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<document id="5F5E4ABE53D4B423E307B3517DE64A74" ID="10.11646/zootaxa.4235.1.1SLASH10244" ID-CLB-Dataset="36483" ID-DOI="10.5281/zenodo.322062" ID-GBIF-Dataset="29139646-2af5-4d4f-a7a4-b03c54cb4af8" ID-ISSN="1175-5326" ID-Zenodo-Dep="322062" ID-ZooBank="AEA565CC-1A14-4987-977A-1C269B0FE2A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487671437888" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ellenrieder, Natalia Von" docDate="2017" docId="A15040439A27FFDEFF39FE42FB7EFC4B" docLanguage="en" docName="zootaxa.4235.1.1SLASH10244.pdf" docOrigin="Zootaxa 4235 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Argia haberi Garrison &amp; von Ellenrieder, n. sp." docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="5D69383B9A32FFC8FFAEFF9EFF8AFFA4" masterDocTitle="New species of the damselfly genus Argia from Mexico, Central America and Ecuador with an emphasis on Costa Rica (Insecta: Odonata: Coenagrionidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="22" updateTime="1698437136683" updateUser="ExternalLinkService">
<mods:mods id="756727BAB973D4AF7517E00F2EB3ABFC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1807199C2B9D2C35A2DA7FF38AA6C3E9">
<mods:title id="CBABBEBDF80A1E088FE8AF246A8A8BF0">New species of the damselfly genus Argia from Mexico, Central America and Ecuador with an emphasis on Costa Rica (Insecta: Odonata: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="2BBFDA3648B0660A80C9F8332E1BADD1" type="personal">
<mods:role id="95DF7AE3D33C57787AF291E89F34988A">
<mods:roleTerm id="8E47EAC62E07D51634295E40FEA90386">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9BF9E94F5B68F15A44FA8BCCD5A6DE0F">Garrison, Rosser W.</mods:namePart>
</mods:name>
<mods:name id="F751BA52B2459E5FB200F5E1E0AE767A" type="personal">
<mods:role id="49EB42B7694236D3C81DC33A2BAED9BF">
<mods:roleTerm id="EB1CC32AAFF6F222F7EC4DFC40B40981">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7864C05770028786D50013411D35ABBF">Ellenrieder, Natalia Von</mods:namePart>
</mods:name>
<mods:typeOfResource id="65954F8785F25CD29DCC43997C0644A0">text</mods:typeOfResource>
<mods:relatedItem id="24964D90803F9F15B9369129E463E01D" type="host">
<mods:titleInfo id="D100FD77B8E68C6467310352D8AB4E7F">
<mods:title id="DFFC2B45EC63077A79F4EB0571B0BBE7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="311C36E1470046F04D8E8E3CADC07868">
<mods:date id="012259E73FCF13375C05C42FBCCB3B54">2017</mods:date>
<mods:detail id="ABD712D97EBE8FACBC3187429D3C09D8" type="volume">
<mods:number id="DE7CDF317999FC56920374C46F762DFF">4235</mods:number>
</mods:detail>
<mods:detail id="D878550F69E53737F6F94DF7001258AC" type="issue">
<mods:number id="9C333CA2BBF3053B29D3D1072B361E3D">1</mods:number>
</mods:detail>
<mods:extent id="7E5D80841D945431E4DDABD3E5BB7AD6" unit="page">
<mods:start id="5601ABA1E139B84F9264DDC0EAF0F081">1</mods:start>
<mods:end id="AC2D326A80866137B72398F048271EB2">93</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="AE08F45EA796A37E52B79692416C67D4">journal article</mods:classification>
<mods:identifier id="509B3A3B586DFD232673DD34550756E9" type="CLB-Dataset">36483</mods:identifier>
<mods:identifier id="09B6F45B64612C83CB252B3A75B0FC68" type="DOI">10.5281/zenodo.322062</mods:identifier>
<mods:identifier id="BDA52403AF7A8AA93E9892EEED8966E6" type="GBIF-Dataset">29139646-2af5-4d4f-a7a4-b03c54cb4af8</mods:identifier>
<mods:identifier id="05A729B9700A5B1319EC58CABE51B133" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BF5A29DEE3012C47F212CFECBD6BDCE7" type="Zenodo-Dep">322062</mods:identifier>
<mods:identifier id="E98BD828BBB9B5D27BFFC3DA1C96D5A9" type="ZooBank">AEA565CC-1A14-4987-977A-1C269B0FE2A8</mods:identifier>
</mods:mods>
<treatment id="A15040439A27FFDEFF39FE42FB7EFC4B" ID-DOI="http://doi.org/10.5281/zenodo.6030013" ID-GBIF-Taxon="127697286" ID-Zenodo-Dep="6030013" LSID="urn:lsid:plazi:treatment:A15040439A27FFDEFF39FE42FB7EFC4B" httpUri="http://treatment.plazi.org/id/A15040439A27FFDEFF39FE42FB7EFC4B" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="61E3A2DE9A27FFDDFF39FE42FE21FD9F" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="2946F1559A27FFDDFF39FE42FD68FE52" blockId="21.[151,1436,476,571]" box="[151,738,476,502]" pageId="21" pageNumber="22">
<heading id="720E46399A27FFDDFF39FE42FD68FE52" bold="true" box="[151,738,476,502]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="1B8D2D479A27FFDDFF39FE42FD68FE52" bold="true" box="[151,738,476,502]" pageId="21" pageNumber="22">
<taxonomicName id="EEF98AD69A27FFDDFF39FE42FD19FE52" ID-CoL="9X22N" authority="Garrison &amp; von Ellenrieder" authorityName="Garrison &amp; von Ellenrieder" box="[151,659,476,502]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="haberi" status="sp. nov.">
<emphasis id="1B8D2D479A27FFDDFF39FE42FEBBFE52" bold="true" box="[151,305,476,502]" italics="true" pageId="21" pageNumber="22">Argia haberi</emphasis>
Garrison &amp; von Ellenrieder
</taxonomicName>
,
<taxonomicNameLabel id="00BE903C9A27FFDDFD31FE42FD68FE52" box="[671,738,476,502]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2946F1559A27FFDDFF39FE60FE21FD9F" blockId="21.[151,1436,476,571]" pageId="21" pageNumber="22">
<figureCitation id="B1C2EDD09A27FFDDFF39FE60FF7BFDB3" box="[151,241,510,535]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="21" pageNumber="22">Figs. 26</figureCitation>
(head, thorax, S1–
<specimenCount id="3FFF3ADC9A27FFDDFE6BFE61FE79FDB3" box="[453,499,511,535]" pageId="21" pageNumber="22" type="male">4 ♂</specimenCount>
); 75 (S7–
<specimenCount id="3FFF3ADC9A27FFDDFDCFFE61FD16FDB3" box="[609,668,511,535]" pageId="21" pageNumber="22" type="male">10 ♂</specimenCount>
); 124 (wings); 140 (genital ligula); 160 (appendages
<specimenCount id="3FFF3ADC9A27FFDDFB5DFE61FA80FDB3" box="[1267,1290,511,535]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
); 171 (map);
<tableCitation id="647BC4EE9A27FFDDFF39FDBCFF60FD9F" box="[151,234,546,571]" captionStart="TABLE 4" captionStartId="20.[119,197,507,527]" captionTargetBox="[119,1761,656,1074]" captionTargetPageId="20" captionText="TABLE 4. Wing counts and measurements (average and standard deviation followed by range in square brackets) for various Argia species treated in this study." httpUri="http://table.plazi.org/id/7D86A1DD9A26FFDCFFD9FE65F9B0FDAB" pageId="21" pageNumber="22" tableUuid="7D86A1DD9A26FFDCFFD9FE65F9B0FDAB">Table 4</tableCitation>
(measurements).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A27FFDDFF39FDF4FCCDFCDB" pageId="21" pageNumber="22" type="etymology">
<paragraph id="2946F1559A27FFDDFF39FDF4FADFFD03" blockId="21.[151,1437,618,2014]" pageId="21" pageNumber="22">
<emphasis id="1B8D2D479A27FFDDFF39FDF4FE9CFD27" bold="true" box="[151,278,618,643]" pageId="21" pageNumber="22">Etymology</emphasis>
. Named
<taxonomicName id="EEF98AD69A27FFDDFED5FDF4FE4CFD27" box="[379,454,618,643]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="haberi">
<emphasis id="1B8D2D479A27FFDDFED5FDF4FE4CFD27" bold="true" box="[379,454,618,643]" italics="true" pageId="21" pageNumber="22">haberi</emphasis>
</taxonomicName>
(Latinized name) in honor of our friend and colleague William A. Haber, in recognition to his valuable contributions to the knowledge of
<collectingRegion id="EB3D3FB79A27FFDDFD6CFD11FC88FD03" box="[706,770,655,679]" country="New Zealand" name="West Coast" pageId="21" pageNumber="22">Costa</collectingRegion>
Rican odonates and his generous help in our studies.
</paragraph>
<paragraph id="2946F1559A27FFDDFF69FD2CFAEFFC92" blockId="21.[151,1437,618,2014]" pageId="21" pageNumber="22">
<emphasis id="1B8D2D479A27FFDDFF69FD2CFE4BFD6F" bold="true" box="[199,449,690,715]" pageId="21" pageNumber="22">Specimens examined</emphasis>
. 4 ♂.
<emphasis id="1B8D2D479A27FFDDFDBEFD2CFDDEFD6F" bold="true" box="[528,596,690,715]" pageId="21" pageNumber="22">Types</emphasis>
.
<materialsCitation id="9991FB089A27FFDDFDC8FD2CFC26FD6F" ID-GBIF-Occurrence="1434659037" box="[614,940,690,715]" country="Costa Rica" location="COSTA RICA" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="1B8D2D479A27FFDDFDC8FD2CFD59FD6F" bold="true" box="[614,723,690,715]" pageId="21" pageNumber="22">
<typeStatus id="F6424FF79A27FFDDFDC8FD2CFD59FD6F" box="[614,723,690,715]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="3FFF3ADC9A27FFDDFD72FD2DFD7EFD6E" box="[732,756,691,714]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
:
<collectingCountry id="51EEB1C59A27FFDDFCADFD2DFC26FD6F" box="[771,940,691,715]" name="Costa Rica" pageId="21" pageNumber="22">COSTA RICA</collectingCountry>
</materialsCitation>
,
<materialsCitation id="9991FB089A27FFDDFC14FD2DFCBAFCB7" ID-GBIF-Occurrence="1434659044" collectingDate="2006-03-27" collectionCode="FSCA" collectorName="F. Sibley" country="Costa Rica" county="Pan American" elevation="1710" latitude="9.471666" location="Tolomuco" longLatPrecision="21" longitude="-83.69667" municipality="Bosque" pageId="21" pageNumber="22" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
<collectingRegion id="EB3D3FB79A27FFDDFC14FD2DFB94FD6F" box="[954,1054,691,715]" country="Costa Rica" name="San Jose" pageId="21" pageNumber="22">San José</collectingRegion>
Prov.: km 118 on
<collectingCounty id="C02789D99A27FFDDFB57FD2DFA16FD6F" box="[1273,1436,690,715]" pageId="21" pageNumber="22">Pan American</collectingCounty>
highway,
<collectingMunicipality id="C9226B2F9A27FFDDFEA9FD49FED1FD4A" box="[263,347,727,750]" pageId="21" pageNumber="22">Bosque</collectingMunicipality>
del
<location id="2C26A78E9A27FFDDFE3DFD48FD8DFD4B" LSID="urn:lsid:plazi:treatment:A15040439A27FFDEFF39FE42FB7EFC4B:2C26A78E9A27FFDDFE3DFD48FD8DFD4B" box="[403,519,726,751]" country="Costa Rica" county="Pan American" latitude="9.471666" longLatPrecision="21" longitude="-83.69667" municipality="Bosque" name="Tolomuco" pageId="21" pageNumber="22" stateProvince="San Jose">Tolomuco</location>
, in seeps and trickles through brushy pasture on forested hillside (
<geoCoordinate id="4CCD97929A27FFDDFABBFD49FA12FD4A" box="[1301,1432,727,751]" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="15" value="9.471666">9°28'18'' N</geoCoordinate>
,
<geoCoordinate id="4CCD97929A27FFDDFF39FD65FEA3FCB6" box="[151,297,763,787]" direction="west" orientation="longitude" pageId="21" pageNumber="22" precision="15" value="-83.69667">83°41'48'' W</geoCoordinate>
,
<quantity id="EE015CB09A27FFDDFE98FD65FE1AFCB7" box="[310,400,763,787]" metricMagnitude="3" metricUnit="m" metricValue="1.71" pageId="21" pageNumber="22" unit="m" value="1710.0">
<elevation id="A2D416669A27FFDDFE98FD65FE1AFCB7" box="[310,400,763,787]" metricMagnitude="3" metricUnit="m" metricValue="1.71" pageId="21" pageNumber="22" unit="m" value="1710.0">1710 m</elevation>
</quantity>
),
<date id="5D47D7959A27FFDDFE08FD65FDADFCB7" box="[422,551,762,787]" pageId="21" pageNumber="22" value="2006-03-27">
<collectingDate id="4D032E7D9A27FFDDFE08FD65FDADFCB7" box="[422,551,762,787]" pageId="21" pageNumber="22" value="2006-03-27">27 iii 2006</collectingDate>
</date>
,
<collectorName id="840C94839A27FFDDFD9AFD65FD13FCB7" box="[564,665,762,787]" pageId="21" pageNumber="22">F. Sibley</collectorName>
leg. [
<collectionCode id="4FE869909A27FFDDFD74FD65FCA3FCB7" box="[730,809,763,787]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="21" pageNumber="22">FSCA</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="9991FB089A27FFDDFC92FD65FBE9FCB7" ID-GBIF-Occurrence="1434659018" box="[828,1123,763,787]" country="Costa Rica" location="COSTA RICA" pageId="21" pageNumber="22" specimenCount="1" typeStatus="paratype">
<typeStatus id="F6424FF79A27FFDDFC92FD65FC24FCB7" box="[828,942,763,787]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="51EEB1C59A27FFDDFC14FD65FBE9FCB7" box="[954,1123,763,787]" name="Costa Rica" pageId="21" pageNumber="22">COSTA RICA</collectingCountry>
</materialsCitation>
,
<materialsCitation id="9991FB089A27FFDDFBC1FD65FE5FFC93" ID-GBIF-Occurrence="1434659036" country="Costa Rica" location="COSTA RICA" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="San Jose" typeStatus="paratype">
<collectingRegion id="EB3D3FB79A27FFDDFBC1FD65FB58FCB7" box="[1135,1234,763,787]" country="Costa Rica" name="San Jose" pageId="21" pageNumber="22">San José</collectingRegion>
Prov.:
<specimenCount id="3FFF3ADC9A27FFDDFA89FD65FADFFCB6" box="[1319,1365,763,787]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
, same data as holotype but [RWG]
</materialsCitation>
;
<materialsCitation id="9991FB089A27FFDDFE4EFC81FC15FC93" ID-GBIF-Occurrence="1434659022" box="[480,927,798,823]" collectionCode="CSCA" country="Costa Rica" location="COSTA RICA" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A27FFDDFE4EFC81FD86FC92" box="[480,524,799,823]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
, same data as holotype but [
<collectionCode id="4FE869909A27FFDDFCE9FC81FC12FC93" box="[839,920,799,823]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="21" pageNumber="22">CSCA</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="9991FB089A27FFDDFC05FC81FAEFFC92" ID-GBIF-Occurrence="1434659024" box="[939,1381,798,823]" country="Costa Rica" location="COSTA RICA" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="3FFF3ADC9A27FFDDFC05FC81FC5DFC92" box="[939,983,799,823]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
, same data as holotype but [TWD].
</materialsCitation>
</paragraph>
<paragraph id="2946F1559A27FFDDFF69FCDDFCCDFCDB" blockId="21.[151,1437,618,2014]" pageId="21" pageNumber="22">
A medium-sized largely blue species with slightly amber wings (
<figureCitation id="B1C2EDD09A27FFDDFC4EFCDCFBCCFCFF" box="[992,1094,834,859]" captionStart="FIGURES 124 – 125" captionStartId="67.[151,264,1874,1897]" captionTargetBox="[157,1419,203,1815]" captionTargetId="figure@67.[151,1436,193,1853]" captionTargetPageId="67" captionText="FIGURES 124 – 125. Male wings, right pair." httpUri="https://zenodo.org/record/322090/files/figure.png" pageId="21" pageNumber="22">Fig. 124</figureCitation>
) and appendages (
<figureCitation id="B1C2EDD09A27FFDDFA83FCDCFA1EFCFF" box="[1325,1428,834,859]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="21" pageNumber="22">Fig. 160</figureCitation>
) superficially similar to those of
<taxonomicName id="EEF98AD69A27FFDDFE52FCF9FD43FCDB" authority="Calvert" authorityName="Calvert" box="[508,713,870,895]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="rhoadsi">
<emphasis id="1B8D2D479A27FFDDFE52FCF9FDFAFCDA" box="[508,624,870,894]" italics="true" pageId="21" pageNumber="22">A. rhoadsi</emphasis>
Calvert
</taxonomicName>
(
<figureCitation id="B1C2EDD09A27FFDDFD76FCF8FCB1FCDB" box="[728,827,870,895]" captionStart="FIGURES 161 – 164" captionStartId="81.[151,264,1912,1935]" captionTargetBox="[151,1422,202,1890]" captionTargetId="figure@81.[151,1436,193,1892]" captionTargetPageId="81" captionText="FIGURES 161 – 164. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, ventral view." httpUri="https://zenodo.org/record/322104/files/figure.png" pageId="21" pageNumber="22">Fig. 161</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A27FFDEFF69FC14FD3AFE53" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="2946F1559A27FFDDFF69FC14FCE5FBF3" blockId="21.[151,1437,618,2014]" pageId="21" pageNumber="22">
<emphasis id="1B8D2D479A27FFDDFF69FC14FDA9FC07" bold="true" box="[199,547,906,931]" pageId="21" pageNumber="22">
Description of male
<typeStatus id="F6424FF79A27FFDDFE12FC14FDA9FC07" box="[444,547,906,931]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
</emphasis>
. Head (
<figureCitation id="B1C2EDD09A27FFDDFDD0FC14FD5EFC07" box="[638,724,906,931]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="21" pageNumber="22">Fig. 26</figureCitation>
): Labium blue lined with black at base and with a small black medial spot connected to basal black, anteclypeus blue; postclypeus blue margined with black at base and with a basal black spot mediolaterally extending from base; base of mandibles and genae blue, antefrons blue; remainder of epicranium black with the following blue: large postocular spot confluent with eye, small triangular spot anterolateral to lateral ocellus, a wash of light brown on posterior most margin of occiput; antennae black, rear of head except for narrow pale margin bordering eye entirely black.
</paragraph>
<paragraph id="2946F1559A27FFDDFF69FBFCFAE8FA42" blockId="21.[151,1437,618,2014]" pageId="21" pageNumber="22">
Prothorax black with following areas pale: posterior margin of anterior lobe, dorsolateral spot on middle lobe, laterobasal 0.10 of posterior margin of pronotum, propleuron except for brown notopleural suture. Pale areas of pterothorax blue, with broad black middorsal stripe about three times as wide as pale antehumeral stripe, the latter narrowing dorsally; black humeral stripe extending from anterior half of mesinfraepisternum and constricted above and widening irregularly with a thick posterior fork at upper fourth, anterior portion of stripe narrow connecting below antealar crest with middorsal stripe above and with a short ventrally directed finger on dorsal portion of obsolete interpleural suture; interpleural suture with narrow black stripe separating paler areas on side of thorax (
<figureCitation id="B1C2EDD09A27FFDDFF31FAC0FF79FAD3" box="[159,243,1374,1399]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="21" pageNumber="22">Fig. 26</figureCitation>
). Wings slightly amber (as in
<figureCitation id="B1C2EDD09A27FFDDFD90FAC0FD11FAD3" box="[574,667,1374,1399]" captionStart="FIGURES 124 – 125" captionStartId="67.[151,264,1874,1897]" captionTargetBox="[157,1419,203,1815]" captionTargetId="figure@67.[151,1436,193,1853]" captionTargetPageId="67" captionText="FIGURES 124 – 125. Male wings, right pair." httpUri="https://zenodo.org/record/322090/files/figure.png" pageId="21" pageNumber="22">Fig. 124</figureCitation>
) with venation black; pterostigma dark brown, surmounting 1.5 cells in Fw, 2 cells in Hw; postnodals Fw 15/15, Hw 13/13; postquadrangular cells Fw 4/4, Hw 3/3; RP2 at Fw 7/7, Hw 6/6. Coxae and trochanters pale except for wash of black on medial portions of coxae and dorsum of trochanters; internal 0.50 base surface of femora and external surface of tibiae pale, remainder of legs and armature black.
</paragraph>
<paragraph id="2946F1559A27FFDDFF69FA6FFEEFF8A2" blockId="21.[151,1437,618,2014]" pageId="21" pageNumber="22">
Abdomen (
<figureCitation id="B1C2EDD09A27FFDDFEE7FA6FFE21F9AE" box="[329,427,1521,1546]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="21" pageNumber="22">Figs. 26</figureCitation>
,
<figureCitation id="B1C2EDD09A27FFDDFE15FA6FFE52F9AE" box="[443,472,1521,1546]" captionStart="FIGURES 73 – 79" captionStartId="55.[151,264,1620,1643]" captionTargetBox="[157,1423,211,1533]" captionTargetId="figure@55.[151,1436,193,1600]" captionTargetPageId="55" captionText="FIGURES 73 – 79. Color pattern of male S 7 – 10 of other species, lateral view." httpUri="https://zenodo.org/record/322078/files/figure.png" pageId="21" pageNumber="22">75</figureCitation>
) mostly black; S1 blue with a basal black ring at basal 0.50 and with black spot extending dorsally to apical 0.70; S2 black with a campanulate dorsal spot at basal 0.60, laterally with black separated below by blue, genital lobe slightly angulate, annulus black; S3 with distal 0.70 black, this receding above allowing for an extension of middorsal blue stripe ending at basal 0.40 of segment; S4–6 similar to S3 but basal blue further restricted basally; on S6, narrow upper offshoot of black at proximal 0.10 almost isolates dorsal blue, thus almost forming a middorsal spot; S7 black with a narrow blue ring; S8–9 blue except for narrow incomplete vertical black spot apically on intersegmental membrane on S8; S10 black, blue dorsally; torus pale, appendages black.
</paragraph>
<paragraph id="2946F1559A27FFDDFF69F88FFD58F8D6" blockId="21.[151,1437,618,2014]" pageId="21" pageNumber="22">
Genital ligula (
<figureCitation id="B1C2EDD09A27FFDDFEDFF88FFE5BF88E" box="[369,465,1809,1834]" captionStart="FIGURES 140 – 141" captionStartId="73.[151,264,1840,1863]" captionTargetBox="[158,1432,233,1749]" captionTargetId="figure@73.[151,1436,193,1819]" captionTargetPageId="73" captionText="FIGURES 140 – 141. Genital ligula of other species. (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322096/files/figure.png" pageId="21" pageNumber="22">Fig. 140</figureCitation>
) with apical segment spatulate broadening basally with short lateral sclerotized lobes, ental surface of genital ligula concave; small inner fold present, sclerotized portion proximal to flexure lacking a microspinulate patch on ental surface on each side.
</paragraph>
<paragraph id="2946F1559A27FFDEFF69F8E0FD0FFF5C" blockId="21.[151,1437,618,2014]" lastBlockId="22.[151,1437,151,1008]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
Torus large, transversely oval, swollen ventrally, occupying 0.50 ventral margin of torifer and not overlapping bilobed epiproct (
<figureCitation id="B1C2EDD09A27FFDDFEC0F83FFE52F81E" box="[366,472,1953,1978]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="21" pageNumber="22">Fig. 160</figureCitation>
c); area around epiproct and base of same black; cercus (
<figureCitation id="B1C2EDD09A27FFDDFB0DF83FFA9DF81E" box="[1187,1303,1953,1978]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="21" pageNumber="22">Figs. 160</figureCitation>
a, c) short, quadrangular, about half as long as paraproct (
<figureCitation id="B1C2EDD09A27FFDDFD02F85BFC84F87A" box="[684,782,1989,2014]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="21" pageNumber="22">Fig. 160</figureCitation>
b), outer margin convex, slightly bilobed apically, medial margin armed with a prominent decumbent tooth, slightly concave medially; medial margin linear, planate dorsomedially; paraproct unilobed, its ventral branch vestigial and much smaller than broadly triangular upper branch, its tip blunt, posterior margin linear.
</paragraph>
<paragraph id="2946F1559A24FFDEFF69FE9AFCB3FEB8" blockId="22.[151,1437,151,1008]" box="[199,825,260,285]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69FE9AFED8FEB9" bold="true" box="[199,338,260,285]" pageId="22" pageNumber="23">Dimensions</emphasis>
. Hw 22.9, abdomen 29.4, total length 37.7.
</paragraph>
<paragraph id="2946F1559A24FFDEFF69FEB9FE2EFEE4" blockId="22.[151,1437,151,1008]" box="[199,420,295,320]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69FEB9FEADFEE4" bold="true" box="[199,295,295,320]" pageId="22" pageNumber="23">Female:</emphasis>
Unknown.
</paragraph>
<paragraph id="2946F1559A24FFDEFF69FED2FD3AFE53" blockId="22.[151,1437,151,1008]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69FED2FE5FFEC1" bold="true" box="[199,469,332,357]" pageId="22" pageNumber="23">
Variation in
<typeStatus id="F6424FF79A24FFDEFEF1FED2FE5FFEC1" box="[351,469,332,357]" pageId="22" pageNumber="23" type="paratype">paratypes</typeStatus>
</emphasis>
. Little variation was observed in the series collected at the
<typeStatus id="F6424FF79A24FFDEFBC5FED3FB11FEC1" box="[1131,1179,333,357]" pageId="22" pageNumber="23">type</typeStatus>
locality. One male has reduced black on labrum; the other
<typeStatus id="F6424FF79A24FFDEFD80FEEEFD04FE2C" box="[558,654,368,392]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
has the black humeral stripe branched as in
<typeStatus id="F6424FF79A24FFDEFB22FEF1FB64FE2C" box="[1164,1262,367,392]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
but stripe is of uniform width medially. Pterostigma surmounting 1–2 cells; postnodals: Fw 15–16, Hw 13–14; postquadrangular cells Fw 4, Hw 3–4; RP2 at Fw 7–8, Hw 6–7.
<emphasis id="1B8D2D479A24FFDEFD1EFE29FCB0FE74" bold="true" box="[688,826,439,464]" pageId="22" pageNumber="23">Dimensions</emphasis>
.
<specimenCount id="3FFF3ADC9A24FFDEFCE5FE26FCE9FE6B" box="[843,867,440,463]" pageId="22" pageNumber="23" type="male">♂</specimenCount>
: Hw 22.9 ± 0.4 [22.5–23.3], abdomen 29.4 ± 1.0 [28.3–30.2], total length 37.6 ± 1.1 [36.8–38.4].
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A24FFDEFF69FD9CFA8FFC9F" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="2946F1559A24FFDEFF69FD9CFA8FFC9F" blockId="22.[151,1437,151,1008]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69FD9CFEB0FDBF" bold="true" box="[199,314,514,539]" pageId="22" pageNumber="23">Diagnosis</emphasis>
. Appendage morphology superficially resembles that of
<taxonomicName id="EEF98AD69A24FFDEFC60FD9DFBCFFDBF" box="[974,1093,515,539]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhoadsi">
<emphasis id="1B8D2D479A24FFDEFC60FD9DFBCFFDBF" box="[974,1093,515,539]" italics="true" pageId="22" pageNumber="23">A. rhoadsi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFBF6FD9CFB36FDBF" box="[1112,1212,514,539]" captionStart="FIGURES 161 – 164" captionStartId="81.[151,264,1912,1935]" captionTargetBox="[151,1422,202,1890]" captionTargetId="figure@81.[151,1436,193,1892]" captionTargetPageId="81" captionText="FIGURES 161 – 164. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, ventral view." httpUri="https://zenodo.org/record/322104/files/figure.png" pageId="22" pageNumber="23">Fig. 161</figureCitation>
), as both possess a quadrate cercus and a unilobate paraproct extending well beyond level of cercus. However, the decumbent tooth is not as prominent and is placed more basally in
<taxonomicName id="EEF98AD69A24FFDEFD0BFDD5FC93FDC7" box="[677,793,587,611]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhoadsi">
<emphasis id="1B8D2D479A24FFDEFD0BFDD5FC93FDC7" box="[677,793,587,611]" italics="true" pageId="22" pageNumber="23">A. rhoadsi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFC86FDD4FC0DFDC7" box="[808,903,586,611]" captionStart="FIGURES 161 – 164" captionStartId="81.[151,264,1912,1935]" captionTargetBox="[151,1422,202,1890]" captionTargetId="figure@81.[151,1436,193,1892]" captionTargetPageId="81" captionText="FIGURES 161 – 164. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, ventral view." httpUri="https://zenodo.org/record/322104/files/figure.png" pageId="22" pageNumber="23">Fig. 161</figureCitation>
) compared with
<taxonomicName id="EEF98AD69A24FFDEFBEBFDD5FB27FDC7" box="[1093,1197,587,611]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="haberi">
<emphasis id="1B8D2D479A24FFDEFBEBFDD5FB27FDC7" box="[1093,1197,587,611]" italics="true" pageId="22" pageNumber="23">A. haberi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFB12FDD4FA94FDC7" box="[1212,1310,586,611]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="22" pageNumber="23">Fig. 160</figureCitation>
). The torus in
<taxonomicName id="EEF98AD69A24FFDEFF19FDF1FEA5FD23" box="[183,303,623,647]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhoadsi">
<emphasis id="1B8D2D479A24FFDEFF19FDF1FEA5FD23" box="[183,303,623,647]" italics="true" pageId="22" pageNumber="23">A. rhoadsi</emphasis>
</taxonomicName>
is more elongate and partially overlaps the epiproct (
<figureCitation id="B1C2EDD09A24FFDEFC0BFDF0FB83FD23" box="[933,1033,622,647]" captionStart="FIGURES 161 – 164" captionStartId="81.[151,264,1912,1935]" captionTargetBox="[151,1422,202,1890]" captionTargetId="figure@81.[151,1436,193,1892]" captionTargetPageId="81" captionText="FIGURES 161 – 164. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view; (d) right cercus, ventral view." httpUri="https://zenodo.org/record/322104/files/figure.png" pageId="22" pageNumber="23">Fig. 161</figureCitation>
c), compared with the shorter nonoverlapping torus in
<taxonomicName id="EEF98AD69A24FFDEFE22FD0DFE72FD0F" box="[396,504,659,683]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="haberi">
<emphasis id="1B8D2D479A24FFDEFE22FD0DFE72FD0F" box="[396,504,659,683]" italics="true" pageId="22" pageNumber="23">A. haberi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFDA2FD0CFDF9FD0F" box="[524,627,658,683]" captionStart="FIGURES 158 – 160" captionStartId="80.[151,264,1826,1849]" captionTargetBox="[189,1398,197,1790]" captionTargetId="figure@80.[188,1398,193,1805]" captionTargetPageId="80" captionText="FIGURES 158 – 160. Male caudal appendages of other species. (a) mediodorsal view; (b) lateral view; (c) dorsal view." httpUri="https://zenodo.org/record/322103/files/figure.png" pageId="22" pageNumber="23">Fig. 160</figureCitation>
c). The greatest difference is in the morphology of the genital ligula: terminal segment rectangular with two flagella and small poorly developed lateral lobes at flexure in
<taxonomicName id="EEF98AD69A24FFDEFA8BFD29FA16FD6B" box="[1317,1436,695,719]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhoadsi">
<emphasis id="1B8D2D479A24FFDEFA8BFD29FA16FD6B" box="[1317,1436,695,719]" italics="true" pageId="22" pageNumber="23">A. rhoadsi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFF31FD44FF75FD57" box="[159,255,730,755]" captionStart="FIGURES 140 – 141" captionStartId="73.[151,264,1840,1863]" captionTargetBox="[158,1432,233,1749]" captionTargetId="figure@73.[151,1436,193,1819]" captionTargetPageId="73" captionText="FIGURES 140 – 141. Genital ligula of other species. (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322096/files/figure.png" pageId="22" pageNumber="23">Fig. 141</figureCitation>
) versus a spatulate terminal segment lacking flagella and with relatively larger lateral lobes in
<taxonomicName id="EEF98AD69A24FFDEFA9DFD45FA16FD57" box="[1331,1436,731,755]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="haberi">
<emphasis id="1B8D2D479A24FFDEFA9DFD45FA16FD57" box="[1331,1436,731,755]" italics="true" pageId="22" pageNumber="23">A. haberi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFF31FD60FE89FCB3" box="[159,259,766,791]" captionStart="FIGURES 140 – 141" captionStartId="73.[151,264,1840,1863]" captionTargetBox="[158,1432,233,1749]" captionTargetId="figure@73.[151,1436,193,1819]" captionTargetPageId="73" captionText="FIGURES 140 – 141. Genital ligula of other species. (a) lateral view; (b) ectal view; (c) entolateral view." httpUri="https://zenodo.org/record/322096/files/figure.png" pageId="22" pageNumber="23">Fig. 140</figureCitation>
). Although males of both species possess amber-colored wings, the pale coloration in
<taxonomicName id="EEF98AD69A24FFDEFB73FD61FAD9FCB3" box="[1245,1363,767,791]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhoadsi">
<emphasis id="1B8D2D479A24FFDEFB73FD61FAD9FCB3" box="[1245,1363,767,791]" italics="true" pageId="22" pageNumber="23">A. rhoadsi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFACAFD60FF3CFC9F" captionStart="FIGURES 27 – 29" captionStartId="46.[151,264,1840,1863]" captionTargetBox="[167,1389,231,1661]" captionTargetId="figure@46.[151,1436,193,1819]" captionTargetPageId="46" captionText="FIGURES 27 – 29. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322069/files/figure.png" pageId="22" pageNumber="23">Figs. 27</figureCitation>
,
<figureCitation id="B1C2EDD09A24FFDEFF6FFCBCFF54FC9F" box="[193,222,802,827]" captionStart="FIGURES 73 – 79" captionStartId="55.[151,264,1620,1643]" captionTargetBox="[157,1423,211,1533]" captionTargetId="figure@55.[151,1436,193,1600]" captionTargetPageId="55" captionText="FIGURES 73 – 79. Color pattern of male S 7 – 10 of other species, lateral view." httpUri="https://zenodo.org/record/322078/files/figure.png" pageId="22" pageNumber="23">76</figureCitation>
) is an aquamarine blue compared to a more intense violaceous blue in
<taxonomicName id="EEF98AD69A24FFDEFC56FCBDFBEAFC9F" box="[1016,1120,803,827]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="haberi">
<emphasis id="1B8D2D479A24FFDEFC56FCBDFBEAFC9F" box="[1016,1120,803,827]" italics="true" pageId="22" pageNumber="23">A. haberi</emphasis>
</taxonomicName>
(
<figureCitation id="B1C2EDD09A24FFDEFBDEFCBCFB47FC9F" box="[1136,1229,802,827]" captionStart="FIGURES 24 – 26" captionStartId="45.[151,264,1836,1859]" captionTargetBox="[160,1408,215,1680]" captionTargetId="figure@45.[151,1436,193,1815]" captionTargetPageId="45" captionText="FIGURES 24 – 26. Color pattern of male head, thorax, and S 1 – 4 of other species. Head, dorsal view, thorax and S 1 – 4, lateral view." httpUri="https://zenodo.org/record/322068/files/figure.png" pageId="22" pageNumber="23">Figs. 26</figureCitation>
,
<figureCitation id="B1C2EDD09A24FFDEFB77FCBCFB73FC9F" box="[1241,1273,802,827]" captionStart="FIGURES 73 – 79" captionStartId="55.[151,264,1620,1643]" captionTargetBox="[157,1423,211,1533]" captionTargetId="figure@55.[151,1436,193,1600]" captionTargetPageId="55" captionText="FIGURES 73 – 79. Color pattern of male S 7 – 10 of other species, lateral view." httpUri="https://zenodo.org/record/322078/files/figure.png" pageId="22" pageNumber="23">75</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A24FFDEFF69FCD8FF7BFC6F" pageId="22" pageNumber="23" type="discussion">
<paragraph id="2946F1559A24FFDEFF69FCD8FC1BFC27" blockId="22.[151,1437,151,1008]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69FCD8FEBEFCFB" bold="true" box="[199,308,838,863]" pageId="22" pageNumber="23">Remarks</emphasis>
. Despite superficially similarity between
<taxonomicName id="EEF98AD69A24FFDEFCBCFCD9FCF6FCFB" box="[786,892,839,863]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="haberi">
<emphasis id="1B8D2D479A24FFDEFCBCFCD9FCF6FCFB" box="[786,892,839,863]" italics="true" pageId="22" pageNumber="23">A. haberi</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF98AD69A24FFDEFC18FCD9FBA6FCFB" box="[950,1068,839,863]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="rhoadsi">
<emphasis id="1B8D2D479A24FFDEFC18FCD9FBA6FCFB" box="[950,1068,839,863]" italics="true" pageId="22" pageNumber="23">A. rhoadsi</emphasis>
</taxonomicName>
, these two species with entirely different genital ligula morphology are probably not closely related.
</paragraph>
<paragraph id="2946F1559A24FFDEFF69FC10FF7BFC6F" blockId="22.[151,1437,151,1008]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69FC10FEA9FC03" bold="true" box="[199,291,910,935]" pageId="22" pageNumber="23">Habitat</emphasis>
. The four males known were found along seeps and trickles through brushy pasture on a forested hillside.
</paragraph>
</subSubSection>
<subSubSection id="61E3A2DE9A24FFDEFF69FC48FB7EFC4B" box="[199,1268,982,1008]" pageId="22" pageNumber="23" type="distribution">
<paragraph id="2946F1559A24FFDEFF69FC48FB7EFC4B" blockId="22.[151,1437,151,1008]" box="[199,1268,982,1008]" pageId="22" pageNumber="23">
<emphasis id="1B8D2D479A24FFDEFF69FC48FED3FC4B" bold="true" box="[199,345,982,1007]" pageId="22" pageNumber="23">Distribution</emphasis>
. So far known only from its
<typeStatus id="F6424FF79A24FFDEFD33FC46FD47FC54" box="[669,717,984,1008]" pageId="22" pageNumber="23">type</typeStatus>
locality in
<collectingRegion id="EB3D3FB79A24FFDEFCE2FC49FC27FC4B" box="[844,941,983,1007]" country="Costa Rica" name="San Jose" pageId="22" pageNumber="23">San José</collectingRegion>
Prov.,
<collectingCountry id="51EEB1C59A24FFDEFC52FC49FBFCFC4B" box="[1020,1142,982,1007]" name="Costa Rica" pageId="22" pageNumber="23">
<collectingRegion id="EB3D3FB79A24FFDEFC52FC49FBB6FC4B" box="[1020,1084,983,1007]" country="New Zealand" name="West Coast" pageId="22" pageNumber="23">Costa</collectingRegion>
Rica
</collectingCountry>
(
<figureCitation id="B1C2EDD09A24FFDEFB2BFC48FB6DFC4B" box="[1157,1255,982,1007]" captionStart="FIGURE 171" captionStartId="85.[151,250,973,995]" captionTargetBox="[203,1335,232,906]" captionTargetId="figure@85.[188,1398,193,952]" captionTargetPageId="85" captionText="FIGURE 171. Distribution of other species." httpUri="https://zenodo.org/record/322108/files/figure.png" pageId="22" pageNumber="23">Fig. 171</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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