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<document id="FC37D104BF67679AA8C4D1A2237E958B" ID-CLB-Dataset="24929" ID-DOI="10.5281/zenodo.3532762" ID-GBIF-Dataset="db23a775-d25c-4253-abda-b5a007110f6d" ID-ISSN="0374-1036" ID-Zenodo-Dep="3532762" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573206962882" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2013" docId="03A3FB753F5C1E65FE30FF31441A0C18" docLanguage="en" docName="ActaEntMusNatPra.53.1.155-176.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Holotrochus yasuniensis Irmler 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="168" masterDocId="FF9A830D3F511E68FF93FFED46660A2F" masterDocTitle="New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)" masterLastPageNumber="176" masterPageNumber="155" pageNumber="168" updateTime="1698764282444" updateUser="ExternalLinkService">
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<mods:title id="74B4C1C0DF1BC5C57616443BEB3216CE">New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)</mods:title>
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<mods:namePart id="446FAF9C5DC0C7651053BC9477590D32">Irmler, Ulrich</mods:namePart>
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<mods:title id="64548BADA2E6B814EA1738BDFBB0AEF5">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="4212A2E66715C007B3DBB02E380A0A24">2013</mods:date>
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<treatment id="03A3FB753F5C1E65FE30FF31441A0C18" ID-DOI="http://doi.org/10.5281/zenodo.4427167" ID-GBIF-Taxon="160218242" ID-Zenodo-Dep="4427167" LSID="urn:lsid:plazi:treatment:03A3FB753F5C1E65FE30FF31441A0C18" httpUri="http://treatment.plazi.org/id/03A3FB753F5C1E65FE30FF31441A0C18" lastPageNumber="168" pageId="13" pageNumber="168">
<subSubSection id="C31019E83F5C1E65FE30FF3145690ADB" box="[419,783,220,244]" pageId="13" pageNumber="168" type="nomenclature">
<paragraph id="8BB54A633F5C1E65FE30FF3145690ADB" blockId="13.[419,783,220,269]" box="[419,783,220,244]" pageId="13" pageNumber="168">
<heading id="D0FDFD0F3F5C1E65FE30FF3145690ADB" bold="true" box="[419,783,220,244]" centered="true" fontSize="10" level="3" pageId="13" pageNumber="168" reason="4">
<emphasis id="B97E96713F5C1E65FE30FF3145690ADB" bold="true" box="[419,783,220,244]" pageId="13" pageNumber="168">
<taxonomicName id="4C0A31E03F5C1E65FE30FF3144D40ADB" ID-CoL="6LYP2" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[419,690,220,244]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="168" phylum="Arthropoda" rank="species" species="yasuniensis" status="sp. nov.">
<emphasis id="B97E96713F5C1E65FE30FF3144D40ADB" bold="true" box="[419,690,220,244]" italics="true" pageId="13" pageNumber="168">Holotrochus yasuniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5C1E65FD29FF3145690ADB" box="[698,783,220,244]" pageId="13" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5C1E65FDB3FF1744F40B22" box="[544,658,250,269]" pageId="13" pageNumber="168" type="description">
<paragraph id="8BB54A633F5C1E65FDB3FF1744F40B22" blockId="13.[419,783,220,269]" box="[544,658,250,269]" pageId="13" pageNumber="168">
(
<figureCitation id="133156E63F5C1E65FDB5FF17443B0B22" box="[550,605,250,269]" captionStart="Figs 811. 8" captionStartId="9.[113,150,1524,1543]" captionTargetBox="[150,1050,427,1501]" captionTargetId="figure@9.[143,1058,419,1513]" captionTargetPageId="9" captionText="Figs 811. 8 Holotrochus yasuniensis sp. nov.; 9 H. mrazi sp. nov.; 10 H. paraguensis sp. nov.; 11 H. hamatus sp. nov. Body parts:A aedeagus; B antenna, C tergite XII, D last sternite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532768" httpUri="https://zenodo.org/record/3532768/files/figure.png" pageId="13" pageNumber="168">Figs 8</figureCitation>
,
<figureCitation id="133156E63F5C1E65FDF5FF1744EA0B22" box="[614,652,250,269]" captionStart="Fig" captionStartId="17.[113,144,1541,1560]" captionTargetBox="[121,1067,217,1507]" captionTargetId="figure@17.[113,1086,211,1515]" captionTargetPageId="17" captionText="Fig. 18. Head, pronotum, and elytra of Holotrochus Erichson,1839 showing shape, punctation, and microsculpture.A H. paraguensis sp. nov.; B H.tortilis sp. nov.; C H.hamatus sp. nov.; D H.yanuniensis sp. nov.; E H. mrazi sp. nov.; F H. lobatus sp. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3532776" httpUri="https://zenodo.org/record/3532776/files/figure.png" pageId="13" pageNumber="168">18D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5C1E65FFE2FEC946CA0BA8" pageId="13" pageNumber="168" type="materials_examined">
<paragraph id="8BB54A633F5C1E65FFE2FEC944400B18" blockId="13.[113,1089,292,391]" box="[113,550,292,311]" pageId="13" pageNumber="168">
<materialsCitation id="3B62403E3F5C1E65FFE2FEC944440B18" ID-GBIF-Occurrence="2449305653" box="[113,546,292,311]" country="Ecuador" location="Yasuni" municipality="Province" pageId="13" pageNumber="168" specimenCount="1" stateProvince="Orellana" typeStatus="holotype">
<emphasis id="B97E96713F5C1E65FFE2FEC9468C0B18" bold="true" box="[113,234,292,311]" pageId="13" pageNumber="168">
<typeStatus id="54B1F4C13F5C1E65FFE2FEC946F80B18" box="[113,158,292,311]" pageId="13" pageNumber="168">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D0AF33F5C1E65FF7CFEC9475E0B18" box="[239,312,292,311]" name="Ecuador" pageId="13" pageNumber="168">Ecuador</collectingCountry>
,
<collectingMunicipality id="6BD1D0193F5C1E65FED3FEC947EA0B18" box="[320,396,292,311]" pageId="13" pageNumber="168">Province</collectingMunicipality>
<collectingRegion id="49CE84813F5C1E65FE02FEC947BB0B18" box="[401,477,292,311]" country="Ecuador" name="Orellana" pageId="13" pageNumber="168">Orellana</collectingRegion>
,
<location id="8ED51CB83F5C1E65FE76FEC944440B18" LSID="urn:lsid:plazi:treatment:03A3FB753F5C1E65FE30FF31441A0C18:8ED51CB83F5C1E65FE76FEC944440B18" box="[485,546,292,311]" country="Ecuador" municipality="Province" name="Yasuni" pageId="13" pageNumber="168" stateProvince="Orellana">Yasuni</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB54A633F5C1E65FFE2FED246CA0BA8" blockId="13.[113,1089,292,391]" pageId="13" pageNumber="168">
<emphasis id="B97E96713F5C1E65FFE2FED2469D0B7E" bold="true" box="[113,251,318,338]" pageId="13" pageNumber="168">Type material.</emphasis>
<materialsCitation id="3B62403E3F5C1E65FE90FED244B50B43" ID-GBIF-Occurrence="2449305657" collectingDate="2011-04-20" collectionCode="VAC, NHMW" collectorName="E. Cientifica Yasuni &amp; J. Torres" country="Ecuador" latitude="-0.6896667" location="Orellana" longLatPrecision="13" longitude="-77.13416" pageId="13" pageNumber="168" specimenCount="1" stateProvince="Orellana" typeStatus="holotype">
<typeStatus id="54B1F4C13F5C1E65FE90FED2473E0B7F" box="[259,344,319,338]" pageId="13" pageNumber="168" type="holotype">HOLOTYPE</typeStatus>
:,
<collectingCountry id="F31D0AF33F5C1E65FE10FED247AB0B7D" box="[387,461,319,338]" name="Ecuador" pageId="13" pageNumber="168">Ecuador</collectingCountry>
,
<collectingRegion id="49CE84813F5C1E65FE4BFED244450B7D" box="[472,547,319,338]" country="Ecuador" name="Orellana" pageId="13" pageNumber="168">Orellana</collectingRegion>
(
<geoCoordinate id="EE3E2CA43F5C1E65FDA2FED344F80B7E" box="[561,670,318,337]" degrees="77" direction="west" minutes="08.05" orientation="longitude" pageId="13" pageNumber="168" precision="9" value="-77.13416">77°08.05W</geoCoordinate>
,
<geoCoordinate id="EE3E2CA43F5C1E65FD3BFED245620B7D" box="[680,772,319,338]" degrees="0" direction="south" minutes="41.38" orientation="latitude" pageId="13" pageNumber="168" precision="9" value="-0.6896667">0°41.38S</geoCoordinate>
),
<collectorName id="26FF2FB53F5C1E65FC86FED245AF0B7D" box="[789,969,319,338]" pageId="13" pageNumber="168">E. Cientifica Yasuni</collectorName>
, collected by fogging,
<date id="FFB46CA33F5C1E65FF53FEB447420B43" box="[192,292,345,364]" pageId="13" pageNumber="168" value="2011-04-20">
<collectingDate id="EFF0954B3F5C1E65FF53FEB447420B43" box="[192,292,345,364]" pageId="13" pageNumber="168" value="2011-04-20">20.04.2011</collectingDate>
</date>
, leg.
<collectorName id="26FF2FB53F5C1E65FEC0FEB747FB0B42" box="[339,413,346,365]" pageId="13" pageNumber="168">J. Torres</collectorName>
(
<collectionCode id="ED1BD2A63F5C1E65FE3AFEB447BE0B43" box="[425,472,345,364]" pageId="13" pageNumber="168">VAC</collectionCode>
, to be deposited in
<collectionCode id="ED1BD2A63F5C1E65FD11FEB444AB0B43" box="[642,717,345,364]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="168" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B62403E3F5C1E65FD4FFEB746CE0BA8" ID-GBIF-Occurrence="2449305665" collectingDate="2011-04-20" collectionCode="UIC" collectorName="E. Cientifica Yasuni &amp; J. Torres" country="Ecuador" latitude="-0.6896667" location="Orellana" longLatPrecision="13" longitude="-77.13416" pageId="13" pageNumber="168" specimenCount="1" stateProvince="Orellana" typeStatus="paratype">
<typeStatus id="54B1F4C13F5C1E65FD4FFEB7454B0B45" box="[732,813,346,365]" pageId="13" pageNumber="168" type="paratype">PARATYPE</typeStatus>
:, with same data as holotype (
<collectionCode id="ED1BD2A63F5C1E65FFEBFE9946C40BA8" box="[120,162,372,391]" pageId="13" pageNumber="168">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5C1E65FFE2FE72448C0818" pageId="13" pageNumber="168" type="diagnosis">
<paragraph id="8BB54A633F5C1E65FFE2FE72448C0818" blockId="13.[113,1089,415,1592]" pageId="13" pageNumber="168">
<emphasis id="B97E96713F5C1E65FFE2FE7246800B98" bold="true" box="[113,230,415,439]" pageId="13" pageNumber="168">Diagnosis.</emphasis>
Concerning the size, punctation and microsculpture of the fore-body,
<taxonomicName id="4C0A31E03F5C1E65FC42FE4D46C00BF8" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="168" phylum="Arthropoda" rank="species" species="yasuniensis" status="sp. nov.">
<emphasis id="B97E96713F5C1E65FC42FE4D46C00BF8" italics="true" pageId="13" pageNumber="168">H. yasuniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5C1E65FF3CFE2D47650BF8" box="[175,259,448,471]" pageId="13" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="4C0A31E03F5C1E65FEEEFE2D44C20BF8" authority="Irmler, 2005" authorityName="Irmler" authorityYear="2005" box="[381,676,447,471]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="168" phylum="Arthropoda" rank="species" species="decumanus">
<emphasis id="B97E96713F5C1E65FEEEFE2D44710BF8" box="[381,535,448,471]" italics="true" pageId="13" pageNumber="168">H. decumanus</emphasis>
<bibRefCitation id="EF9B37923F5C1E65FD8CFE5244C20BF8" author="IRMLER U." box="[543,676,447,471]" journalOrPublisher="Studies on Neotropical Fauna and Environment" pageId="13" pageNumber="168" pagination="49 - 54" part="40" refId="ref10251" refString="IRMLER U. 2005 a: New species of the genus Mimogonia (Coleoptera: Staphylinidae, Osoriinae) from the Neotropical region. Studies on Neotropical Fauna and Environment 40: 49 - 54." title="New species of the genus Mimogonia (Coleoptera: Staphylinidae, Osoriinae) from the Neotropical region" type="journal article" year="2005">Irmler, 2005</bibRefCitation>
</taxonomicName>
from Central America that belongs to the
<taxonomicName id="4C0A31E03F5C1E65FF04FE0D47440BD8" authorityName="Sharp" authorityYear="1887" box="[151,290,480,503]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="168" phylum="Arthropoda" rank="species" species="rufopygus">
<emphasis id="B97E96713F5C1E65FF04FE0D47440BD8" box="[151,290,480,503]" italics="true" pageId="13" pageNumber="168">H. rufopygus</emphasis>
</taxonomicName>
group.
<taxonomicName id="4C0A31E03F5C1E65FEE2FE0D44100BD8" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[369,630,480,503]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="168" phylum="Arthropoda" rank="species" species="yasuniensis" status="sp. nov.">
<emphasis id="B97E96713F5C1E65FEE2FE0D44100BD8" box="[369,630,480,503]" italics="true" pageId="13" pageNumber="168">Holotrochus yasuniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5C1E65FDEFFE0D44AB0BD8" box="[636,717,480,503]" pageId="13" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
is characterised by the shape of the pronotum that seems to be unique in the Neotropical
<taxonomicName id="4C0A31E03F5C1E65FD08FDED45790838" authorityName="Erichson" authorityYear="1839" box="[667,799,512,535]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B97E96713F5C1E65FD08FDED45790838" box="[667,799,512,535]" italics="true" pageId="13" pageNumber="168">Holotrochus</emphasis>
</taxonomicName>
species. It can be identified easily by the slight lateral emargination in the anterior half.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5C1E65FFE2FDD247970FF8" pageId="13" pageNumber="168" type="description">
<paragraph id="8BB54A633F5C1E65FFE2FDD245B60878" blockId="13.[113,1089,415,1592]" box="[113,976,575,599]" pageId="13" pageNumber="168">
<emphasis id="B97E96713F5C1E65FFE2FDD2469A0878" bold="true" box="[113,252,575,599]" pageId="13" pageNumber="168">Description.</emphasis>
Length:
<quantity id="4CF2E7863F5C1E65FECFFDD247D70878" box="[348,433,575,599]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="13" pageNumber="168" unit="mm" value="5.1">5.1 mm</quantity>
. Colouration black; legs and antennae light brown.
</paragraph>
<paragraph id="8BB54A633F5C1E65FF03FDB2446F0938" blockId="13.[113,1089,415,1592]" pageId="13" pageNumber="168">
Head
<quantity id="4CF2E7863F5C1E65FF43FDB247560858" box="[208,304,607,631]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="168" unit="mm" value="0.5">0.50 mm</quantity>
long,
<quantity id="4CF2E7863F5C1E65FEE4FD8D47B10858" box="[375,471,608,632]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="168" unit="mm" value="0.81">0.81 mm</quantity>
wide; eyes large and not prominent, slightly longer than temples; sides of clypeus narrowed to anterior edge in smooth convex curve; vertex evenly punctate; punctation on fore-head denser than posteriorly; supraocular area from base of antennae to posterior edge of eyes impunctate; several setae on supraocular carina and pair of setae on neck; neck setae in equal distance to eyes as interstice between setae; weak net-like microsculpture; surface slightly shiny.
</paragraph>
<paragraph id="8BB54A633F5C1E65FF03FCCD44DD09B8" blockId="13.[113,1089,415,1592]" pageId="13" pageNumber="168">Antennae as long as head and half of pronotum combined; first antennomere rectangular and thick, second oval and slightly smaller than first; conical third two times as long as second; fourth and fifth antennomere quadrate; following antennomeres slightly increasing in width; penultimate antennomere nearly twice as wide as long.</paragraph>
<paragraph id="8BB54A633F5C1E65FF03FC4D44C60EB8" blockId="13.[113,1089,415,1592]" pageId="13" pageNumber="168">
Pronotum
<quantity id="4CF2E7863F5C1E65FE93FC4D47390998" box="[256,351,928,952]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="13" pageNumber="168" unit="mm" value="0.89">0.89 mm</quantity>
long,
<quantity id="4CF2E7863F5C1E65FE37FC4D44650998" box="[420,515,928,952]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="13" pageNumber="168" unit="mm" value="1.17">1.17 mm</quantity>
wide; widest at middle; slightly narrowed to anterior and posterior angles; in anterior half smoothly emarginate; in posterior half slightly concave, but nearly straight; anterior angles distinctly prominent; thus, anterior edge widely emarginate; sides and most parts of anterior edge margined; in dorsal aspect, lateral margin visible throughout its total length; at anterior angles, margin distinctly thicker than laterally and at anterior margin; punctation moderately dense and deep; slightly deeper than on head; interstices between punctures irregular, but, on average, twice as wide as diameter of punctures; weak net-like microsculpture; surface slightly shiny.
</paragraph>
<paragraph id="8BB54A633F5C1E65FF03FB72473D0F38" blockId="13.[113,1089,415,1592]" pageId="13" pageNumber="168">
Elytra
<quantity id="4CF2E7863F5C1E65FF4AFB4D475F0E98" box="[217,313,1184,1208]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="168" unit="mm" value="1.2">1.20 mm</quantity>
long,
<quantity id="4CF2E7863F5C1E65FE13FB7247860E98" box="[384,480,1183,1207]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="13" pageNumber="168" unit="mm" value="1.26">1.26 mm</quantity>
wide; with even punctation; punctures larger and partly denser than on pronotum; on average, distance between punctures as wide as diameter of punctures; net-like microsculpture weak; meshes of microsculpture wider than on pronotum; surface slightly shiny.
</paragraph>
<paragraph id="8BB54A633F5C1E65FF03FAF245C70F58" blockId="13.[113,1089,415,1592]" pageId="13" pageNumber="168">Abdomen with sparser and weaker punctation than on fore-body; laterally and ventrally pubescent; net-like microsculpture denser than on fore-body; thus, surface matte; abdominal tergite VIII with smooth oval depression, depression without distinct border.</paragraph>
<paragraph id="8BB54A633F5C1E65FF03FA9247970FF8" blockId="13.[113,1089,415,1592]" pageId="13" pageNumber="168">Aedeagus with similarly long basal and apical part; apical part in nearly rectangular angle to basal part; apical part carinate from base of parameres to acute apex: parameres sinuate; endophallus indistinct as short tube.</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5C1E65FFE2FA0D473B0C38" pageId="13" pageNumber="168" type="etymology">
<paragraph id="8BB54A633F5C1E65FFE2FA0D473B0C38" blockId="13.[113,1089,415,1592]" pageId="13" pageNumber="168">
<emphasis id="B97E96713F5C1E65FFE2FA0D46940FD7" bold="true" box="[113,242,1504,1528]" pageId="13" pageNumber="168">Etymology.</emphasis>
The specific name is an adjective derived from the region of Yasuni where the species was collected.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5C1E65FFE2F9F2441A0C18" box="[113,636,1567,1592]" pageId="13" pageNumber="168" type="distribution">
<paragraph id="8BB54A633F5C1E65FFE2F9F2441A0C18" blockId="13.[113,1089,415,1592]" box="[113,636,1567,1592]" pageId="13" pageNumber="168">
<emphasis id="B97E96713F5C1E65FFE2F9F247620C18" bold="true" box="[113,260,1567,1591]" pageId="13" pageNumber="168">Distribution.</emphasis>
Known only from the
<typeStatus id="54B1F4C13F5C1E65FE64F9CD44430C17" box="[503,549,1568,1592]" pageId="13" pageNumber="168">type</typeStatus>
locality
</paragraph>
</subSubSection>
</treatment>
</document>