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<document id="1189A7E6FC8B3D80FFAE27315FDA0E57" ID-CLB-Dataset="9409" ID-DOI="10.11646/zootaxa.4890.3.1" ID-GBIF-Dataset="2565fffa-2704-4101-8fdf-2b863ef19be5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4306124" ID-ZooBank="4D7E2CD3-5350-479F-905A-C2671C414CBC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607097313496" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2020" docId="AE6287F45021FFE0A7FFFB6B7FA8F9C6" docLanguage="en" docName="zootaxa.4890.3.1.pdf" docOrigin="Zootaxa 4890 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nipponophloeostiba longilobata Shavrin 2020, sp.n." docType="treatment" docVersion="10" lastPageNumber="319" masterDocId="525BFF8C5031FFF2A768FFAA7D46FFB5" masterDocTitle="New species and records of Paraphloeostiba Steel, 1960 from China and Laos and descriptions of four new species of related genera (Coleoptera: Staphylinidae Omaliinae: Omaliini)" masterLastPageNumber="329" masterPageNumber="301" pageNumber="317" updateTime="1698847096712" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="30EDCA48FCE60252B9A7FF46D51D6F25" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7B79C7E48CB5D52B66AD73CE5382356D">New species and records of Paraphloeostiba Steel, 1960 from China and Laos and descriptions of four new species of related genera (Coleoptera: Staphylinidae Omaliinae: Omaliini)</mods:title>
</mods:titleInfo>
<mods:name id="1DCB221BCB5AF47E440C62600E935E9B" type="personal">
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<mods:namePart id="8340959430412EFEF6793260B0806935">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="96A7735F9135ED1E3D9AA358FE98626D">Zootaxa</mods:title>
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<mods:part id="DD490C3F08521106BE98209BDB3B49EF">
<mods:date id="30A5876DF567E3919E90BB63412A361A">2020</mods:date>
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<mods:number id="06D3F843BEAF6336938CEE343B66CE96">2020-12-03</mods:number>
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<mods:number id="FF903EE11880CDFCEF9F6266BBCC23B8">4890</mods:number>
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<mods:classification id="0EF3D5BB6403E6F6A5849A1A55C74106">journal article</mods:classification>
<mods:identifier id="30BEE2C19142A1BE3F218C82289BE8B1" type="CLB-Dataset">9409</mods:identifier>
<mods:identifier id="5BA0690CCD61601C55C770ECCB804A4A" type="DOI">10.11646/zootaxa.4890.3.1</mods:identifier>
<mods:identifier id="86B16AA94835F8AB55B3AC8D5A004C6B" type="GBIF-Dataset">2565fffa-2704-4101-8fdf-2b863ef19be5</mods:identifier>
<mods:identifier id="BC7628BC8AFD5566DA785919303C1762" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D35F77EA2FFA6F4575017645233F8656" type="Zenodo-Dep">4306124</mods:identifier>
<mods:identifier id="29136F24968B2120E619B120BE538E50" type="ZooBank">4D7E2CD3-5350-479F-905A-C2671C414CBC</mods:identifier>
</mods:mods>
<treatment id="AE6287F45021FFE0A7FFFB6B7FA8F9C6" ID-DOI="http://doi.org/10.5281/zenodo.4332507" ID-GBIF-Taxon="170712168" ID-Zenodo-Dep="4332507" LSID="urn:lsid:plazi:treatment:AE6287F45021FFE0A7FFFB6B7FA8F9C6" httpUri="http://treatment.plazi.org/id/AE6287F45021FFE0A7FFFB6B7FA8F9C6" lastPageId="18" lastPageNumber="319" pageId="16" pageNumber="317">
<subSubSection id="6ED165695021FFE2A7FFFB6B7F1BFB69" box="[151,605,1217,1244]" pageId="16" pageNumber="317" type="nomenclature">
<paragraph id="267436E25021FFE2A7FFFB6B7F1BFB69" blockId="16.[151,605,1217,1280]" box="[151,605,1217,1244]" pageId="16" pageNumber="317">
<heading id="7D3C818E5021FFE2A7FFFB6B7F1BFB69" bold="true" box="[151,605,1217,1244]" fontSize="11" level="1" pageId="16" pageNumber="317" reason="1">
<emphasis id="14BFEAF05021FFE2A7FFFB6B7F1BFB69" bold="true" box="[151,605,1217,1244]" pageId="16" pageNumber="317">
<taxonomicName id="E1CB4D615021FFE2A7FFFB6B7F5CFB69" ID-CoL="8Y6P8" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[151,538,1217,1244]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="14BFEAF05021FFE2A7FFFB6B7F5CFB69" bold="true" box="[151,538,1217,1244]" italics="true" pageId="16" pageNumber="317">Nipponophloeostiba longilobata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0F8C578B5021FFE2A54AFB687F1BFB69" box="[546,605,1218,1244]" pageId="16" pageNumber="317" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6ED165695021FFE2A7FFFB4C7C68FAB5" box="[151,302,1254,1280]" pageId="16" pageNumber="317" type="description">
<paragraph id="267436E25021FFE2A7FFFB4C7C68FAB5" blockId="16.[151,605,1217,1280]" box="[151,302,1254,1280]" pageId="16" pageNumber="317">
(
<figureCitation id="BEF02A675021FFE2A7F7FB4C7C60FAB5" box="[159,294,1254,1280]" captionStart="FIGURES 39–42" captionStartId="17.[151,264,1853,1877]" captionTargetBox="[160,1425,424,1815]" captionTargetId="figure-146@17.[151,1436,417,1830]" captionTargetPageId="17" captionText="FIGURES 39–42.Aedeagus (Figs. 39–40) and abdominal segments (Figs. 41–42) of Nipponophloeostiba longilobata: 39—aedeagus, parameral view, 40—aedeagus, lateral view, 41—male tergite VIII (dorsal view), 42—male sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306150" httpUri="https://zenodo.org/record/4306150/files/figure.png" pageId="16" pageNumber="317">Figs. 39–42</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="6ED165695021FFE2A7FFFA84787EFA25" pageId="16" pageNumber="317" type="materials_examined">
<paragraph id="267436E25021FFE2A7FFFA84787EFA25" blockId="16.[151,1437,1325,2036]" pageId="16" pageNumber="317">
<emphasis id="14BFEAF05021FFE2A7FFFA847C85FAF2" bold="true" box="[151,451,1325,1352]" pageId="16" pageNumber="317">Type material examined:</emphasis>
<materialsCitation id="96A33CBF5021FFE2A6A2FA84787EFA25" ID-GBIF-Occurrence="2989989338" collectingDate="2009-05-30" collectionCode="V, NHMB" elevation="1420" latitude="19.636667" location="Phonsavan" longLatPrecision="127" longitude="103.33667" municipality="Khouang" pageId="16" pageNumber="317" specimenCount="1" typeStatus="holotype">
<typeStatus id="F97088405021FFE2A6A2FA847F74FAFD" box="[458,562,1326,1352]" pageId="16" pageNumber="317" type="holotype">Holotype</typeStatus>
Ƌ (dissected; right elytron slightly damaged): ‘LAOS—Xieng
<collectingMunicipality id="C610AC985021FFE2A396FA84782EFAFD" box="[1278,1384,1326,1352]" pageId="16" pageNumber="317">Khouang</collectingMunicipality>
, |
<locationDeviation id="37C62FC05021FFE2A2E8FA847DA1FAD9" bearing="45.0" distance="30000.0" pageId="16" pageNumber="317">
<quantity id="E1339B075021FFE2A2E8FA847DFDFAD9" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="317" unit="km" value="30.0">30 km</quantity>
NE
</locationDeviation>
<location id="231460395021FFE2A786FAF87C2CFAD9" LSID="urn:lsid:plazi:treatment:AE6287F45021FFE0A7FFFB6B7FA8F9C6:231460395021FFE2A786FAF87C2CFAD9" box="[238,362,1362,1388]" latitude="19.636667" longLatPrecision="127" longitude="103.33667" municipality="Khouang" name="Phonsavan" pageId="16" pageNumber="317">Phonsavan</location>
, |
<location id="231460395021FFE2A6EAFAF87F6DFAD9" LSID="urn:lsid:plazi:treatment:AE6287F45021FFE0A7FFFB6B7FA8F9C6:231460395021FFE2A6EAFAF87F6DFAD9" box="[386,555,1362,1388]" latitude="19.636667" longLatPrecision="127" longitude="103.33667" municipality="Khouang" name="Phou Sane Mt." pageId="16" pageNumber="317">Phou Sane Mt.</location>
,
<geoCoordinate id="43FF50255021FFE2A55EFAF87FE8FAD9" box="[566,686,1362,1388]" degrees="19" direction="north" minutes="38.2" orientation="latitude" pageId="16" pageNumber="317" precision="92" value="19.636667">19°38.2’N</geoCoordinate>
, |
<geoCoordinate id="43FF50255021FFE2A5AEFAF87E0CFAD9" box="[710,842,1362,1388]" degrees="103" direction="east" minutes="20.2" orientation="longitude" pageId="16" pageNumber="317" precision="92" value="103.33667">103°20.2’E</geoCoordinate>
,
<quantity id="E1339B075021FFE2A43DFAF87EEAFAD9" box="[853,940,1362,1388]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="16" pageNumber="317" unit="m" value="1420.0">
<elevation id="ADE6D1D15021FFE2A43DFAF87EEAFAD9" box="[853,940,1362,1388]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="16" pageNumber="317" unit="m" value="1420.0">1420 m</elevation>
</quantity>
, 10– |
<date id="527510225021FFE2A49EFAF8792DFAD9" box="[1014,1131,1362,1388]" pageId="16" pageNumber="317" value="2009-05-30">
<collectingDate id="4231E9CA5021FFE2A49EFAF8792DFAD9" box="[1014,1131,1362,1388]" pageId="16" pageNumber="317" value="2009-05-30">30.V.2009</collectingDate>
</date>
, leg. Brancucci’ &lt;printed&gt;, ‘
<typeStatus id="F97088405021FFE2A7F3FADC7C72FA25" box="[155,308,1398,1424]" pageId="16" pageNumber="317" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="E1CB4D615021FFE2A620FADC7F6DFA25" authorityName="Watanabe" authorityYear="1962" box="[328,555,1398,1424]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="14BFEAF05021FFE2A620FADC7F6DFA25" box="[328,555,1398,1424]" italics="true" pageId="16" pageNumber="317">
<location id="231460395021FFE2A620FADC7F6DFA25" LSID="urn:lsid:plazi:treatment:AE6287F45021FFE0A7FFFB6B7FA8F9C6:231460395021FFE2A620FADC7F6DFA25" box="[328,555,1398,1424]" latitude="19.636667" longLatPrecision="127" longitude="103.33667" municipality="Khouang" name="Nipponophloeostiba" pageId="16" pageNumber="317">Nipponophloeostiba</location>
</emphasis>
</taxonomicName>
|
<taxonomicName id="E1CB4D615021FFE2A557FADC7F86FA25" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[575,704,1398,1424]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="14BFEAF05021FFE2A557FADC7F86FA25" box="[575,704,1398,1424]" italics="true" pageId="16" pageNumber="317">longilobata</emphasis>
</taxonomicName>
<emphasis id="14BFEAF05021FFE2A5AFFADC7FB8FA25" bold="true" box="[711,766,1398,1424]" pageId="16" pageNumber="317">
<taxonomicNameLabel id="0F8C578B5021FFE2A5AFFADC7FB8FA25" box="[711,766,1398,1424]" pageId="16" pageNumber="317" rank="species">
<specimenCount id="30CDFD6B5021FFE2A5AFFADC7FAEFA25" box="[711,744,1398,1424]" pageId="16" pageNumber="317" type="generic">sp.</specimenCount>
n.
</taxonomicNameLabel>
</emphasis>
|
<location id="231460395021FFE2A47AFADC7ECFFA25" LSID="urn:lsid:plazi:treatment:AE6287F45021FFE0A7FFFB6B7FA8F9C6:231460395021FFE2A47AFADC7ECFFA25" box="[786,905,1398,1424]" latitude="19.636667" longLatPrecision="127" longitude="103.33667" municipality="Khouang" name="Shavrin A." pageId="16" pageNumber="317">Shavrin A.</location>
<collectionCode id="40DAAE275021FFE2A4E0FADC7ED8FA25" box="[904,926,1398,1424]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="317" type="Museum">V</collectionCode>
. des. 2020’ &lt;red, printed&gt; (
<collectionCode id="40DAAE275021FFE2A3BAFADC786AFA25" box="[1234,1324,1398,1424]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="16" pageNumber="317">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6ED165695021FFE0A7AFFA337E93FBA4" lastPageId="18" lastPageNumber="319" pageId="16" pageNumber="317" type="description">
<paragraph id="267436E25021FFE2A7AFFA33781DFA6D" blockId="16.[151,1437,1325,2036]" pageId="16" pageNumber="317">
<emphasis id="14BFEAF05021FFE2A7AFFA337C17FA06" bold="true" box="[199,337,1433,1459]" pageId="16" pageNumber="317">Description</emphasis>
. Measurements: HW: 0.38; HL: 0.25; AL: 0.40; OL: 0.09; TM: 0.07; PL: 0.32; PW: 0.40; ESL: 0.41; EW: 0.48; AW: 0.43; MTbL: 0.22; MTrL: 0.12 (MTrL 1–4: 0.05; MTrL 5: 0.07); AedL: 0.33; TL: 2.07.
</paragraph>
<paragraph id="267436E25021FFE2A7AFFA4879D1F8F5" blockId="16.[151,1437,1325,2036]" pageId="16" pageNumber="317">Head, antennomeres 6–11, pronotum, scutellum, parascutellar portion of elytra, and abdomen reddish-brown; elytra yellow-brown; mouthparts, antennomeres 1–5, intersegmental membranes between abdominal tergites, and legs yellow. Head with sparse and fine punctation, denser in middle portion; neck without visible punctures; pronotum with sparse, fine punctation, similar to that of head, sparser in middle, mediobasal portion without punctures; scutellum without punctation; elytra with fine punctation, denser than that on pronotum, deeper along suture, and indistinct in apical third; abdomen with very indistinct, sparse and fine punctures. Head with dense, relatively coarse microsculpture: transverse in apical and longitudinal in middle and lateral portions; neck with transverse meshes, with wide, impunctate middle portion; pronotum with dense, longitudinal microreticulation, coarser than that of head; elytra with longitudinal, regular sculpture, similar to that of pronotum, but slightly coarser; abdomen with distinct, dense, and transverse microsculpture, denser and coarser on abdominal tergite VII.</paragraph>
<paragraph id="267436E25021FFE3A7AFF8E07821FE3C" blockId="16.[151,1437,1325,2036]" lastBlockId="17.[151,1437,151,393]" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317">Head wide, 1.5 times as broad as long, with deep, moderately long, straight anteocellar foveae extending to level of apical third of eyes; middle portion between anteocellar foveae with distinct, transverse impression; basal margin of head straight; posterior portions of head between ocelli and temples with narrow, straight carina, almost reaching posterior margin of eye; postantennal impressions indistinct; temples (postocular parts) long, parallelsided, not expanded laterad, slightly shorter than longitudinal length of eyes. Eyes large, moderately convex. Ocelli small, located very close to posterior constriction, distance between ocelli about one and a half times as long as distance between ocellus and posterior margin of eye. Neck separated from head by narrow, deep constriction. Maxillary palpi long; preapical palpomere slightly longer than wide; apical palpomere narrow, from basal portion gradually narrowing apicad, about 1.4 times as long as preceding palpomere. Antennae reaching apical third of pronotum when reclined; basal antennomere wide, about 1.6 times as long as wide, antennomere 2 small, slightly longer than wide, 3 narrow, about as long as 2, 4–5 small, transverse, about as broad as 3, 6–10 strongly transverse, significantly wider than 5, 7–10 slightly wider than 6, apical antennomere about as broad as, and about twice as long as 10.</paragraph>
<caption id="72B4666A5020FFE3A7FFF8977C11F828" ID-DOI="http://doi.org/10.5281/zenodo.4306150" ID-Zenodo-Dep="4306150" httpUri="https://zenodo.org/record/4306150/files/figure.png" pageId="17" pageNumber="318" startId="17.[151,264,1853,1877]" targetBox="[160,1425,424,1815]" targetPageId="17">
<paragraph id="267436E25020FFE3A7FFF8977C11F828" blockId="17.[151,1437,1853,1950]" pageId="17" pageNumber="318">
<emphasis id="14BFEAF05020FFE3A7FFF8977C12F8E0" bold="true" box="[151,340,1853,1877]" pageId="17" pageNumber="318">FIGURES 39–42.</emphasis>
Aedeagus (Figs. 39–40) and abdominal segments (Figs. 41–42) of
<taxonomicName id="E1CB4D615020FFE3A49EF8977878F8E0" authorityName="Shavrin" authorityYear="2020" box="[1014,1342,1853,1877]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="longilobata">
<emphasis id="14BFEAF05020FFE3A49EF8977878F8E0" box="[1014,1342,1853,1877]" italics="true" pageId="17" pageNumber="318">Nipponophloeostiba longilobata</emphasis>
</taxonomicName>
: 39—aedeagus, parameral view, 40—aedeagus, lateral view, 41—male tergite VIII (dorsal view), 42—male sternite VIII (ventral view). Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="267436E25023FFE0A7AFFF3D7913FEF4" blockId="18.[151,1437,151,1041]" pageId="18" pageNumber="319">Pronotum subtrapezoidal, slightly wider than long and wider than head, widest in apical third, from about middle strongly narrowed posteriad; latero-basal margins slightly concave in front of rounded posterior angles; anterior angles widely rounded; anterior margin slightly concave in middle, with two very small, convex tubercles at level of lateral margins of neck; posterior margin rounded; median part with longitudinal narrow band; lateral portions narrowly bordered and flattened at margins, laterobasal portions distinctly impressed.</paragraph>
<paragraph id="267436E25023FFE0A7AFFEE17840FE3C" blockId="18.[151,1437,151,1041]" pageId="18" pageNumber="319">Elytra moderately short, slightly wider than long, 1.2 times as long as pronotum, relatively parallel-sided, with slightly concave lateral margins in middle; posterior margin slightly rounded. Wings fully developed.</paragraph>
<paragraph id="267436E25023FFE0A7AFFE397C5EFE64" blockId="18.[151,1437,151,1041]" pageId="18" pageNumber="319">Metatarsus 1.8 times as long as metatibia; apical metatarsomere distinctly longer than remaining four combined tarsomeres.</paragraph>
<paragraph id="267436E25023FFE0A7AFFE7179F1FDAC" blockId="18.[151,1437,151,1041]" pageId="18" pageNumber="319">Abdomen parallel-sided, narrower than combined width of elytra, with two small, round wing-folding patches in middle of abdominal tergite IV and indistinct palisade fringe at apical margin of tergite VII.</paragraph>
<paragraph id="267436E25023FFE0A7AFFD897C6FFD44" blockId="18.[151,1437,151,1041]" pageId="18" pageNumber="319">
Male. Posterior margins of abdominal tergite VIII (
<figureCitation id="BEF02A675023FFE0A474FD897E37FD88" box="[796,881,547,573]" captionStart="FIGURES 39–42" captionStartId="17.[151,264,1853,1877]" captionTargetBox="[160,1425,424,1815]" captionTargetId="figure-146@17.[151,1436,417,1830]" captionTargetPageId="17" captionText="FIGURES 39–42.Aedeagus (Figs. 39–40) and abdominal segments (Figs. 41–42) of Nipponophloeostiba longilobata: 39—aedeagus, parameral view, 40—aedeagus, lateral view, 41—male tergite VIII (dorsal view), 42—male sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306150" httpUri="https://zenodo.org/record/4306150/files/figure.png" pageId="18" pageNumber="319">Fig. 41</figureCitation>
) and sternite VIII (
<figureCitation id="BEF02A675023FFE0A33EFD8979EDFD88" box="[1110,1195,547,573]" captionStart="FIGURES 39–42" captionStartId="17.[151,264,1853,1877]" captionTargetBox="[160,1425,424,1815]" captionTargetId="figure-146@17.[151,1436,417,1830]" captionTargetPageId="17" captionText="FIGURES 39–42.Aedeagus (Figs. 39–40) and abdominal segments (Figs. 41–42) of Nipponophloeostiba longilobata: 39—aedeagus, parameral view, 40—aedeagus, lateral view, 41—male tergite VIII (dorsal view), 42—male sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306150" httpUri="https://zenodo.org/record/4306150/files/figure.png" pageId="18" pageNumber="319">Fig. 42</figureCitation>
) indistinctly sinuate. Aedeagus long, with wide basal portion, gradually narrowed apically, and with moderately narrow, long median lobe (
<figureCitation id="BEF02A675023FFE0A7BFFDC17C6DFD30" box="[215,299,619,645]" captionStart="FIGURES 39–42" captionStartId="17.[151,264,1853,1877]" captionTargetBox="[160,1425,424,1815]" captionTargetId="figure-146@17.[151,1436,417,1830]" captionTargetPageId="17" captionText="FIGURES 39–42.Aedeagus (Figs. 39–40) and abdominal segments (Figs. 41–42) of Nipponophloeostiba longilobata: 39—aedeagus, parameral view, 40—aedeagus, lateral view, 41—male tergite VIII (dorsal view), 42—male sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306150" httpUri="https://zenodo.org/record/4306150/files/figure.png" pageId="18" pageNumber="319">Fig. 39</figureCitation>
); parameres wide, distinctly extending beyond apex of median lobe, distinctly widened toward apical portions, with five long apical, two preapical, four setae along inner edge, and several very small setae on middle surface of each paramere; internal sac with two long structures consisting of fine, small spines. Aedeagus laterally as in
<figureCitation id="BEF02A675023FFE0A7BAFD7D7C63FD44" box="[210,293,727,753]" captionStart="FIGURES 39–42" captionStartId="17.[151,264,1853,1877]" captionTargetBox="[160,1425,424,1815]" captionTargetId="figure-146@17.[151,1436,417,1830]" captionTargetPageId="17" captionText="FIGURES 39–42.Aedeagus (Figs. 39–40) and abdominal segments (Figs. 41–42) of Nipponophloeostiba longilobata: 39—aedeagus, parameral view, 40—aedeagus, lateral view, 41—male tergite VIII (dorsal view), 42—male sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306150" httpUri="https://zenodo.org/record/4306150/files/figure.png" pageId="18" pageNumber="319">Fig. 40</figureCitation>
.
</paragraph>
<paragraph id="267436E25023FFE0A7AFFD517CD7FCA0" blockId="18.[151,1437,151,1041]" box="[199,401,763,789]" pageId="18" pageNumber="319">Female unknown.</paragraph>
<paragraph id="267436E25023FFE0A7AFFCB47E93FBA4" blockId="18.[151,1437,151,1041]" pageId="18" pageNumber="319">
<emphasis id="14BFEAF05023FFE0A7AFFCB47CEEFC8C" bold="true" box="[199,424,798,825]" pageId="18" pageNumber="319">Comparative notes</emphasis>
. Regarding t body size and the laterally not widened postocular parts of head,
<taxonomicName id="E1CB4D615023FFE0A260FCB57DEBFCE9" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="14BFEAF05023FFE0A260FCB5785DFC8D" box="[1288,1307,799,824]" italics="true" pageId="18" pageNumber="319">N</emphasis>
.
<emphasis id="14BFEAF05023FFE0A240FCB57DEBFCE9" italics="true" pageId="18" pageNumber="319">longilobata</emphasis>
</taxonomicName>
<emphasis id="14BFEAF05023FFE0A7DCFCE97DADFCE8" bold="true" box="[180,235,835,861]" pageId="18" pageNumber="319">
<taxonomicNameLabel id="0F8C578B5023FFE0A7DCFCE97DADFCE8" box="[180,235,835,861]" pageId="18" pageNumber="319" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="E1CB4D615023FFE0A616FCE97FD3FCE8" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[382,661,835,861]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="thayeri">
<emphasis id="14BFEAF05023FFE0A616FCE97CD7FCE9" box="[382,401,835,860]" italics="true" pageId="18" pageNumber="319">N</emphasis>
.
<emphasis id="14BFEAF05023FFE0A6F6FCE97CABFCE8" box="[414,493,835,861]" italics="true" pageId="18" pageNumber="319">thayeri</emphasis>
<bibRefCitation id="425A4B135023FFE0A69DFCE97FD3FCE8" author="Shavrin, A. V." box="[501,661,835,861]" pageId="18" pageNumber="319" pagination="196 - 200" refId="ref17748" refString="Shavrin, A. V. (2020) Nipponophloeostiba thayeri sp. n., a new species from Sabah, Malaysia (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4802 (1), 196 - 200. https: // doi. org / 10.11646 / zootaxa. 4802.1.14" type="journal article" year="2020">Shavrin, 2020</bibRefCitation>
</taxonomicName>
, a species recently described from
<collectingRegion id="E40FF8005023FFE0A373FCE97924FCE8" box="[1051,1122,835,861]" country="Malaysia" name="Sabah" pageId="18" pageNumber="319">Sabah</collectingRegion>
, Borneo,
<collectingCountry id="5EDC76725023FFE0A3A6FCE97873FCE8" box="[1230,1333,835,861]" name="Malaysia" pageId="18" pageNumber="319">Malaysia</collectingCountry>
(
<bibRefCitation id="425A4B135023FFE0A22BFCE97D92FC34" author="Shavrin, A. V." pageId="18" pageNumber="319" pagination="196 - 200" refId="ref17748" refString="Shavrin, A. V. (2020) Nipponophloeostiba thayeri sp. n., a new species from Sabah, Malaysia (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4802 (1), 196 - 200. https: // doi. org / 10.11646 / zootaxa. 4802.1.14" type="journal article" year="2020">Shavrin 2020</bibRefCitation>
). It differs from this species by paler coloration, slightly less prominent eyes, shorter elytra, a wider median lobe of the aedeagus and parameres, and the internal structures of the aedeagus. The shape of the median lobe is somewhat similar to
<taxonomicName id="E1CB4D615023FFE0A6E4FC057FB6FC7C" authority="Watanabe, 1962" authorityName="Watanabe" authorityYear="1962" box="[396,752,943,969]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="verrucifera">
<emphasis id="14BFEAF05023FFE0A6E4FC057CD9FC7D" box="[396,415,943,968]" italics="true" pageId="18" pageNumber="319">N</emphasis>
.
<emphasis id="14BFEAF05023FFE0A6D8FC057F68FC7C" box="[432,558,943,969]" italics="true" pageId="18" pageNumber="319">verrucifera</emphasis>
Watanabe, 1962
</taxonomicName>
from Izu islands and Yaku-shima Island,
<collectingCountry id="5EDC76725023FFE0A3B3FC05785DFC7C" box="[1243,1307,943,969]" name="Japan" pageId="18" pageNumber="319">Japan</collectingCountry>
(
<bibRefCitation id="425A4B135023FFE0A244FC057DA6FC58" author="Watanabe, Y." pageId="18" pageNumber="319" pagination="77 - 80" refId="ref18369" refString="Watanabe, Y. (1962 b) Descriptions of a new genus and species of Omaliinae from the Izu islands, Japan (Coleoptera: Staphylinidae). Journal of Agricultural Science, Tokyo Nogyo Daigaku, 8 (2), 77 - 80." type="journal article" year="1962">Watanabe 1962b</bibRefCitation>
,
<bibRefCitation id="425A4B135023FFE0A783FC797C61FC58" author="Watanabe, Y." box="[235,295,979,1005]" pageId="18" pageNumber="319" pagination="59 - 391" refId="ref18416" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">1990</bibRefCitation>
), but the parameres of
<taxonomicName id="E1CB4D615023FFE0A54BFC797F82FC58" authority="Shavrin, 2020" authorityName="Shavrin" authorityYear="2020" box="[547,708,979,1005]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="longilobata" status="sp. nov.">
<emphasis id="14BFEAF05023FFE0A54BFC797F70FC59" box="[547,566,979,1004]" italics="true" pageId="18" pageNumber="319">N</emphasis>
.
<emphasis id="14BFEAF05023FFE0A52BFC797F82FC58" box="[579,708,979,1005]" italics="true" pageId="18" pageNumber="319">longilobata</emphasis>
</taxonomicName>
<emphasis id="14BFEAF05023FFE0A5A2FC797E44FC58" bold="true" box="[714,770,979,1005]" pageId="18" pageNumber="319">
<taxonomicNameLabel id="0F8C578B5023FFE0A5A2FC797E44FC58" box="[714,770,979,1005]" pageId="18" pageNumber="319" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
are wider, with longer and more numerous apical setae. For additional details see the key to the species of
<taxonomicName id="E1CB4D615023FFE0A5F6FC5D7EC7FBA4" authorityName="Watanabe" authorityYear="1962" box="[670,897,1015,1041]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="14BFEAF05023FFE0A5F6FC5D7EC7FBA4" box="[670,897,1015,1041]" italics="true" pageId="18" pageNumber="319">Nipponophloeostiba</emphasis>
</taxonomicName>
below:
</paragraph>
</subSubSection>
<subSubSection id="6ED165695023FFE0A7FFFB9678DBFAC3" pageId="18" pageNumber="319" type="key">
<key id="525A61D35023FFE0A7FFFB9678DBFAC3" pageId="18" pageNumber="319">
<keyStep id="9D3F2E475023FFE0A7FFFB9678DAFB73" pageId="18" pageNumber="319">
<paragraph id="267436E25023FFE0A7FFFB9678DAFB3E" blockId="18.[151,1437,1084,1399]" pageId="18" pageNumber="319">
<keyLead id="9D3A95D75023FFE0A7FFFB9678DAFB3E" pageId="18" pageNumber="319">
1 Postocular portions of head distinctly widened lateroposteriad, with short posterior carina not reaching posterior margin of eye. Aedeagus as illustrated by
<bibRefCitation id="425A4B135023FFE0A680FBF37FCEFBDB" author="Watanabe, Y." box="[488,648,1113,1135]" pageId="18" pageNumber="319" pagination="59 - 391" refId="ref18416" refString="Watanabe, Y. (1990) A taxonomic study on the subfamily Omaliinae from Japan (Coleoptera, Staphylinidae). Memoirs of the Tokyo University of Agriculture, 31, 59 - 391." type="journal article" year="1990">Watanabe (1990</bibRefCitation>
: figures 86–87). Head and pronotum reddish-brown, elytra and abdomen yellowbrown. Body larger:
<quantity id="E1339B075023FFE0A6C1FBDC7F71FB39" box="[425,567,1142,1164]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="18" pageNumber="319" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.30–2.70 mm</quantity>
.
<collectingCountry id="5EDC76725023FFE0A529FBDD7F31FB39" box="[577,631,1143,1164]" name="Japan" pageId="18" pageNumber="319">Japan</collectingCountry>
........................................................
<taxonomicName id="E1CB4D615023FFE0A27CFBDD78DAFB3E" ID-CoL="47HB5" authorityName="Watanabe" authorityYear="1962" box="[1300,1436,1142,1164]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="verrucifera">
<emphasis id="14BFEAF05023FFE0A27CFBDD78DAFB3E" box="[1300,1436,1142,1164]" italics="true" pageId="18" pageNumber="319">N. verrucifera</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="267436E25023FFE0A7FFFB3E78DAFB73" blockId="18.[151,1437,1084,1399]" pageId="18" pageNumber="319">
<keyLead id="9D3A95D75023FFE0A7FFFB3E78DAFB73" pageId="18" pageNumber="319">- Postocular portions of head not widened lateroposteriad, with long posterior carina reaching posterior margin of eye. Coloration different. Body smaller................................................................................. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="9D3F2E475023FFE0A7FFFB6478DBFAC3" pageId="18" pageNumber="319">
<paragraph id="267436E25023FFE0A7FFFB6478DAFAAB" blockId="18.[151,1437,1084,1399]" pageId="18" pageNumber="319">
<keyLead id="9D3A95D75023FFE0A7FFFB6478DAFAAB" pageId="18" pageNumber="319">
2 Head as broad as pronotum. Elytra moderately long, significantly longer than pronotum. Median lobe and parameres narrow (
<bibRefCitation id="425A4B135023FFE0A780FB467C29FAB4" author="Shavrin, A. V." box="[232,367,1260,1282]" pageId="18" pageNumber="319" pagination="196 - 200" refId="ref17748" refString="Shavrin, A. V. (2020) Nipponophloeostiba thayeri sp. n., a new species from Sabah, Malaysia (Coleoptera: Staphylinidae: Omaliinae: Omaliini). Zootaxa, 4802 (1), 196 - 200. https: // doi. org / 10.11646 / zootaxa. 4802.1.14" type="journal article" year="2020">Shavrin 2020</bibRefCitation>
: figure 2). Body darker: head and abdomen brown, pronotum reddish-brown. Body length:
<quantity id="E1339B075023FFE0A3B5FB41782DFAB4" box="[1245,1387,1259,1281]" metricMagnitude="-3" metricUnit="m" metricValue="1.985" metricValueMax="2.22" metricValueMin="1.75" pageId="18" pageNumber="319" unit="mm" value="1.985" valueMax="2.22" valueMin="1.75">1.75–2.22 mm</quantity>
.
<collectingCountry id="5EDC76725023FFE0A21CFB467C5CFAAA" name="Malaysia" pageId="18" pageNumber="319">Malaysia</collectingCountry>
........................................................................................
<taxonomicName id="E1CB4D615023FFE0A254FAA078DAFAAB" ID-CoL="8Y6P9" authorityName="Shavrin" authorityYear="2020" box="[1340,1436,1289,1311]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="thayeri">
<emphasis id="14BFEAF05023FFE0A254FAA078DAFAAB" box="[1340,1436,1289,1311]" italics="true" pageId="18" pageNumber="319">N. thayeri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="267436E25023FFE0A7FFFA8D78DBFAC3" blockId="18.[151,1437,1084,1399]" pageId="18" pageNumber="319">
<keyLead id="9D3A95D75023FFE0A7FFFA8D78DBFAC3" pageId="18" pageNumber="319">
- Head slightly narrower than pronotum. Elytra short, slightly longer than pronotum. Median lobe and parameres moderately wide (
<figureCitation id="BEF02A675023FFE0A676FAEE7C21FAEF" box="[286,359,1348,1370]" captionStart="FIGURES 39–42" captionStartId="17.[151,264,1853,1877]" captionTargetBox="[160,1425,424,1815]" captionTargetId="figure-146@17.[151,1436,417,1830]" captionTargetPageId="17" captionText="FIGURES 39–42.Aedeagus (Figs. 39–40) and abdominal segments (Figs. 41–42) of Nipponophloeostiba longilobata: 39—aedeagus, parameral view, 40—aedeagus, lateral view, 41—male tergite VIII (dorsal view), 42—male sternite VIII (ventral view). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4306150" httpUri="https://zenodo.org/record/4306150/files/figure.png" pageId="18" pageNumber="319">Fig. 39</figureCitation>
). Body paler: head, pronotum and abdomen reddish-brown, elytra yellow-brown, with darker parascutellar portion. Body length:
<quantity id="E1339B075023FFE0A6FEFACB7CA8FAC3" box="[406,494,1377,1398]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="18" pageNumber="319" unit="mm" value="2.07">2.07 mm</quantity>
.
<collectingCountry id="5EDC76725023FFE0A69FFACB7F60FAC3" box="[503,550,1377,1398]" name="Laos" pageId="18" pageNumber="319">Laos</collectingCountry>
..............................................................
<taxonomicName id="E1CB4D615023FFE0A27AFAC878DBFAC3" ID-CoL="8Y6P8" authorityName="Shavrin" authorityYear="2020" box="[1298,1437,1377,1399]" class="Insecta" family="Staphylinidae" genus="Nipponophloeostiba" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="longilobata">
<emphasis id="14BFEAF05023FFE0A27AFAC878DBFAC3" box="[1298,1437,1377,1399]" italics="true" pageId="18" pageNumber="319">N. longilobata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="6ED165695023FFE0A7AFFA0F7DAFF99E" pageId="18" pageNumber="319" type="distribution">
<paragraph id="267436E25023FFE0A7AFFA0F7D93FA51" blockId="18.[151,1436,1445,1652]" pageId="18" pageNumber="319">
<emphasis id="14BFEAF05023FFE0A7AFFA0F7C27FA0A" bold="true" box="[199,353,1445,1471]" pageId="18" pageNumber="319">Distribution.</emphasis>
The new species is known only from the
<typeStatus id="F97088405023FFE0A42DFA0C7E33FA75" box="[837,885,1446,1472]" pageId="18" pageNumber="319">type</typeStatus>
locality in Phou Sane mountain range, northern
<collectingCountry id="5EDC76725023FFE0A7FFFA607D97FA51" box="[151,209,1482,1508]" name="Laos" pageId="18" pageNumber="319">Laos</collectingCountry>
.
</paragraph>
<paragraph id="267436E25023FFE0A7AFFA477DAFF99E" blockId="18.[151,1436,1445,1652]" pageId="18" pageNumber="319">
<emphasis id="14BFEAF05023FFE0A7AFFA477C0DF9B2" bold="true" box="[199,331,1517,1543]" pageId="18" pageNumber="319">Bionomics.</emphasis>
The specimen was collected on the wing, most likely with a Malaise trap, at an elevation of
<quantity id="E1339B075023FFE0A20CFA477DEBF999" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="18" pageNumber="319" unit="m" value="1420.0">1420 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="6ED165695023FFE0A7AFF99F7FA8F9C6" pageId="18" pageNumber="319" type="etymology">
<paragraph id="267436E25023FFE0A7AFF99F7FA8F9C6" blockId="18.[151,1436,1445,1652]" pageId="18" pageNumber="319">
<emphasis id="14BFEAF05023FFE0A7AFF99F7C08F9FA" bold="true" box="[199,334,1589,1615]" pageId="18" pageNumber="319">Etymology.</emphasis>
The specific epithet is a combination of Latin adjective
<emphasis id="14BFEAF05023FFE0A4AAF99F794BF9FA" box="[962,1037,1589,1615]" italics="true" pageId="18" pageNumber="319">longus</emphasis>
, -
<emphasis id="14BFEAF05023FFE0A34BF99C7977F9FA" box="[1059,1073,1590,1615]" italics="true" pageId="18" pageNumber="319">a</emphasis>
, -
<emphasis id="14BFEAF05023FFE0A320F99C792CF9FA" box="[1096,1130,1590,1615]" italics="true" pageId="18" pageNumber="319">um</emphasis>
(long) and
<emphasis id="14BFEAF05023FFE0A383F99F787BF9FA" box="[1259,1341,1589,1615]" italics="true" pageId="18" pageNumber="319">lobatus</emphasis>
, -
<emphasis id="14BFEAF05023FFE0A23DF99C7825F9FA" box="[1365,1379,1590,1615]" italics="true" pageId="18" pageNumber="319">a</emphasis>
, -
<emphasis id="14BFEAF05023FFE0A212F99C78DAF9FA" box="[1402,1436,1590,1615]" italics="true" pageId="18" pageNumber="319">um</emphasis>
(lobed). It refers to elongate and narrow median lobe.
</paragraph>
</subSubSection>
</treatment>
</document>
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