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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1164.103184" ID-Pensoft-Pub="1313-2970-1164-23" ID-Pensoft-UUID="62BD8FF233655E5288FA80F56FEA3299" ID-ZooBank="CC06E1473B074F478AFB08520503A404" ModsDocID="1313-2970-1164-23" checkinTime="1685200798416" checkinUser="pensoft" docAuthor="Shepard, William D. &amp; Barr, Cheryl B." docDate="2023" docId="18DD8E0E120352A1904BDA2807D155CD" docLanguage="en" docName="ZooKeys 1164: 23-61" docOrigin="ZooKeys 1164" docPubDate="2023-05-26" docSource="http://dx.doi.org/10.3897/zookeys.1164.103184" docTitle="Tychepsephus felix Waterhouse 1876" docType="treatment" docVersion="1" id="62BD8FF233655E5288FA80F56FEA3299" lastPageNumber="23" masterDocId="62BD8FF233655E5288FA80F56FEA3299" masterDocTitle="A revision of the Chilean water penny genus Tychepsephus Waterhouse, 1876 (Coleoptera, Psephenidae, Eubriinae), with description of a second species and two larval morphotypes, and notes on other Chilean Psephenidae" masterLastPageNumber="61" masterPageNumber="23" pageNumber="23" updateTime="1685200798416" updateUser="pensoft">
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<mods:title id="910A16FA36990B64D732269E3B5848F4">A revision of the Chilean water penny genus Tychepsephus Waterhouse, 1876 (Coleoptera, Psephenidae, Eubriinae), with description of a second species and two larval morphotypes, and notes on other Chilean Psephenidae</mods:title>
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<mods:namePart id="8BFDB499BB7838D36046A0963BD30AC4">Shepard, William D.</mods:namePart>
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<mods:affiliation id="03FCD82A9D377ED6D8C480B4D8424A9E">Essig Museum of Entomology, University of California, 1101 Valley Life Sciences Bldg., Berkeley, CA 94720, USA</mods:affiliation>
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<mods:namePart id="345D5F4F9CEC5E082F1AD43C6C2E05DB">Barr, Cheryl B.</mods:namePart>
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<mods:affiliation id="295B3AFC8CA015F3F75D0967A2165FEB">Essig Museum of Entomology, University of California, 1101 Valley Life Sciences Bldg., Berkeley, CA 94720, USA</mods:affiliation>
<mods:nameIdentifier id="8E3E407A231396155F46902B9C7BCB0F" type="email">cbarr@berkeley.edu</mods:nameIdentifier>
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<mods:date id="B9F5825C4F44E846B987102D20C73504">2023</mods:date>
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<mods:number id="36CA806962D0E19C3EEF21BBBBA7DE25">1164</mods:number>
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<treatment id="18DD8E0E120352A1904BDA2807D155CD" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD" httpUri="http://treatment.plazi.org/id/18DD8E0E120352A1904BDA2807D155CD" lastPageNumber="23" pageId="0" pageNumber="23">
<subSubSection id="4BFEFD0878C7ABF1DFD63DF5244CA60D" pageId="0" pageNumber="23" type="nomenclature">
<paragraph id="A2B9F99A3A011410B0739A2A54EE66F6" pageId="0" pageNumber="23">
<taxonomicName id="DB9D07CEB7B1894A95521A5F54641E27" LSID="18DD8E0E-1203-52A1-904B-DA2807D155CD" authority="Waterhouse, 1876" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">Tychepsephus felix Waterhouse, 1876</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="43C6BC4305D3F60DB14AA9A148757E88" pageId="0" pageNumber="23" type="description">
<paragraph id="B11D40302F5C439A1FD0B73534E5B394" pageId="0" pageNumber="23">
<figureCitation id="8F40758A104050848118EEF3A2B99E9C" captionStart="Figure 7" captionStartId="F4" captionText="Figure 7. Map of Chile with enlarged section showing the geographic distribution of species-verified adult Tychepsephus specimens: T. cekalovici, yellow triangles; T. felix, violet squares; both species, red circles." figureDoi="10.3897/zookeys.1164.103184.figure7" httpUri="https://binary.pensoft.net/fig/855188" pageId="0" pageNumber="23">Figs 7</figureCitation>
<figureCitation id="EEE3853FA59CD22FF798C706AD6F90D1" captionStart="Figures 8–11" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">, 8</figureCitation>
<figureCitation id="E74BAD43845479B44A5ED7DA2EED9E76" captionStart="Figure 12" captionStartId="F6" captionText="Figure 12. Tychepsephus felix, female type specimen a dorsal view b ventral view c specimen data labels. Images provided by Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1164.103184.figure12" httpUri="https://binary.pensoft.net/fig/855190" pageId="0" pageNumber="23">, 12</figureCitation>
<figureCitation id="48E21334576456C8E95ECE9B903FAD32" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">, 13, 14</figureCitation>
<figureCitation id="43C7B7779F609514A52049BF28458531" captionStart="Figures 15, 16" captionStartId="F8" captionText="Figures 15, 16. Tychepsephus felix, female 15 habitus a dorsal view b ventral view; length 4.2 mm 16 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23">, 15, 16</figureCitation>
<figureCitation id="862298E6F0CB2EF2E3C9D9F71877670C" captionStart="Figure 17" captionStartId="F9" captionText="Figure 17. Chilectopria grandis, female type specimen a dorsal view b lateral view c ventral view d specimen data labels. Images provided by Christophe Rivier (MNHN)." figureDoi="10.3897/zookeys.1164.103184.figure17" httpUri="https://binary.pensoft.net/fig/855193" pageId="0" pageNumber="23">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A3C7BE47CD03110FE1149990D2B2F22A" pageId="0" pageNumber="23" type="reference_group">
<paragraph id="3272B5BA0138C160A580653429B5F97F" pageId="0" pageNumber="23">
<taxonomicName id="79F919B8A541CCECD956743BA021EA04" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">Tychepsephus felix</taxonomicName>
Waterhouse, 1876: 16 (original description).
<bibRefCitation id="DAED3CD0D927C40C9A7C78A374D907FF" author="Philippi, F" journalOrPublisher="4 th edn. Academic Press, Salt Lake City" pageId="0" pageNumber="23" refId="B17" refString="Philippi, F, 1887. Catalogo de los Coleopteros de Chile. Anales de la Universidad de Chile 71(1): 79-674, 675-806." title="Catalogo de los Coleopteros de Chile. Anales de la Universidad de Chile 71 (1): 79 - 674, 675 - 806." year="1887">Philippi (1887</bibRefCitation>
: 665, catalog);
<bibRefCitation id="6C62BF8996499DA4CEEFC0757A62D16B" DOI="https://doi.org/10.5479/si.03629236.185.2" author="Blackwelder, RE" journalOrPublisher="Coleopterists Bulletin" pageId="0" pageNumber="23" refId="B4" refString="Blackwelder, RE, 1944. Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum 185. United States Government Printing Office, Washington, D. C., 189-341. https://doi.org/10.5479/si.03629236.185.2" title="Checklist of the Coleopterous Insects of Mexico, Central America, the West Indies, and South America. Part 2. Bulletin of the United States National Museum 185. United States Government Printing Office, Washington, D. C., 189 - 341." url="https://doi.org/10.5479/si.03629236.185.2" year="1944">Blackwelder (1944</bibRefCitation>
: 274, catalog);
<bibRefCitation id="429AB81A129B1F228134D5C8B56B3478" author="Artigas, J" journalOrPublisher="Gayana (Concepcion)" pageId="0" pageNumber="23" pagination="3 - 11" refId="B1" refString="Artigas, J, 1963. Descripcion de la larva de un psephenido Chileno (Coleoptera - Psephenidae). Gayana (Concepcion) 1963 (8): 3 - 11" title="Descripcion de la larva de un psephenido Chileno (Coleoptera - Psephenidae)." volume="1963" year="1963">Artigas (1963</bibRefCitation>
: 8, larval description);
<bibRefCitation id="FEFF43AEA7D6F428CB96B015E2C3AF17" author="Moroni, BJC" journalOrPublisher="Sociedad Chilena de Entomologia" pageId="0" pageNumber="23" pagination="169 - 175" refId="B15" refString="Moroni, BJC, 1985. Addenda y corrigenda al elenco sistematico, sinonimico y distribucion de Coleopteros acuaticos Chilenos. Sociedad Chilena de Entomologia 12: 169 - 175" title="Addenda y corrigenda al elenco sistematico, sinonimico y distribucion de Coleopteros acuaticos Chilenos." volume="12" year="1985">Moroni (1985</bibRefCitation>
: 173, taxonomy, distribution);
<bibRefCitation id="943275EE4174D5C9CC45DEF55E75A1F6" DOI="https://doi.org/10.1093/aesa/80.6.865" author="Ashworth, AC" journalOrPublisher="Annals of the Entomological Society of America" pageId="0" pageNumber="23" pagination="865 - 895" refId="B2" refString="Ashworth, AC, Hoganson, JW, 1987. Coleoptera bioassociations along an elevational gradient in the Lake Region of southern Chile, and comments on the postglacial development of the fauna. Annals of the Entomological Society of America 80 (6): 865 - 895, DOI: https://doi.org/10.1093/aesa/80.6.865" title="Coleoptera bioassociations along an elevational gradient in the Lake Region of southern Chile, and comments on the postglacial development of the fauna." url="https://doi.org/10.1093/aesa/80.6.865" volume="80" year="1987">Ashworth and Hoganson (1987</bibRefCitation>
: 879, habitat, distribution);
<bibRefCitation id="503A6FCCF764D70CA95672A69103BE49" author="Elgueta, DM" journalOrPublisher="Revista Chilena de Entomologia" pageId="0" pageNumber="23" pagination="5 - 60" refId="B7" refString="Elgueta, DM, Arriagada, SG, 1989. Estado actual del conocimiento de los Coleopteros de Chile (Insects: Coleoptera). Revista Chilena de Entomologia 17: 5 - 60" title="Estado actual del conocimiento de los Coleopteros de Chile (Insects: Coleoptera)." volume="17" year="1989">Elgueta and Arriagada (1989</bibRefCitation>
: 16, literature review);
<bibRefCitation id="2CEEADF9F9B208F4C4F83DCE0B60B681" DOI="https://doi.org/10.4067/S0717-65382006000100012" author="Jerez, V" journalOrPublisher="Gayana (Concepcion)" pageId="0" pageNumber="23" pagination="72 - 81" refId="B10" refString="Jerez, V, Moroni, J, 2006. Diversidad de Coleopteros acuaticos en Chile. Gayana (Concepcion) 70 (1): 72 - 81, DOI: https://doi.org/10.4067/S0717-65382006000100012" title="Diversidad de Coleopteros acuaticos en Chile." url="https://doi.org/10.4067/S0717-65382006000100012" volume="70" year="2006">Jerez and Moroni (2006</bibRefCitation>
: 76, taxonomy, checklist);
<bibRefCitation id="489A2F09ED517E3AF0EBC730F40C61B2" DOI="https://doi.org/10.1111/j.1365-3113.2006.00374.x" author="Lee, C-F" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="23" pagination="502 - 538" refId="B13" refString="Lee, C-F, Sato, M, Shepard, WD, Jaech, MA, 2007. Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters. Systematic Entomology 32 (3): 502 - 538, DOI: https://doi.org/10.1111/j.1365-3113.2006.00374.x" title="Phylogeny of Psephenidae (Coleoptera: Byrrhoidea) based on larval, pupal and adult characters." url="https://doi.org/10.1111/j.1365-3113.2006.00374.x" volume="32" year="2007">Lee et al. (2007</bibRefCitation>
: 527, phylogenetics),
<bibRefCitation id="F4FA4894E091DFFA8089AD3C91F8FE9A" DOI="https://doi.org/10.5772/intechopen.89077" author="Zarges, CV" editor="Devlin, A" journalOrPublisher="IntechOpen, London" pageId="0" pageNumber="23" pagination="1 - 30" refId="B27" refString="Zarges, CV, Fierro, P, Olmos, V, 2019. Freshwater invertebrates of southwestern South America: Diversity, biogeography, and threats. In: Devlin, A, Pan, J, Shah, MM, Eds., Inland Waters - Dynamics and Ecology. IntechOpen, London: 1 - 30, DOI: https://doi.org/10.5772/intechopen.89077" title="Freshwater invertebrates of southwestern South America: Diversity, biogeography, and threats." url="https://doi.org/10.5772/intechopen.89077" volumeTitle="Inland Waters - Dynamics and Ecology." year="2019">Zarges et al. (2019</bibRefCitation>
: 16, habitat).
</paragraph>
<paragraph id="9F18E1D78C02D601F0FCD09CF4082883" pageId="0" pageNumber="23">
<taxonomicName id="EEAD51ABAAC67E8C7F68562CEE605D8A" class="Insecta" family="Psephenidae" genus="Ectopria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectopria (Chilectopria) grandis" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="grandis" subGenus="Chilectopria">Ectopria (Chilectopria) grandis</taxonomicName>
Pic, 1947: 4, syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="523C238B01C8FFCE2DCB56B5171176EB" pageId="0" pageNumber="23" type="type locality">
<paragraph id="F09C0E638EC471F0ABB0043058BD8633" pageId="0" pageNumber="23">Type locality.</paragraph>
<paragraph id="17E396D0BF714BBBD64AD4EF180D3CBF" pageId="0" pageNumber="23">
The type locality was listed as only
<normalizedToken id="0096F2C7E5D2769E69478304FBD522FA" originalValue="“Chili”">&quot;Chili&quot;</normalizedToken>
on both the type specimen and in the species description. The female holotype specimen is housed in the NHMUK.
</paragraph>
</subSubSection>
<subSubSection id="3B4C228749395D81CADA766E4544C7A7" pageId="0" pageNumber="23" type="type material">
<paragraph id="60D87CB7899DF2E10BF007872ED3EEDD" pageId="0" pageNumber="23">Type material.</paragraph>
<paragraph id="47065280B0059927E680981990E3E239" pageId="0" pageNumber="23">
<taxonomicName id="F6EC3B270D6640C75E7AB287ACE0A5C5" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">
<emphasis id="66BC2B6194CF386280EABF0199B94B60" italics="true" pageId="0" pageNumber="23">Tychepsephus felix</emphasis>
</taxonomicName>
,
<emphasis id="F86BB184FAB499DAFC6E25A7D6498C61" bold="true" italics="true" pageId="0" pageNumber="23">Holotype</emphasis>
female, pinned. Chile: &quot;Type [white, circular label with red border] // Chili [blue, circular label // 668 //
<taxonomicName id="6512445DB69F2E07C4CE82EEB6120B12" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">Tychepsephus felix</taxonomicName>
, C. Waterh. (Type.) // NHMUK015011475&quot; (Fig.
<figureCitation id="5809FBB690004A74FAB7D0DA8E9BE263" captionStart="Figure 12" captionStartId="F6" captionText="Figure 12. Tychepsephus felix, female type specimen a dorsal view b ventral view c specimen data labels. Images provided by Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1164.103184.figure12" httpUri="https://binary.pensoft.net/fig/855190" pageId="0" pageNumber="23">12</figureCitation>
). Deposited in the NHMUK.
<taxonomicName id="C4DFFD290AA626A2036D3732AD742BB2" class="Insecta" family="Psephenidae" genus="Ectopria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectopria (Chilectopria) grandis" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="grandis" subGenus="Chilectopria">Ectopria (Chilectopria) grandis</taxonomicName>
syn. nov.,
<emphasis id="EB01D8F9E8B8A1538ABD536D537B9210" bold="true" italics="true" pageId="0" pageNumber="23">Holotype</emphasis>
female, pinned. &quot;Chili [green label] // Dascillide? [pale brown label] // type [pale brown label] // Museum Paris / Coll. M.Pic [blue-green label] // Chilectopria / s.g. grandis / n sp [pale brown label] // Chilectopria / grandis Pic // TYPE [red label] // HOLOTYPE [red label] // Tychepsephus / felix ♀ / W D Shepard // MNHN, Paris / EC17127&quot; (Fig.
<figureCitation id="BC2C505DE00CDBE51B6C98E1F89BD014" captionStart="Figure 17" captionStartId="F9" captionText="Figure 17. Chilectopria grandis, female type specimen a dorsal view b lateral view c ventral view d specimen data labels. Images provided by Christophe Rivier (MNHN)." figureDoi="10.3897/zookeys.1164.103184.figure17" httpUri="https://binary.pensoft.net/fig/855193" pageId="0" pageNumber="23">17</figureCitation>
). Deposited in the MNHN.
</paragraph>
<caption id="0D16DE3F6758F6177642A62B95F699AC" doi="10.3897/zookeys.1164.103184.figure12" httpUri="https://binary.pensoft.net/fig/855190" pageId="0" pageNumber="23" start="Figure 12" startId="F6">
<paragraph id="1286973EADB24C7B11645C89B9CFF929" pageId="0" pageNumber="23">
<emphasis id="BA7ED649E9F7C8D0F372667A24FAC1A7" bold="true" pageId="0" pageNumber="23">Figure 12.</emphasis>
<taxonomicName id="F6C2C8F4CFE90C0BE8F53DE3906036D3" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">
<emphasis id="BE4ECC444B3171BCC7A29C459B546A69" italics="true" pageId="0" pageNumber="23">Tychepsephus felix</emphasis>
</taxonomicName>
, female type specimen
<emphasis id="884A2D559A3DDAE98E548AA71021C8B6" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="3FB8F38CF088926C5C58A8B00BE782A4" bold="true" pageId="0" pageNumber="23">b</emphasis>
ventral view
<emphasis id="741EB5E98500324395C5B36DB721A685" bold="true" pageId="0" pageNumber="23">c</emphasis>
specimen data labels. Images provided by Keita Matsumoto (NHMUK).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EBF9D795B679F3421B35DFF45C357969" pageId="0" pageNumber="23" type="materials_examined">
<paragraph id="60E05BFBD41C94BAFFD85033289374FE" pageId="0" pageNumber="23">Other material examined.</paragraph>
<paragraph id="43F2DE31E2AE07FB2D622E54B00B26F4" pageId="0" pageNumber="23">
Non-types (33).
<materialsCitation id="933735BB23FA3DC00A8313E71637DAAF" collectingDate="2002-12-30" collectorName="Region X, William D. Shepard" country="Chile" location="Rio Colegual" specimenCount="5" specimenCount-female="3" specimenCount-male="2">
<collectingCountry id="92BF22509F3EE126889728C109A6E8D0" name="Chile">Chile</collectingCountry>
:
<collectorName id="DECCCA2BADF5A52F62FCC29357DC23A0">Region X</collectorName>
,
<locationDeviation id="AE0D454E5A7890490D6CC51A6945C66F" location="Nueva Braunau">
<quantity id="B0086A400032732E2E7A35CF36F2197A" metricMagnitude="3" metricUnit="m" metricValue="3.0" unit="km" value="3.0">3 km</quantity>
W of Nueva Braunau
</locationDeviation>
,
<location id="5A6ADE3B1D4FAE325FF9DE1A60E29AEB" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:5A6ADE3B1D4FAE325FF9DE1A60E29AEB" country="Chile" name="Rio Colegual">Rio Colegual</location>
,
<collectingDate id="39667EBE0BF1E010C175A9BDD7780083" value="2002-12-30">30 XII 2002</collectingDate>
(WDS-A-1502) [on reverse],
<collectorName id="222A5579BA9EF1D1C67BF1EDBB829325">William D. Shepard</collectorName>
, leg. (EMEC, 5,
<specimenCount id="601905FEE1C473C192422F5401EFE50D" type="male">2♂</specimenCount>
<specimenCount id="B9CEF40F1DDC2435643ECFA0A6CB0504" type="female">3♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="A101B1BECF614C22A17E6D4282D0A333" collectingDate="2003-01-08" collectorName="Region X, William D. Shepard" country="Chile" elevation="198" location="Pte. Colegual" specimenCount="11" specimenCount-female="6" specimenCount-male="5">
<collectingCountry id="F22C37237DCD386EDCF565ADD6DE8CB5" name="Chile">Chile</collectingCountry>
:
<collectorName id="A37A068E067A4E14A2196192EB926AEC">Region X</collectorName>
,
<locationDeviation id="29F6B3F64856E911195372E551743307" location="Loncotoro">
<quantity id="189828F823016FAB68322C807F4E860A" metricMagnitude="3" metricUnit="m" metricValue="9.0" unit="km" value="9.0">9 km</quantity>
E Loncotoro
</locationDeviation>
,
<location id="61FFF54F02B7C5FF7A4C11BDD43FFF91" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:61FFF54F02B7C5FF7A4C11BDD43FFF91" country="Chile" name="Pte. Colegual">Pte. Colegual</location>
2, 650' [
<quantity id="B758BFA7333F399051C6F6DFC62BEFDA" metricMagnitude="2" metricUnit="m" metricValue="1.98" unit="m" value="198.0">
<elevation id="CF2C32E23878B1E6539317884940B051" metricMagnitude="2" metricUnit="m" metricValue="1.98" unit="m" value="198.0">198 m</elevation>
</quantity>
],
<collectingDate id="7E29C438A9ECCFD49C491B4FF5B2DE34" value="2003-01-08">8 I 2003</collectingDate>
(WDS-A-1519) [on reverse],
<collectorName id="83D3A55E2C8A1E93E2AD89FD657847C9">William D. Shepard</collectorName>
, leg. (11; EMEC, 7,
<specimenCount id="11E6F0BA0578CEC4F816BC8169126CF4" type="male">3♂</specimenCount>
<specimenCount id="5732983E685F80A6275157BB677AAFA5" type="female">4♀</specimenCount>
; MNNC, 4,
<specimenCount id="57F673B0EAE294C08D81AEBDB4549135" type="male">2♂</specimenCount>
<specimenCount id="E53AB7556DD98C4B17C07D319D3FD84D" type="female">2♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="91D79E2D899D1C9C40D446D3FD291322" collectingDate="2003-01-08" collectorName="C. B. Barr" country="Chile" county="Llanquehue" elevation="213" latitude="-41.275166" location="Region" longLatPrecision="11" longitude="-73.108665" municipality="Rio Colegual" specimenCount="5" specimenCount-female="1" specimenCount-male="4">
<collectingCountry id="0A449C2FCDB0D010920CDD025FBEE728" name="Chile">Chile</collectingCountry>
:
<normalizedToken id="6314C6EB5EB7947E3732FBA1CC0E43F6" originalValue="Región">
<location id="D9A322761CD81F7343F7FFC79641F1FE" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:D9A322761CD81F7343F7FFC79641F1FE" country="Chile" county="Llanquehue" latitude="-41.275166" longLatPrecision="11" longitude="-73.108665" municipality="Rio Colegual" name="Region">Region</location>
</normalizedToken>
X
<location id="8220A255D0FF6A7EC9348D0776612C5D" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:8220A255D0FF6A7EC9348D0776612C5D" country="Chile" county="Llanquehue" latitude="-41.275166" longLatPrecision="11" longitude="-73.108665" municipality="Rio Colegual" name="Lagos">Lagos</location>
,
<collectingMunicipality id="51005FB0E62569C34C758FA2C6E802C7">
<normalizedToken id="40C2FADDC357C6204CDE7ADC187FBB64" originalValue="Río">Rio</normalizedToken>
Colegual
</collectingMunicipality>
8 rd. km W
<collectingCounty id="05EEC11C1B64EB40AB46D15F16AEC19A">Llanquehue</collectingCounty>
, elev.700' [
<quantity id="008FF78F934EC4DC3D8D2A17F0122AC6" metricMagnitude="2" metricUnit="m" metricValue="2.13" unit="m" value="213.0">
<elevation id="BDB929F1373ABFB700DD3B83F8AE1FC3" metricMagnitude="2" metricUnit="m" metricValue="2.13" unit="m" value="213.0">213 m</elevation>
</quantity>
],
<geoCoordinate id="9D2AD1931547A559CC8C10310FC95BE7" degrees="41" direction="south" minutes="16.51" orientation="latitude" precision="9" value="-41.275166">41°16.51'S</geoCoordinate>
,
<geoCoordinate id="B20EB125950066B13F32EB0AFC604EA0" degrees="73" direction="west" minutes="06.52" orientation="longitude" precision="9" value="-73.108665">73°06.52'W</geoCoordinate>
,
<collectingDate id="10CA18DD00117824107C2299233E2C29" value="2003-01-08">8 Jan. 2003</collectingDate>
,
<collectorName id="98420676E1D37AAE07CE2C7005C78BDB">C. B. Barr</collectorName>
, sweeping willows and other riparian vegetation (EMEC, 5,
<specimenCount id="5512A87ADC83FCEA762EEB42C4BEE3C4" type="male">4♂</specimenCount>
<specimenCount id="EA01BA005ED3BD75E6BE8176324929A3" type="female">1♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="F25063BF3ABC1E5D46CAB46DB6B5F815" collectingDate="2002-12-31" collectorName="Region X, William D. Shepard" country="Chile" elevation="128" location="Pte. Hondo" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F441D6D7B1E42150EA3FD7CB7BBA6585" name="Chile">Chile</collectingCountry>
:
<collectorName id="9052F6316DEC5634B396B558D6BC51F4">Region X</collectorName>
,
<locationDeviation id="C2B4446DF86F3AE630F0B3F466860134" location="Correntoso">
<quantity id="403C3DD4A11F81CFB2114F2FA6D53DB2" metricMagnitude="3" metricUnit="m" metricValue="8.0" unit="km" value="8.0">8 km</quantity>
SW Correntoso
</locationDeviation>
,
<location id="4DFF341F65387CA7DA8807F55D72F65F" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:4DFF341F65387CA7DA8807F55D72F65F" country="Chile" name="Pte. Hondo">Pte. Hondo</location>
[
<normalizedToken id="A273676F23AF4236F3E0052173819336" originalValue="Río">Rio</normalizedToken>
Oroco], 420' [
<quantity id="C8CCECF58DB722C78A737B9A2595AA27" metricMagnitude="2" metricUnit="m" metricValue="1.28" unit="m" value="128.0">
<elevation id="9DCDE942BFFC4E2311065652DAD757FF" metricMagnitude="2" metricUnit="m" metricValue="1.28" unit="m" value="128.0">128 m</elevation>
</quantity>
],
<collectingDate id="A309F280B776F6AE3CFEFD35BD179CE8" value="2002-12-31">31 XII 2002</collectingDate>
(WDS-A-1504) [on reverse],
<collectorName id="7EB3B562D494F4AC426B01A77B913660">William D. Shepard</collectorName>
, leg. (EMEC,
<specimenCount id="79FD4E2B7CE68C25A8F35F4DAFCEA9CC" type="female">1♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="BA5615E641D609EE8E3BEEC775B651B2" collectingDate="1905-12" collectorName="R. Thaxter" country="Chile" location="Corral" specimenCount="1" specimenCount-male="1">
<collectingCountry id="B07AB4F5DFAC2D6F3DC620B80F199411" name="Chile">Chile</collectingCountry>
:
<location id="AA27D4785C2439FAE2A7FB5BB68025E4" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:AA27D4785C2439FAE2A7FB5BB68025E4" country="Chile" name="Corral">Corral</location>
,
<collectingDate id="8200D043F7B10E2289FA0237DAAD580E" value="1905-12">Dec. 1905</collectingDate>
,
<collectorName id="2E169E8EB6EB13D0643017FC17F9872D">R. Thaxter</collectorName>
, MCZ (MCZC,
<specimenCount id="CBB59389D6B061F10A4D7D0205B0ABB0" type="male">1♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="247B8A4724321DF1CDC618B3C32153D4" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="F. C. Bowditch Coll." country="Chile" location="Chile" specimenCount="1" specimenCount-male="1">
<collectingCountry id="55996E8AD77B0E5EFD89C83E77F08691" name="Chile">Chile</collectingCountry>
:
<collectorName id="08575167FD889F254AEA8BDC5DDD7331">F. C. Bowditch Coll.</collectorName>
,
<normalizedToken id="2EC5BA20A53616B1B7E8735C35FB0DEA" originalValue="“Bradytoma”">&quot;Bradytoma&quot;</normalizedToken>
, MCZ (MCZC,
<specimenCount id="851021E8B10E8838638CA431E8B6D3E5" type="male">1♂</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="949FD68E27EEE337FC5D4391E82684A2" collectingDate="1996-11-11" collectorName="Tomas Cekalovic" country="Chile" location="Concepcion Pr" municipality="Estero Nonguen" specimenCount="1" specimenCount-female="1">
<collectingCountry id="489135DA0BB150A6D46A997DB861E44B" name="Chile">Chile</collectingCountry>
:
<location id="1AC53F9591287F8C885843284934A3D6" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:1AC53F9591287F8C885843284934A3D6" country="Chile" municipality="Estero Nonguen" name="Concepcion Pr">
<normalizedToken id="F0EEE361E1CA55372A91409BC128D49F" originalValue="Concepción">Concepcion</normalizedToken>
Pr
</location>
,
<collectingMunicipality id="1869B78C5A698917ADBF3FE1D518C458">Estero Nonguen</collectingMunicipality>
,
<collectingDate id="B0D77C688BE1A8A4E881FE2FD8A76E2A" value="1996-11-11">11 Noviembre 1996</collectingDate>
,
<collectorName id="77AF496E8F01B47A256C61E1442D8FB1">
Tomas
<normalizedToken id="B4FA1169DD3B7D35F0D410CE13589674" originalValue="Cekálovic">Cekalovic</normalizedToken>
</collectorName>
(EMEC,
<specimenCount id="321DD3B4BFD24F148F04D632B783BB7C" type="female">1♀</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="0C4977252AFE26E16EF0541E163CB4FB" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Chile" location="Germain" specimenCount="1">
<collectingCountry id="B2BD639154CD2827C18B7C26B82EE897" name="Chile">Chile</collectingCountry>
: Chili,
<location id="5E3E7F489298F3F1B458D10D5423D7E7" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:5E3E7F489298F3F1B458D10D5423D7E7" country="Chile" name="Germain">Germain</location>
, Sharp Coll. 1905-313, NHMUK015011806 (NHMUK, 1)
</materialsCitation>
;
<materialsCitation id="476468979D70524082E5A410CED55E88" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Tycepsephus, C. O. Waterh" country="Chile" location="Sharp Coll." specimenCount="1">
<collectingCountry id="320C17010CB4FB2B8BD1E8128012F325" name="Chile">Chile</collectingCountry>
:
<location id="BE38E97C0F9B62E280A188DD01B73546" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:BE38E97C0F9B62E280A188DD01B73546" country="Chile" name="Sharp Coll.">Sharp Coll.</location>
1905-313,
<collectorName id="40AF7C0326BFBBB8BB7BD23DEF789D6E">Tycepsephus</collectorName>
[sic] felix,
<collectorName id="5FFA0D7F311BB41D4B118C2F7CD2DF4A">C.O. Waterh</collectorName>
</materialsCitation>
.,
<materialsCitation id="805A2CB4A447A9B4917AA02C621D2848" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Chili" location="Germain" specimenCount="1">
<collectingCountry id="8551D56FB25AFEE6F11A8942B144180A" name="Chile">Chili</collectingCountry>
-
<location id="BE97C8C6295137E9CABE94FF90C37447" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:BE97C8C6295137E9CABE94FF90C37447" country="Chili" name="Germain">Germain</location>
, NHMUK015011809 (NHMUK, 1)
</materialsCitation>
;
<materialsCitation id="06BC0BAE106644C3986F2DB82663EF8D" collectingDate="1926-12-16" country="Chile" location="Puerto Varas" specimenCount="1">
<collectingCountry id="F23419F35951BC9CD117BE77EB655322" name="Chile">Chile</collectingCountry>
:
<location id="DA435E4FB1A6F7ED146A2617B9DE0FE5" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:DA435E4FB1A6F7ED146A2617B9DE0FE5" country="Chile" name="Puerto Varas">Puerto Varas</location>
,
<collectingDate id="8C54CD5AE1729C3DD9BAB5B5B239F1FE" value="1926-12-16">16.xii.1926</collectingDate>
, S
</materialsCitation>
.
<materialsCitation id="47E3E0CC0351C3217C725576DC0EB896" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="F. &amp; M. Edwards" country="Chile" location="Llanquihue prov." specimenCount="1">
<collectingCountry id="49104BBD43D28C09E8A61DBAB5F4D88A" name="Chile">Chile</collectingCountry>
:
<location id="D4C09972631567158B98D4124DE2AD0C" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:D4C09972631567158B98D4124DE2AD0C" country="Chile" name="Llanquihue prov.">Llanquihue prov.</location>
,
<collectorName id="339CD82B0E4667BD2245577D40953F6E">F. &amp; M. Edwards</collectorName>
, B.M.1927-63, NHMUK015011807 (NHMUK, 1)
</materialsCitation>
;
<materialsCitation id="A5456969D1C1DB660A338D978C48574C" collectingDate="1926-12-04" collectingDateMax="1926-12-10" collectingDateMin="1926-12-04" country="Chile" location="Casa Pangue" specimenCount="1">
<collectingCountry id="45AB4483455F92FEB2F1582A743ABE62" name="Chile">Chile</collectingCountry>
:
<location id="5289F35E2DA0B36B29894D7ADD1BD305" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:5289F35E2DA0B36B29894D7ADD1BD305" country="Chile" name="Casa Pangue">Casa Pangue</location>
,
<collectingDate id="8FA618FD5095D03DA0E8B9586D0FC0D9" value="1926-12-04" valueMax="1926-12-10" valueMin="1926-12-04">4-10.xii.1926</collectingDate>
, S
</materialsCitation>
.
<materialsCitation id="786F5C63F255E19DE32DFF4812B175E1" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="F. &amp; M. Edwards, Tychepsephus, Waterh." country="Chile" determinerName="M. I. Russell" location="Llanquihue prov." specimenCount="1">
<collectingCountry id="C6104CA08BD95BAFBFA34C53AB3B98CB" name="Chile">Chile</collectingCountry>
:
<location id="16EBC08856377FF15237B8C099487EFC" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:16EBC08856377FF15237B8C099487EFC" country="Chile" name="Llanquihue prov.">Llanquihue prov.</location>
,
<collectorName id="C1816316C7D07C560F12951DB5D6F055">F. &amp; M. Edwards</collectorName>
, B.M.1927-63,
<collectorName id="666A41121FF2CC561C880642DEEDAB13">Tychepsephus</collectorName>
felix
<collectorName id="506BDDFEF2338A2609C1B3F819080C67">Waterh.</collectorName>
,
<determinerName id="8830F452B84DE52CCFA3E5B0DD5FE435">M.I. Russell</determinerName>
det. 1973, NHMUK015011808 (NHMUK, 1)
</materialsCitation>
;
<materialsCitation id="F0CA87248CDC4D08536E4FC76CD0DC36" collectingDate="1926-12-17" collectingDateMax="1926-12-19" collectingDateMin="1926-12-17" country="Chile" location="Ancud." specimenCount="1">
<collectingCountry id="042EC480F5217F9B7FB2DBC07EF2115A" name="Chile">Chile</collectingCountry>
:
<location id="49B31685817866F9C6AC190EEEF6DE3A" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:49B31685817866F9C6AC190EEEF6DE3A" country="Chile" name="Ancud.">Ancud.</location>
<collectingDate id="A6491705E62AC895F4F0A341071CDF35" value="1926-12-17" valueMax="1926-12-19" valueMin="1926-12-17">17-19.xii.1926</collectingDate>
, S
</materialsCitation>
.
<materialsCitation id="0BAFCCEA8CFA15A9A5FBC82EDC0E805D" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="F. &amp; M. Edwards" country="Chile" location="Chiloe" specimenCount="1">
<collectingCountry id="85DDEA289959BBFB8586D5514CFF0262" name="Chile">Chile</collectingCountry>
:
<location id="C713A45D6ECB1EA01863A9AB952FE571" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:C713A45D6ECB1EA01863A9AB952FE571" country="Chile" name="Chiloe">Chiloe</location>
I.,
<collectorName id="BDF07351356D6F95A828E51BB95E71AF">F. &amp; M. Edwards</collectorName>
, B.M.1927-63, NHMUK015011810 (NHMUK, 1)
</materialsCitation>
;
<collectingCountry id="7B4A78D26FF8C5C03D9848E62D2273DD" name="Chile">Chile</collectingCountry>
: as above, NHMUK015011811 (NHMUK, 1);
<materialsCitation id="B63CDC566D736A07601964781C7F449D" collectingDate="1926-12-19" country="Chile" location="Ancud" specimenCount="1">
<collectingCountry id="8FFFD803BC371C7CF4CFC44E0A3114E2" name="Chile">Chile</collectingCountry>
:
<location id="96B7BC479153A2A65BEDCAB87309ED69" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:96B7BC479153A2A65BEDCAB87309ED69" country="Chile" name="Ancud">Ancud</location>
,
<collectingDate id="5B3955F6ADAEDEC36FC1C16FB84BF828" value="1926-12-19">19.xii.1926</collectingDate>
, S
</materialsCitation>
.
<materialsCitation id="F25030815F1136185263409920A41525" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="F. &amp; M. Edwards, Waterh." country="Chile" determinerName="M. I. Russell" location="Chiloe" specimenCount="1">
<collectingCountry id="C6DE2E8D3BA0ECDE9A3D4CB3F85E71F4" name="Chile">Chile</collectingCountry>
:
<location id="57320CC466B9937E9F40BBD2448E8597" LSID="urn:lsid:plazi:treatment:18DD8E0E120352A1904BDA2807D155CD:57320CC466B9937E9F40BBD2448E8597" country="Chile" name="Chiloe">Chiloe</location>
I.,
<collectorName id="730E73FC79F6F47CE2A2161D5B1746C8">F. &amp; M. Edwards</collectorName>
, B.M.1927-63, NHMUK015011812 (NHMUK, 1); as above,
<taxonomicName id="18E5D578A5B56117A706D6C8F1194A05" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">Tychepsephus felix</taxonomicName>
<collectorName id="2F75A8B3974F300C4B44B7C17BDD3813">Waterh.</collectorName>
,
<determinerName id="37B0BBFE1E57DBC733D7F0B2C296EC1F">M.I. Russell</determinerName>
det. 1973, NHMUK015011813 (NHMUK, 1)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B1381D1C03370F4E655611DB841BFB0A" pageId="0" pageNumber="23" type="differential diagnosis">
<paragraph id="A38893FB59143142578F8957C9351FF5" pageId="0" pageNumber="23">Differential diagnosis.</paragraph>
<paragraph id="B7FFDA03B14CD309F33CF3B5935E77E9" pageId="0" pageNumber="23">
Males of
<taxonomicName id="C674812DE78BA5521A47CD53ACCA360A" class="Insecta" kingdom="Animalia" lsidName="T. felix" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="2320ACA6971B110F3CD634C35C6A843B" italics="true" pageId="0" pageNumber="23">T. felix</emphasis>
</taxonomicName>
(Figs
<figureCitation id="B806228F85B6B5DA5975FCA868F7901C" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">13</figureCitation>
,
<figureCitation id="393D96690DE3CC3182EC10A9CAFC9EAA" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">14</figureCitation>
) are much larger (4.6-5.2 mm long) than those of
<taxonomicName id="41A15389030846D921C7DE675C73F0E8" class="Insecta" kingdom="Animalia" lsidName="T. cekalovici" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="6F63B665459DD6EE1602CC72CD9D9E82" italics="true" pageId="0" pageNumber="23">T. cekalovici</emphasis>
</taxonomicName>
sp. nov. (3.3-3.9 mm long); the pronotal cuticle is dark brown to black with pale lateral margins, with no yellow markings on disc; the elytral cuticle is dark brown or red-brown, with setal patterning only; the depressed frontal area between the eyes does not have an inverted Y-shaped sulcus; and abdominal ventrite 3 has a median, golden yellow setal patch that is not distinctly raised and does not extend the entire length of the ventrite.
</paragraph>
<caption id="A02DFCB33E49BA50E167C3947D517F2E" doi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23" start="Figures 13, 14" startId="F7">
<paragraph id="2658E5051D42C047CC4CE86B1D905A03" pageId="0" pageNumber="23">
<emphasis id="690A997354C7C9C4A11810463A16E322" bold="true" pageId="0" pageNumber="23">Figures 13, 14.</emphasis>
<taxonomicName id="77E2E94940004F5D7BB13BCD5E4BE396" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">
<emphasis id="8CD85F43F57A05F9195740A0910D3EBB" italics="true" pageId="0" pageNumber="23">Tychepsephus felix</emphasis>
</taxonomicName>
, male
<emphasis id="ED204C00D9A8BD2B1FCFA29299ABA5AF" bold="true" pageId="0" pageNumber="23">13</emphasis>
habitus
<emphasis id="3EC7DF53DD23EDE22F164FF359DEAD9F" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="2BD1F8984431418E8F6EE85370A49E61" bold="true" pageId="0" pageNumber="23">b</emphasis>
ventral view; length 5.0 mm
<emphasis id="250F2C0222EF24F45C76B67726E5C572" bold="true" pageId="0" pageNumber="23">14</emphasis>
aedeagus
<emphasis id="C6C8813A073E4B011280595A2F0BB5B9" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="95B6CFE3FC7ED7E19FA1DFED67951242" bold="true" pageId="0" pageNumber="23">b</emphasis>
lateral view
<emphasis id="D17A44EBE4893366B3789D7BC200A3D8" bold="true" pageId="0" pageNumber="23">c</emphasis>
ventral view.
</paragraph>
</caption>
<paragraph id="70EE3136FC9F3F5EF72415E78EE565E6" pageId="0" pageNumber="23">
In contrast, males of
<taxonomicName id="837BEDA312F42314320B606586973656" class="Insecta" kingdom="Animalia" lsidName="T. cekalovici" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="B90F095DBB5EE7A3B7425C1AEA73C3E9" italics="true" pageId="0" pageNumber="23">T. cekalovici</emphasis>
</taxonomicName>
sp. nov. (Figs
<figureCitation id="6A21233EA581EE3789A65E88AD65D284" captionStart="Figures 18, 19" captionStartId="F10" captionText="Figures 18, 19. Tychepsephus cekalovici sp. nov., male 18 habitus a dorsal view b ventral view; length 3.2 mm 19 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures18-19" httpUri="https://binary.pensoft.net/fig/855194" pageId="0" pageNumber="23">18</figureCitation>
-
<figureCitation id="A82F3F74FFF07B4304F4C491718CF165" captionStart="Figure 20" captionStartId="F11" captionText="Figure 20. Tychepsephus cekalovici sp. nov., male dorsal habitus showing color pattern variation a length 3.3 mm b length 3.5 mm c length 3.5 mm." figureDoi="10.3897/zookeys.1164.103184.figure20" httpUri="https://binary.pensoft.net/fig/855195" pageId="0" pageNumber="23">20</figureCitation>
) are considerably smaller than males of
<taxonomicName id="75BFBA9D41B8952EC66B796C8FADAF4A" class="Insecta" kingdom="Animalia" lsidName="T. felix" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="DB94920C8CEDC5E9A09736855A3F06CC" italics="true" pageId="0" pageNumber="23">T. felix</emphasis>
</taxonomicName>
; the pronotal cuticle is dark brown to black with pale lateral and basal margins, and a mediobasal yellow spot anterior and adjacent to the scutellar shield; the elytral cuticle is usually yellow-brown with dark markings in a zig-zag pattern, but may be mostly plain, without patterning; the depression between the eyes has a narrow, inverted Y-shaped sulcus; and abdominal ventrite 3 has a prominent, raised, golden yellow setal patch extending the full length of the ventrite.
</paragraph>
<paragraph id="ACCE9AAFC169705398684B3D46441DF0" pageId="0" pageNumber="23">
The aedeagi (Figs
<figureCitation id="37B17AA0BFFDA684A4CA0A884D708922" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">14</figureCitation>
,
<figureCitation id="708C1AE09FE41A5A7DA2052B607C5FA2" captionStart="Figures 18, 19" captionStartId="F10" captionText="Figures 18, 19. Tychepsephus cekalovici sp. nov., male 18 habitus a dorsal view b ventral view; length 3.2 mm 19 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures18-19" httpUri="https://binary.pensoft.net/fig/855194" pageId="0" pageNumber="23">19</figureCitation>
) of the two species are clearly different. In
<taxonomicName id="1C7507B56B9D47FA214DCA8F60D9A057" class="Insecta" kingdom="Animalia" lsidName="T. felix" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="2B79B886C0034D08552FC5F6843D9AEE" italics="true" pageId="0" pageNumber="23">T. felix</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="ED8705280E49919020210476197923B8" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">14</figureCitation>
), the parameres have straight lateral margins and only slightly curved medial margins, and the apices are narrow. In
<taxonomicName id="CC95089A9FCE089D83BCD390B2A40977" class="Insecta" kingdom="Animalia" lsidName="T. cekalovici" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="EBC55C054B4383162A1D3C09DA30043D" italics="true" pageId="0" pageNumber="23">T. cekalovici</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="0F86F57D2A734CB10D1B46269A56673F" captionStart="Figures 18, 19" captionStartId="F10" captionText="Figures 18, 19. Tychepsephus cekalovici sp. nov., male 18 habitus a dorsal view b ventral view; length 3.2 mm 19 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures18-19" httpUri="https://binary.pensoft.net/fig/855194" pageId="0" pageNumber="23">19</figureCitation>
) the parameres have curved lateral and medial margins, and the apices are broad.
</paragraph>
<paragraph id="149D84904B69D7A8DDFC23E4B632E254" pageId="0" pageNumber="23">
Females of
<taxonomicName id="76FD16CFC967A8A4EE1DDF52B13F0CA8" class="Insecta" kingdom="Animalia" lsidName="T. felix" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="3220A3E1F773AC756677343823A2F145" italics="true" pageId="0" pageNumber="23">T. felix</emphasis>
</taxonomicName>
(Figs
<figureCitation id="F39BF1D4DB8D22F736F8A0F89B9DCAA4" captionStart="Figures 15, 16" captionStartId="F8" captionText="Figures 15, 16. Tychepsephus felix, female 15 habitus a dorsal view b ventral view; length 4.2 mm 16 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23">15</figureCitation>
,
<figureCitation id="CAF6B93C827418E539D031D17D5D2FA2" captionStart="Figures 15, 16" captionStartId="F8" captionText="Figures 15, 16. Tychepsephus felix, female 15 habitus a dorsal view b ventral view; length 4.2 mm 16 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23">16</figureCitation>
) are much larger (4.3-5.7 mm long) than those of
<taxonomicName id="71A77C7C9E8EB6D65B81750FC4782AB6" class="Insecta" kingdom="Animalia" lsidName="T. cekalovici" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="70D9C26D61295C457A2B6CF710C4F040" italics="true" pageId="0" pageNumber="23">T. cekalovici</emphasis>
</taxonomicName>
(3.3-3.9 mm long); the elytral cuticle is brown, without yellow patterning except for variable, slightly paler areas near the base; and the pronotal disc has no yellow markings.
<taxonomicName id="0F5803D2A47F6DEFA42004256DD27FA8" authorityName="Shepard &amp; Barr" authorityYear="2023" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus cekalovici" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="cekalovici">
<emphasis id="78AFB84D0C85F4D75EF814DC654D675E" italics="true" pageId="0" pageNumber="23">Tychepsephus cekalovici</emphasis>
</taxonomicName>
sp. nov. females (Figs
<figureCitation id="1B620E3B3D98174E3F8ECF18FFEE854A" captionStart="Figures 21–23" captionStartId="F12" captionText="Figures 21 - 23. Tychepsephus cekalovici sp. nov., female 21 habitus a dorsal view b ventral view; length 4.4 mm 22 dorsal color pattern variation; length 4.3 mm 23 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures21-23" httpUri="https://binary.pensoft.net/fig/855196" pageId="0" pageNumber="23">21-23</figureCitation>
) usually have elytra with transverse yellow bands in a zig-zag pattern, but those without may be distinguished by a mediobasal yellow spot on the pronotum anterior to the scutellar shield. The ovipositors are similar.
</paragraph>
<caption id="FFBB795B4983EE67B2243375156B0964" doi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23" start="Figures 15, 16" startId="F8">
<paragraph id="BEE0F6B1E03081A132FE9D6821EACA59" pageId="0" pageNumber="23">
<emphasis id="295A3F2ABF1B546CDFB1BCF17E9EDFA2" bold="true" pageId="0" pageNumber="23">Figures 15, 16.</emphasis>
<taxonomicName id="41096786BEB01D18DC30B89DC48320A8" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">
<emphasis id="441BEAFE19ACEDD660D1AA59DF2A8DE3" italics="true" pageId="0" pageNumber="23">Tychepsephus felix</emphasis>
</taxonomicName>
, female
<emphasis id="2738128E6BF69A07B77DFE3C01A87CC3" bold="true" pageId="0" pageNumber="23">15</emphasis>
habitus
<emphasis id="4ED695DB48D493216E6AAA88E466C4C4" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="FF02F61412EF84CB7AFB62178749B0A4" bold="true" pageId="0" pageNumber="23">b</emphasis>
ventral view; length 4.2 mm
<emphasis id="A40AD87C08D195C294E64856223002C0" bold="true" pageId="0" pageNumber="23">16</emphasis>
ovipositor, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0EC25EA4F4550983C9B6A2769F13497A" pageId="0" pageNumber="23" type="description">
<paragraph id="C2392E20C8326BFB74D8D7596E84F6AA" pageId="0" pageNumber="23">Original type description, female</paragraph>
<paragraph id="70AFE4E41F63D1B8E0D64DC0ABF346FB" pageId="0" pageNumber="23">
<emphasis id="A9BC3A52DEB8D4C780D11F685413C8FB" bold="true" pageId="0" pageNumber="23">
(Fig.
<figureCitation id="47D8443C3F171F8CC19A02FD7A02D0C7" captionStart="Figure 12" captionStartId="F6" captionText="Figure 12. Tychepsephus felix, female type specimen a dorsal view b ventral view c specimen data labels. Images provided by Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1164.103184.figure12" httpUri="https://binary.pensoft.net/fig/855190" pageId="0" pageNumber="23">12</figureCitation>
).
</emphasis>
<bibRefCitation id="FFC1367A1529683849EC62139A140A9C" DOI="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" author="Waterhouse, CO" journalOrPublisher="Transactions of the Royal Entomological Society" pageId="0" pageNumber="23" pagination="11 - 25" refId="B25" refString="Waterhouse, CO, 1876. III. On various new genera and species of Coleoptera. Transactions of the Royal Entomological Society 1876 (1): 11 - 25, DOI: https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" title="III. On various new genera and species of Coleoptera." url="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" volume="1876" year="1876">Waterhouse (1876)</bibRefCitation>
described the species in Latin, as was the custom at the time. An English translation follows:
</paragraph>
<paragraph id="905D7ABA416CB8AF219C4B87636EABBA" pageId="0" pageNumber="23">Ovate, convex, glossy, dark pitch black, bronzy; fine, short, grey pubescence. Head yellow, rather wide, narrow between antennae, eyes prominent, antennal bases yellow. Thorax slightly convex, densely finely punctate, length twice width, suddenly narrowed anteriorly, frontal margin slightly lobed in middle, both sides sinuate, anterior angles somewhat prominent, sides slightly rounded behind middle, posterior angles almost right-angled, edges narrowly yellow. Scutellum yellow, apex acute. Elytral bases slightly wider than thorax, enlarged posteriorly, arched at apex, narrowed, convex, more clearly finely punctured, dorsum depressed; humeri obtuse, with edges narrowly yellow. Venter with densely grey pubescence, tarsi pitch black-yellow. Length 2.75 lin., width 2 lin.</paragraph>
<paragraph id="76242DBCD206BFF89FDDD5F593C17661" pageId="0" pageNumber="23">
<bibRefCitation id="B12AE62685BBB3A230D59136CAD3E2D1" DOI="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" author="Waterhouse, CO" journalOrPublisher="Transactions of the Royal Entomological Society" pageId="0" pageNumber="23" pagination="11 - 25" refId="B25" refString="Waterhouse, CO, 1876. III. On various new genera and species of Coleoptera. Transactions of the Royal Entomological Society 1876 (1): 11 - 25, DOI: https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" title="III. On various new genera and species of Coleoptera." url="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" volume="1876" year="1876">Waterhouse (1876)</bibRefCitation>
added two sentences, in English: &quot;The thorax is at the base nearly straight next to the scutellum, but is broadly sinuate on each side, so that at first sight it appears only bisinuate. Epipleural fold of the elytra is broad at the base, gradually narrowing to the apex, channeled posteriorly.&quot;
</paragraph>
</subSubSection>
<subSubSection id="215028860DDDF896FB31D8FA8DBA449A" pageId="0" pageNumber="23" type="materials_examined">
<paragraph id="383C134FA33B64562E6836030BE4B0A2" pageId="0" pageNumber="23">Redescription based on new material.</paragraph>
<paragraph id="451F9CD4403868AA4AE07E969A4EBDB1" pageId="0" pageNumber="23">
<emphasis id="6A63348A18AFE519053012184391F665" bold="true" pageId="0" pageNumber="23">Male</emphasis>
(Figs
<figureCitation id="61C8B828B640AAD37E1C5EF83DA5F8E1" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">13</figureCitation>
,
<figureCitation id="8F91745F762FF8D86917DD5A907BCD97" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">14</figureCitation>
)
<emphasis id="89C4583B03F457806364CC24BACEF401" bold="true" pageId="0" pageNumber="23">
.
<emphasis id="24A09977D21743B72E25495FACD7012B" italics="true" pageId="0" pageNumber="23">Body</emphasis>
</emphasis>
: covered with short black setae and thick, moderately long yellow setae; yellow setae forming patterns on elytra. Cuticle with closely spaced punctures, punctures finer ventrally. Pronotum black with yellow margins. Elytra dark brown or red-brown with yellow lateral margins. Length
<quantity id="C0C61EA9C184F5478CEF1353B4E21329" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.2" metricValueMin="4.6" unit="mm" value="4.9" valueMax="5.2" valueMin="4.6">4.6-5.2 mm</quantity>
(
<emphasis id="49C2C754810F0B90B5E0D20C02B4715F" italics="true" pageId="0" pageNumber="23">n</emphasis>
= 6), width
<quantity id="916DF209403A0DF45FE9BEA5A72F657C" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8-3.5 mm</quantity>
(
<emphasis id="5BABB0AF9A1F16FCD262D0E402B77B56" italics="true" pageId="0" pageNumber="23">n</emphasis>
= 5).
<emphasis id="AB989168028C98CB71FBA69C85F348B7" bold="true" italics="true" pageId="0" pageNumber="23">Head</emphasis>
: covered with moderately long, yellow setae. Vertex between eyes wider than diameter of an eye. Frons deflexed at 90° angle from vertex, with a contiguous pair of broad, moderately deep depressions between eyes. Frontoclypeal suture absent. Clypeus trapezoidal, longer than wide, widest apically; clypeus noticeably raised; anterolateral angles curved beneath antennal bases. Maxillary palpus with four palpomeres; palpomeres 1-3 yellow to yellow-brown; 4 yellow-brown to dark brown, obliquely hatchet-shaped, weakly curved at apex. Labial palpus with three palpomeres; palpomeres 1-2 yellow; 3 yellow-brown to dark brown, with apex truncate to weakly curved; glossae and paraglossae split, apically acicular. Antenna weakly serrate, with 11 antennomeres; 1 longest, cylindrical, yellow; 2 shortest, yellow-brown; 3-11 dark brown, each widest apically. Antennal base encircled by raised margin. Eye large, bulbous, finely faceted.
<emphasis id="1D3C06D741D47D25CA416EB26A961833" bold="true" italics="true" pageId="0" pageNumber="23">Pronotum</emphasis>
: twice as wide as long, widest just anterior to base; anterior margin convex between anterior angles; anterior angles obtuse, broadly rounded, projecting anteriorly, clasping eyes; lateral margins finely sculptured with shallow notches, narrowly explanate, straight from anterior angles to basal 1/3 then curved to posterior angles; posterior angles square; posterior margin crenulate, nearly straight between posterior angles almost to scutellar shield, then curved posteriorly, straight adjacent to scutellar shield; disc depressed near anterior angles, convex at middle, flattened basomedially.
<emphasis id="44C041F0A285FDE1558B7D11E6B18287" bold="true" italics="true" pageId="0" pageNumber="23">Scutellar shield</emphasis>
: pentagonal, flat, depressed between elytra; densely covered by moderately long, yellow setae; anterior margin crenulate; apex acute.
<emphasis id="0A4FC41F47EC837C4F607049C0EB6720" bold="true" italics="true" pageId="0" pageNumber="23">Elytra</emphasis>
: conjointly longer than wide, widest at ~ 1/3 the distance from apices, wider than pronotum; each with anterior margin crenulate; lateral margin smooth, narrowly explanate; humerus prominent with adjacent medial depression; area adjacent to lateral margin in anterior 1/3 with a wide, moderately deep depression; disc much more convex at posterior 1/2. Setae largely absent from oval area posterior to middle and round area at apical 1/4 near suture; setae sparse near base. Epipleuron widest below humerus, narrowed posteriorly; channeled adjacent to abdomen, channel becoming deeper towards apex.
<emphasis id="6133FFD1AB13137803A7D48E37E0C166" bold="true" italics="true" pageId="0" pageNumber="23">Metathoracic wings</emphasis>
: macropterous.
<emphasis id="98DD640C228404EDBE942E85D16B1C0E" bold="true" italics="true" pageId="0" pageNumber="23">Prosternum</emphasis>
: wider than long; cuticle brown, densely setose; anterior margin projecting to cover mouthparts with a short chin piece; prosternal process long, narrow, extending well past procoxae to anterior third of mesosternum, elevated above rest of prosternum; disc with shallow, longitudinal, median ridge.
<emphasis id="3EECBEBC0C9F4CFED9DE9407D2DCC589" bold="true" italics="true" pageId="0" pageNumber="23">Mesoventrite</emphasis>
: brown; wider than long, extending between mesocoxae; mesoventral cavity deep to receive prosternal process.
<emphasis id="DB440AB28D9F0014C03737D36303327F" bold="true" italics="true" pageId="0" pageNumber="23">Metaventrite</emphasis>
: wider than long, cuticle black; disc convex lateral to discrimen; deep fossa at junction of discrimen and metakatepisternal suture; posterior margin weakly sinuate anterior to metacoxae.
<emphasis id="0F1BFE5C1244E4F7000B8D3D972B3B5F" bold="true" italics="true" pageId="0" pageNumber="23">Legs</emphasis>
: completely covered with dense, yellow setae; similar, except procoxa and mesocoxa globular, metacoxa widely transverse; femur and tibia usually darker than tarsus; tibia and tarsus long and narrow, tarsus longer than tibia; tarsus densely covered with pale setae; claws long, slender.
<emphasis id="E014F028081636B1C67DE8C5EBF7220B" bold="true" italics="true" pageId="0" pageNumber="23">Abdomen</emphasis>
: brown, densely setose; with five weakly convex ventrites; ventrites 1-4 short, ventrite 5 longest; ventrites 1-3 with lateral margins notched medially; ventrite 3 with a dense patch of coarse, golden yellow setae covering the posteromedial 1/2-2/3 of the disc.
<emphasis id="3858E5A57C59F2525BACC2ACEB1C8C89" bold="true" italics="true" pageId="0" pageNumber="23">Aedeagus</emphasis>
: (Fig.
<figureCitation id="E4BAE250929A4D7C793F007AFF08E1DB" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">14</figureCitation>
) more than twice as long as wide; phallobase vase-shaped, narrow at base, wide at apex; parameres stout, much longer than penis, lateral margins straight at basal 2/3 then gradually curved inward at apical 1/3, medial margins only slightly curved, parameres each narrowed toward a rounded apex; penis tapered, apex broadly rounded, basally with two apophyses.
<emphasis id="668C90834DD48A8DBC4B17778130D851" bold="true" pageId="0" pageNumber="23">Female</emphasis>
(Figs
<figureCitation id="99D5CB1A03EB32C8942FEA74B568B5D6" captionStart="Figures 15, 16" captionStartId="F8" captionText="Figures 15, 16. Tychepsephus felix, female 15 habitus a dorsal view b ventral view; length 4.2 mm 16 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23">15</figureCitation>
,
<figureCitation id="0DBFEA8E380A44E72E07838E8B5BC7B9" captionStart="Figures 15, 16" captionStartId="F8" captionText="Figures 15, 16. Tychepsephus felix, female 15 habitus a dorsal view b ventral view; length 4.2 mm 16 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23">16</figureCitation>
) similar to male except: frons with an inverted Y-shaped suture between eyes; setation of dorsum less dense except for scutellar shield; cuticle of humeri and elytral bases sometimes paler than disc; tarsi less densely setose; abdominal ventrite 3 without a dense patch of yellow setae.
<emphasis id="C93873A017B7E10272D5B20CF1933120" bold="true" italics="true" pageId="0" pageNumber="23">Ovipositor</emphasis>
: (Fig.
<figureCitation id="4F8FAA19C7440A32A845AE41CFD8A238" captionStart="Figures 15, 16" captionStartId="F8" captionText="Figures 15, 16. Tychepsephus felix, female 15 habitus a dorsal view b ventral view; length 4.2 mm 16 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23">16</figureCitation>
) with baculus nearly twice as long as gonocoxite; baculus almost twice as long as wide, strap-like, wider apically; each proximal gonocoxite triangular, distal gonocoxites separate basally then converging to meet apically, apices obliquely truncate; each gonostylus long and narrow, half as long as distal gonocoxite.
</paragraph>
</subSubSection>
<subSubSection id="AFA4B304247E478979E8BA47D464C254" pageId="0" pageNumber="23" type="variation">
<paragraph id="44D456F1FCD40F80CDE1B2F63AB3A492" pageId="0" pageNumber="23">Variation.</paragraph>
<paragraph id="FDF10E0BB46DBF18C1A61AECC83BAB46" pageId="0" pageNumber="23">
The sizes of males and females overlap: males, 4.6-5.2 mm long (
<emphasis id="778205251F6B36E59437283972E28AC7" italics="true" pageId="0" pageNumber="23">n</emphasis>
= 6), 2.8-3.5 mm wide (
<emphasis id="C5C011B6F550CF3747CFF5C587000C5B" italics="true" pageId="0" pageNumber="23">n</emphasis>
= 5); females 4.3-5.7 mm long (
<emphasis id="5BCC68435C38F7B90424C6820B64F8E2" italics="true" pageId="0" pageNumber="23">n</emphasis>
= 10), 2.9-3.7 wide (
<emphasis id="9ACA0EE962DCDA6E7CFEE700688AE6C1" italics="true" pageId="0" pageNumber="23">n</emphasis>
= 5); but the largest specimens are female. Most males (Fig.
<figureCitation id="77D9B52E5EA6BB0C89361CE31A51DBB3" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">13a</figureCitation>
) have an elytral pattern with moderately dense setae surrounding mostly glabrous areas; most females (Fig.
<figureCitation id="D410F14BC958152CC6791F49FDE35091" captionStart="Figures 15, 16" captionStartId="F8" captionText="Figures 15, 16. Tychepsephus felix, female 15 habitus a dorsal view b ventral view; length 4.2 mm 16 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23">15a</figureCitation>
) have no pattern and are more sparsely setose. Females (Fig.
<figureCitation id="9605E289680AACD69E54462E9FA52A3E" captionStart="Figures 15, 16" captionStartId="F8" captionText="Figures 15, 16. Tychepsephus felix, female 15 habitus a dorsal view b ventral view; length 4.2 mm 16 ovipositor, dorsal view." figureDoi="10.3897/zookeys.1164.103184.figures15-16" httpUri="https://binary.pensoft.net/fig/855192" pageId="0" pageNumber="23">15b</figureCitation>
) lack the dense patch of yellow setae on abdominal ventrite 3 that is present in males (Fig.
<figureCitation id="5076DF703B7BDE4441617A4BC75DA82A" captionStart="Figures 13, 14" captionStartId="F7" captionText="Figures 13, 14. Tychepsephus felix, male 13 habitus a dorsal view b ventral view; length 5.0 mm 14 aedeagus a dorsal view b lateral view c ventral view." figureDoi="10.3897/zookeys.1164.103184.figures13-14" httpUri="https://binary.pensoft.net/fig/855191" pageId="0" pageNumber="23">13b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="462125DDF7FD7B0035AC721062E11E40" pageId="0" pageNumber="23" type="etymology">
<paragraph id="E6E97A3B89796BB4E59A73A3DA078D18" pageId="0" pageNumber="23">Etymology.</paragraph>
<paragraph id="76BF427476531F3A34E5271BC89AD0C2" pageId="0" pageNumber="23">
<bibRefCitation id="6B0CF068CA2A88D481847F4615922B95" DOI="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" author="Waterhouse, CO" journalOrPublisher="Transactions of the Royal Entomological Society" pageId="0" pageNumber="23" pagination="11 - 25" refId="B25" refString="Waterhouse, CO, 1876. III. On various new genera and species of Coleoptera. Transactions of the Royal Entomological Society 1876 (1): 11 - 25, DOI: https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" title="III. On various new genera and species of Coleoptera." url="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" volume="1876" year="1876">Waterhouse (1876)</bibRefCitation>
did not explain the trivial name. However,
<emphasis id="2F7B7EDF8FBB10A69E9062C860E567C3" italics="true" pageId="0" pageNumber="23">felix</emphasis>
(L.) means happy or prosperous.
</paragraph>
</subSubSection>
<subSubSection id="6F645B5CC76062C1476E8A5CCB8BA7A2" pageId="0" pageNumber="23" type="distribution">
<paragraph id="B549563E36007A5F41E3FC722FAC41F7" pageId="0" pageNumber="23">Geographic distribution.</paragraph>
<paragraph id="2E091A185E56A02FE5D0A993F902B4A7" pageId="0" pageNumber="23">
<taxonomicName id="40D91F6B254B4186634B39DD7396A6B5" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">
<emphasis id="A1CD3A2C74F053C509B5F034EFE6CD98" italics="true" pageId="0" pageNumber="23">Tychepsephus felix</emphasis>
</taxonomicName>
is known only from Chile. Adults have been collected mainly in
<normalizedToken id="B27A75A3832D27046CDF09ECC16449EE" originalValue="Región">Region</normalizedToken>
X (Los Lagos), but also in regions VIII (
<normalizedToken id="407B559693A8E13BC9C3E3977D65D804" originalValue="Bío">Bio</normalizedToken>
<normalizedToken id="C7EAC19C65D3F48AC205C75F4179CEA9" originalValue="Bío">Bio</normalizedToken>
), IX (
<normalizedToken id="03644324BDA48B1348E30EA5AF1100AA" originalValue="Araucanía">Araucania</normalizedToken>
), and XIV (Los
<normalizedToken id="5E9FCFFAE6E39335ECF65F9491F1CE9B" originalValue="Ríos">Rios</normalizedToken>
) (M. Elgueta, in litt.), in both the Andean and the coastal mountain areas (Fig.
<figureCitation id="75F5A75203F5176279210F857B485E20" captionStart="Figure 7" captionStartId="F4" captionText="Figure 7. Map of Chile with enlarged section showing the geographic distribution of species-verified adult Tychepsephus specimens: T. cekalovici, yellow triangles; T. felix, violet squares; both species, red circles." figureDoi="10.3897/zookeys.1164.103184.figure7" httpUri="https://binary.pensoft.net/fig/855188" pageId="0" pageNumber="23">7</figureCitation>
). The greatest number of adults collected by the authors was at
<normalizedToken id="60E6962DB4DD798BCEE2775DA56512CD" originalValue="Río">Rio</normalizedToken>
Colegual west of Puerto Varas (Fig.
<figureCitation id="2F480F96C6C555155936D89EDF390FDD" captionStart="Figures 8–11" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C86B03B09EC02662B127E0BC38FDB992" pageId="0" pageNumber="23" type="habitat">
<paragraph id="E4ADB3861927E24B3C5F9792EF0D578C" pageId="0" pageNumber="23">Habitat.</paragraph>
<paragraph id="3AFCDBC071EB9C1A375A14DFBFEA1EB7" pageId="0" pageNumber="23">
<taxonomicName id="9C1FD5897D36A8503DCE17F5E1A4116F" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">
<emphasis id="CED323064A6C93F3F66F5B491B52EAED" italics="true" pageId="0" pageNumber="23">Tychepsephus felix</emphasis>
</taxonomicName>
adults were found in habitats as described for the genus. Specimens were collected by sweeping marginal vegetation along streams during the austral summer (Fig.
<figureCitation id="D4297B68CCB216B76B09CFED616EEA31" captionStart="Figures 8–11" captionStartId="F5" captionText="Figures 8 - 11. Tychepsephus habitat 8 Rio Colegual, T. cekalovici type locality a stream with riffle-pool morphology flowing through a mixture of forest and pasture land b streamside shrubby vegetation, adult habitat 9 Rio Contaco 10 stream at Puente El Avion 11 tributary Rio Blanco." figureDoi="10.3897/zookeys.1164.103184.figures8-11" httpUri="https://binary.pensoft.net/fig/855189" pageId="0" pageNumber="23">8b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8440263700DA63D3F04983F56A7A7CE1" pageId="0" pageNumber="23" type="associated dryopoid taxa">
<paragraph id="0EBCD29F01227A9927D2D32AAF0E800D" pageId="0" pageNumber="23">Associated dryopoid taxa.</paragraph>
<paragraph id="2CC06A509F28B05BF5C1298458A42F4F" pageId="0" pageNumber="23">
<taxonomicName id="95FDEE7A739FE5A23079B6205DA637AF" authorityName="Curtis" authorityYear="1830" class="Insecta" family="Elmidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="family">Elmidae</taxonomicName>
:
<taxonomicName id="0A58EDA34671761875221257F8D011EC" authorityName="LeConte" authorityYear="1861" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Larainae">Larainae</taxonomicName>
:
<taxonomicName id="B2EA8C1F71A6D58BB945FA2F27EDF4F4" class="Insecta" family="Elmidae" genus="Hydora" kingdom="Animalia" lsidName="Hydora annectans" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="annectans">
<emphasis id="2F771C9003929B38A128B42E0D8B675F" italics="true" pageId="0" pageNumber="23">Hydora annectans</emphasis>
</taxonomicName>
Spangler &amp; Brown, 1981,
<taxonomicName id="72C60358D1A3D595AD9F0392B0776F7B" lsidName="H. lenta" pageId="0" pageNumber="23" rank="species" species="lenta">
<emphasis id="C3A39423FD045BDCC107293FB7FD0E61" italics="true" pageId="0" pageNumber="23">H. lenta</emphasis>
</taxonomicName>
Spangler &amp; Brown, 1981;
<taxonomicName id="61E8825AC34CA26AB71C1C867E157F3E" authorityName="Curtis" authorityYear="1830" lsidName="" pageId="0" pageNumber="23" rank="subFamily" subFamily="Elminae">Elminae</taxonomicName>
:
<taxonomicName id="60E5C2E473CAD8FDEF9B8C35F333583D" authorityName="Carter &amp; Zeck" authorityYear="1929" class="Insecta" family="Elmidae" genus="Austrolimnius" kingdom="Animalia" lsidName="Austrolimnius" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="399DDBC2FC35942760B02E96CF3D942E" italics="true" pageId="0" pageNumber="23">Austrolimnius</emphasis>
</taxonomicName>
Carter &amp; Zeck, 1929,
<taxonomicName id="154B5EB27D5C320F5FC0C56E934B4A1E" authorityName="Spangler &amp; Staines" authorityYear="2004" class="Insecta" family="Elmidae" genus="Luchoelmis" kingdom="Animalia" lsidName="Luchoelmis" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="B0E9F567B289A913620656583F4EF64D" italics="true" pageId="0" pageNumber="23">Luchoelmis</emphasis>
</taxonomicName>
Spangler &amp; Staines, 2004,
<taxonomicName id="E144F1A58809D44F1B40EF4417F71553" class="Insecta" family="Elmidae" genus="Neoelmis" kingdom="Animalia" lsidName="Neoelmis sissicollis" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="sissicollis">
<emphasis id="59B58EE3D989FB9F21F608A0B8F327FC" italics="true" pageId="0" pageNumber="23">Neoelmis sissicollis</emphasis>
</taxonomicName>
(Germain, 1892). Both
<taxonomicName id="3AE79BC98740D8ECD369059D197DB509" class="Insecta" kingdom="Animalia" lsidName="T. felix" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="45C50BBB39468BBF8867D67A28EA5BF1" italics="true" pageId="0" pageNumber="23">T. felix</emphasis>
</taxonomicName>
and
<taxonomicName id="17F5DFF17C6D1BE031858D5C10F6DB31" class="Insecta" kingdom="Animalia" lsidName="T. cekalovici" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="cekalovici">
<emphasis id="90DEF85611EC08ED728F35E9FBAECD5B" italics="true" pageId="0" pageNumber="23">T. cekalovici</emphasis>
</taxonomicName>
sp. nov. occurred at
<normalizedToken id="C4D50E086542332497C2DA9076FB2B66" originalValue="Río">Rio</normalizedToken>
Colegual.
</paragraph>
</subSubSection>
<subSubSection id="4DCC3A50B57CBA61097D30BFFDA39822" pageId="0" pageNumber="23" type="type remarks">
<paragraph id="02A7480650CC184630D5A6E34F79852D" pageId="0" pageNumber="23">Type remarks.</paragraph>
<paragraph id="4477C35363BE4DA6D070F696E05827C7" pageId="0" pageNumber="23">
The species was described by
<bibRefCitation id="3B5685F672251B0A491A359882DA349A" DOI="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" author="Waterhouse, CO" journalOrPublisher="Transactions of the Royal Entomological Society" pageId="0" pageNumber="23" pagination="11 - 25" refId="B25" refString="Waterhouse, CO, 1876. III. On various new genera and species of Coleoptera. Transactions of the Royal Entomological Society 1876 (1): 11 - 25, DOI: https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" title="III. On various new genera and species of Coleoptera." url="https://doi.org/10.1111/j.1365-2311.1876.tb01122.x" volume="1876" year="1876">Waterhouse (1876)</bibRefCitation>
from a single female type (Fig.
<figureCitation id="0B620A2EFE5A8268F7C8B4D0C7B107F1" captionStart="Figure 12" captionStartId="F6" captionText="Figure 12. Tychepsephus felix, female type specimen a dorsal view b ventral view c specimen data labels. Images provided by Keita Matsumoto (NHMUK)." figureDoi="10.3897/zookeys.1164.103184.figure12" httpUri="https://binary.pensoft.net/fig/855190" pageId="0" pageNumber="23">12</figureCitation>
) which is in the NHMUK along with eight non-type specimens subsequently acquired. The type was originally pinned, breaking the right elytron just posterior to the pin. Subsequently, the specimen was removed from the pin and card-mounted. Missing parts include the following: left antenna without antennomeres 3-11; left foreleg; right hind leg; tarsi on all legs except left hind leg; and tarsomere 5 on the left hind leg. One leg, without the tarsus, is in a gelatin capsule pinned below the carded specimen.
</paragraph>
<paragraph id="FBFF0181FD9AA341E0ADF76D8B3F0F75" pageId="0" pageNumber="23">
The female type of
<taxonomicName id="A0CA91EADE3F347C8CECACF391DAF00C" class="Insecta" family="Psephenidae" genus="Ectopria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectopria (Chilectopria) grandis" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="grandis" subGenus="Chilectopria">Ectopria (Chilectopria) grandis</taxonomicName>
Pic, 1947 (Fig.
<figureCitation id="E61680EF6AFB6FFE6AA20BFD8459E34D" captionStart="Figure 17" captionStartId="F9" captionText="Figure 17. Chilectopria grandis, female type specimen a dorsal view b lateral view c ventral view d specimen data labels. Images provided by Christophe Rivier (MNHN)." figureDoi="10.3897/zookeys.1164.103184.figure17" httpUri="https://binary.pensoft.net/fig/855193" pageId="0" pageNumber="23">17</figureCitation>
), housed at the MNHN, was examined and found to be synonymous with
<taxonomicName id="958AA674FF347968A0CCBE0A2E8A946B" authorityName="Waterhouse" authorityYear="1876" family="Psephenidae" genus="Tychepsephus" kingdom="Animalia" lsidName="Tychepsephus felix" order="Coleoptera" pageId="0" pageNumber="23" rank="species" species="felix">
<emphasis id="440FACF93B39610466777EB5EDEE1B83" italics="true" pageId="0" pageNumber="23">Tychepsephus felix</emphasis>
</taxonomicName>
Waterhouse, 1876. Pic described
<taxonomicName id="37D5AB4E56D5B9CA02682F6B37A0293B" authorityName="Pic" authorityYear="1947" class="Insecta" family="Psephenidae" genus="Chilectopria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chilectopria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="E7B974E31222A5A723CBE064F0438124" italics="true" pageId="0" pageNumber="23">Chilectopria</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName id="F21E2F5F28A616AF7890BA15993391D4" authorityName="LeConte" authorityYear="1853" class="Insecta" family="Psephenidae" genus="Ectopria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectopria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="F0859347B43E4F5B97DF85A8F326B01C" italics="true" pageId="0" pageNumber="23">Ectopria</emphasis>
</taxonomicName>
: &quot;
<taxonomicName id="4E302F98906B6F5F7B415CE93DF5EE5D" authorityName="Pic" authorityYear="1947" class="Insecta" family="Psephenidae" genus="Chilectopria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chilectopria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="9065086A6A7091880025CCD262907BF0" italics="true" pageId="0" pageNumber="23">Chilectopria</emphasis>
</taxonomicName>
s. g. de
<taxonomicName id="72381B84BC98D4441848BB2FAF021467" class="Insecta" family="Psephenidae" genus="Ectopria." kingdom="Animalia" lsidName="Ectopria." order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="2EFCC1BA4C4987113630D590EBD48BF5" italics="true" pageId="0" pageNumber="23">Ectopria.</emphasis>
</taxonomicName>
&quot; Subsequently,
<taxonomicName id="28B1BEC1B9A7B71749C2DB136D1FD5AA" authorityName="Pic" authorityYear="1947" class="Insecta" family="Psephenidae" genus="Chilectopria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chilectopria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="596106EFED09580F643741C0E7CB4DDA" italics="true" pageId="0" pageNumber="23">Chilectopria</emphasis>
</taxonomicName>
sometimes has been cited incorrectly as a genus, rather than a subgenus, likely because the heading and description occur on different pages (
<bibRefCitation id="6DEAD54BDE71A60D39502ABD3C5CAF0F" DOI="https://doi.org/10.5962/bhl.title.107929" author="Pic, M" journalOrPublisher="Moulins" pageId="0" pageNumber="23" pagination="1 - 16" refId="B18" refString="Pic, M, 1947. Diversites entomologiques (10 Juin 1947). Les Impremeries Reunies. Moulins 1: 1 - 16, DOI: https://doi.org/10.5962/bhl.title.107929" title="Diversites entomologiques (10 Juin 1947). Les Impremeries Reunies." url="https://doi.org/10.5962/bhl.title.107929" volume="1" year="1947">Pic 1947</bibRefCitation>
: 3-4). Contributing to this error, the type specimen also lacks &quot;
<taxonomicName id="9AA5161482285888172CB6E287380434" authorityName="LeConte" authorityYear="1853" class="Insecta" family="Psephenidae" genus="Ectopria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectopria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="244D73393E3B80421EB56C7D912069E4" italics="true" pageId="0" pageNumber="23">Ectopria</emphasis>
</taxonomicName>
&quot; on the identification label. The genus
<taxonomicName id="6D74921459B21241C57E2F98E8B24753" authorityName="LeConte" authorityYear="1853" class="Insecta" family="Psephenidae" genus="Ectopria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ectopria" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis id="8162DE48146798C0C63CF5B11408E0A5" italics="true" pageId="0" pageNumber="23">Ectopria</emphasis>
</taxonomicName>
is a Holarctic element.
</paragraph>
<caption id="88342AAEE14552C3E8EE31C8C276E442" doi="10.3897/zookeys.1164.103184.figure17" httpUri="https://binary.pensoft.net/fig/855193" pageId="0" pageNumber="23" start="Figure 17" startId="F9">
<paragraph id="D885FA8BEB0324DFA546544B65577F33" pageId="0" pageNumber="23">
<emphasis id="32233F2C78F000542C84723643EB225A" bold="true" pageId="0" pageNumber="23">Figure 17.</emphasis>
<taxonomicName id="8BA2E9E6D39412F048949D909F896FA7" class="Insecta" family="Psephenidae" genus="Chilectopria" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Chilectopria grandis" order="Coleoptera" pageId="0" pageNumber="23" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="2A3CAF3308865D8197383458484C7407" italics="true" pageId="0" pageNumber="23">Chilectopria grandis</emphasis>
</taxonomicName>
, female type specimen
<emphasis id="4F5A043D5FAF8149C1E4DF619805BBA3" bold="true" pageId="0" pageNumber="23">a</emphasis>
dorsal view
<emphasis id="A894BEBE19C54FBE18CDB65ADBE16F15" bold="true" pageId="0" pageNumber="23">b</emphasis>
lateral view
<emphasis id="8271B91E5E6B63F04E11426B2C8D5FDC" bold="true" pageId="0" pageNumber="23">c</emphasis>
ventral view
<emphasis id="6DDD33CDE986640E3893A729F086BB65" bold="true" pageId="0" pageNumber="23">d</emphasis>
specimen data labels. Images provided by Christophe Rivier (MNHN).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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