143 lines
13 KiB
XML
143 lines
13 KiB
XML
<document ID-DOI="10.11646/zootaxa.5278.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFBCFFF5FF6D6C62CA1FFDA8" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eopachylosticta Malaise 1945" docType="treatment" docVersion="2" lastPageNumber="25" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="24" updateTime="1683183299905" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Archibald, S. Bruce</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
|
||
<mods:affiliation>Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
|
||
<mods:nameIdentifier type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Rasnitsyn, Alexandr P.</mods:namePart>
|
||
<mods:affiliation>A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-05-04</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5278</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>38</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/zootaxa.5278.1.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.5278.1.1</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7894826</mods:identifier>
|
||
<mods:identifier type="ZooBank">BE0A07DB-170A-4B72-8178-9B8144C1FBF6</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7894884" ID-Zenodo-Dep="7894884" LSID="urn:lsid:plazi:treatment:A148D509FFBCFFF5FF6D6C62CA1FFDA8" httpUri="http://treatment.plazi.org/id/A148D509FFBCFFF5FF6D6C62CA1FFDA8" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
|
||
<subSubSection box="[151,609,1847,1874]" pageId="23" pageNumber="24" type="nomenclature">
|
||
<paragraph blockId="23.[151,609,1847,1874]" box="[151,609,1847,1874]" pageId="23" pageNumber="24">
|
||
<heading bold="true" box="[151,609,1847,1874]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
|
||
<emphasis bold="true" box="[151,609,1847,1874]" pageId="23" pageNumber="24">
|
||
Genus
|
||
<taxonomicName authority="Malaise, 1945" authorityName="Malaise" authorityYear="1945" box="[238,609,1847,1874]" class="Insecta" family="Cimbicidae" genus="Eopachylosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis bold="true" box="[238,430,1847,1874]" italics="true" pageId="23" pageNumber="24">Eopachylosticta</emphasis>
|
||
<bibRefCitation author="Malaise, R." box="[438,609,1847,1873]" pageId="23" pageNumber="24" pagination="1 - 288" refId="ref18378" refString="Malaise, R. (1945) Tenthredinoidoa of south-eastern Asia with a general zoogeographical review. Opuscula Entomologica Suppl, 4 (288), 1 - 288. https: // doi. org / 10.1080 / 11035894509446460" type="journal article" year="1945">Malaise, 1945</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,534,1916,1940]" pageId="23" pageNumber="24" type="reference_group">
|
||
<paragraph blockId="23.[151,534,1916,1940]" box="[151,534,1916,1940]" pageId="23" pageNumber="24">
|
||
<treatmentCitationGroup box="[151,534,1916,1940]" pageId="23" pageNumber="24">
|
||
<treatmentCitation author="Malaise, R." box="[151,530,1916,1940]" page="14" pageId="23" pageNumber="24" year="1945">
|
||
<taxonomicName authority="Malaise, 1945, p. 14" authorityName="Malaise" authorityPageNumber="14" authorityYear="1945" box="[151,530,1916,1940]" class="Insecta" family="Cimbicidae" genus="Eopachylosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[151,314,1916,1940]" italics="true" pageId="23" pageNumber="24">Eopachylosticta</emphasis>
|
||
<bibRefCitation author="Malaise, R." box="[321,467,1916,1940]" pageId="23" pageNumber="24" pagination="1 - 288" refId="ref18378" refString="Malaise, R. (1945) Tenthredinoidoa of south-eastern Asia with a general zoogeographical review. Opuscula Entomologica Suppl, 4 (288), 1 - 288. https: // doi. org / 10.1080 / 11035894509446460" type="journal article" year="1945">Malaise, 1945</bibRefCitation>
|
||
, p. 14
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="type_taxon">
|
||
<paragraph blockId="23.[151,624,1982,2007]" box="[151,624,1982,2007]" pageId="23" pageNumber="24">
|
||
<emphasis bold="true" box="[151,291,1982,2007]" pageId="23" pageNumber="24">
|
||
<typeStatus box="[151,204,1983,2007]" pageId="23" pageNumber="24">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName authority="Cockerell, 1925" authorityName="Cockerell" authorityYear="1925" box="[298,621,1982,2007]" class="Insecta" family="Cimbicidae" genus="Amasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="byrami">
|
||
<emphasis box="[298,449,1983,2007]" italics="true" pageId="23" pageNumber="24">Amasis byrami</emphasis>
|
||
<bibRefCitation author="Cockerell, T. D. A." box="[455,621,1982,2007]" pageId="23" pageNumber="24" pagination="1 - 15" refId="ref17014" refString="Cockerell, T. D. A. (1925) Fossil insects in the United States National Museum. Proceedings of the United States National Museum, 64, 1 - 15. [https: // www. biodiversitylibrary. org / item / 32410 # page / 7 / mode / 1 up]" type="journal article" year="1925">Cockerell, 1925</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1436,150,609]" pageId="24" pageNumber="25">
|
||
<emphasis bold="true" box="[151,391,150,177]" pageId="24" pageNumber="25">Emended diagnosis.</emphasis>
|
||
Because of lack of sufficiently preserved wings and incompletely preserved body, the
|
||
<typeStatus box="[1388,1436,151,177]" pageId="24" pageNumber="25">type</typeStatus>
|
||
and only fossil lacks many characters reliably used as diagnostic in
|
||
<taxonomicName authorityName="Curtis" authorityYear="1825" box="[957,1090,187,213]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="family">Cimbicidae</taxonomicName>
|
||
. However, generic distinction in
|
||
<taxonomicName authorityName="Benson" authorityYear="1938" box="[182,366,223,249]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="subFamily" subFamily="Pachylostictinae">Pachylostictinae</taxonomicName>
|
||
is currently based in part on colour pattern (
|
||
<bibRefCitation author="Vilhelmsen, L." box="[884,1148,223,249]" pageId="24" pageNumber="25" pagination="103 - 127" refId="ref19825" refString="Vilhelmsen, L. (2019) Giant sawflies and their kin: morphological phylogeny of Cimbicidae (Hymenoptera). Systematic Entomology, 44, 103 - 127. https: // doi. org / 10.1111 / syen. 12314" type="journal article" year="2019">
|
||
Vilhelmsen
|
||
<emphasis box="[1022,1081,223,249]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
) which is available here, allowing at least a tentative genus diagnosis until better preserved fossils are found. The genus is distinct from others of
|
||
<taxonomicName authorityName="Benson" authorityYear="1938" box="[182,366,295,321]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="subFamily" subFamily="Pachylostictinae">Pachylostictinae</taxonomicName>
|
||
by its dark head and thorax in contrast to its distinctly paler (somewhat infuscate) abdomen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="24" pageNumber="25" type="discussion">
|
||
<paragraph blockId="24.[151,1436,150,609]" pageId="24" pageNumber="25">
|
||
<emphasis bold="true" box="[199,315,331,357]" pageId="24" pageNumber="25">Remarks.</emphasis>
|
||
<taxonomicName authorityName="Malaise" authorityYear="1945" box="[321,501,331,357]" class="Insecta" family="Cimbicidae" genus="Eopachylosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[321,501,331,357]" italics="true" pageId="24" pageNumber="25">Eopachylosticta</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName authorityName="Benson" authorityYear="1938" box="[670,854,331,357]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="subFamily" subFamily="Pachylostictinae">Pachylostictinae</taxonomicName>
|
||
by a combination of its antennae with five segments and the median longitudinal mesonotal sulcus and notauli present and well developed (
|
||
<bibRefCitation author="Vilhelmsen, L." box="[1123,1320,367,393]" pageId="24" pageNumber="25" pagination="103 - 127" refId="ref19825" refString="Vilhelmsen, L. (2019) Giant sawflies and their kin: morphological phylogeny of Cimbicidae (Hymenoptera). Systematic Entomology, 44, 103 - 127. https: // doi. org / 10.1111 / syen. 12314" type="journal article" year="2019">Vilhelmsen 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1436,150,609]" pageId="24" pageNumber="25">
|
||
Preservation of the
|
||
<typeStatus box="[425,473,403,429]" pageId="24" pageNumber="25">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName authorityName="Cockerell" authorityYear="1925" baseAuthorityName="Cockerell" baseAuthorityYear="1925" box="[630,898,403,429]" class="Insecta" family="Cimbicidae" genus="Eopachylosticta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="byrami">
|
||
<emphasis box="[630,898,403,429]" italics="true" pageId="24" pageNumber="25">Eopachylosticta byrami</emphasis>
|
||
</taxonomicName>
|
||
suffers from the aggressive hydrochemistry of the Green River depositional environment. This is a common problem for fossil insects in parts of this formation, where the wings, legs and other body parts which were pale in life may become extremely faint or invisible as fossils. Therefore, our description and interpretation of this fossil is limited to distinguishing the genus from others of
|
||
<taxonomicName authorityName="Benson" authorityYear="1938" box="[182,369,547,573]" class="Insecta" family="Cimbicidae" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="24" phylum="Arthropoda" rank="subFamily" subFamily="Pachylostictinae">Pachylostictinae</taxonomicName>
|
||
, setting aside more detailed analysis until better preserved material is available.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1436,150,609]" box="[199,615,582,609]" pageId="24" pageNumber="25">
|
||
<emphasis bold="true" box="[199,396,582,608]" pageId="24" pageNumber="25">Species included</emphasis>
|
||
.
|
||
<typeStatus box="[409,465,583,609]" pageId="24" pageNumber="25">Type</typeStatus>
|
||
species only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |