]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="17" pageNumber="82">Fig. 5M</figureCitation>
|
||
). (5) Female and male
|
||
<taxonomicName id="4C687FA6FFF576347FCEFC8FFEC697CE" authorityName="Lang" authorityYear="1949" box="[212,322,1679,1705]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="pilosus">
|
||
<emphasis id="B91CD837FFF576347FCEFC8FFEC697CE" box="[212,322,1679,1705]" italics="true" pageId="17" pageNumber="82">E. pilosus</emphasis>
|
||
</taxonomicName>
|
||
exhibited a subcuticular reinforcement structure at the base of the laterodorsal tube plus an external cylindrical cuticular sheath surrounding the basal part of the tube (
|
||
<figureCitation id="135318A0FFF576347C92FCACFC6797A9" box="[904,995,1715,1742]" captionStart="FIGURE 5" captionStartId="7.[151,250,1594,1618]" captionTargetBox="[151,1355,151,1575]" captionTargetId="figure-17@7.[151,1355,151,1575]" captionTargetPageId="7" captionText="FIGURE 5. Female specimens of species of Echinoderes, dorsal view (A, B, H, I, L–N), lateral view (C, K), and optical section (D–G, J), DIC. A–E. Echinoderes goku sp. nov. A, B. paratype ZMB 12335, segments 1–5 (A) and 6–9 (B). C. Paratype ZMB 12344, segments 2–7. D, E. Paratypes ZMB 12342 (D) and ZMB 12264 (E), segments 10 and 11 (D) and 10 (E). F–N. Segments 10 and 11. F, G. Echinoderes capitatus, ZMB 11913 (F) and 11921 (G). H, I. Echinoderes meteorensis, paratypes ZMB 11571b (H) and ZMB 11572b (I). J. Echinoderes unispinosus, paratype ZMB 11589b. K, L. Echinoderes apex, holotype ZMB 11561, paratype ZMB 11562a. M. Echinoderes ferrugineus, ZMB 11929. N. Echinoderes pilosus, paralectotype SMNH 9256; black arrow marks cuticular sheath around tube. Abbreviations: co, female-specific sublateral, subcuticular conical structure; gc1, type-1 gland cell outlet; ldt, laterodorsal tube; ltas, lateral accessory terminal spine; lts, lateral terminal spine; lvs, lateroventral spine; lvt, lateroventral tube; mds, middorsal spine; re, cuticular reinforcement; s, segment; sdt, subdorsal tube; slt, sublateral tube; sph, laterodorsal subcuticular spherical structure; vlt, ventrolateral tube. Digits after abbreviations indicate the corresponding segment number. Arrowheads mark type-1 sensory spots. White arrows point to type-1 gland cell outlets. Scale bar in B 50 μm, valid for A–C. Scale bar in D 20 µm, valid for D–M. Scale bar in N 20 µm." figureDoi="http://doi.org/10.5281/zenodo.6036938" httpUri="https://zenodo.org/record/6036938/files/figure.png" pageId="17" pageNumber="82">Fig. 5N</figureCitation>
|
||
). (6) Female
|
||
<taxonomicName id="4C687FA6FFF576347B61FCACFA1297AA" baseAuthorityName="Adrianov" baseAuthorityYear="1989" box="[1147,1430,1715,1741]" class="Cyclorhagida" family="Echinoderidae" genus="Cephalorhyncha" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="asiatica">
|
||
<emphasis id="B91CD837FFF576347B61FCACFA1297AA" box="[1147,1430,1715,1741]" italics="true" pageId="17" pageNumber="82">Cephalorhyncha asiatica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C687FA6FFF576347F8DFCC8FE119796" box="[151,405,1751,1777]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="applicitus">
|
||
<emphasis id="B91CD837FFF576347F8DFCC8FE119796" box="[151,405,1751,1777]" italics="true" pageId="17" pageNumber="82">Echinoderes applicitus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C687FA6FFF576347EBBFCC7FDA19796" box="[417,549,1751,1777]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="bathyalis">
|
||
<emphasis id="B91CD837FFF576347EBBFCC7FDA19796" box="[417,549,1751,1777]" italics="true" pageId="17" pageNumber="82">E. bathyalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C687FA6FFF576347D2BFCC7FD439796" box="[561,711,1751,1777]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="multiporus">
|
||
<emphasis id="B91CD837FFF576347D2BFCC7FD439796" box="[561,711,1751,1777]" italics="true" pageId="17" pageNumber="82">E. multiporus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C687FA6FFF576347DC9FCC7FCB29796" box="[723,822,1752,1777]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="pterus">
|
||
<emphasis id="B91CD837FFF576347DC9FCC7FCB29796" box="[723,822,1752,1777]" italics="true" pageId="17" pageNumber="82">E. pterus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C687FA6FFF576347C58FCC8FBAD9796" box="[834,1065,1751,1777]" class="Cyclorhagida" family="Echinoderidae" genus="Fissuroderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="higgins">
|
||
<emphasis id="B91CD837FFF576347C58FCC8FBAD9796" box="[834,1065,1751,1777]" italics="true" pageId="17" pageNumber="82">Fissuroderes higgins</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C687FA6FFF576347B2CFCC7FB229796" authorityName="Neuhaus & Blasche" authorityYear="2006" box="[1078,1190,1751,1777]" class="Cyclorhagida" family="Echinoderidae" genus="Fissuroderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="thermoi">
|
||
<emphasis id="B91CD837FFF576347B2CFCC7FB229796" box="[1078,1190,1751,1777]" italics="true" pageId="17" pageNumber="82">F. thermoi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C687FA6FFF576347BFAFCC8FE859672" authorityName="Sorensen" authorityYear="2008" class="Cyclorhagida" family="Echinoderidae" genus="Polacanthoderes" kingdom="Animalia" order="Echinorhagata" pageId="17" pageNumber="82" phylum="Kinorhyncha" rank="species" species="martinezi">
|
||
<emphasis id="B91CD837FFF576347BFAFCC8FE859672" italics="true" pageId="17" pageNumber="82">Polacanthoderes martinezi</emphasis>
|
||
</taxonomicName>
|
||
did not show any subcuticular structures, males were not studied.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |