treatments-xml/data/A1/0D/48/A10D4838C340374525B4F9C3197AFC39.xml
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<document id="52258E65AF9164FB765420BC2D0C754C" ID-CLB-Dataset="55793" ID-DOI="10.1080/00222933.2013.791936" ID-GBIF-Dataset="6591e69b-631a-4c4d-bf83-db8c70d1ad05" ID-ISSN="1464-5262" ID-Zenodo-Dep="5198102" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" checkinTime="1628884845165" checkinUser="felipe" docAuthor="Zhou, Yu-Lingzi &amp; Zhou, Hong-Zhang" docDate="2013" docId="A10D4838C340374525B4F9C3197AFC39" docLanguage="en" docName="JNatHist.47.45-46.2869-2904.pdf" docOrigin="Journal of Natural History 47 (45 - 46)" docSource="http://dx.doi.org/10.1080/00222933.2013.791936" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Liotesba punctiventris Sharp 1889" docType="treatment" docVersion="10" lastPageNumber="2896" masterDocId="5D343040C35B37582413FFC11B0DFF84" masterDocTitle="Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species" masterLastPageNumber="2904" masterPageNumber="2869" pageNumber="2894" updateTime="1708086164713" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:title id="B277FA361AC352BF0984468682188004">Taxonomy and phylogeny of the genus Liotesba Scheerpeltz (Coleoptera: Staphylinidae: Staphylininae: Xantholinini) with descriptions of four new species</mods:title>
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<mods:namePart id="566EA6E339CF02CE3CCCAD8783489380">Zhou, Yu-Lingzi</mods:namePart>
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<paragraph id="291BF92EC340374325B4F9C31835F99E" blockId="27.[423,824,1538,1562]" box="[423,824,1538,1562]" pageId="27" pageNumber="2894">
<heading id="72534E42C340374325B4F9C31835F99E" box="[423,824,1538,1562]" centered="true" fontSize="10" level="2" pageId="27" pageNumber="2894" reason="2">
<taxonomicName id="EEA482ADC340374325B4F9C31835F99E" ID-CoL="3V87Q" authority="(Sharp, 1889)" authorityName="Sharp" authorityYear="1889" box="[423,824,1538,1562]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="2894" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="1BD0253CC340374325B4F9C31994F99E" bold="true" box="[423,665,1538,1562]" italics="true" pageId="27" pageNumber="2894">Liotesba punctiventris</emphasis>
(
<bibRefCitation id="4D3584DFC340374326BBF9C31822F99E" author="Sharp DS" box="[680,815,1538,1562]" pageId="27" pageNumber="2894" pagination="249 - 267" refId="ref18004" refString="Sharp DS. 1889. The Staphylinidae of Japan. Ann Mag Nat History. 6: 249 - 267." type="journal article" year="1889">Sharp, 1889</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C3403744249CF9831F81FF7B" lastPageId="28" lastPageNumber="2895" pageId="27" pageNumber="2894" type="reference_group">
<paragraph id="291BF92EC3403744249CF9831F81FF7B" blockId="27.[143,1103,1602,1690]" lastBlockId="28.[211,1172,167,255]" lastPageId="28" lastPageNumber="2895" pageId="27" pageNumber="2894">
<treatmentCitationGroup id="09B4DE00C3403744249CF9831F81FF7B" lastPageId="28" lastPageNumber="2895" pageId="27" pageNumber="2894">
<treatmentCitation id="A805DF3FC3403743249CF9831A42F9DE" author="Sharp" box="[143,335,1602,1626]" page="252" pageId="27" pageNumber="2894" year="1889">
<bibRefCitation id="4D3584DFC3403743249CF9831A42F9DE" author="Sharp DS" box="[143,335,1602,1626]" pageId="27" pageNumber="2894" pagination="249 - 267" refId="ref18004" refString="Sharp DS. 1889. The Staphylinidae of Japan. Ann Mag Nat History. 6: 249 - 267." type="journal article" year="1889">Sharp, 1889: 252</bibRefCitation>
</treatmentCitation>
(orig.
<taxonomicName id="EEA482ADC3403743258DF98319BAF9DE" ID-CoL="5C695" authorityName="Sharp" authorityYear="1889" box="[414,695,1602,1626]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="2894" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="1BD0253CC3403743258DF98319BAF9DE" box="[414,695,1602,1626]" italics="true" pageId="27" pageNumber="2894">Xantholinus punctiventris</emphasis>
</taxonomicName>
;
<typeStatus id="F61F478CC340374326D4F98219FAF9DF" box="[711,759,1603,1627]" pageId="27" pageNumber="2894">type</typeStatus>
locality: Ichiuchi);
<treatmentCitation id="A805DF3FC340374327CBF9831A92F9FE" author="Bernhauer &amp; Schubert" page="306" pageId="27" pageNumber="2894" year="1914">
<bibRefCitation id="4D3584DFC340374327CBF9831A92F9FE" author="Bernhauer M &amp; Schubert K" pageId="27" pageNumber="2894" pagination="289 - 408" refId="ref16960" refString="Bernhauer M, Schubert K. 1914. Staphylinidae IV. In: Schenkling S, editor. Coleopterorum Catalogus. Berlin (Germany): Junk; 289 - 408." type="book chapter" year="1914">Bernhauer and Schubert, 1914: 306</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="A805DF3FC34037432622F9A3180AF9FE" author="Bordoni A" box="[561,775,1634,1658]" page="171" pageId="27" pageNumber="2894" year="1997">
<bibRefCitation id="4D3584DFC34037432622F9A3180AF9FE" author="Bordoni A" box="[561,775,1634,1658]" pageId="27" pageNumber="2894" pagination="167 - 179" refId="ref16992" refString="Bordoni A. 1997. A revision of the tribe Xantholinini from Japan, 1 (Coleoptera, Staphylinidae). 116 o contribution to the knowledge of the Staphylinidae. Jpn J System Entomol. 3: 167 - 179." type="journal article" year="1997">Bordoni, 1997: 171</bibRefCitation>
</treatmentCitation>
(
<taxonomicName id="EEA482ADC34037432704F9A3187BF9FE" ID-CoL="8VYYF" box="[791,886,1634,1658]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="2894" phylum="Arthropoda" rank="genus">
<emphasis id="1BD0253CC34037432704F9A3187BF9FE" box="[791,886,1634,1658]" italics="true" pageId="27" pageNumber="2894">Liotesba</emphasis>
</taxonomicName>
; characters;
<collectingCountry id="51B3B9BEC34037432015F9A21F46F9FE" box="[1030,1099,1635,1658]" name="Japan" pageId="27" pageNumber="2894">Japan</collectingCountry>
, Honshu, Mikurajima Is., Kyushu);
<treatmentCitation id="A805DF3FC34037432604F94219F4F91E" author="Herman LH" box="[535,761,1666,1690]" page="3692" pageId="27" pageNumber="2894" year="2001">
<bibRefCitation id="4D3584DFC34037432604F94219F4F91E" author="Herman LH" box="[535,761,1666,1690]" pageId="27" pageNumber="2894" pagination="3021 - 3839" refId="ref17732" refString="Herman LH. 2001. Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3) Staphylininae: Staphylinini (Quediina, Staphylinina, Tanygnathinina, Xanthopygina), Xantholinini. Staphylinidae incertae sedis. Fossils, Protactinae. Bull Am Museum Nat History. 265: 3021 - 3839." type="journal article" year="2001">Herman, 2001: 3692</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="A805DF3FC3403743279BF9431F42F91E" author="Bordoni A" box="[904,1103,1666,1690]" page="9" pageId="27" pageNumber="2894" year="2003">
<bibRefCitation id="4D3584DFC3403743279BF9431F42F91E" author="Bordoni A" box="[904,1103,1666,1690]" pageId="27" pageNumber="2894" pagination="9 - 10" refId="ref17073" refString="Bordoni A. 2003 a. Contribution to the knowledge of the Xantholinini from Japan, II. Description of the male of Liotesba punctiventris (Sharp) (Coleoptera Staphylinidae): 138 o Contribution to the knowledge of the Staphylinidae. Jpn J Syst Entomol. 9: 9 - 10." type="journal article" year="2003">Bordoni, 2003a: 9</bibRefCitation>
</treatmentCitation>
(first recorded characters of male;
<collectingRegion id="EB6037CCC34737442642FF6619F5FF3B" box="[593,760,167,191]" country="Japan" name="Fukuoka" pageId="28" pageNumber="2895">Fukuoka Pref.</collectingRegion>
, Mt Tachibana, T. Ueno, Y. Hoshina and H. Goto leg.;
<date id="5D1ADFEEC3473744258FFF061937FF5B" box="[412,570,199,224]" pageId="28" pageNumber="2895" value="1996-05-06" valueMax="1996-05-09" valueMin="1996-05-06">69 May 1996</date>
; flight intercept trap);
<treatmentCitation id="A805DF3FC34737442722FF061F07FF5B" author="Smetana A" box="[817,1034,199,223]" page="691" pageId="28" pageNumber="2895" year="2004">
<bibRefCitation id="4D3584DFC34737442722FF061F07FF5B" author="Smetana A" box="[817,1034,199,223]" pageId="28" pageNumber="2895" pagination="687 - 698" refId="ref18057" refString="Smetana A. 2004. Staphylinidae, tribe Xantholinini. In: Lobl I, Smetana A, editors. Catalogue of Palaearctic Coleoptera, Vol. 2. Stenstrup: Apollo Books; p. 687 - 698." type="book chapter" year="2004">Smetana, 2004: 691</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="A805DF3FC347374424C0FF261A91FF7B" author="Bordoni A" box="[211,412,231,255]" page="9" pageId="28" pageNumber="2895" year="2007">
<bibRefCitation id="4D3584DFC347374424C0FF261A91FF7B" author="Bordoni A" box="[211,412,231,255]" pageId="28" pageNumber="2895" pagination="7 - 18" refId="ref17357" refString="Bordoni A. 2007 a. Contribution to the knowledge of the Xantholinini of China. XI. New species from Northern Yunnan, Guizhou, Hubei and Anhui (Coleoptera Staphylinidae). Bollettino della Societa Entomol Italiana. 139: 7 - 18." type="journal article" year="2007">Bordoni, 2007a: 9</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="51B3B9BEC347374425B8FF261AF9FF7B" box="[427,500,231,255]" name="China" pageId="28" pageNumber="2895">China</collectingCountry>
,
<collectingRegion id="EB6037CCC347374425EDFF26194AFF7B" box="[510,583,231,255]" country="China" name="Hubei" pageId="28" pageNumber="2895">Hubei</collectingRegion>
,
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31
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2
<superScript id="DED15466C34737442696FF221987FF70" attach="left" box="[645,650,227,244]" fontSize="7" pageId="28" pageNumber="2895"></superScript>
N
</geoCoordinate>
,
<geoCoordinate id="4C909FE9C347374426A7FF261818FF7B" box="[692,789,227,255]" degrees="110" direction="east" minutes="5" orientation="longitude" pageId="28" pageNumber="2895" precision="925" value="110.083336">
110
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5
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E
</geoCoordinate>
);
<treatmentCitation id="A805DF3FC3473744273AFF261F0CFF7B" author="Bordoni A" box="[809,1025,231,255]" page="71" pageId="28" pageNumber="2895" year="2007">
<bibRefCitation id="4D3584DFC3473744273AFF261F0CFF7B" author="Bordoni A" box="[809,1025,231,255]" pageId="28" pageNumber="2895" pagination="67 - 78" refId="ref17403" refString="Bordoni A. 2007 b. Xantholinini (Coleoptera staphylinidae). Supplement to the catalogue Palaearctic coleoptera, volume 2. Bollettino della Societa Entomologica Italiana. 139: 67 - 78." type="journal article" year="2007">Bordoni, 2007b: 71</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C347374424C0FEE618CBFEFB" pageId="28" pageNumber="2895" type="materials_examined">
<paragraph id="291BF92EC347374424C0FEE618CBFEFB" blockId="28.[211,1172,295,383]" pageId="28" pageNumber="2895">
<emphasis id="1BD0253CC347374424C0FEE61ABDFEBB" bold="true" box="[211,432,295,319]" pageId="28" pageNumber="2895">Material examined.</emphasis>
<materialsCitation id="99CCF373C347374425AEFEE618CFFEFB" ID-GBIF-Occurrence="3757895302" collectingDate="1881-05-01" collectionCode="V, BMNH" collectorName="D. S. Ichiuchi &amp; G. Lewis" country="Japan" location="Ichiuchi" pageId="28" pageNumber="2895" specimenCode="IV- 2" specimenCount="1" typeStatus="holotype">
female, with labels as follows: “
<typeStatus id="F61F478CC34737442721FEE618AFFEBB" box="[818,930,295,319]" pageId="28" pageNumber="2895" type="holotype">Holotype</typeStatus>
”/“
<taxonomicName id="EEA482ADC347374427D4FEE61A21FEDB" authorityName="Sharp" authorityYear="1889" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="2895" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="1BD0253CC347374427D4FEE61A21FEDB" italics="true" pageId="28" pageNumber="2895">Xantholinus punctiventris</emphasis>
</taxonomicName>
type
<collectorName id="84519CF8C3473744257FFE86190FFEDB" box="[364,514,327,351]" pageId="28" pageNumber="2895">D.S. Ichiuchi</collectorName>
,
<date id="5D1ADFEEC3473744261DFE861962FEDB" box="[526,623,327,351]" pageId="28" pageNumber="2895" value="1881-05-01">
<collectingDate id="4D5E2606C3473744261DFE861962FEDB" box="[526,623,327,351]" pageId="28" pageNumber="2895" value="1881-05-01">1.5.1881</collectingDate>
</date>
”/“
<collectingCountry id="51B3B9BEC34737442682FE8619D5FEDB" box="[657,728,327,351]" name="Japan" pageId="28" pageNumber="2895">Japan</collectingCountry>
,
<collectorName id="84519CF8C347374426F7FE861847FEDB" box="[740,842,327,351]" pageId="28" pageNumber="2895">G. Lewis</collectorName>
, 1910-320”/“
<location id="2C7BAFF5C347374427F3FE861F4FFEDB" LSID="urn:lsid:plazi:treatment:A10D4838C340374525B4F9C3197AFC39:2C7BAFF5C347374427F3FE861F4FFEDB" box="[992,1090,327,351]" country="Japan" name="Ichiuchi" pageId="28" pageNumber="2895">Ichiuchi</location>
, 30.
<specimenCode id="79025155C3473744207EFE861BEFFEFB" collectionCode="IV-" pageId="28" pageNumber="2895">IV- 2</specimenCode>
.
<collectionCode id="4FB561EBC347374424F6FEA61BF6FEFB" box="[229,251,359,383]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="28" pageNumber="2895" type="Museum">V</collectionCode>
.81”/“
<taxonomicName id="EEA482ADC3473744252CFEA6184DFEFB" authority=", Det. A. Bordoni, 1996" authorityName=", Det. A. Bordoni" authorityYear="1996" box="[319,832,359,383]" class="Insecta" family="Staphylinidae" genus="Liotesba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="2895" phylum="Arthropoda" rank="species" species="punctiventris">
<emphasis id="1BD0253CC3473744252CFEA6193CFEFB" box="[319,561,359,383]" italics="true" pageId="28" pageNumber="2895">Liotesba punctiventris</emphasis>
, Det. A. Bordoni, 1996
</taxonomicName>
”. (
<collectionCode id="4FB561EBC3473744274DFEA618B6FEFB" box="[862,955,359,383]" pageId="28" pageNumber="2895">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61BEAAA5C347374524C0FE7E197AFC39" lastPageId="29" lastPageNumber="2896" pageId="28" pageNumber="2895" type="description">
<paragraph id="291BF92EC347374424C0FE7E1A5FFE52" blockId="28.[211,1171,447,638]" box="[211,338,447,470]" pageId="28" pageNumber="2895">
<heading id="72534E42C347374424C0FE7E1A5FFE52" box="[211,338,447,470]" fontSize="10" level="3" pageId="28" pageNumber="2895" reason="5">
<emphasis id="1BD0253CC347374424C0FE7E1A5FFE52" box="[211,338,447,470]" italics="true" pageId="28" pageNumber="2895">Description</emphasis>
</heading>
</paragraph>
<paragraph id="291BF92EC347374424C0FE2718A3FD9A" blockId="28.[211,1171,447,638]" pageId="28" pageNumber="2895">
<emphasis id="1BD0253CC347374424C0FE271A7DFE7A" bold="true" box="[211,368,486,510]" pageId="28" pageNumber="2895">Measurement.</emphasis>
BL =
<quantity id="EE5C54CBC347374425DAFE27193CFE7A" box="[457,561,486,510]" metricMagnitude="-3" metricUnit="m" metricValue="8.85" pageId="28" pageNumber="2895" unit="mm" value="8.85">8.85 mm</quantity>
, FL =
<quantity id="EE5C54CBC3473744269EFE2719FBFE7A" box="[653,758,486,510]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="28" pageNumber="2895" unit="mm" value="4.5">4.50 mm</quantity>
, HL =
<quantity id="EE5C54CBC34737442745FE2718B3FE7A" box="[854,958,486,510]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="28" pageNumber="2895" unit="mm" value="1.29">1.29 mm</quantity>
, HW =
<quantity id="EE5C54CBC34737442034FE271F82FE7A" box="[1063,1167,486,510]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="28" pageNumber="2895" unit="mm" value="1.05">1.05 mm</quantity>
, PL =
<quantity id="EE5C54CBC34737442504FDC71A71FD9A" box="[279,380,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="28" pageNumber="2895" unit="mm" value="1.44">1.44 mm</quantity>
, PW =
<quantity id="EE5C54CBC347374425C0FDC7193AFD9A" box="[467,567,518,542]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" pageId="28" pageNumber="2895" unit="mm" value="0.99">0.99 mm</quantity>
, EL =
<quantity id="EE5C54CBC34737442694FDC719E1FD9A" box="[647,748,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="28" pageNumber="2895" unit="mm" value="1.62">1.62 mm</quantity>
, EW =
<quantity id="EE5C54CBC34737442756FDC718A4FD9A" box="[837,937,518,542]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="28" pageNumber="2895" unit="mm" value="1.2">1.20 mm</quantity>
.
</paragraph>
<paragraph id="291BF92EC347374424EDFDE718D8FDFA" blockId="28.[211,1171,447,638]" pageId="28" pageNumber="2895">Body nearly cylindrical and medium-sized. Head, pronotum and abdomen dark brown. Elytra and legs reddish brown. Antennae brown, antennomere I and III darker in colour, antennomere II reddish. Maxillary and labial palpi brown.</paragraph>
<paragraph id="291BF92EC347374424C0FD6419DEFB98" blockId="28.[211,1172,677,1053]" pageId="28" pageNumber="2895">
<emphasis id="1BD0253CC347374424C0FD641A1AFD39" bold="true" box="[211,279,677,701]" pageId="28" pageNumber="2895">Head.</emphasis>
Subrectangular (HL to HW ratio 1.2), tempora straight, posterior angles rounded. Dorsal integument of head shiny, rather smooth; extensively with micropunctures, and also with few sparsely scattered punctures in medium size toward lateral and posterior margin; deflexed portion of tempora with a row of large punctures. Each side of cranium with anterolateral puncture near antennal insertion, midlateral puncture approximately a distance of three punctures diameter to dorsal margin of eye, temporal puncture at the posterior one-fifth, and occipital puncture at the lateral one-third. Frontal furrows relatively long and deep, anteocular furrows absent. Eye medium in size, shorter than half the length of tempora (eye: tempora = 0.30:
<quantity id="EE5C54CBC347374427D9FC641F3DFC39" box="[970,1072,933,957]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="28" pageNumber="2895" unit="mm" value="0.68">0.68 mm</quantity>
), slightly protruding laterad; one large and one small puncture next to dorsal margin of eye. Epistoma narrow, and acute. Distance between antennal insertions
<quantity id="EE5C54CBC347374427C2FC251F3AFC78" box="[977,1079,996,1020]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="2895" unit="mm" value="0.3">0.30 mm</quantity>
, shorter than distance from antenna to eye (
<quantity id="EE5C54CBC34737442673FBC519CAFB98" box="[608,711,1028,1052]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="28" pageNumber="2895" unit="mm" value="0.37">0.37 mm</quantity>
).
</paragraph>
<paragraph id="291BF92EC347374424C0FB85187AFB38" blockId="28.[211,1171,1092,1212]" pageId="28" pageNumber="2895">
<emphasis id="1BD0253CC347374424C0FB851A4FFBD8" bold="true" box="[211,322,1092,1116]" pageId="28" pageNumber="2895">Antennae.</emphasis>
Scape stout, thickened apically, slightly longer than three subsequent antennomeres combined,
<quantity id="EE5C54CBC347374425A9FBA5192CFBF8" box="[442,545,1124,1148]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="28" pageNumber="2895" unit="mm" value="0.48">0.48 mm</quantity>
; antennomere II and III elongate, and of same length,
<quantity id="EE5C54CBC347374424C0FB451A36FB18" box="[211,315,1156,1180]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="2895" unit="mm" value="0.14">0.14 mm</quantity>
; IV and V of equal in length,
<quantity id="EE5C54CBC3473744268DFB451808FB18" box="[670,773,1156,1180]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="2895" unit="mm" value="0.06">0.06 mm</quantity>
; last antennomere medium length,
<quantity id="EE5C54CBC347374424C0FB651A35FB38" box="[211,312,1188,1212]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="28" pageNumber="2895" unit="mm" value="0.17">0.17 mm</quantity>
, as long as two preceding antennomeres combined.
</paragraph>
<paragraph id="291BF92EC347374424C0FB22199EFAFF" blockId="28.[211,1172,1251,1404]" pageId="28" pageNumber="2895">
<emphasis id="1BD0253CC347374424C0FB221A50FB7F" bold="true" box="[211,349,1251,1275]" pageId="28" pageNumber="2895">Mouthparts.</emphasis>
Labrum transverse, with a pair of median teeth, and a lateral tooth at each side, which slightly protruding. Mandibles falciform, right one with a broad tooth on inner edge. Maxillary palpus elongate, second segment longest, last segment fusiform, with subtruncated apex. Labial palpus slender, each segment of same length, last segment slightly dilated subapically.
</paragraph>
<paragraph id="291BF92EC347374424C0FA6218D0FA3F" blockId="28.[211,989,1443,1467]" box="[211,989,1443,1467]" pageId="28" pageNumber="2895">
<emphasis id="1BD0253CC347374424C0FA621A18FA3F" bold="true" box="[211,277,1443,1467]" pageId="28" pageNumber="2895">Neck.</emphasis>
Wide (
<quantity id="EE5C54CBC34737442576FA621AC1FA3F" box="[357,460,1443,1467]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="28" pageNumber="2895" unit="mm" value="0.48">0.48 mm</quantity>
), slightly narrower than half the width of head.
</paragraph>
<paragraph id="291BF92EC347374424C0FA2319A6F91E" blockId="28.[211,1172,1506,1690]" pageId="28" pageNumber="2895">
<emphasis id="1BD0253CC347374424C0FA231A45FA7E" bold="true" box="[211,328,1506,1530]" pageId="28" pageNumber="2895">Pronotum.</emphasis>
Subrectangular and distinctly oblong (PL to PW ratio 1.5), slightly longer, narrower than head. Widened anteriad, lateral margins sinuate, anterior angles well defined, posterior angles broadly rounded. Integument extensively covered with micropunctures, with two large punctures on each anterolateral corner, and one large puncture on each posterolateral corner, and also with other irregular punctures near anterior and lateral margins of pronotum.
</paragraph>
<paragraph id="291BF92EC3463745249CFF66183FFF3B" blockId="29.[143,818,167,191]" box="[143,818,167,191]" pageId="29" pageNumber="2896">
<emphasis id="1BD0253CC3463745249CFF661A36FF3B" bold="true" box="[143,315,167,191]" pageId="29" pageNumber="2896">Mesoscutellum.</emphasis>
Shiny, scattered with two pairs of punctures.
</paragraph>
<paragraph id="291BF92EC3463745249CFF261F42FEFB" blockId="29.[143,1103,231,383]" pageId="29" pageNumber="2896">
<emphasis id="1BD0253CC3463745249CFF261BD0FF7B" bold="true" box="[143,221,231,255]" pageId="29" pageNumber="2896">Elytra.</emphasis>
Subrectangular, elongate (EL to EW ratio 1.4), distinctly longer and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, hind margin rounded. Integument shiny and flattened, without microsculpture; each elytron symmetrically with two or three rows of smaller punctures near median suture, a row of median punctures on centre, also with three rows of punctures on deflexed portion.
</paragraph>
<paragraph id="291BF92EC3463745249CFE67180AFE7A" blockId="29.[143,1103,422,510]" pageId="29" pageNumber="2896">
<emphasis id="1BD0253CC3463745249CFE671BC1FE3A" bold="true" box="[143,204,422,446]" pageId="29" pageNumber="2896">Legs.</emphasis>
First four segments of protarsi dilated, heart-shaped; last segment of both mesotarsi and metatarsi slender. Last segment of protarsi and mesotarsi shorter than each IIIV combined, metatarsi subequal to IIIV combined.
</paragraph>
<paragraph id="291BF92EC3463745249CFDE71BEAFD59" blockId="29.[143,1103,550,733]" pageId="29" pageNumber="2896">
<emphasis id="1BD0253CC3463745249CFDE71BF3FDBA" bold="true" box="[143,254,550,574]" pageId="29" pageNumber="2896">Abdomen.</emphasis>
Cylindrical, broadest at segment VI. Tergites IIIVII shiny, each segment covered with distinct transverse microstriae, and with large, densely distributed punctures, distance between punctures less than a punctures diameter. Each tergite with a basal impression near anterior margin, bearing transverse microstriae and larger punctures. All abdominal sternites shiny, with microstriae and punctures as those on tergites.
</paragraph>
<paragraph id="291BF92EC3463745249CFCC41BF1FCB9" blockId="29.[143,1103,773,829]" pageId="29" pageNumber="2896">
<emphasis id="1BD0253CC3463745249CFCC41A11FC99" bold="true" box="[143,284,773,797]" pageId="29" pageNumber="2896">Distribution.</emphasis>
<collectingCountry id="51B3B9BEC3463745253CFCC41A79FC99" box="[303,372,773,797]" name="China" pageId="29" pageNumber="2896">China</collectingCountry>
(
<collectingRegion id="EB6037CCC3463745259CFCC41AD7FC99" box="[399,474,773,797]" country="China" name="Hubei" pageId="29" pageNumber="2896">Hubei</collectingRegion>
);
<collectingCountry id="51B3B9BEC346374525E9FCC71931FC99" box="[506,572,774,797]" name="Japan" pageId="29" pageNumber="2896">Japan</collectingCountry>
(
<collectingRegion id="EB6037CCC34637452645FCC419CFFC99" box="[598,706,773,797]" country="Japan" name="Fukuoka" pageId="29" pageNumber="2896">Fukuoka</collectingRegion>
, Kyushu, Mikurajima, Honshu, Ichiuchi).
</paragraph>
<paragraph id="291BF92EC3463745249CFCA4197AFC39" blockId="29.[143,1103,869,957]" pageId="29" pageNumber="2896">
<emphasis id="1BD0253CC3463745249CFCA41BF5FCF9" bold="true" box="[143,248,869,893]" pageId="29" pageNumber="2896">Remarks.</emphasis>
This species can be distinguished from other congeners by the unique coloration of the antennae, acute epistoma, distinctly oblong pronotum and slightly protruding teeth of labrum in combination.
</paragraph>
</subSubSection>
</treatment>
</document>