246 lines
26 KiB
XML
246 lines
26 KiB
XML
<document id="559C0A0793B0226B1C44B946E74F5933" ID-CLB-Dataset="27116" ID-DOI="http://dx.doi.org/10.3897/zookeys.840.33326" ID-GBIF-Dataset="363f2c7c-0635-46c8-b598-9e9aa014b6d2" ID-PMC="PMC6482118" ID-Pensoft-Pub="1313-2970-840-35" ID-PubMed="31065227" ID-ZooBank="C607C2FDC17644FABBC582D561C6434C" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-840-35" ModsDocOrigin="ZooKeys 840" ModsDocTitle="Conchological and molecular analysis of the “non-scaly” Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)" checkinTime="1555526082661" checkinUser="pensoft" docAuthor="Zacaery Khalik, Mohd, P. Hendriks, Kasper, J. Vermeulen, Jaap & Menno Schilthuizen," docDate="2019" docId="C635D90329018AEC41FE14ACE0DCDA76" docLanguage="en" docName="ZooKeys 840: 35-86" docOrigin="ZooKeys 840" docSource="http://dx.doi.org/10.3897/zookeys.840.33326" docTitle="Georissa borneensis Smith 1895" docType="treatment" docVersion="6" lastPageNumber="44" masterDocId="647C1D35F570C105FFFCFFCDB169212D" masterDocTitle="Conchological and molecular analysis of the " non-scaly " Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)" masterLastPageNumber="86" masterPageNumber="35" pageNumber="43" updateTime="1701381596384" updateUser="ExternalLinkService">
|
||
<mods:mods id="C35E2F09BD1FED1A15EF63ECAC5A72C1" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="B72050D6C9DC570A0E4D9BCB8AFB67B1">
|
||
<mods:title id="F85C96068B8A54CFAB30870D7181DA60">Conchological and molecular analysis of the " non-scaly " Bornean Georissa with descriptions of three new species (Gastropoda, Neritimorpha, Hydrocenidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FD5C11BCD16BB27CC870DAC483FA3A03" type="personal">
|
||
<mods:role id="5AF36EC45292F1281A8FBF76B464E501">
|
||
<mods:roleTerm id="DBE962AE4EAD7FFDE3F53665F29A7A16">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="92C1C9F338AA785FF9CD41598E6902D5">Zacaery Khalik, Mohd</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F4879ADF665E8909205CCD5734E64702" type="personal">
|
||
<mods:role id="6F940C5A5E4D9D302EE343EE34704D81">
|
||
<mods:roleTerm id="9D8624D6832B961A9B63B1FEFD022CCB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="23CA64E28AD41EA706C7A63724ADD230">P. Hendriks, Kasper</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="9286683C64B339C22258449792635455" type="personal">
|
||
<mods:role id="15B5275EF1EAADC3CF7C8FBAD377BAB0">
|
||
<mods:roleTerm id="98C1FAD837AFD81BAB2293C70A69E0F3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1101A10082A8C5F24E5FF22F57EF42AB">J. Vermeulen, Jaap</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FFCE4DC04842A7866E1648AC19560885" type="personal">
|
||
<mods:role id="E1223B8A7150D55FABBE1120D991A5F2">
|
||
<mods:roleTerm id="27F44BC31F64A7C7598EE34D5CB5B32D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9DC30429ECB3520CACB1747EBFA474D6">Menno Schilthuizen,</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="76FB9E001C7ACC6ED8424C95401EAF97">text</mods:typeOfResource>
|
||
<mods:relatedItem id="3237EB0F303398D05434EB2CE394FB9E" type="host">
|
||
<mods:titleInfo id="647E8BB7DC0CD792236702AADD5DC8D2">
|
||
<mods:title id="E8F650B46371B318448A827B472EAA0C">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="86C7AA47FCA2A1EB999264D078FF4AC4">
|
||
<mods:date id="0411B5DAF898608879E0979C0FB9A7F5">2019</mods:date>
|
||
<mods:detail id="C92ED26A12C1CD32D35FE3AB7C48C22A" type="volume">
|
||
<mods:number id="ABB7A3397A4F016B37376805C70203F2">840</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="34F70F9CDD161DE16EDB1B050D7F0533" unit="page">
|
||
<mods:start id="BBBA1DE4FA513EF73E8FC5CB233B291D">35</mods:start>
|
||
<mods:end id="F5BC7EB809B6F93DCAB119FCA7B2D1A9">86</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="39113D6091A98E68AD59D4ECD6CC88ED">
|
||
<mods:url id="825150CF1D90C4BC2A205EAD52C98DB3">http://dx.doi.org/10.3897/zookeys.840.33326</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="7865B875A389EBE81629F69728FDA61A">journal article</mods:classification>
|
||
<mods:identifier id="133767284EFB23BEFAAF4A4F5D0DF829" type="DOI">http://dx.doi.org/10.3897/zookeys.840.33326</mods:identifier>
|
||
<mods:identifier id="3EA77DA7D3D36A57B0B4475A588CED9F" type="Pensoft-Pub">1313-2970-840-35</mods:identifier>
|
||
<mods:identifier id="CF8A3D816467509034A712C0B70ED577" type="ZooBank">C607C2FDC17644FABBC582D561C6434C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="C635D90329018AEC41FE14ACE0DCDA76" ID-GBIF-Taxon="156787938" LSID="urn:lsid:plazi:treatment:C635D90329018AEC41FE14ACE0DCDA76" httpUri="http://treatment.plazi.org/id/C635D90329018AEC41FE14ACE0DCDA76" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43">
|
||
<subSubSection id="59AA21002C45D3119BF0D6F6CD66DA79" pageId="8" pageNumber="43" type="nomenclature">
|
||
<paragraph id="58D5CE19F69B901A699680032C7739A7" pageId="8" pageNumber="43">
|
||
<taxonomicName id="4C66E1C64D64D70605BFDD4259A5927C" ID-CoL="7RDTH" authority="Smith, 1895" authorityName="Smith" authorityYear="1895" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa borneensis" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="borneensis">Georissa borneensis Smith, 1895</taxonomicName>
|
||
Figures 1A, 5
|
||
<normalizedToken id="ECD96F1A3A5EF88728904D9CF2972A09" originalValue="A–K">A-K</normalizedToken>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EED91A1273BFD2FA924401D6C3928C4E" pageId="8" pageNumber="43" type="reference_group">
|
||
<paragraph id="02C63F7CD40670A399F1DE8E4F643C3F" pageId="8" pageNumber="43">
|
||
<taxonomicName id="E0DFF6F0D6FBEFB71B696EC26F8C58BE" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa borneensis" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="borneensis">Georissa borneensis</taxonomicName>
|
||
Smith, 1895: 126, plate IV fig. 18;
|
||
<bibRefCitation id="7A3BB89FA482C3AF163471E08A509FA6" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance 1983</bibRefCitation>
|
||
: 122, figs 18, 61-62.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0ECF1EB085E91F5EA5710DE991B5F303" pageId="8" pageNumber="43" type="type locality">
|
||
<paragraph id="4F34BE6772C0BA80D2417D1265B917FC" pageId="8" pageNumber="43">Type locality.</paragraph>
|
||
<paragraph id="F0B8D90D9C20367B0F5561994AFB9061" pageId="8" pageNumber="43">Gomanton, N.E. Borneo.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE7D339B08C5B7DE8C79F6DD7C996182" pageId="8" pageNumber="43" type="type material">
|
||
<paragraph id="CFF30D75ECBDF7792A0DE1EA0F018A38" pageId="8" pageNumber="43">Type material.</paragraph>
|
||
<paragraph id="D799730499AA0997894E766077D34F97" pageId="8" pageNumber="43">
|
||
Lectotype (Designation by
|
||
<bibRefCitation id="E304AD33307CAC63F25550CC03B9513D" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance 1983</bibRefCitation>
|
||
) (Fig. 1A). Gomanton, N. Borneo: NHMUK 1894.7.20.61 (glued on paper). Paralectotypes. Gomanton, N. Borneo: NHMUK 1894.7.20.62, NHMUK 1894.21.54-57 (glued on paper).
|
||
</paragraph>
|
||
<caption id="A523CED47DAF2E6808E86754C3529648" pageId="8" pageNumber="43">
|
||
<paragraph id="A3D1A0EAA9193E3A8906F1D9B81503F1" pageId="8" pageNumber="43">
|
||
Figure 1. The types specimens of the
|
||
<normalizedToken id="BD0C39083A352FEB42F409BA187EAAC2" originalValue="“non-scaly”">"non-scaly"</normalizedToken>
|
||
<taxonomicName id="E1A16ECCA22A34438437CA6E4A3FEECA" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
|
||
of Borneo from NHMUK. A
|
||
<taxonomicName id="BD56E75ADA785F8FDB05CEC5342EA501" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa borneensis" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="borneensis">Georissa borneensis</taxonomicName>
|
||
Smith, 1895 B
|
||
<taxonomicName id="BD195707E8BD3461F5D24CBA014C1E2F" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa everetti" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="everetti">Georissa everetti</taxonomicName>
|
||
Smith, 1895 C
|
||
<taxonomicName id="EAA730DBF67A5357EC7582A1FB65A0A1" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa williamsi" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="williamsi">Georissa williamsi</taxonomicName>
|
||
Godwin-Austen, 1889 D
|
||
<taxonomicName id="4C651766E37F54AC241E62EFF0943F2E" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa hungerfordi" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="hungerfordi">Georissa hungerfordi</taxonomicName>
|
||
Godwin-Austen, 1889 E
|
||
<taxonomicName id="DD771884FE529C6FBFEFDCA940DD2F53" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa gomantonensis" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="gomantonensis">Georissa gomantonensis</taxonomicName>
|
||
Smith, 1893 F
|
||
<taxonomicName id="BC481A52149CF847F5ED042C555C4A30" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa similis" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="similis">Georissa similis</taxonomicName>
|
||
Smith, 1893 G
|
||
<taxonomicName id="64B77660256A8BBBC0116EB591CDA87E" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa xesta" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="xesta">Georissa xesta</taxonomicName>
|
||
Thompson & Dance, 1983 H
|
||
<taxonomicName id="48E7CE273858C5FB9C7E2BAEA3837495" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa bangueyensis" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="bangueyensis">Georissa bangueyensis</taxonomicName>
|
||
Smith, 1895 I
|
||
<taxonomicName id="5BB563CB1CA95C304FDD06C550B94D71" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa flavescens" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="flavescens">Georissa flavescens</taxonomicName>
|
||
Smith, 1895. Scale bar: 1 mm. Photos by NHMUK.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="5FD18214132F834EDCDBFDBD2B4ED5B9" pageId="8" pageNumber="43">
|
||
<paragraph id="C2EBC363AE8FC4367D6D6C191BA124B4" pageId="8" pageNumber="43">
|
||
Figure 2. A Maximum likelihood phylogenetic reconstruction with ultrafast bootstrapping (5000 replicates) B bayesian phylogenetic reconstruction with posterior probabilities, constructed using MrBayes analysis. Phylogenetic analyses were conducted using concatenated sequence alignments of partial 16S and CO1 mtDNA. The analyses consist of 69 ingroup taxa (11 taxa representing the
|
||
<normalizedToken id="F34FB0CCB877222AA937953303A0A09F" originalValue="“scaly”">"scaly"</normalizedToken>
|
||
group and 58 taxa representing the
|
||
<normalizedToken id="3067430098FA928A69495843E6E48922" originalValue="“non-scaly”">"non-scaly"</normalizedToken>
|
||
group), and
|
||
<taxonomicName id="60A64A025A0F9C1A717CDA87A7191B02" class="Gastropoda" family="Phenacolepadidae" genus="Bathynerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bathynerita naticoidea" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="naticoidea">Bathynerita naticoidea</taxonomicName>
|
||
as an outgroup.
|
||
<normalizedToken id="4320CE91BC7954C3E43FDA4F156FDFE8" originalValue="“Scaly”">"Scaly"</normalizedToken>
|
||
taxa in the phylogenies are with the red branches.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7B3556B0C3DCC4F703275F7CA22D495B" pageId="8" pageNumber="43">
|
||
<paragraph id="68D521E2320A06ACDD71F6376A5E37E1" pageId="8" pageNumber="43">
|
||
Figure 3. The distribution of eight species of the
|
||
<normalizedToken id="C036DB2C8F9B5A678C6A2361ED94F296" originalValue="“non-scaly”">"non-scaly"</normalizedToken>
|
||
<taxonomicName id="5E756BA2AB3D0F69117C1B602B3EB070" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
|
||
of Malaysian Borneo, based on studied materials.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C70B7DFBE1FB2978E3E602F420CAB4F8" pageId="8" pageNumber="43">
|
||
<paragraph id="E3252750291EA6A1F5CE51A2D8A727FB" pageId="8" pageNumber="43">
|
||
Figure 4. The distribution of another eight species of the
|
||
<normalizedToken id="FB9BADA146309068E92233DD04B2C4CB" originalValue="“non-scaly”">"non-scaly"</normalizedToken>
|
||
<taxonomicName id="061E005CD3C24734BFAD6F1943A88E32" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="genus">Georissa</taxonomicName>
|
||
of Malaysian Borneo, based on studied materials.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="08F962BBB72EC6499C5FE4B72B053632" pageId="8" pageNumber="43" type="materials_examined">
|
||
<paragraph id="BE3C28B6237AC639431873F5444181C8" pageId="8" pageNumber="43">Other material.</paragraph>
|
||
<paragraph id="03DF5688D3BE524296D8FB2096941463" pageId="8" pageNumber="43">
|
||
N. Borneo: RMNH/MOL 152748, ZMA/MOLL 315546 (Fig. 5). Gomanton, N. Borneo: MFN 47552, MFN 47942. Kinabatangan valley, Gomantong Hill 30 km South of Sandakan, Sandakan Province, Sabah (
|
||
<geoCoordinate id="EA65CAC75A84CCCF51DF93FEA7474277" direction="north" orientation="latitude" precision="9" value="5.32">05°19.20'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2E574391FE7277A27363D155D97C3F0C" direction="east" orientation="longitude" precision="9" value="118.06">118°3.60'E</geoCoordinate>
|
||
): JJV 1613.
|
||
</paragraph>
|
||
<caption id="EDDACBCEA8C851CC555F1B70C1C41109" pageId="8" pageNumber="43">
|
||
<paragraph id="67A4C0F7A9F1A192307C6DE27C625326" pageId="8" pageNumber="43">
|
||
Figure 5.
|
||
<taxonomicName id="4D91275475681CF4DCAE98F037EF91C4" class="Gastropoda" family="Hydrocenidae" genus="Georissa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Georissa borneensis" order="Cycloneritida" pageId="8" pageNumber="43" phylum="Mollusca" rank="species" species="borneensis">Georissa borneensis</taxonomicName>
|
||
Smith, 1895.
|
||
<normalizedToken id="250C7B2CA9FB65B86253139A531B1DC0" originalValue="A–K">A-K</normalizedToken>
|
||
ZMA/MOLL 315546 A, D shell apertural view B shell side view C shell rear view E, F shell cross-section from 3D model G, H operculum frontal and ventral view from 3D model I shell top view J protoconch side view K close up of protoconch from top at 1000
|
||
<normalizedToken id="E0D63C9A65F98C1E4CD0195E69067082" originalValue="×">x</normalizedToken>
|
||
magnification. Scale bars: 500
|
||
<normalizedToken id="F5F4E25FF72EC422F06C4BC3F5778FD3" originalValue="µm">µm</normalizedToken>
|
||
(
|
||
<normalizedToken id="EA62EA84ABFBC52080D1FC7528395EB4" originalValue="A–I">A-I</normalizedToken>
|
||
); 200
|
||
<normalizedToken id="A6BD99835AEB92834221FEE2B7CFEE8B" originalValue="µm">µm</normalizedToken>
|
||
(J); 10
|
||
<normalizedToken id="7A2256FBBA17CF1E52591B429477BBF7" originalValue="µm">µm</normalizedToken>
|
||
(K).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="4606FBB3C83C0EE88E0FC5D933729199" pageId="8" pageNumber="43" type="description">
|
||
<paragraph id="5ABF3FB7876AF1D546588CD5426372CE" pageId="8" pageNumber="43">Description.</paragraph>
|
||
<paragraph id="DDCE2ECF66E60BC16F06EB63640B055C" pageId="8" pageNumber="43">
|
||
Protoconch. Colour: white to pale orange, darker than the rest of the shell. Sculpture pattern: irregular sculpture pattern, from base to apex end with no specific sculptural shape. Mesh width: 2.5-8.0
|
||
<normalizedToken id="6417BB4AB6DDED815A592BCB8B7E2554" originalValue="µm">µm</normalizedToken>
|
||
. Teleoconch. Colour: white to pale orange. First whorl: flat, convex close to the suture. Subsequent whorls: flat, convex and angular at the periphery. Suture: clearly impressed. Shoulder: narrow. Number of whorls: 2
|
||
<normalizedToken id="33FEEE3EEBB01A351A8A2551DBFA36BF" originalValue="¾–">3/4-</normalizedToken>
|
||
3
|
||
<normalizedToken id="47388C95217C75AEDE94605A16D150C7" originalValue="½">1/2</normalizedToken>
|
||
. SH: 1.91-2.23 mm. SW: 1.65-1.82 mm. SI: 1.12-1.28. Shell sculpture. Radial sculpture: present, weak or flattened, densely sculpted on the whorls, ca. two to three ribs per 0.1 mm. Spiral sculpture: present, but thin and weak, only visible under high magnification (>
|
||
<normalizedToken id="B77DD19596C53F6A3033F558336194A7" originalValue="×">x</normalizedToken>
|
||
100 magnification), strongest at the first whorl, weaker at subsequent whorls. Aperture. Shape: semi-elliptic, straight parietal side, palatal edge contiguous with the body whorl, palatal side tilted and angular, basal side convex. AH: 0.82-1.07 mm. AW: 1.00-1.09 mm. AI: 0.75-1.02.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="84EC13031F6C48EDB623F62000BDAFB6" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43" type="diagnosis">
|
||
<paragraph id="736B39822B3666D6530FA44FA541CA8D" pageId="8" pageNumber="43">Diagnosis.</paragraph>
|
||
<paragraph id="EEE5F4B10B25F1E3CAB9422E8D9781ED" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43">
|
||
The flat whorls that are strongly convex at the periphery, giving the shell an angular shape, are diagnostic. The sculpture of
|
||
<taxonomicName id="E1337718696C4634658E74BEB9CE7A01" lsidName="G. borneensis" pageId="8" pageNumber="43" rank="species" species="borneensis">G. borneensis</taxonomicName>
|
||
resembles that of
|
||
<taxonomicName id="23AED78EA2EACCC8B4A947ACA46E67C8" lsidName="G. similis" pageId="8" pageNumber="43" rank="species" species="similis">G. similis</taxonomicName>
|
||
and
|
||
<taxonomicName id="C1C2031D0DE0F63A01958A16FCFD3AA0" lsidName="G. corrugata" pageId="8" pageNumber="43" rank="species" species="corrugata">G. corrugata</taxonomicName>
|
||
, but is weaker and more flattened than in those species. The spiral sculpture of
|
||
<taxonomicName id="31BDCD39787CA4CDD878F74969B1584D" lsidName="G. corrugata" pageId="8" pageNumber="43" rank="species" species="corrugata">G. corrugata</taxonomicName>
|
||
is also more irregular. The adult shell
|
||
<taxonomicName id="ABCEEAEC96F8A02AB8DA91707925B8A9" lsidName="G. borneensis" pageId="8" pageNumber="43" rank="species" species="borneensis">G. borneensis</taxonomicName>
|
||
is larger than in adult
|
||
<taxonomicName id="6AA09204B354D49B2B288F06CF604294" lsidName="G. similis" pageId="8" pageNumber="43" rank="species" species="similis">G. similis</taxonomicName>
|
||
and
|
||
<taxonomicName id="BE9BE57135EB6FC0D95DFA1AE431B2E0" lsidName="G. corrugata" pageId="8" pageNumber="43" rank="species" species="corrugata">G. corrugata</taxonomicName>
|
||
. Additonally, the base to apex end sculpture of the protoconch of
|
||
<taxonomicName id="86CB79F8C9FAD5D49DC8AC4E75F5684C" lsidName="G. borneensis" pageId="8" pageNumber="43" rank="species" species="borneensis">G. borneensis</taxonomicName>
|
||
is distinct compared to these species (
|
||
<taxonomicName id="E2677E2120FDAA6CBB0192A113C4AF79" lsidName="G. similis" pageId="9" pageNumber="44" rank="species" species="similis">
|
||
<pageBreakToken id="24E89315E2C79E68D4B276FF91C2D4A1" pageId="9" pageNumber="44" start="start">G</pageBreakToken>
|
||
. similis
|
||
</taxonomicName>
|
||
has a rounded protoconch sculpture and
|
||
<taxonomicName id="993FAAFF1196047C4FD5D8D993F982CE" lsidName="G. corrugata" pageId="9" pageNumber="44" rank="species" species="corrugata">G. corrugata</taxonomicName>
|
||
has straight-line protoconch sculpture).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C6507D993DDBEACF969F39578B503F2C" pageId="9" pageNumber="44" type="distribution">
|
||
<paragraph id="EFA9EB0D527A4B35E20FCD1C4E874F71" pageId="9" pageNumber="44">Distribution.</paragraph>
|
||
<paragraph id="1297EA3D18AA3FF35FEB9732178F2613" pageId="9" pageNumber="44">The species is known only from Gomantong hill in the Kinabatangan region of Sabah.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A627C0178BB1445DCF7AA5F44B078163" pageId="9" pageNumber="44" type="discussion">
|
||
<paragraph id="3969DB79DABAB487169389FBF9D30DED" pageId="9" pageNumber="44">Discussion.</paragraph>
|
||
<paragraph id="DCF84A255C16F928012325EC487A8852" pageId="9" pageNumber="44">
|
||
The identification of
|
||
<taxonomicName id="36C1ABAF0AF8ABA22F7854B2AD8C37B7" lsidName="G. borneensis" pageId="9" pageNumber="44" rank="species" species="borneensis">G. borneensis</taxonomicName>
|
||
can be confusing when we refer to the sketches by
|
||
<bibRefCitation id="C7805DBC1D9AE9E7F34D646EF9EE4AE5" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance (1983</bibRefCitation>
|
||
: figs 61, 62). These appear to reflect the radial sculpture of
|
||
<taxonomicName id="473D3D5526ECE6F78E55441D475E9E4F" lsidName="G. corrugata" pageId="9" pageNumber="44" rank="species" species="corrugata">G. corrugata</taxonomicName>
|
||
, which is wavy/irregular and strongly sculpted.
|
||
<bibRefCitation id="1ED6091FFB3C224057D1E0A7D447EF64" author="Smith, EA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="41" pageNumber="76" pagination="97 - 127" title="On a collection of land-shells from Sarawak, British North Borneo, Palawan, and other neighboring islands." url="https://doi.org/10.1111/j.1469-7998.1895.tb07883.x" volume="63" year="1895">Smith (1895)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="5588101D2D38B3DAAAE2889C0136C2CF" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance (1983)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="0019064CDFEAE044CB706C2050DE0F70" lsidName="G. borneensis" pageId="9" pageNumber="44" rank="species" species="borneensis">G. borneensis</taxonomicName>
|
||
referring to the absence of spiral sculpture, which, however, is present but only visible under high magnification. The bright red colour of the peristome as described by
|
||
<bibRefCitation id="3732A220EB6061C25158758BB264E745" author="Smith, EA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="41" pageNumber="76" pagination="97 - 127" title="On a collection of land-shells from Sarawak, British North Borneo, Palawan, and other neighboring islands." url="https://doi.org/10.1111/j.1469-7998.1895.tb07883.x" volume="63" year="1895">Smith (1895)</bibRefCitation>
|
||
could not be observed by us, probably due to the faded condition of the shells. We also find there is no association based on colouration of
|
||
<taxonomicName id="60EE8218A8887D4B4B68096DB028470E" lsidName="G. borneensis" pageId="9" pageNumber="44" rank="species" species="borneensis">G. borneensis</taxonomicName>
|
||
with
|
||
<taxonomicName id="B01852E2C9FCA5F21E09AAB60394C714" lsidName="G. pyrrhoderma" pageId="9" pageNumber="44" rank="species" species="pyrrhoderma">G. pyrrhoderma</taxonomicName>
|
||
and the
|
||
<normalizedToken id="BBC593B154C14BDC2D23AF8127EF84C3" originalValue="“williamsi”">"williamsi"</normalizedToken>
|
||
.
|
||
<bibRefCitation id="3F712430DD6980D5F2E4D061656CE191" author="Thompson, FG" journalOrPublisher="Bulletin of the Florida State Museum Biological Sciences" pageId="41" pageNumber="76" pagination="101 - 152" title="Non-marine mollusks of Borneo. II Pulmonata: Pupillidae, Clausiliidae. III Prosobranchia: Hydrocenidae, Helicinidae." url="https://www.floridamuseum.ufl.edu/files/2114/7180/1931/Vol-29-No-3.PDF" volume="29" year="1983">Thompson and Dance (1983)</bibRefCitation>
|
||
grouped the
|
||
<normalizedToken id="6782B2EDC8A9D8F3E83516971D657ACC" originalValue="‘borneensis’">'borneensis'</normalizedToken>
|
||
based on their reddish shell colour, while
|
||
<normalizedToken id="B3C7F43B28452736FE1988C66ECA4F30" originalValue="‘williamsi’">'williamsi'</normalizedToken>
|
||
with their light brown colour, of which we find these colours are often a variation within these groups of species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |