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<document id="4EA43DDEE6C03AFA491D5E7A181F8256" ID-CLB-Dataset="42325" ID-DOI="10.11646/zootaxa.3911.4.2" ID-GBIF-Dataset="83770aa6-52d7-4e4e-85d2-4126ca5c1cf0" ID-ISSN="1175-5326" ID-Zenodo-Dep="68799" ID-ZooBank="EDF9E69E-7898-4CF8-B447-EFF646FE3B44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460945716228" checkinUser="plazi" docAuthor="Taylor, Robert W." docDate="2015" docId="660EEE4F6717FF87FF59F8DFFE88FF1F" docLanguage="en" docName="zt03911p520.pdf" docOrigin="Zootaxa 3911 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Myrmecia haskinsorum Taylor, 2015, sp.n." docType="treatment" docVersion="14" lastPageNumber="501" masterDocId="9A3796376712FF8FFFCEFF91FFA9FC42" masterDocTitle="Ants with Attitude: Australian Jack-jumpers of the Myrmecia pilosula species complex, with descriptions of four new species (Hymenoptera: Formicidae: Myrmeciinae)" masterLastPageNumber="520" masterPageNumber="493" pageNumber="498" updateTime="1732577186238" updateUser="ExternalLinkService">
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<mods:title id="09C56568B27977F3B24402EE2A34A88A">Ants with Attitude: Australian Jack-jumpers of the Myrmecia pilosula species complex, with descriptions of four new species (Hymenoptera: Formicidae: Myrmeciinae)</mods:title>
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<mods:name id="02EAADDAF3DD007BFCE3CE9A1B027BA2" type="personal">
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<mods:roleTerm id="90F4261D6D9121BB4D1378A64A43E8B4">Author</mods:roleTerm>
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<mods:namePart id="8C92D667B23B09AA590495403359EEB6">Taylor, Robert W.</mods:namePart>
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<mods:title id="F63C6FFE06D46BFD1B01EF70C20D29D8">Zootaxa</mods:title>
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<mods:part id="5E1970F18015D7328EB3801A608920E0">
<mods:date id="341C5E822BE9E9CCAB1BE1B04051D11A">2015</mods:date>
<mods:detail id="257198F9EDC978BEDF96CD89CC61132E" type="volume">
<mods:number id="4DD090E07D9A22D3E91F72599A3D6952">3911</mods:number>
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<mods:classification id="56A3C04AB356396122BF61E5F5252CC9">journal article</mods:classification>
<mods:identifier id="44FE006B1A17B30EC9704B29305F303B" type="CLB-Dataset">42325</mods:identifier>
<mods:identifier id="D5E448C7B88ABCA5AAB34D09DD312904" type="DOI">10.11646/zootaxa.3911.4.2</mods:identifier>
<mods:identifier id="53157DB6E4F5D118EAA3523C5088099F" type="GBIF-Dataset">83770aa6-52d7-4e4e-85d2-4126ca5c1cf0</mods:identifier>
<mods:identifier id="5447A95FA4700A5D7D354B09E8D7E98B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="407AF1B2A037C3CC2DC5F9D6294788D7" type="Zenodo-Dep">68799</mods:identifier>
<mods:identifier id="944C3DCDE8A631CA818E4CC00C65D054" type="ZooBank">EDF9E69E-7898-4CF8-B447-EFF646FE3B44</mods:identifier>
</mods:mods>
<treatment id="660EEE4F6717FF87FF59F8DFFE88FF1F" ID-DOI="http://doi.org/10.5281/zenodo.6109278" ID-GBIF-Taxon="119591870" ID-Zenodo-Dep="6109278" LSID="urn:lsid:plazi:treatment:660EEE4F6717FF87FF59F8DFFE88FF1F" httpUri="http://treatment.plazi.org/id/660EEE4F6717FF87FF59F8DFFE88FF1F" lastPageId="8" lastPageNumber="501" pageId="5" pageNumber="498" scope_family="Formicidae" scope_order="Hymenoptera" scope_subFamily="Myrmeciinae">
<subSubSection id="A6BD0CD26717FF89FF59F8DFFE75FCBA" lastPageId="6" lastPageNumber="499" pageId="5" pageNumber="498" type="nomenclature">
<paragraph id="EE185F596717FF8AFF59F8DFFE50FB2A" blockId="5.[151,505,1870,1930]" box="[151,505,1870,1896]" pageId="5" pageNumber="498">
<heading id="B550E8356717FF8AFF59F8DFFE50FB2A" bold="true" box="[151,505,1870,1896]" fontSize="11" level="1" pageId="5" pageNumber="498" reason="1">
<emphasis id="DCD3834B6717FF8AFF59F8DFFE50FB2A" bold="true" box="[151,505,1870,1896]" pageId="5" pageNumber="498">
<taxonomicName id="29A724DA6717FF8AFF59F8DFFE1EFB2A" ID-CoL="CMC76" box="[151,439,1870,1896]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="haskinsorum" status="sp. nov.">
<emphasis id="DCD3834B6717FF8AFF59F8DFFE1EFB2A" bold="true" box="[151,439,1870,1896]" italics="true" pageId="5" pageNumber="498">Myrmecia haskinsorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C7E03E306717FF8AFE73F8DFFE50FB2A" box="[445,505,1870,1896]" pageId="5" pageNumber="498" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EE185F596717FF8AFF59F8E0FEA3FBC8" blockId="5.[151,505,1870,1930]" box="[151,266,1905,1930]" pageId="5" pageNumber="498">
(
<figureCitation id="769C43DC6717FF8AFF51F8E0FEA8FBC8" box="[159,257,1905,1930]" captionStart="FIGURES 1 – 3" captionStartId="7.[151,264,1957,1980]" captionTargetBox="[264,1323,195,1934]" captionTargetId="figure@7.[264,1323,195,1934]" captionTargetPageId="7" captionText="FIGURES 1 – 3. Myrmecia haskinsorum, Jamberoo, NSW. Standard views. HW 2.46 mm, WL 3.64 mm, PW 1.57 mm." httpUri="https://zenodo.org/record/398906/files/figure.png" pageId="5" pageNumber="498">Figs 1–3</figureCitation>
)
</paragraph>
<paragraph id="EE185F596717FF89FF59F82BFE75FCBA" blockId="5.[151,1436,1977,2038]" lastBlockId="6.[151,1437,151,1977]" lastPageId="6" lastPageNumber="499" pageId="5" pageNumber="498">
<taxonomicName id="29A724DA6717FF8AFF59F82BFE08FB90" box="[151,417,1977,2002]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6717FF8AFF59F82BFE08FB90" box="[151,417,1977,2002]" italics="true" pageId="5" pageNumber="498">Myrmecia haskinsorum</emphasis>
</taxonomicName>
is a species essentially of high elevations, known from the Snowy Mountains, Victorian Alps and upland Tasmania, with some Tasmanian records from lower elevations. In most localities of record nests are regularly covered or brushed by winter snow (as also in sympatric populations of
<taxonomicName id="29A724DA6714FF89FB80FF08FB64FCF2" box="[1102,1229,151,176]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B6714FF89FB80FF08FB64FCF2" box="[1102,1229,151,176]" italics="true" pageId="6" pageNumber="499">M. pilosula</emphasis>
</taxonomicName>
(Eastern Race) on mainland
<collectingCountry id="96B01FC96714FF89FEC9FF2DFEC7FC97" box="[263,366,188,213]" name="Australia" pageId="6" pageNumber="499">Australia</collectingCountry>
and
<taxonomicName id="29A724DA6714FF89FE68FF2CFD81FC97" box="[422,552,188,213]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B6714FF89FE68FF2CFD81FC97" box="[422,552,188,213]" italics="true" pageId="6" pageNumber="499">M. pilosula</emphasis>
</taxonomicName>
(Western Race) on Tasmania). This species was discussed as
<taxonomicName id="29A724DA6714FF89FB28FF2CFA35FC97" box="[1254,1436,188,213]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6714FF89FB28FF2CFA35FC97" box="[1254,1436,188,213]" italics="true" pageId="6" pageNumber="499">M. haskinsorum</emphasis>
</taxonomicName>
by
<bibRefCitation id="8A3622A86714FF89FF74FF4EFE7EFCBA" box="[186,471,223,248]" pageId="6" pageNumber="499" refString="Hirai, H., Yamamoto, M. T., Ogura, K., Satta, Y., Yamada, M., Taylor, R. W. &amp; Imai, H. T. (1994) Multiplication of 28 s rDNA and NOR activity in chromosome evolution among ants of the Myrmecia pilosula species complex (Hymenoptera: Formicidae: Myrmeciinae). Chromosoma, 103, 171 - 178. [Berlin] http: // dx. doi. org / 10.1007 / BF 00368009" type="journal article">
Imai, Taylor
<emphasis id="DCD3834B6714FF89FE85FF70FE2AFCBA" box="[331,387,223,248]" italics="true" pageId="6" pageNumber="499">et al.</emphasis>
(1994)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A6BD0CD26714FF89FF09FE95FB6AFF57" pageId="6" pageNumber="499" type="materials_examined">
<paragraph id="EE185F596714FF89FF09FE95FE9CFD02" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="499">
<emphasis id="DCD3834B6714FF89FF09FE95FECEFD5F" bold="true" box="[199,359,260,285]" pageId="6" pageNumber="499">
<typeStatus id="311CE1FB6714FF89FF09FE95FEABFD5F" box="[199,258,260,285]" pageId="6" pageNumber="499">Type</typeStatus>
locality.
</emphasis>
<collectionCode id="88B6C79C6714FF89FEBEFE94FE19FD5E" box="[368,432,261,284]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="499">NEW</collectionCode>
<collectionCode id="88B6C79C6714FF89FE77FE95FD28FD5E" box="[441,641,260,284]" pageId="6" pageNumber="499">
SOUTH
<collectingCountry id="96B01FC96714FF89FDD1FE95FD28FD5E" box="[543,641,260,284]" name="United Kingdom" pageId="6" pageNumber="499">WALES</collectingCountry>
</collectionCode>
: Dead Horse Gap (-36 31, 148 15), Snowy Mountains National Park, near Thredbo.
</paragraph>
<paragraph id="EE185F596714FF89FF09FEDDFC77FDCA" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="499">
<emphasis id="DCD3834B6714FF89FF09FEDDFE22FD27" bold="true" box="[199,395,332,357]" pageId="6" pageNumber="499">
<typeStatus id="311CE1FB6714FF89FF09FEDDFEABFD27" box="[199,258,332,357]" pageId="6" pageNumber="499">Type</typeStatus>
deposition.
</emphasis>
<typeStatus id="311CE1FB6714FF89FE5FFEDDFE50FD27" box="[401,505,332,357]" pageId="6" pageNumber="499" type="holotype">Holotype</typeStatus>
and
<typeStatus id="311CE1FB6714FF89FDFEFEDCFD32FD27" box="[560,667,333,357]" pageId="6" pageNumber="499" type="paratype">paratypes</typeStatus>
in
<collectionCode id="88B6C79C6714FF89FD0EFEDDFCAEFD26" LSID="urn:lsid:biocol.org:col:32981" box="[704,775,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="6" pageNumber="499">ANIC</collectionCode>
,
<typeStatus id="311CE1FB6714FF89FCDCFEDCFCD4FD27" box="[786,893,333,357]" pageId="6" pageNumber="499" type="paratype">paratypes</typeStatus>
or type-compared vouchers in
<collectionCode id="88B6C79C6714FF89FB15FEDDFA86FD26" box="[1243,1327,332,356]" httpUri="http://grbio.org/cool/ymbs-hrkx" name="Albany Museum" pageId="6" pageNumber="499">AMSA</collectionCode>
,
<collectionCode id="88B6C79C6714FF89FAF5FEDDFA30FD26" box="[1339,1433,332,356]" pageId="6" pageNumber="499">MVMA</collectionCode>
,
<collectionCode id="88B6C79C6714FF89FF59FEE1FF47FDCA" box="[151,238,368,392]" pageId="6" pageNumber="499">QMBA</collectionCode>
,
<collectionCode id="88B6C79C6714FF89FF37FEE1FEE4FDCA" LSID="urn:lsid:biocol.org:col:34982" box="[249,333,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="6" pageNumber="499">SAMA</collectionCode>
,
<collectionCode id="88B6C79C6714FF89FE96FEE1FE1DFDCA" box="[344,436,368,392]" pageId="6" pageNumber="499">WAMA</collectionCode>
,
<collectionCode id="88B6C79C6714FF89FE71FEE1FDBAFDCA" box="[447,531,368,392]" pageId="6" pageNumber="499">TMHA</collectionCode>
) and in
<collectionCode id="88B6C79C6714FF89FDA3FEE1FD6DFDC5" box="[621,708,368,391]" pageId="6" pageNumber="499">BMNH</collectionCode>
,
<collectionCode id="88B6C79C6714FF89FD1EFEE1FCB5FDCA" box="[720,796,368,392]" pageId="6" pageNumber="499">CASC</collectionCode>
,
<collectionCode id="88B6C79C6714FF89FCE9FEE1FCD0FDCA" box="[807,889,368,392]" pageId="6" pageNumber="499">MCZC</collectionCode>
,
<collectionCode id="88B6C79C6714FF89FC4AFEE1FC73FDCA" box="[900,986,368,392]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="499">MHNG</collectionCode>
.
</paragraph>
<paragraph id="EE185F596714FF89FF09FE05FB6AFF57" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="499">
<emphasis id="DCD3834B6714FF89FF09FE05FDE2FDEF" bold="true" box="[199,587,404,429]" pageId="6" pageNumber="499">Material examined, distribution.</emphasis>
<collectionCode id="88B6C79C6714FF89FD9FFE04FD38FDEE" box="[593,657,405,428]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="6" pageNumber="499">NEW</collectionCode>
<collectionCode id="88B6C79C6714FF89FD59FE05FCF4FDEE" box="[663,861,404,428]" pageId="6" pageNumber="499">
SOUTH
<collectingCountry id="96B01FC96714FF89FD35FE05FCF4FDEE" box="[763,861,404,428]" name="United Kingdom" pageId="6" pageNumber="499">WALES</collectingCountry>
</collectionCode>
: Alpine Creek [-35 55, 148 33],
<date id="9A1979996714FF89FB0BFE05FA9AFDEF" box="[1221,1331,404,429]" pageId="6" pageNumber="499" value="1935-11-04">4/xi/1935</date>
, G.F. Hill (
<collectionCode id="88B6C79C6714FF89FF51FE29FF5EFD92" box="[159,247,440,464]" pageId="6" pageNumber="499">QMBA</collectionCode>
, ANIC—some labeled “
<taxonomicName id="29A724DA6714FF89FDC1FE28FCA3FD92" box="[527,778,439,464]" class="Insecta" family="Formicidae" genus="Promyrmecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B6714FF89FDC1FE28FCA3FD92" box="[527,778,439,464]" italics="true" pageId="6" pageNumber="499">Promyrmecia pilosula</emphasis>
</taxonomicName>
” by John Clark); near Corang River Bridge (-35 12, 150 03) HI87–152, HI99–14; Dead Horse Gap [-36 31, 148 15], HI87–233, 234, HI91–046, 0 47, HI99–017;
<date id="9A1979996714FF89FAE4FE4DFA31FDB7" box="[1322,1432,476,501]" pageId="6" pageNumber="499" value="2006-01-27">27/i/2006</date>
, SB&amp;SM, 7 samples (
<collectionCode id="88B6C79C6714FF89FE4DFD91FE4DFE5A" box="[387,484,512,536]" pageId="6" pageNumber="499">AAVAS</collectionCode>
); Eucumbene [-36 0 8, 148 36],
<date id="9A1979996714FF89FC88FE6EFC15FE5A" box="[838,956,511,536]" pageId="6" pageNumber="499" value="1959-02-22">22/ii/1959</date>
, EFR; Jamberoo [-34 39, 150 46],
<quantity id="295FF2BC6714FF89FA8CFD91FAD2FE5A" box="[1346,1403,511,536]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="499" unit="km" value="7.0">7 km</quantity>
W
<quantity id="295FF2BC6714FF89FF59FDB5FF4AFE7E" box="[151,227,548,573]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="499" unit="m" value="600.0">600 m</quantity>
,
<date id="9A1979996714FF89FF3DFDB5FEFAFE7F" box="[243,339,548,573]" pageId="6" pageNumber="499" value="1977-11-26">26/xi/77</date>
, PSW acc 2900, 2907 (PSW collection); Pipers Creek [-33 3, 149 55],
<date id="9A1979996714FF89FB55FDB5FABEFE7F" box="[1179,1303,548,573]" pageId="6" pageNumber="499" value="1960-11-10">10/xi/1960</date>
, EFR; near Smiggins Holes [-33 59, 148 48],
<quantity id="295FF2BC6714FF89FDE8FDD6FD29FE1D" box="[550,640,583,608]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="499" unit="m" value="1650.0">1650 m</quantity>
,
<date id="9A1979996714FF89FD40FDD6FD49FE22" box="[654,736,583,608]" pageId="6" pageNumber="499" value="1975-01-27">27/i/75</date>
, PSW acc 607 (PSW collection). AUSTRALIAN CAPITAL TERRITORY: Mt Ginini, [-35 32, 148 46], HI99–016. VICTORIA: Benambra [-36 57, 147 42],
<quantity id="295FF2BC6714FF89FB14FDFDFA8DFEC6" box="[1242,1316,620,644]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="499" unit="m" value="700.0">700 m</quantity>
,
<date id="9A1979996714FF89FAFFFDFDFA35FEC7" box="[1329,1436,620,645]" pageId="6" pageNumber="499" value="2006-01-25">25/i/2006</date>
SB&amp;SM (
<collectionCode id="88B6C79C6714FF89FECBFD01FECFFEEA" box="[261,358,656,680]" pageId="6" pageNumber="499">AAVAS</collectionCode>
); Ben Cairn [-37 43, 145 37],
<date id="9A1979996714FF89FD7CFD01FC8EFEEA" box="[690,807,656,680]" pageId="6" pageNumber="499" value="1937-05-14">14/v/1937</date>
, TG; Mt Buffalo, near The Chalet ski lodge [-36 43, 146 49], HI87–224, 225, TASMANIA: near Bothwell [-42 23, 147 00], HI87–197–199; Epping Forest [-41 46, 147 21], BBL; Great Lake [-41 53, 146 44],
<date id="9A1979996714FF89FDEAFD49FD3EFEB2" box="[548,663,728,752]" pageId="6" pageNumber="499" value="1949-10-28">28/x/1949</date>
, TG; Miena [-41 59, 146 43], HI91–027, 0 41, HI 99–022; Murderers Hill [-41 58, 146 40], HI87–200; Tunbridge [-42 0 8, 147 25], BBL. All
<collectionCode id="88B6C79C6714FF89FC72FD6DFBA8FF56" LSID="urn:lsid:biocol.org:col:32981" box="[956,1025,764,788]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="6" pageNumber="499">ANIC</collectionCode>
unless indicated.
</paragraph>
</subSubSection>
<subSubSection id="A6BD0CD26714FF89FF09FC8EFCD2FBFA" pageId="6" pageNumber="499" type="description">
<paragraph id="EE185F596714FF89FF09FC8EFDCBF94F" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="499">
<emphasis id="DCD3834B6714FF89FF09FC8EFE0DFF7A" bold="true" box="[199,420,799,824]" pageId="6" pageNumber="499">Worker diagnosis.</emphasis>
General features as illustrated and in key couplet 1 above. The most easily recognized species of the
<taxonomicName id="29A724DA6714FF89FEFBFCD4FDA1FF1F" box="[309,520,836,861]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B6714FF89FEFBFCD4FDA1FF1F" box="[309,520,836,861]" italics="true" pageId="6" pageNumber="499">Myrmecia pilosula</emphasis>
</taxonomicName>
complex. Distinguished by the almost total lack of pilosity interrupting the dorsal body profile, by the lack of reddish-orange apical color-marking on the dark brown anterior femora, and the absence of brightly differentiated reddish-orange tibiae and tarsi on the more or less uniformly dark brown, apically lightly infuscated, middle and hind legs. The body often has a distinctive and sometimes very noticeable diffuse bluish cast in live specimens, recalling the bloom of grape skin. This feature seems unique to
<taxonomicName id="29A724DA6714FF89FB22FC45FAD2FFAF" box="[1260,1403,980,1005]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6714FF89FB22FC45FAD2FFAF" box="[1260,1403,980,1005]" italics="true" pageId="6" pageNumber="499">haskinsorum</emphasis>
</taxonomicName>
in
<taxonomicName id="29A724DA6714FF89FF59FC68FEA0F852" box="[151,265,1017,1040]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="genus">
<emphasis id="DCD3834B6714FF89FF59FC68FEA0F852" box="[151,265,1017,1040]" italics="true" pageId="6" pageNumber="499">Myrmecia</emphasis>
</taxonomicName>
and is lost in dry or spirit-preserved specimens. It has been observed in the Snowy Mountains, Mt Buffalo and central Tasmania, and appears less prominent at lower elevations of record. Labrum yellowish in the center with bilateral dark brown markings. Terminal two or three antennomeres usually darkly infuscated. Mesonotum (longitudinally striate-rugose) and petiolar dorsum (more or less transversely rugose) distinctly and quite strongly sculptured, with little evidence of allometric reduction in smaller specimens. Pubescence silvery, without yellowish reflections; very fine, short and sparse except on the clypeus, coxae, femora, postpetiole and gaster. A very few short hairs may be visible in profile view of the head and pronotum, and a few minute bristles on the mesonotum, less often on the petiole.
</paragraph>
<paragraph id="EE185F596714FF89FF09FA86FDDCF9DE" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="499">
<emphasis id="DCD3834B6714FF89FF09FA86FEF0F972" bold="true" box="[199,345,1303,1328]" pageId="6" pageNumber="499">Dimensions.</emphasis>
(
<typeStatus id="311CE1FB6714FF89FEA4FA86FE7FF972" box="[362,470,1303,1328]" pageId="6" pageNumber="499" type="holotype">Holotype</typeStatus>
, smallest available specimen (Mt Buffalo), largest available specimen (Mt Buffalo) (mm): TL = 11.6, 11.4, 12.0; HW = 2.36, 2.34, 2.46; HL = 2.15, 2.13, 2.22; CI = 109, 110, 111; EL = 0.92, 0.87, 0.92; OI = 39, 37, 41; SL = 1.94, 1.90, 1.93; SI = 82, 81, 78; PW = 1.48, 1.45, 1.48; WL = 3.64, 3.60, 3.78; PetW = 0.86, 0.85, 0.88; PpetW = 1.36, 1.33, 1.41.
</paragraph>
<paragraph id="EE185F596714FF89FF09FA36FB82FAFE" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="499">
<emphasis id="DCD3834B6714FF89FF09FA36FE08F982" bold="true" box="[199,417,1447,1472]" pageId="6" pageNumber="499">Elevational range.</emphasis>
The AAVAS Dead Horse Gap sites were recorded by GPS at
<quantity id="295FF2BC6714FF89FB94FA36FABCF982" box="[1114,1301,1447,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.5805" metricValueMax="1.608" metricValueMin="1.553" pageId="6" pageNumber="499" unit="m" value="1580.5" valueMax="1608.0" valueMin="1553.0">1,553 to 1,608m</quantity>
, the highest confirmed for the
<taxonomicName id="29A724DA6714FF89FEA6FA5CFE43F9A7" box="[360,490,1484,1509]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B6714FF89FEA6FA5CFE43F9A7" box="[360,490,1484,1509]" italics="true" pageId="6" pageNumber="499">M. pilosula</emphasis>
</taxonomicName>
complex,
<quantity id="295FF2BC6714FF89FDADFA5DFD0AF9A7" box="[611,675,1484,1509]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="6" pageNumber="499" unit="m" value="620.0">620m</quantity>
short of Mt Kosciusko summit.
<bibRefCitation id="8A3622A86714FF89FBD5FA5DFB64F9A6" author="Wheeler" box="[1051,1229,1484,1509]" pageId="6" pageNumber="499" refString="Wheeler, W. M. (1933) Colony-founding among Ants with an account of some primitive Australian species. Cambridge, Massachusetts, Harvard University Press, 179 pp." type="book" year="1933">Wheeler (1933)</bibRefCitation>
probably referred partly to
<taxonomicName id="29A724DA6714FF89FF30FA60FE1DFA4A" box="[254,436,1519,1544]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6714FF89FF30FA60FE1DFA4A" box="[254,436,1519,1544]" italics="true" pageId="6" pageNumber="499">M. haskinsorum</emphasis>
</taxonomicName>
when reporting “
<taxonomicName id="29A724DA6714FF89FDB2FA60FD54FA4A" box="[636,765,1519,1544]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B6714FF89FDB2FA60FD54FA4A" box="[636,765,1519,1544]" italics="true" pageId="6" pageNumber="499">M. pilosula</emphasis>
</taxonomicName>
” from Mt Kosciusko (see below). Elevations for other key localities are: Corang River Bridge, ca
<quantity id="295FF2BC6714FF89FDAFF985FD08FA6F" box="[609,673,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="6" pageNumber="499" unit="m" value="580.0">580m</quantity>
(GPS reading), the lowest mainland record for
<taxonomicName id="29A724DA6714FF89FB1FF985FAC9FA6F" box="[1233,1376,1556,1581]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6714FF89FB1FF985FAC9FA6F" box="[1233,1376,1556,1581]" italics="true" pageId="6" pageNumber="499">haskinsorum</emphasis>
</taxonomicName>
); Mt Buffalo, ca
<quantity id="295FF2BC6714FF89FED1F9A9FE29FA12" box="[287,384,1592,1616]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="499" unit="m" value="1300.0">1,300 m</quantity>
; and in Tasmania: Tunbridge, ca
<quantity id="295FF2BC6714FF89FCCCF9A9FCEBFA12" box="[770,834,1592,1616]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="499" unit="m" value="200.0">200m</quantity>
(The lowest-elevation confirmed Tasmanian record) and Murderers Hill, ca
<quantity id="295FF2BC6714FF89FE54F9CDFE5FFA36" box="[410,502,1628,1652]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="499" unit="m" value="1100.0">1,100 m</quantity>
(the highest-elevation Tasmanian record). Targeted search at the summit area of Mt William (= Mt Duwil –
<quantity id="295FF2BC6714FF89FE66F9EEFDA8FADA" box="[424,513,1663,1688]" metricMagnitude="3" metricUnit="m" metricValue="1.167" pageId="6" pageNumber="499" unit="m" value="1167.0">1,167m</quantity>
), the highest peak in the Grampian Ranges, VIC, by four collectors including the author, failed to find
<taxonomicName id="29A724DA6714FF89FE4DF934FD9EFAFF" box="[387,567,1700,1725]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6714FF89FE4DF934FD9EFAFF" box="[387,567,1700,1725]" italics="true" pageId="6" pageNumber="499">M. haskinsorum</emphasis>
</taxonomicName>
, which perhaps does not range that far west.
</paragraph>
<paragraph id="EE185F596714FF89FF09F956FCD2FBFA" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="499">
<emphasis id="DCD3834B6714FF89FF09F956FE92FAA2" bold="true" box="[199,315,1735,1760]" pageId="6" pageNumber="499">Variation.</emphasis>
The mainland and Tasmanian populations were presumably geographically isolated after dispersal of ancestral stock across intervening low elevation areas during a past cool climate episode prior to the separation of Tasmania (ca 14,000ybp) (since
<taxonomicName id="29A724DA6714FF89FDFBF89EFD6DFB6A" box="[565,708,1807,1832]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6714FF89FDFBF89EFD6DFB6A" box="[565,708,1807,1832]" italics="true" pageId="6" pageNumber="499">haskinsorum</emphasis>
</taxonomicName>
is present there). Diagnostic morphological characters are not significantly variable among available specimens. The Snowy Mountains and Mt Ginini populations are those most likely to have vestigial dorsal mesosomal pilosity. The body color is generally a little more blackish in
<taxonomicName id="29A724DA6714FF89FAC3F8C6FA35FB32" box="[1293,1436,1879,1904]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6714FF89FAC3F8C6FA35FB32" box="[1293,1436,1879,1904]" italics="true" pageId="6" pageNumber="499">haskinsorum</emphasis>
</taxonomicName>
than other
<taxonomicName id="29A724DA6714FF89FED5F8EDFEDCFBD7" box="[283,373,1916,1941]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B6714FF89FED5F8EDFEDCFBD7" box="[283,373,1916,1941]" italics="true" pageId="6" pageNumber="499">pilosula</emphasis>
</taxonomicName>
-group species, and the brown sections of the legs darker in hue. Specimens from lower elevations tend to have the leg color lighter brown, as in
<figureCitation id="769C43DC6714FF89FCDDF831FCDEFBFA" box="[787,887,1952,1977]" captionStart="FIGURES 1 – 3" captionStartId="7.[151,264,1957,1980]" captionTargetBox="[264,1323,195,1934]" captionTargetId="figure@7.[264,1323,195,1934]" captionTargetPageId="7" captionText="FIGURES 1 – 3. Myrmecia haskinsorum, Jamberoo, NSW. Standard views. HW 2.46 mm, WL 3.64 mm, PW 1.57 mm." httpUri="https://zenodo.org/record/398906/files/figure.png" pageId="6" pageNumber="499">Figs 1–3</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="BAD80FD16715FF88FF59F834FAE4FBFE" box="[151,1357,1957,1980]" httpUri="https://zenodo.org/record/398906/files/figure.png" pageId="7" pageNumber="500" targetBox="[264,1323,195,1934]" targetPageId="7">
<paragraph id="EE185F596715FF88FF59F834FAE4FBFE" blockId="7.[151,1357,1957,1980]" box="[151,1357,1957,1980]" pageId="7" pageNumber="500">
<emphasis id="DCD3834B6715FF88FF59F834FE93FBF9" bold="true" box="[151,314,1957,1980]" pageId="7" pageNumber="500">FIGURES 1–3.</emphasis>
<taxonomicName id="29A724DA6715FF88FE8EF836FD09FBFE" authority="Jamberoo" authorityName="Jamberoo" box="[320,672,1958,1980]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="500" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B6715FF88FE8EF836FD87FBFE" box="[320,558,1958,1980]" italics="true" pageId="7" pageNumber="500">Myrmecia haskinsorum</emphasis>
, Jamberoo
</taxonomicName>
, NSW. Standard views. HW 2.46 mm, WL 3.64 mm, PW 1.57 mm.
</paragraph>
</caption>
<subSubSection id="A6BD0CD2671AFF87FF09FF06FEA1FE5A" pageId="8" pageNumber="501" type="etymology">
<paragraph id="EE185F59671AFF87FF09FF06FC63FC96" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="501">
<emphasis id="DCD3834B671AFF87FF09FF06FEE7FCF2" bold="true" box="[199,334,151,176]" pageId="8" pageNumber="501">Etymology.</emphasis>
Named for Caryl P. Haskins and his wife Edna Haskins in recognition of their important pioneering studies on the biology of
<taxonomicName id="29A724DA671AFF87FDFFFF2CFD0AFC96" box="[561,675,189,212]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="genus">
<emphasis id="DCD3834B671AFF87FDFFFF2CFD0AFC96" box="[561,675,189,212]" italics="true" pageId="8" pageNumber="501">Myrmecia</emphasis>
</taxonomicName>
and other Australian ants.
</paragraph>
<paragraph id="EE185F59671AFF87FF09FF4EFC1DFDEE" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="501">
<emphasis id="DCD3834B671AFF87FF09FF4EFEE3FCBA" bold="true" box="[199,330,223,248]" pageId="8" pageNumber="501">Karyology.</emphasis>
Chromosome numbers vary between 2n=12 and 2n=24, the lower counts resulting mainly from centric fission, centric fusion and AM inversion (
<bibRefCitation id="8A3622A8671AFF87FD0EFE95FC7CFD5E" author="Imai" box="[704,981,260,285]" pageId="8" pageNumber="501" refString="Hirai, H., Yamamoto, M. T., Ogura, K., Satta, Y., Yamada, M., Taylor, R. W. &amp; Imai, H. T. (1994) Multiplication of 28 s rDNA and NOR activity in chromosome evolution among ants of the Myrmecia pilosula species complex (Hymenoptera: Formicidae: Myrmeciinae). Chromosoma, 103, 171 - 178. [Berlin] http: // dx. doi. org / 10.1007 / BF 00368009" type="journal article" year="1994">
Imai, Taylor
<emphasis id="DCD3834B671AFF87FC9DFE94FC3BFD5F" box="[851,914,260,285]" italics="true" pageId="8" pageNumber="501">et al.,</emphasis>
1994
</bibRefCitation>
: 147,
<figureCitation id="769C43DC671AFF87FBD7FE95FBFCFD5F" box="[1049,1109,260,285]" captionStart="FIGURES 4 – 6" captionStartId="9.[151,264,1954,1977]" captionTargetBox="[278,1308,193,1931]" captionTargetId="figure@9.[278,1308,193,1933]" captionTargetPageId="9" captionText="FIGURES 4 – 6. Myrmecia banksi, Holotype, South Nowra, NSW. Standard views. HW 2.66 mm, WL 3.86 mm, PW 1.72 mm." httpUri="https://zenodo.org/record/398907/files/figure.png" pageId="8" pageNumber="501">Fig 6</figureCitation>
). The karyotypes of colonies from Corang River Bridge and Mt. Buffalo, respectively with 2n=23/24 and 2n=20, were considered by Imai, Taylor
<emphasis id="DCD3834B671AFF87FF29FEDCFE89FD27" box="[231,288,332,357]" italics="true" pageId="8" pageNumber="501">et al.</emphasis>
to be closely related to those from Tasmania with 2n=18. The Dead Horse Gap colonies have 2n=
<date id="9A197999671AFF87FAB7FEDDFF70FDCA" pageId="8" pageNumber="501" value="1918-12-15">18/ 15/12</date>
, and are karyotypically most similar to a Tasmanian colony with 2n=17. Four fusions with different arm combinations were detected from colony HI87–234 (Dead Horse Gap).
</paragraph>
<paragraph id="EE185F59671AFF87FF09FE26FEA1FE5A" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="501">
<emphasis id="DCD3834B671AFF87FF09FE26FE0DFD92" bold="true" box="[199,420,439,464]" pageId="8" pageNumber="501">Field associations.</emphasis>
Sympatric with the Eastern Race of
<taxonomicName id="29A724DA671AFF87FCAFFE28FC4CFD92" box="[865,997,439,464]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B671AFF87FCAFFE28FC4CFD92" box="[865,997,439,464]" italics="true" pageId="8" pageNumber="501">M. pilosula</emphasis>
</taxonomicName>
and
<taxonomicName id="29A724DA671AFF87FBEBFE28FB11FD92" box="[1061,1208,439,464]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="croslandi">
<emphasis id="DCD3834B671AFF87FBEBFE28FB11FD92" box="[1061,1208,439,464]" italics="true" pageId="8" pageNumber="501">M. croslandi</emphasis>
</taxonomicName>
near Corang River Bridge, with Eastern Race
<taxonomicName id="29A724DA671AFF87FE1EFE4DFD82FDB7" box="[464,555,476,501]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B671AFF87FE1EFE4DFD82FDB7" box="[464,555,476,501]" italics="true" pageId="8" pageNumber="501">pilosula</emphasis>
</taxonomicName>
in the Snowy Mountains and on Mt Buffalo, and with the Western Race in Tasmania.
</paragraph>
</subSubSection>
<subSubSection id="A6BD0CD2671AFF87FF09FDB5FE88FF1F" pageId="8" pageNumber="501" type="discussion">
<paragraph id="EE185F59671AFF87FF09FDB5FCA5FE8E" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="501">
<emphasis id="DCD3834B671AFF87FF09FDB5FEE5FE7F" bold="true" box="[199,332,548,573]" pageId="8" pageNumber="501">Discussion.</emphasis>
<bibRefCitation id="8A3622A8671AFF87FE9AFDB5FDF6FE7F" author="Wheeler" box="[340,607,548,573]" pageId="8" pageNumber="501" refString="Wheeler, W. M. (1933) Colony-founding among Ants with an account of some primitive Australian species. Cambridge, Massachusetts, Harvard University Press, 179 pp." type="book" year="1933">Wheeler (1933: 56–57)</bibRefCitation>
, who confused the two races of
<taxonomicName id="29A724DA671AFF87FC1FFDB4FBFAFE7F" box="[977,1107,548,573]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B671AFF87FC1FFDB4FBFAFE7F" box="[977,1107,548,573]" italics="true" pageId="8" pageNumber="501">M. pilosula</emphasis>
</taxonomicName>
and
<taxonomicName id="29A724DA671AFF87FB42FDB4FAEBFE7F" box="[1164,1346,548,573]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B671AFF87FB42FDB4FAEBFE7F" box="[1164,1346,548,573]" italics="true" pageId="8" pageNumber="501">M. haskinsorum</emphasis>
</taxonomicName>
, almost certainly referred to
<taxonomicName id="29A724DA671AFF87FE4AFDD6FDBAFE22" box="[388,531,583,608]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="haskinsorum">
<emphasis id="DCD3834B671AFF87FE4AFDD6FDBAFE22" box="[388,531,583,608]" italics="true" pageId="8" pageNumber="501">haskinsorum</emphasis>
</taxonomicName>
when describing “
<taxonomicName id="29A724DA671AFF87FD23FDD6FCEEFE22" box="[749,839,583,608]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="pilosula">
<emphasis id="DCD3834B671AFF87FD23FDD6FCEEFE22" box="[749,839,583,608]" italics="true" pageId="8" pageNumber="501">pilosula</emphasis>
</taxonomicName>
” workers and queens on Mount Kosciusco as “dull black, but when alive and in bright sunlight, with a distinct bluish tinge, probably due to a fugitive pigment”. Also, at least in part, when stating that on Kosciusko (“
<emphasis id="DCD3834B671AFF87FD1EFD00FCCBFEEA" box="[720,866,655,680]" italics="true" pageId="8" pageNumber="501">
<taxonomicName id="29A724DA671AFF87FD1EFD00FCF0FEEA" box="[720,857,655,680]" class="Insecta" family="Formicidae" genus="Myrmecia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="pilosula">M. pilosula</taxonomicName>
”
</emphasis>
) was “very abundant and the dominant species at elevations from
<quantity id="295FF2BC671AFF87FE83FD25FE54FE8F" box="[333,509,692,717]" metricMagnitude="3" metricUnit="m" metricValue="1.524" metricValueMax="1.8288" metricValueMin="1.2192" pageId="8" pageNumber="501" unit="ft" value="5000.0" valueMax="6000.0" valueMin="4000.0">4000 to 6000 ft</quantity>
.” (c.a.
<quantity id="295FF2BC671AFF87FD84FD25FD56FE8E" box="[586,767,692,716]" metricMagnitude="3" metricUnit="m" metricValue="1.529" metricValueMax="1.838" metricValueMin="1.22" pageId="8" pageNumber="501" unit="m" value="1529.0" valueMax="1838.0" valueMin="1220.0">
<date id="9A197999671AFF87FD84FD25FD77FE8E" box="[586,734,692,716]" pageId="8" pageNumber="501" value="1220" valueMax="1838">1220 to 1838</date>
m
</quantity>
).
</paragraph>
<paragraph id="EE185F59671AFF87FF09FD46FE88FF1F" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="501">
<emphasis id="DCD3834B671AFF87FF09FD46FE1CFEB2" bold="true" box="[199,437,727,752]" pageId="8" pageNumber="501">Research prospects.</emphasis>
Future potential research topics include (1) genetical and morphological comparison of the mainland and Tasmanian populations, which have a defined approximate time of geographical and phylogenetic separation; (2) investigation of the nature and role of the fugitive bluish flush in high alpine populations.
</paragraph>
</subSubSection>
</treatment>
</document>
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