520 lines
47 KiB
XML
520 lines
47 KiB
XML
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<mods:title id="3EA9BB8DDB92A691F99D91509D79976B">Integration of nuclear and mitochondrial gene sequences and morphology reveals unexpected diversity in the forest cobra (Naja melanoleuca) species complex in Central and West Africa (Serpentes: Elapidae)</mods:title>
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<mods:namePart id="1988838287D76979BDB37524E3C869BF">Wüster, Wolfgang</mods:namePart>
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<mods:namePart id="B6A2C741DC70AB816037B1AA3AE94F8B">Nagy, Zoltán T.</mods:namePart>
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<mods:namePart id="25A82DE0464F263C2AD274E75CB6AA37">Hall, Cara</mods:namePart>
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<mods:namePart id="63902A6625539323F03108A1896A7530">Wüster, Catharine E.</mods:namePart>
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<mods:namePart id="9B10A7B2E2B350959453B466B1A87365">Broadley, Donald G.</mods:namePart>
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<mods:title id="C65F50A02A442464C17F441FBE946C5C">Zootaxa</mods:title>
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<mods:date id="C1D72C230ED874B5FB1417C4B9B49CCA">2018</mods:date>
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<mods:number id="E0D408A216563377F5FC8A489B0E4CDE">2018-08-01</mods:number>
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<mods:number id="D67F84BF328E2F2F7CAFA3894ED91976">4455</mods:number>
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<treatment id="03ADDC33D47FFA77FF5931CCD6BCFD3D" ID-DOI="http://doi.org/10.5281/zenodo.5950782" ID-GBIF-Taxon="148691099" ID-Zenodo-Dep="5950782" LSID="urn:lsid:zoobank.org:act:90A2780C-7963-457E-A209-59495A80142A" httpUri="http://treatment.plazi.org/id/03ADDC33D47FFA77FF5931CCD6BCFD3D" lastPageId="15" pageId="13">
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<taxonomicName id="4C0416A6D47FFA75FF5931CCD60DF93C" authority="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley, 2018" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[151,535,1679,1706]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="savannula" status="sp. nov." subGenus="Boulengerina">
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<emphasis id="B970B137D47FFA75FF5931CCD4C8F93C" bold="true" box="[151,210,1679,1705]" italics="true" pageId="13">Naja</emphasis>
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(
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<emphasis id="B970B137D47FFA75FF2D31CCD593F93C" bold="true" box="[227,393,1679,1705]" italics="true" pageId="13">Boulengerina</emphasis>
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)
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<emphasis id="B970B137D47FFA75FE5431CCD60DF93C" bold="true" box="[410,535,1679,1705]" italics="true" pageId="13">savannula</emphasis>
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<taxonomicNameLabel id="A2430C4CD47FFA75FDD031D3D666F93F" box="[542,636,1680,1706]" pageId="13" rank="species">sp. nov.</taxonomicNameLabel>
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Broadley, Trape, Chirio & Wüster
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBB6D25D47FFA75FF59319BD779F979" blockId="13.[151,1436,1750,1987]" box="[151,867,1750,1773]" pageId="13">
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<taxonomicName id="4C0416A6D47FFA75FF59319BD555F979" authorityName="Hallowell" authorityYear="1857" box="[151,335,1750,1773]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="melanoleuca">
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<emphasis id="B970B137D47FFA75FF59319BD555F979" box="[151,335,1750,1773]" italics="true" pageId="13">Naia melanoleuca</emphasis>
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(not Hallowell)
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<treatmentCitation id="0AA54B34D47FFA75FE363195D6C9F979" author="" box="[504,723,1750,1772]" page="376" pageId="13">
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<bibRefCitation id="EF9510D4D47FFA75FE363195D6B9F979" box="[504,675,1750,1772]" pageId="13" refId="ref12241" refString="Boulenger, G. A. (1896) Catalogue of the Snakes in the British Museum (Natural History) III. British Museum (Natural History), London, 727 pp. https: // doi. org / 10.5962 / bhl. title. 54273" type="book">Boulenger, 1896</bibRefCitation>
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: 376
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(part, var. D).
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</paragraph>
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<taxonomicName id="4C0416A6D47FFA75FF5931B5D4D3F89E" box="[151,201,1782,1803]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="genus">
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<emphasis id="B970B137D47FFA75FF5931B5D4D3F89E" box="[151,201,1782,1803]" italics="true" pageId="13">Naja</emphasis>
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“banded form”
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<bibRefCitation id="EF9510D4D47FFA75FEA231B6D5E6F89E" box="[364,508,1781,1803]" pageId="13" refId="ref14133" refString="Hughes, B. (2013) Snakes of Benin, West Africa. Bulletin de la Societe Herpetologique de France, 144, 101 - 159." type="journal article">Hughes, 2013</bibRefCitation>
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: 128
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</treatmentCitation>
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.
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</paragraph>
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<subSubSection id="C31E3EAED47FFA75FF593056D7B1F8DC" pageId="13" type="reference_group">
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<paragraph id="8BBB6D25D47FFA75FF593056D047F8BF" blockId="13.[151,1436,1750,1987]" box="[151,1117,1812,1834]" pageId="13">
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<treatmentCitationGroup id="AB144A0BD47FFA75FF593056D047F8BF" box="[151,1117,1812,1834]" pageId="13">
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<taxonomicName id="4C0416A6D47FFA75FF593056D4D2F8BF" box="[151,200,1813,1834]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="genus">
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<emphasis id="B970B137D47FFA75FF593056D4D2F8BF" box="[151,200,1813,1834]" italics="true" pageId="13">Naja</emphasis>
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</taxonomicName>
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sp. 1
|
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<taxonomicName id="4C0416A6D47FFA75FECA3057D64AF8BF" authority="Hallowell, 1857" authorityName="Hallowell" authorityYear="1857" box="[260,592,1812,1834]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="melanoleuca">
|
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cf.
|
||
<emphasis id="B970B137D47FFA75FEEB3057D5BCF8BF" box="[293,422,1812,1834]" italics="true" pageId="13">melanoleuca</emphasis>
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<bibRefCitation id="EF9510D4D47FFA75FE623057D64AF8BF" box="[428,592,1812,1834]" pageId="13" refId="ref13793" refString="Hallowell, E. (1857) Notes of a collection of reptiles from the Gaboon country, West Africa, recently presented to the Academy of Natural Sciences of Philadelphia, by Dr. Henry A. Ford. Proceedings of Academy of Natural Sciences, Philadelphia, 9, 48 - 72." type="journal article">Hallowell, 1857</bibRefCitation>
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</taxonomicName>
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(yellow banded dorsum)
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<treatmentCitation id="0AA54B34D47FFA75FC9D3056D042F8BF" author="" box="[851,1112,1812,1834]" page="318" pageId="13">
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<bibRefCitation id="EF9510D4D47FFA75FC9D3056D034F8BF" box="[851,1070,1812,1834]" pageId="13" refId="ref16483" refString="Trape, J. - F. & Balde, C. (2014) A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa, 3900 (3), 301 - 338. https: // doi. org / 10.11646 / zootaxa. 3900.3.1" type="journal article">Trape & Baldé, 2014</bibRefCitation>
|
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:318
|
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BBB6D25D47FFA75FF593077D7B1F8DC" blockId="13.[151,1436,1750,1987]" box="[151,939,1843,1865]" pageId="13">
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<taxonomicName id="4C0416A6D47FFA75FF593077D4D2F8DC" box="[151,200,1844,1865]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="genus">
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<emphasis id="B970B137D47FFA75FF593077D4D2F8DC" box="[151,200,1844,1865]" italics="true" pageId="13">Naja</emphasis>
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</taxonomicName>
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||
sp. 1
|
||
<taxonomicName id="4C0416A6D47FFA75FECA3070D5BDF8DC" authorityName="Hallowell" authorityYear="1857" box="[260,423,1843,1865]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="13" phylum="Chordata" rank="species" species="melanoleuca">
|
||
cf.
|
||
<emphasis id="B970B137D47FFA75FEEB3070D5BDF8DC" box="[293,423,1843,1865]" italics="true" pageId="13">melanoleuca</emphasis>
|
||
</taxonomicName>
|
||
(banded savanna form)
|
||
<treatmentCitation id="0AA54B34D47FFA75FD523077D7BDF8DC" author="" box="[668,935,1843,1865]" page="336" pageId="13">
|
||
<bibRefCitation id="EF9510D4D47FFA75FD523077D76FF8DC" box="[668,885,1843,1865]" pageId="13" refId="ref16483" refString="Trape, J. - F. & Balde, C. (2014) A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa, 3900 (3), 301 - 338. https: // doi. org / 10.11646 / zootaxa. 3900.3.1" type="journal article">Trape & Baldé, 2014</bibRefCitation>
|
||
: 336
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47FFA75FF593012D56BF856" pageId="13" type="materials_examined">
|
||
<paragraph id="8BBB6D25D47FFA75FF593012D702F813" blockId="13.[151,1436,1750,1987]" pageId="13">
|
||
<materialsCitation id="3B6C6778D47FFA75FF593012D702F813" ID-GBIF-Occurrence="1917075148" collectingDate="2007-03-31" collectionCode="MNHN" collectorName="Laurent Chirio & Nienie, W" country="Benin" elevation="272" latitude="11.0592" location="Benin" longLatPrecision="1" longitude="2.20488" pageId="13" specimenCode="MNHN 2018.0002" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54BFD387D47FFA75FF593012D4EEF8F2" box="[151,244,1873,1895]" pageId="13" type="holotype">
|
||
<emphasis id="B970B137D47FFA75FF593012D4EEF8F2" box="[151,244,1873,1895]" italics="true" pageId="13">Holotype</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<specimenCode id="DBA2C55ED47FFA75FECB3011D5D0F8F2" box="[261,458,1873,1895]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13">MNHN 2018.0002</specimenCode>
|
||
, an adult male collected by
|
||
<collectorName id="26F108F3D47FFA75FD353011D78FF8F2" box="[763,917,1873,1895]" pageId="13">Laurent Chirio</collectorName>
|
||
on
|
||
<collectingDate id="EFFEB20DD47FFA75FC0C3012D02FF8F2" box="[962,1077,1873,1895]" pageId="13" value="2007-03-31">
|
||
<date id="FFBA4BE5D47FFA75FC0C3012D02FF8F2" box="[962,1077,1873,1895]" pageId="13" value="2007-03-31">
|
||
31/03/
|
||
<quantity id="4CFCC0C0D47FFA75FBCF3012D02FF8F2" box="[1025,1077,1873,1895]" metricMagnitude="1" metricUnit="m" metricValue="5.097779999999999" pageId="13" unit="in" value="2007.0">2007</quantity>
|
||
</date>
|
||
</collectingDate>
|
||
in
|
||
<collectorName id="26F108F3D47FFA75FB933012D0D3F8F2" box="[1117,1225,1873,1895]" pageId="13">Niénié, W</collectorName>
|
||
Biosphere Reserve,
|
||
<collectingCountry id="F3132DB5D47FFA75FF093033D51CF813" box="[199,262,1904,1926]" name="Benin" pageId="13">Benin</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE300BE2D47FFA75FEDE3033D589F813" box="[272,403,1904,1926]" direction="north" orientation="latitude" pageId="13" precision="1" value="11.0592">11.05920 °N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE300BE2D47FFA75FE533033D608F813" box="[413,530,1904,1926]" direction="east" orientation="longitude" pageId="13" precision="1" value="2.20488">2.20488 °E</geoCoordinate>
|
||
,
|
||
<elevation id="00298A16D47FFA75FDD23033D6A5F813" box="[540,703,1904,1926]" metricMagnitude="2" metricUnit="m" metricValue="2.7199999999999998" pageId="13" unit="m" value="272.0">
|
||
elevation
|
||
<quantity id="4CFCC0C0D47FFA75FDB13033D6A5F813" box="[639,703,1904,1926]" metricMagnitude="2" metricUnit="m" metricValue="2.7199999999999998" pageId="13" unit="m" value="272.0">272 m</quantity>
|
||
</elevation>
|
||
(
|
||
<figureCitation id="133F71A0D47FFA75FD033033D711F813" box="[717,779,1904,1926]" captionStart="FIGURE 7" captionStartId="14.[151,250,1937,1959]" captionTargetBox="[152,1435,1060,1916]" captionTargetId="figure@14.[152,1435,1060,1916]" captionTargetPageId="14" captionText="FIGURE 7. Naja (Boulengerina) savannula sp. nov. Top row and bottom left: holotype, MNHN 2018.0002. Bottom right: live specimen from Kindia, Guinea, showing conspicuous, broad dorsal bands and ventral banding, including narrow accessory bands (not vouchered). Photos J.-F. Trape." httpUri="https://zenodo.org/record/1456985/files/figure.png" pageId="13">Fig. 7</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47FFA75FF5930D3D56BF856" blockId="13.[151,1436,1750,1987]" pageId="13">
|
||
<materialsCitation id="3B6C6778D47FFA75FF5930D3D56BF856" ID-GBIF-Occurrence="1917075153" collectionCode="BMNH" country="Nigeria" location="Zaria" municipality="Amadou Bello University Hospital" pageId="13" specimenCode="BMNH 1975.657" specimenCount="1" stateProvince="A" typeStatus="paratype">
|
||
<typeStatus id="54BFD387D47FFA75FF5930D3D4EEF830" box="[151,244,1936,1957]" pageId="13" type="paratype">
|
||
<emphasis id="B970B137D47FFA75FF5930D3D4EEF830" box="[151,244,1936,1957]" italics="true" pageId="13">Paratype</emphasis>
|
||
</typeStatus>
|
||
:
|
||
<specimenCode id="DBA2C55ED47FFA75FECC30CCD5ACF831" box="[258,438,1934,1956]" collectionCode="BMNH" pageId="13">BMNH 1975.657</specimenCode>
|
||
, a male specimen collected on the grounds of
|
||
<collectingMunicipality id="6BDFF75FD47FFA75FC5A30CDD0E1F831" box="[916,1275,1934,1956]" pageId="13">Amadou Bello University Hospital</collectingMunicipality>
|
||
,
|
||
<location id="8EDB3BFED47FFA75FAC830CDD124F831" LSID="urn:lsid:plazi:treatment:03ADDC33D47FFA77FF5931CCD6BCFD3D:8EDB3BFED47FFA75FAC830CDD124F831" box="[1286,1342,1934,1956]" country="Nigeria" municipality="Amadou Bello University Hospital" name="Zaria" pageId="13" stateProvince="A">Zaria</location>
|
||
,
|
||
<collectingCountry id="F3132DB5D47FFA75FA8730CDD182F831" box="[1353,1432,1934,1956]" name="Nigeria" pageId="13">Nigeria</collectingCountry>
|
||
, by D.
|
||
<collectingRegion id="49C0A3C7D47FFA75FF3130EDD50FF856" box="[255,277,1966,1987]" country="Nigeria" name="Rivers" pageId="13">A</collectingRegion>
|
||
. Warrell.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47CFA76FF5937D4D4C8FEF0" pageId="14" type="diagnosis">
|
||
<paragraph id="8BBB6D25D47CFA76FF5937D4D4C8FEF0" blockId="14.[151,1437,151,1007]" pageId="14">
|
||
<emphasis id="B970B137D47CFA76FF5937D4D509FF25" bold="true" box="[151,275,151,176]" pageId="14">Diagnosis:</emphasis>
|
||
DistiNguishAblE frOm OthEr spEciEs Of thE
|
||
<taxonomicName id="4C0416A6D47CFA76FCEA37DAD7C6FF25" authorityName="Hallowell" authorityYear="1857" box="[804,988,152,176]" class="Reptilia" family="Elapidae" genus="Naia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="melanoleuca">
|
||
<emphasis id="B970B137D47CFA76FCEA37DAD7C6FF25" box="[804,988,152,176]" italics="true" pageId="14">N. melanoleuca</emphasis>
|
||
</taxonomicName>
|
||
cOmplEx by thE prEsENcE Of 3
|
||
<collectingRegion id="49C0A3C7D47CFA76FAA737DBD165FF25" box="[1385,1407,152,176]" country="South Sudan" name="Lakes" pageId="14">tO</collectingRegion>
|
||
8 sEmidividEd yEllOwish Or whitish bANds ON thE ANtEriOr dOrsAl fOrEbOdy, bEcOmiNg uNifOrm blAck cAudAd; vENtEr yEllOw with 2
|
||
<collectingRegion id="49C0A3C7D47CFA76FEF537A3D54BFF6D" box="[315,337,224,248]" country="South Sudan" name="Lakes" pageId="14">tO</collectingRegion>
|
||
8 blAck bANds. GENErAlly highEr mEAN subcAudAl scAlE cOuNts thAN thE OthEr spEciEs. GENEticAlly diAgNOsAblE thrOugh pOssEssiON Of uNiquE mitOchONdriAl hAplOtypEs (cytOchrOmE
|
||
<emphasis id="B970B137D47CFA76FB883647D04EFE89" box="[1094,1108,260,284]" italics="true" pageId="14">b</emphasis>
|
||
: GENBANk
|
||
<accessionNumber id="9457F0C6D47CFA76FB153647D145FE88" box="[1243,1375,260,285]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH337597" pageId="14">MH337597</accessionNumber>
|
||
–602; ND4:
|
||
<accessionNumber id="9457F0C6D47CFA76FF10366BD57AFED5" box="[222,352,296,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH337403" pageId="14">MH337403</accessionNumber>
|
||
–408) ANd uNiquE PRLR ANd UBN1 hAplOtypEs (PRLR:
|
||
<accessionNumber id="9457F0C6D47CFA76FC273664D076FED5" box="[1001,1132,295,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH337501" pageId="14">MH337501</accessionNumber>
|
||
–504; UBN1:
|
||
<accessionNumber id="9457F0C6D47CFA76FAC33664D194FED5" box="[1293,1422,295,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH337532" pageId="14">MH337532</accessionNumber>
|
||
– 535).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47CFA76FF09362CD0F4FC7A" pageId="14" type="description">
|
||
<paragraph id="8BBB6D25D47CFA76FF09362CD6E5FE38" blockId="14.[151,1437,151,1007]" pageId="14">
|
||
<emphasis id="B970B137D47CFA76FF09362CD5F0FE1D" bold="true" box="[199,490,367,392]" pageId="14">
|
||
Description of
|
||
<typeStatus id="54BFD387D47CFA76FEB2362CD5FFFE1D" box="[380,485,367,392]" pageId="14" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
DimENsiONs:
|
||
<collectingRegion id="49C0A3C7D47CFA76FD423633D6BAFE1D" box="[652,672,368,392]" country="Nigeria" name="Rivers" pageId="14">A</collectingRegion>
|
||
mAlE spEcimEN, sNOut–vENt lENgth
|
||
<quantity id="4CFCC0C0D47CFA76FBF4362CD0B3FE12" box="[1082,1193,367,392]" metricMagnitude="0" metricUnit="m" metricValue="1.815" pageId="14" unit="mm" value="1815.0">1815 mm</quantity>
|
||
, tAil lENgth
|
||
<quantity id="4CFCC0C0D47CFA76FAF8362CD182FE12" box="[1334,1432,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="3.76" pageId="14" unit="mm" value="376.0">376 mm</quantity>
|
||
, tOtAl lENgth
|
||
<quantity id="4CFCC0C0D47CFA76FED336D7D590FE39" box="[285,394,404,428]" metricMagnitude="0" metricUnit="m" metricValue="2.191" pageId="14" unit="mm" value="2191.0">2191 mm</quantity>
|
||
, rAtiO tOtAl lENgth: tAil lENgth 5.8.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47CFA76FF0936F4D741FD8D" blockId="14.[151,1437,151,1007]" pageId="14">
|
||
BOdy scAlAtiON: 25 scAlE rOws ArOuNd hOOd, 19 ArOuNd midbOdy, 15 ONE hEAd lENgth AhEAd Of thE vENt, All smOOth ANd ObliquE. DOrsAl scAlEs smOOth, ObliquE. VErtEbrAl rOw
|
||
<collectingRegion id="49C0A3C7D47CFA76FC47369FD7B7FE61" box="[905,941,476,500]" country="Cameroon" name="North" pageId="14">NOt</collectingRegion>
|
||
ENlArgEd. 213 vENtrAls (ANd ONE iNcOmplEtE vENtrAl bEfOrE thE ANAl), 69 subcAudAls, All dividEd, ANAl siNglE.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47CFA76FF093565D0DBFD3E" blockId="14.[151,1437,151,1007]" pageId="14">
|
||
HEAd scAlAtiON: 7/7 suprAlAbiAls, 3rd & 4th cONtAct Orbit, 6th lArgEst; 8/8 iNfrAlAbiAls, first 4 cONtAct ANtEriOr
|
||
<collectingRegion id="49C0A3C7D47CFA76FAA23565D186FDAA" box="[1388,1436,550,575]" country="Myanmar" name="Chin" pageId="14">chiN</collectingRegion>
|
||
shiElds; pOstEriOr
|
||
<collectingRegion id="49C0A3C7D47CFA76FE933509D597FDF6" box="[349,397,586,611]" country="Myanmar" name="Chin" pageId="14">chiN</collectingRegion>
|
||
shiElds iN NArrOw cONtAct At thEir ANtEriOr ENd, sEpArAtEd by mEdiAN scAlE AlONg mOst Of thEir lENgth; 1/1 prEOculAr, twicE As lONg As high; 3/3 pOstOculArs; 1/1 ANtEriOr tEmpOrAl; 3/3 pOstEriOr tEmpOrAls; rOstrAl brOAdEr thAN high, clEArly visiblE frOm AbOvE; 7 tEmpOrAls ANd NuchAls cONtActiNg thE pAriEtAls.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47CFA76FF0935F4D578FC32" blockId="14.[151,1437,151,1007]" pageId="14">
|
||
<materialsCitation id="3B6C6778D47CFA76FF0935F4D050FC82" ID-GBIF-Occurrence="1917075144" collectingDate="1940-05" county="Of" location="At" municipality="Of" pageId="14" specimenCount="1" stateProvince="tO">
|
||
PAttErN: uppEr sidE
|
||
<collectingCounty id="62DA15A9D47CFA76FE6935F5D5A4FD5A" box="[423,446,694,719]" pageId="14">Of</collectingCounty>
|
||
hEAd brOwN, lips ANd vENtrAl sidE
|
||
<collectingMunicipality id="6BDFF75FD47CFA76FC9E35F5D77DFD5A" box="[848,871,694,719]" pageId="14">Of</collectingMunicipality>
|
||
thE hEAd pAlE crEAmy yEllOwish with EAch lAbiAl scAlE lAtErAlly bOrdErEd with blAck. NEck brOwN, dOrsum bEcOmiNg grAduAlly blAck tOwArds rEAr. TwO vEry distiNct pAlE dOrsAl crOssbANds
|
||
<location id="8EDB3BFED47CFA76FE5635BCD5B6FC82" LSID="urn:lsid:plazi:treatment:03ADDC33D47FFA77FF5931CCD6BCFD3D:8EDB3BFED47CFA76FE5635BCD5B6FC82" box="[408,428,767,791]" county="Of" municipality="Of" name="At" pageId="14" stateProvince="tO">At</location>
|
||
thE lEvEl
|
||
<location id="8EDB3BFED47CFA76FDD735BDD62AFC82" LSID="urn:lsid:plazi:treatment:03ADDC33D47FFA77FF5931CCD6BCFD3D:8EDB3BFED47CFA76FDD735BDD62AFC82" box="[537,560,766,791]" county="Of" municipality="Of" name="Of" pageId="14" stateProvince="tO">Of</location>
|
||
vENtrAls (V) 24
|
||
<collectingRegion id="49C0A3C7D47CFA76FD2835BCD6E6FC82" box="[742,764,767,791]" country="South Sudan" name="Lakes" pageId="14">tO</collectingRegion>
|
||
28, twO
|
||
<location id="8EDB3BFED47CFA76FC9235BDD7AAFC82" LSID="urn:lsid:plazi:treatment:03ADDC33D47FFA77FF5931CCD6BCFD3D:8EDB3BFED47CFA76FC9235BDD7AAFC82" box="[860,944,766,791]" county="Of" municipality="Of" name="OthEr At" pageId="14" stateProvince="tO">OthEr At</location>
|
||
lEvEl
|
||
<location id="8EDB3BFED47CFA76FC3A35BDD011FC82" LSID="urn:lsid:plazi:treatment:03ADDC33D47FFA77FF5931CCD6BCFD3D:8EDB3BFED47CFA76FC3A35BDD011FC82" box="[1012,1035,766,791]" county="Of" municipality="Of" name="Of" pageId="14" stateProvince="tO">Of</location>
|
||
<collectingDate id="EFFEB20DD47CFA76FBDC3443D050FC82" box="[1042,1098,766,791]" pageId="14" value="1940-05">
|
||
<date id="FFBA4BE5D47CFA76FBDC3443D050FC82" box="[1042,1098,766,791]" pageId="14" value="1940-05">V 40</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="3B6C6778D47CFA76FB9735BDD171FC82" ID-GBIF-Occurrence="1917075151" box="[1113,1387,766,791]" collectingDate="1958-05" location="Of" municipality="At" pageId="14" specimenCount="1" stateProvince="tO">
|
||
45, thrEE
|
||
<collectingMunicipality id="6BDFF75FD47CFA76FB7135BCD0C9FC82" box="[1215,1235,767,791]" pageId="14">At</collectingMunicipality>
|
||
lEvEl
|
||
<location id="8EDB3BFED47CFA76FAD935BDD134FC82" LSID="urn:lsid:plazi:treatment:03ADDC33D47FFA77FF5931CCD6BCFD3D:8EDB3BFED47CFA76FAD935BDD134FC82" box="[1303,1326,766,791]" municipality="At" name="Of" pageId="14" stateProvince="tO">Of</location>
|
||
<collectingDate id="EFFEB20DD47CFA76FAFA3443D171FC82" box="[1332,1387,766,791]" pageId="14" value="1958-05">
|
||
<date id="FFBA4BE5D47CFA76FAFA3443D171FC82" box="[1332,1387,766,791]" pageId="14" value="1958-05">V 58</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="3B6C6778D47CFA76FAB535BDD60AFCAE" ID-GBIF-Occurrence="1917075152" collectingDate="1977-05" location="Of" municipality="At" pageId="14" specimenCount="1" stateProvince="tO">
|
||
64, thrEE lEss distiNct
|
||
<collectingMunicipality id="6BDFF75FD47CFA76FEAE3460D56EFCAE" box="[352,372,803,827]" pageId="14">At</collectingMunicipality>
|
||
lEvEl
|
||
<location id="8EDB3BFED47CFA76FE773461D5CAFCAE" LSID="urn:lsid:plazi:treatment:03ADDC33D47FFA77FF5931CCD6BCFD3D:8EDB3BFED47CFA76FE773461D5CAFCAE" box="[441,464,802,827]" municipality="At" name="Of" pageId="14" stateProvince="tO">Of</location>
|
||
<collectingDate id="EFFEB20DD47CFA76FE163460D60AFCAE" box="[472,528,803,827]" pageId="14" value="1977-05">
|
||
<date id="FFBA4BE5D47CFA76FE163460D60AFCAE" box="[472,528,803,827]" pageId="14" value="1977-05">V 77</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="3B6C6778D47CFA76FDEE3460D001FCAE" ID-GBIF-Occurrence="1917075147" box="[544,1051,802,827]" collectingDate="1998-05" location="Of" municipality="At" pageId="14" specimenCount="1" stateProvince="tO">
|
||
83, ANd thrEE slightly mArkEd
|
||
<collectingMunicipality id="6BDFF75FD47CFA76FCA23460D79AFCAE" box="[876,896,803,827]" pageId="14">At</collectingMunicipality>
|
||
lEvEl
|
||
<location id="8EDB3BFED47CFA76FC0B3461D7C6FCAE" LSID="urn:lsid:plazi:treatment:03ADDC33D47FFA77FF5931CCD6BCFD3D:8EDB3BFED47CFA76FC0B3461D7C6FCAE" box="[965,988,802,827]" municipality="At" name="Of" pageId="14" stateProvince="tO">Of</location>
|
||
<collectingDate id="EFFEB20DD47CFA76FC2D3460D001FCAE" box="[995,1051,803,827]" pageId="14" value="1998-05">
|
||
<date id="FFBA4BE5D47CFA76FC2D3460D001FCAE" box="[995,1051,803,827]" pageId="14" value="1998-05">V 98</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="3B6C6778D47CFA76FBE53460D743FCCA" ID-GBIF-Occurrence="1917075154" collectingDate="1920-05-22" location="tO" pageId="14" specimenCount="1" stateProvince="tO">
|
||
104. VENtrAls 1
|
||
<collectingRegion id="49C0A3C7D47CFA76FB163460D0F4FCAE" box="[1240,1262,803,827]" country="South Sudan" name="Lakes" pageId="14">tO</collectingRegion>
|
||
11, 13–19, 23– 31, 38–47 ANd 57–66 pAlE crEAmy yEllOwish,
|
||
<collectingDate id="EFFEB20DD47CFA76FD6E340BD71EFCCA" box="[672,772,839,863]" pageId="14" value="1920-05-22">
|
||
<date id="FFBA4BE5D47CFA76FD6E340BD71EFCCA" box="[672,772,839,863]" pageId="14" value="1920-05-22">V 20–22</date>
|
||
</collectingDate>
|
||
ANd 32
|
||
</materialsCitation>
|
||
–
|
||
<materialsCitation id="3B6C6778D47CFA76FCA63404D578FC32" ID-GBIF-Occurrence="1917075155" collectingDate="2012-05" location="tO" pageId="14" specimenCount="1" stateProvince="tO">
|
||
37 blAck;
|
||
<collectingDate id="EFFEB20DD47CFA76FC14340BD008FCCA" box="[986,1042,839,863]" pageId="14" value="2012-05">
|
||
<date id="FFBA4BE5D47CFA76FC14340BD008FCCA" box="[986,1042,839,863]" pageId="14" value="2012-05">V 12</date>
|
||
</collectingDate>
|
||
ANd 48–56 yEllOwish spOttEd with blAck spOts. FrOm vENtrAl 67
|
||
<collectingRegion id="49C0A3C7D47CFA76FDCD3428D603FC16" box="[515,537,875,899]" country="South Sudan" name="Lakes" pageId="14">tO</collectingRegion>
|
||
thE vENt, vENtrAls spOttEd with blAck grAduAlly bEcOmE cOmplEtEly blAck. SubcAudAls blAck.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47CFA76FF0934F1D0F4FC7A" blockId="14.[151,1437,151,1007]" pageId="14">
|
||
<emphasis id="B970B137D47CFA76FF0934F1D55BFC5E" bold="true" box="[199,321,946,971]" pageId="14">Variation:</emphasis>
|
||
DOrsAl scAlE rOws ON NEck 19–25, At midbOdy 19, bEfOrE vENt 12–15; vENtrAls 211–233, subcAudAls 63–77 (
|
||
<tableCitation id="C686589ED47CFA76FF243495D558FC7A" box="[234,322,982,1007]" captionStart="TABLE 8" captionStartId="16.[147,224,712,735]" captionText="TABLE 8. Variation in meristic characters in the species of the N. melanoleuca complex." pageId="14">TAblE 8</tableCitation>
|
||
). DOrsAl sEmi-dividEd yEllOw bANds 3–8; vENtrAl priNcipAl blAck bANds 2–8 (
|
||
<figureCitation id="133F71A0D47CFA76FB553495D0FAFC7A" box="[1179,1248,982,1007]" captionStart="FIGURE 7" captionStartId="14.[151,250,1937,1959]" captionTargetBox="[152,1435,1060,1916]" captionTargetId="figure@14.[152,1435,1060,1916]" captionTargetPageId="14" captionText="FIGURE 7. Naja (Boulengerina) savannula sp. nov. Top row and bottom left: holotype, MNHN 2018.0002. Bottom right: live specimen from Kindia, Guinea, showing conspicuous, broad dorsal bands and ventral banding, including narrow accessory bands (not vouchered). Photos J.-F. Trape." httpUri="https://zenodo.org/record/1456985/files/figure.png" pageId="14">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7B3DADD47CFA76FF5930D2D65EF871" httpUri="https://zenodo.org/record/1456985/files/figure.png" pageId="14" startId="14.[151,250,1937,1959]" targetBox="[152,1435,1060,1916]" targetPageId="14">
|
||
<paragraph id="8BBB6D25D47CFA76FF5930D2D65EF871" blockId="14.[151,1436,1937,2020]" pageId="14">
|
||
<emphasis id="B970B137D47CFA76FF5930D2D508F832" bold="true" box="[151,274,1937,1959]" pageId="14">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C0416A6D47CFA76FED630D1D642F832" authority="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley, 2018" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[280,600,1937,1959]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="14" phylum="Chordata" rank="species" species="savannula" status="sp. nov." subGenus="Boulengerina">
|
||
<emphasis id="B970B137D47CFA76FED630D1D553F832" box="[280,329,1938,1959]" italics="true" pageId="14">Naja</emphasis>
|
||
(
|
||
<emphasis id="B970B137D47CFA76FE9630D2D5F8F832" box="[344,482,1937,1959]" italics="true" pageId="14">Boulengerina</emphasis>
|
||
)
|
||
<emphasis id="B970B137D47CFA76FE3E30D2D642F832" box="[496,600,1937,1959]" italics="true" pageId="14">savannula</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2430C4CD47CFA76FD9030D2D6B5F832" box="[606,687,1937,1959]" pageId="14" rank="species">
|
||
<emphasis id="B970B137D47CFA76FD9030D2D6B5F832" bold="true" box="[606,687,1937,1959]" pageId="14">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Top row and bottom left: holotype, MNHN 2018.0002. Bottom right: live specimen from Kindia, Guinea, showing conspicuous, broad dorsal bands and ventral banding, including narrow accessory bands (not vouchered). Photos J.-F. Trape.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31E3EAED47DFA77FF0937D4D0B5FE88" pageId="15" type="etymology">
|
||
<paragraph id="8BBB6D25D47DFA77FF0937D4D5B7FF6D" blockId="15.[151,1437,151,680]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF0937D4D54BFF25" bold="true" box="[199,337,151,176]" pageId="15">Etymology:</emphasis>
|
||
ThE NAmE is dErivEd frOm thE cONtrActiON Of its sAvANNA hAbitAt ANd ANNulAtEd cOlOur pAttErN ANd wAs cOiNEd by BArry HughEs iN AN uNpublishEd 1968 mANuscript. WE hAvE rEtAiNEd this NAmE At thE rEquEst Of Our cOllEAguE BArry HughEs.
|
||
</paragraph>
|
||
<paragraph id="8BBB6D25D47DFA77FF093647D0B5FE88" blockId="15.[151,1437,151,680]" box="[199,1199,260,285]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF093647D584FE88" bold="true" box="[199,414,260,285]" pageId="15">Largest recorded:</emphasis>
|
||
1825+405 =
|
||
<quantity id="4CFCC0C0D47DFA77FDFA3647D6BBFE89" box="[564,673,260,284]" metricMagnitude="0" metricUnit="m" metricValue="2.23" pageId="15" unit="mm" value="2230.0">2230 mm</quantity>
|
||
, frOm MEdiNA DjikOyE,
|
||
<collectingCountry id="F3132DB5D47DFA77FC613647D012FE88" box="[943,1032,260,285]" name="Senegal" pageId="15">SENEgAl</collectingCountry>
|
||
(IRD 6155.S).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47DFA77FF093664D727FED5" box="[199,829,295,320]" pageId="15" type="description">
|
||
<paragraph id="8BBB6D25D47DFA77FF093664D727FED5" blockId="15.[151,1437,151,680]" box="[199,829,295,320]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF093664D5EEFED5" bold="true" box="[199,500,295,320]" pageId="15">Suggested common name</emphasis>
|
||
: WEst AfricAN bANdEd cObrA.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31E3EAED47DFA77FF09360FD6BCFD3D" pageId="15" type="distribution">
|
||
<paragraph id="8BBB6D25D47DFA77FF09360FD6BCFD3D" blockId="15.[151,1437,151,680]" pageId="15">
|
||
<emphasis id="B970B137D47DFA77FF09360FD578FEF0" bold="true" box="[199,354,332,357]" pageId="15">Distribution:</emphasis>
|
||
<collectingCountry id="F3132DB5D47DFA77FEA7360FD5D8FEF0" box="[361,450,332,357]" name="Senegal" pageId="15">SENEgAl</collectingCountry>
|
||
ANd
|
||
<collectingCountry id="F3132DB5D47DFA77FE38360FD655FEF0" box="[502,591,332,357]" name="Gambia" pageId="15">GAmbiA</collectingCountry>
|
||
<collectingRegion id="49C0A3C7D47DFA77FD98360FD698FEF1" box="[598,642,332,356]" country="Cameroon" name="East" pageId="15">EAst</collectingRegion>
|
||
<collectingRegion id="49C0A3C7D47DFA77FD49360FD687FEF1" box="[647,669,332,356]" country="South Sudan" name="Lakes" pageId="15">tO</collectingRegion>
|
||
<collectingRegion id="49C0A3C7D47DFA77FD6A360FD719FEF0" box="[676,771,332,357]" country="Ghana" name="Northern" pageId="15">NOrthErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D47DFA77FCC7360FD764FEF1" box="[777,894,332,356]" name="Cameroon" pageId="15">CAmErOON</collectingCountry>
|
||
(
|
||
<figureCitation id="133F71A0D47DFA77FC42360FD7CAFEF0" box="[908,976,332,357]" captionStart="FIGURE 6" captionStartId="13.[151,250,1552,1574]" captionTargetBox="[157,1421,845,1502]" captionTargetId="figure@13.[151,1436,807,1530]" captionTargetPageId="13" captionText="FIGURE 6. Distribution of the five species of the N. melanoleuca complex. A full list of voucher specimens beyond those included in multivariate analyses and Appendix 2 is available on request from the first author." httpUri="https://zenodo.org/record/1456983/files/figure.png" pageId="15">Fig. 6</figureCitation>
|
||
).
|
||
<taxonomicName id="4C0416A6D47DFA77FC2A360ED08FFEF1" authorityName="Wüster & Chirio & Trape & Ineich & Jackson & Greenbaum & Barron & Kusamba & Nagy & Storey & Hall & Wüster & Barlow & Broadley" authorityYear="2018" box="[996,1173,332,356]" class="Reptilia" family="Elapidae" genus="Naja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="15" phylum="Chordata" rank="species" species="savannula">
|
||
<emphasis id="B970B137D47DFA77FC2A360ED08FFEF1" box="[996,1173,332,356]" italics="true" pageId="15">Naja savannula</emphasis>
|
||
</taxonomicName>
|
||
AppEArs
|
||
<collectingRegion id="49C0A3C7D47DFA77FB39360FD117FEF1" box="[1271,1293,332,356]" country="South Sudan" name="Lakes" pageId="15">tO</collectingRegion>
|
||
bE rEstrictEd
|
||
<collectingRegion id="49C0A3C7D47DFA77FF593633D4B7FE1D" box="[151,173,368,392]" country="South Sudan" name="Lakes" pageId="15">tO</collectingRegion>
|
||
gAllEry fOrEst ArEAs iN AfricA iN GuiNEAN FOrEst/SAvANNA MOsAic, ExtENdiNg NOrthwArds iNtO
|
||
<collectingRegion id="49C0A3C7D47DFA77FB3C3633D133FE12" box="[1266,1321,368,391]" country="Cameroon" name="West" pageId="15">WEst</collectingRegion>
|
||
SudANiAN SAvANNA (
|
||
<bibRefCitation id="EF9510D4D47DFA77FECB36D7D583FE39" box="[261,409,404,429]" pageId="15" refId="ref13069" refString="Chirio, L. (2003) Inventaire des reptiles de la region de la Reserve de Biosphere Transfrontaliere du W (Niger / Benin / Burkina Faso: Afrique de l'Ouest). Bulletin de la Societe Herpetologique de France, 132, 13 - 41." type="journal article">ChiriO, 2003</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9510D4D47DFA77FE6B36D7D5FAFE39" box="[421,480,404,428]" pageId="15" refId="ref13117" refString="Chirio, L. (2013) Inventaire des reptiles de la region de Sangaredi (Guinee maritime). Bulletin de la Societe Herpetologique de France, 144, 67 - 100." type="journal article">2013</bibRefCitation>
|
||
; MONAstEriO
|
||
<emphasis id="B970B137D47DFA77FDBB36D6D6B4FE39" box="[629,686,404,428]" italics="true" pageId="15">et al.</emphasis>
|
||
, 2016). Our rEcOrds ArE mOstly frOm lAtitudEs 10–14°N, ExcEpt iN thE
|
||
<collectingCountry id="F3132DB5D47DFA77FF7136F4D536FE45" box="[191,300,439,464]" name="Benin" pageId="15">DAhOmEy</collectingCountry>
|
||
GAp iN
|
||
<collectingRegion id="49C0A3C7D47DFA77FE4A36FBD638FE45" box="[388,546,439,464]" country="Ghana" name="Eastern" pageId="15">
|
||
<collectingRegion id="49C0A3C7D47DFA77FE4A36FBD5C9FE45" box="[388,467,440,464]" country="American Samoa" name="Eastern" pageId="15">EAstErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D47DFA77FE1736F4D638FE45" box="[473,546,439,464]" name="Ghana" pageId="15">GhANA</collectingCountry>
|
||
</collectingRegion>
|
||
ANd
|
||
<collectingCountry id="F3132DB5D47DFA77FD9636F4D685FE45" box="[600,671,439,464]" name="Benin" pageId="15">BENiN</collectingCountry>
|
||
, whErE thE spEciEs ApprOAchEs thE Gulf Of
|
||
<collectingCountry id="F3132DB5D47DFA77FBB936F4D0D2FE45" box="[1143,1224,439,464]" name="Guinea-Bissau" pageId="15">GuiNEA</collectingCountry>
|
||
<collectingRegion id="49C0A3C7D47DFA77FB0036FBD108FE45" box="[1230,1298,440,464]" country="Tanzania" name="Pwani" pageId="15">COAst</collectingRegion>
|
||
. ThE
|
||
<collectingRegion id="49C0A3C7D47DFA77FA8336FBD186FE45" box="[1357,1436,440,464]" country="Ghana" name="Eastern" pageId="15">EAstErN</collectingRegion>
|
||
ExtENt Of thE rANgE is pOOrly uNdErstOOd.
|
||
<collectingRegion id="49C0A3C7D47DFA77FDA2369FD69AFE61" box="[620,640,476,500]" country="Nigeria" name="Rivers" pageId="15">A</collectingRegion>
|
||
spEcimEN frOm MArgui WANdAlA district,
|
||
<collectingRegion id="49C0A3C7D47DFA77FB94369FD0A3FE60" box="[1114,1209,476,501]" country="Ghana" name="Northern" pageId="15">NOrthErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D47DFA77FB0F369FD12CFE61" box="[1217,1334,476,500]" name="Cameroon" pageId="15">CAmErOON</collectingCountry>
|
||
(ApprOx.
|
||
<geoCoordinate id="EE300BE2D47DFA77FF5936BCD4FDFD8D" box="[151,231,511,536]" direction="north" orientation="latitude" pageId="15" precision="5555" value="10.5">10.5°N</geoCoordinate>
|
||
<geoCoordinate id="EE300BE2D47DFA77FF2036BCD525FD8D" box="[238,319,511,536]" direction="east" orientation="longitude" pageId="15" precision="5555" value="13.6">13.6°E</geoCoordinate>
|
||
; MNHN 1962.0022) AppEArs
|
||
<collectingRegion id="49C0A3C7D47DFA77FD423543D6B8FD8D" box="[652,674,512,536]" country="South Sudan" name="Lakes" pageId="15">tO</collectingRegion>
|
||
bE AssigNAblE
|
||
<collectingRegion id="49C0A3C7D47DFA77FC893543D747FD8D" box="[839,861,512,536]" country="South Sudan" name="Lakes" pageId="15">tO</collectingRegion>
|
||
this spEciEs, ANd ONE Of us (JFT) rEcENtly cOllEctEd
|
||
<collectingRegion id="49C0A3C7D47DFA77FF593567D4B9FDA9" box="[151,163,548,572]" country="Nigeria" name="Rivers" pageId="15">A</collectingRegion>
|
||
spEcimEN frOm MbOurA,
|
||
<quantity id="4CFCC0C0D47DFA77FE1E3567D606FDA8" box="[464,540,548,573]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="15" unit="km" value="20.0">20 km</quantity>
|
||
SW BAïbOkOum,
|
||
<collectingRegion id="49C0A3C7D47DFA77FD3F3567D7B4FDA8" box="[753,942,548,573]" country="Chad" name="Logone Oriental" pageId="15">
|
||
LOgONE
|
||
<collectingRegion id="49C0A3C7D47DFA77FC9C3567D7B4FDA8" box="[850,942,548,573]" country="Republic of the Congo" name="Orientale" pageId="15">OriENtAl</collectingRegion>
|
||
</collectingRegion>
|
||
PrOviNcE,
|
||
<collectingCountry id="F3132DB5D47DFA77FBE13567D070FDA8" box="[1071,1130,548,573]" name="Chad" pageId="15">ChAd</collectingCountry>
|
||
(
|
||
<geoCoordinate id="EE300BE2D47DFA77FBB33567D0FAFDA8" box="[1149,1248,548,573]" direction="north" orientation="latitude" pageId="15" precision="55" value="7.598">7.598°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE300BE2D47DFA77FB3E3567D146FDA8" box="[1264,1372,548,573]" direction="east" orientation="longitude" pageId="15" precision="55" value="15.596">15.596°E</geoCoordinate>
|
||
; IRD 2281.N), ANd thErE is
|
||
<collectingRegion id="49C0A3C7D47DFA77FE5B350BD5BBFDF5" box="[405,417,584,608]" country="Nigeria" name="Rivers" pageId="15">A</collectingRegion>
|
||
rEcENt rEcOrd frOm NEAr
|
||
<collectingRegion id="49C0A3C7D47DFA77FD093504D73BFDF5" box="[711,801,583,608]" country="Niger" name="Niamey" pageId="15">NiAmEy</collectingRegion>
|
||
,
|
||
<collectingRegion id="49C0A3C7D47DFA77FCE13504D775FDF5" box="[815,879,583,608]" country="Nigeria" name="Niger" pageId="15">
|
||
<collectingCountry id="F3132DB5D47DFA77FCE13504D775FDF5" box="[815,879,583,608]" name="Niger" pageId="15">NigEr</collectingCountry>
|
||
</collectingRegion>
|
||
(LC, uNpublishEd dAtA). It sEEms likEly thAt thE spEciEs hAs
|
||
<collectingRegion id="49C0A3C7D47DFA77FEE7352FD52FFD11" box="[297,309,620,644]" country="Nigeria" name="Rivers" pageId="15">A</collectingRegion>
|
||
widEr distributiON iN
|
||
<collectingRegion id="49C0A3C7D47DFA77FD8B352FD6BEFD10" box="[581,676,620,645]" country="Ghana" name="Northern" pageId="15">NOrthErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D47DFA77FD7C352FD731FD11" box="[690,811,620,644]" name="Cameroon" pageId="15">CAmErOON</collectingCountry>
|
||
, ExtrEmE
|
||
<collectingRegion id="49C0A3C7D47DFA77FC6B352FD01FFD10" box="[933,1029,620,645]" country="Zambia" name="Southern" pageId="15">sOuthErN</collectingRegion>
|
||
<collectingCountry id="F3132DB5D47DFA77FBDD352FD054FD10" box="[1043,1102,620,645]" name="Chad" pageId="15">ChAd</collectingCountry>
|
||
ANd pOssibly EvEN ExtrEmE NOrthwEstErN
|
||
<collectingCountry id="F3132DB5D47DFA77FEFE35CCD651FD3D" box="[304,587,655,680]" name="Central African Republic" pageId="15">
|
||
<collectingRegion id="49C0A3C7D47DFA77FEFE35CCD598FD3D" box="[304,386,655,680]" country="Ghana" name="Central" pageId="15">CENtrAl</collectingRegion>
|
||
AfricAN REpublic
|
||
</collectingCountry>
|
||
(
|
||
<collectingCountry id="F3132DB5D47DFA77FD9435D3D680FD3D" box="[602,666,656,680]" name="Central African Republic" pageId="15">CAR</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |