177 lines
21 KiB
XML
177 lines
21 KiB
XML
<document id="0A2BC0287D2D8EE691549F02DC621AD3" ID-CLB-Dataset="55566" ID-DOI="10.5281/zenodo.5305057" ID-GBIF-Dataset="c7ccba22-dcb8-4e60-b5fb-a8439522e453" ID-ISSN="0253-116X" ID-Zenodo-Dep="5305057" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630111141298" checkinUser="marcus" docAuthor="Assing, V." docDate="2014" docId="184887E9FFF2BD581F9CA9A3933D025C" docLanguage="en" docName="LinzerbiolBeitr.2014.46.1.461-479.pdf" docOrigin="Linzer biologische Beiträge 46 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Orsunius cuneatus Assing 2014, nov.sp." docType="treatment" docVersion="9" lastPageNumber="474" masterDocId="E471FF91FFFFBD551F2EA97490250160" masterDocTitle="On Orsunius II. Eight new species and additional records (Coleoptera: Staphylinidae: Paederinae: Medonina)" masterLastPageNumber="479" masterPageNumber="461" pageNumber="474" updateTime="1699155222530" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1E5745946422C822A5C95944B475DFC6">On Orsunius II. Eight new species and additional records (Coleoptera: Staphylinidae: Paederinae: Medonina)</mods:title>
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<mods:name id="548503F681AB7028DC05F780D69A76CA" type="personal">
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<mods:namePart id="CE87BB5EFB2A624D46B516622E063758">Assing, V.</mods:namePart>
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<mods:title id="2E9D6DF24369E7FC8C4883220816D33A">Linzer biologische Beiträge</mods:title>
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<mods:part id="1CD322F4C541E5C12A3E4509C4B033E0">
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<mods:date id="70CD66CF9A764EAB003689671DE54C6B">2014</mods:date>
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<mods:number id="5A349FC2F7978CA3395D653BFB02A4BF">2014-07-31</mods:number>
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<mods:number id="91519AA09BD51A44BA29F10D0585C49F">46</mods:number>
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<mods:identifier id="91F9EB784335E05638DA10912F3E3FB4" type="DOI">10.5281/zenodo.5305057</mods:identifier>
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<treatment id="184887E9FFF2BD581F9CA9A3933D025C" ID-DOI="http://doi.org/10.5281/zenodo.6539889" ID-GBIF-Taxon="195300190" ID-Zenodo-Dep="6539889" LSID="urn:lsid:plazi:treatment:184887E9FFF2BD581F9CA9A3933D025C" httpUri="http://treatment.plazi.org/id/184887E9FFF2BD581F9CA9A3933D025C" lastPageNumber="474" pageId="13" pageNumber="474">
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<subSubSection id="D8FB6574FFF2BD581F9CA9A391FF018E" box="[178,474,215,238]" pageId="13" pageNumber="474" type="nomenclature">
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<paragraph id="905E36FFFFF2BD581F9CA9A391FF018E" blockId="13.[178,773,215,238]" box="[178,474,215,238]" pageId="13" pageNumber="474">
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<heading id="CB168193FFF2BD581F9CA9A391FF018E" bold="true" box="[178,474,215,238]" fontSize="9" level="2" pageId="13" pageNumber="474" reason="3">
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<emphasis id="A295EAEDFFF2BD581F9CA9A391FF018E" bold="true" box="[178,474,215,238]" pageId="13" pageNumber="474">
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<taxonomicName id="57E14D7CFFF2BD581F9CA9A391A6018E" ID-CoL="74XHT" authority="Assing, 2014" authorityName="Assing" authorityYear="2014" box="[178,387,215,238]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="474" phylum="Arthropoda" rank="species" species="cuneatus" status="sp. nov.">
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<emphasis id="A295EAEDFFF2BD581F9CA9A391A6018E" bold="true" box="[178,387,215,238]" italics="true" pageId="13" pageNumber="474">Orsunius cuneatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B9A65796FFF2BD581EA4A9A391FF018E" box="[394,474,215,238]" pageId="13" pageNumber="474" rank="species">nov.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8FB6574FFF2BD581ECFA9A39320018E" box="[481,773,215,238]" pageId="13" pageNumber="474" type="description">
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<paragraph id="905E36FFFFF2BD581ECFA9A39320018E" blockId="13.[178,773,215,238]" box="[481,773,215,238]" pageId="13" pageNumber="474">
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<heading id="CB168193FFF2BD581ECFA9A39320018E" bold="true" box="[481,773,215,238]" fontSize="9" level="2" pageId="13" pageNumber="474" reason="3">
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(
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<figureCitation id="08DA2A7AFFF2BD581EC6A9A39245018E" box="[488,608,215,238]" captionStart="Figs 36-43" captionStartId="14.[178,218,1484,1504]" captionTargetBox="[192,1078,212,1464]" captionTargetId="figure-2@14.[178,1091,212,1474]" captionTargetPageId="14" captionText="Figs 36-43: Orsunius cuneatus nov.sp. (36-39) and O. heissi nov.sp. (40-43): (36, 40) habitus; (37, 41) forebody; (38, 42) aedeagus in lateral view; (39, 43) aedeagus in ventral view. Scale bars: 36- 37, 40-41: 1.0 mm; 38-39, 42-43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5305071" httpUri="https://zenodo.org/record/5305071/files/figure.png" pageId="13" pageNumber="474">Figs 36-39</figureCitation>
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,
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<figureCitation id="08DA2A7AFFF2BD581D45A9A39289018E" box="[619,684,215,238]" captionStart="Figs 44-48" captionStartId="15.[178,218,779,799]" captionTargetBox="[196,1048,218,767]" captionTargetId="figure-2@15.[178,1091,212,768]" captionTargetPageId="15" captionText="Figs 44-48: Orsunius cuneatus nov.sp. (44-46) and O. heissi nov.sp. (47-48): (44) male sternite VII; (45, 47) male sternite VIII; (46, 48) posterior portion of male sternite VIII. Scale bars: 44-45, 47: 0.2 mm; 46, 48: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305073" httpUri="https://zenodo.org/record/5305073/files/figure.png" pageId="13" pageNumber="474">44-46</figureCitation>
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,
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<figureCitation id="08DA2A7AFFF2BD581D99A9A392D9018E" box="[695,764,215,238]" captionStart="Map 2" captionStartId="2.[178,223,1075,1095]" captionTargetBox="[178,1092,212,1064]" captionTargetId="figure-2@2.[178,1092,212,1064]" captionTargetPageId="2" captionText="Map 2: Distributions of Orsunius gladiator (white and black circles), O. latexcisus (black circle), O. flavovirgatus (black circle), O. heissi (black star), O. stimulans (black and grey triangles), O. granulosus (white and grey triangles), O. cuneatus (white diamond), and O. spathulatus (black square)." figureDoi="http://doi.org/10.5281/zenodo.5305061" httpUri="https://zenodo.org/record/5305061/files/figure.png" pageId="13" pageNumber="474">Map 2</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8FB6574FFF2BD581F9CA98B93FE0020" pageId="13" pageNumber="474" type="materials_examined">
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<paragraph id="905E36FFFFF2BD581F9CA98B93FE0020" blockId="13.[178,1086,255,320]" pageId="13" pageNumber="474">
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Type material
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<materialsCitation id="20893CA2FFF2BD581E82A98B91AC0020" ID-GBIF-Occurrence="3766395309" collectingDate="2001-04-24" collectingDateMax="2001-04-29" collectingDateMin="2001-04-24" collectionCode="NHMB" collectorName="Pacholatko" country="Laos-C" determinerName="V. Assing" elevation="200" latitude="18.116667" location="Ban Khoun Ngeun" longLatPrecision="1276" longitude="104.48333" municipality="Ban Khoun Ngeun" pageId="13" pageNumber="474" specimenCount="1" stateProvince="Khammouan" typeStatus="holotype">
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<typeStatus id="4F5A885DFFF2BD581E82A98B91DD0073" box="[428,504,255,275]" pageId="13" pageNumber="474" type="holotype">Holotype</typeStatus>
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: "
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<collectingCountry id="E8F6766FFFF2BD581D1DA98B925F0073" box="[563,634,255,275]" pageId="13" pageNumber="474">Laos-C</collectingCountry>
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,
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<collectingRegion id="5225F81DFFF2BD581DA9A98B92D90073" box="[647,764,255,275]" country="Laos" name="Khammouan" pageId="13" pageNumber="474">Khammouan</collectingRegion>
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pr.,
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<collectingMunicipality id="703AAC85FFF2BD581C03A98B93C40073" box="[813,993,255,275]" pageId="13" pageNumber="474">Ban Khoun Ngeun</collectingMunicipality>
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, ~
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<quantity id="57199B1AFFF2BD581CD7A98B941C0073" box="[1017,1081,255,275]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="474" unit="m" value="200.0">
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<elevation id="1BCCD1CCFFF2BD581CD7A98B941C0073" box="[1017,1081,255,275]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="474" unit="m" value="200.0">200 m</elevation>
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</quantity>
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,
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<geoCoordinate id="F5D55038FFF2BD581F9CA86290D8004A" box="[178,253,278,298]" degrees="18" direction="north" minutes="07" orientation="latitude" pageId="13" pageNumber="474" precision="925" value="18.116667">18°07'N</geoCoordinate>
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<geoCoordinate id="F5D55038FFF2BD581E2AA862917C004A" box="[260,345,278,298]" degrees="104" direction="east" minutes="29" orientation="longitude" pageId="13" pageNumber="474" precision="925" value="104.48333">104°29'E</geoCoordinate>
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,
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<collectorName id="3D145329FFF2BD581E4AA86291EC004A" box="[356,457,278,298]" pageId="13" pageNumber="474">Pacholátko</collectorName>
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leg.,
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<date id="E45F103FFFF2BD581ED0A86292A7004A" box="[510,642,278,298]" pageId="13" pageNumber="474" value="2001-04-24" valueMax="2001-04-29" valueMin="2001-04-24">
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<collectingDate id="F41BE9D7FFF2BD581ED0A86292A7004A" box="[510,642,278,298]" pageId="13" pageNumber="474" value="2001-04-24" valueMax="2001-04-29" valueMin="2001-04-24">24.-29.iv.2001</collectingDate>
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</date>
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/ Holotypus
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<originalDetermination id="B3FF8E03FFF2BD581C32A86293DC004A" box="[796,1017,278,298]" pageId="13" pageNumber="474">
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<taxonomicName id="57E14D7CFFF2BD581C32A86293E2004A" authority="Assing, 2014" authorityName="Assing" authorityYear="2014" box="[796,967,278,298]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="474" phylum="Arthropoda" rank="species" species="cuneatus" status="sp. nov.">
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<emphasis id="A295EAEDFFF2BD581C32A86293E2004A" box="[796,967,278,298]" italics="true" pageId="13" pageNumber="474">Orsunius cuneatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B9A65796FFF2BD581CE3A86293DC004A" box="[973,1017,278,298]" pageId="13" pageNumber="474" rank="species">sp.n.</taxonomicNameLabel>
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</originalDetermination>
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det.
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<determinerName id="6582DA3BFFF2BD581B09A86290D40020" pageId="13" pageNumber="474">V. Assing</determinerName>
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2013" (
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<collectionCode id="F6F0AE3AFFF2BD581E14A85891A60020" box="[314,387,300,320]" pageId="13" pageNumber="474">NHMB</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="20893CA2FFF2BD581EBCA85893FE0020" ID-GBIF-Occurrence="3766395308" box="[402,987,300,320]" collectingDate="2001-04-24" collectingDateMax="2001-04-29" collectingDateMin="2001-04-24" collectionCode="NHMB, cAss" collectorName="Pacholatko" country="Laos-C" determinerName="V. Assing" elevation="200" latitude="18.116667" location="Ban Khoun Ngeun" longLatPrecision="1276" longitude="104.48333" municipality="Ban Khoun Ngeun" pageId="13" pageNumber="474" specimenCount="6" stateProvince="Khammouan" typeStatus="paratype">
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<typeStatus id="4F5A885DFFF2BD581EBCA85891CF0020" box="[402,490,300,320]" pageId="13" pageNumber="474" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="86E7FD76FFF2BD581ED8A85892240020" box="[502,513,300,320]" pageId="13" pageNumber="474">6</specimenCount>
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, 12: same data as holotype (
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<collectionCode id="F6F0AE3AFFF2BD581C7EA85893BD0020" box="[848,920,300,320]" pageId="13" pageNumber="474">NHMB</collectionCode>
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,
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<collectionCode id="F6F0AE3AFFF2BD581C8FA85893F40020" box="[929,977,300,320]" pageId="13" pageNumber="474">cAss</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8FB6574FFF2BD581F9CA83B933000E2" pageId="13" pageNumber="474" type="etymology">
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<paragraph id="905E36FFFFF2BD581F9CA83B933000E2" blockId="13.[178,1086,335,386]" pageId="13" pageNumber="474">Etymology: The specific epithet (Latin, adjective: wedge-shaped) refers to the shape of the ventral process of the aedeagus in ventral view.</paragraph>
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</subSubSection>
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<subSubSection id="D8FB6574FFF2BD581F9CA8E5939B0321" pageId="13" pageNumber="474" type="description">
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<paragraph id="905E36FFFFF2BD581F9CA8E5938800C7" blockId="13.[178,941,401,423]" box="[178,941,401,423]" pageId="13" pageNumber="474">
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Description: External characters (
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<figureCitation id="08DA2A7AFFF2BD581D48A8E592F000C7" box="[614,725,401,423]" captionStart="Figs 36-43" captionStartId="14.[178,218,1484,1504]" captionTargetBox="[192,1078,212,1464]" captionTargetId="figure-2@14.[178,1091,212,1474]" captionTargetPageId="14" captionText="Figs 36-43: Orsunius cuneatus nov.sp. (36-39) and O. heissi nov.sp. (40-43): (36, 40) habitus; (37, 41) forebody; (38, 42) aedeagus in lateral view; (39, 43) aedeagus in ventral view. Scale bars: 36- 37, 40-41: 1.0 mm; 38-39, 42-43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5305071" httpUri="https://zenodo.org/record/5305071/files/figure.png" pageId="13" pageNumber="474">Figs 36-37</figureCitation>
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) as in
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<taxonomicName id="57E14D7CFFF2BD581C36A8E6938200C7" authorityName="Assing" authorityYear="2014" box="[792,935,401,423]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="474" phylum="Arthropoda" rank="species" species="granulosus">
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<emphasis id="A295EAEDFFF2BD581C36A8E6938200C7" box="[792,935,401,423]" italics="true" pageId="13" pageNumber="474">O. granulosus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="905E36FFFFF2BD581FE9A8C3939B0321" blockId="13.[178,1086,438,578]" pageId="13" pageNumber="474">
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: sternite VII (
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<figureCitation id="08DA2A7AFFF2BD581E4CA8C2919500AC" box="[354,432,438,460]" captionStart="Figs 44-48" captionStartId="15.[178,218,779,799]" captionTargetBox="[196,1048,218,767]" captionTargetId="figure-2@15.[178,1091,212,768]" captionTargetPageId="15" captionText="Figs 44-48: Orsunius cuneatus nov.sp. (44-46) and O. heissi nov.sp. (47-48): (44) male sternite VII; (45, 47) male sternite VIII; (46, 48) posterior portion of male sternite VIII. Scale bars: 44-45, 47: 0.2 mm; 46, 48: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305073" httpUri="https://zenodo.org/record/5305073/files/figure.png" pageId="13" pageNumber="474">Fig. 44</figureCitation>
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) of similar shape as in
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<taxonomicName id="57E14D7CFFF2BD581D9CA8C3936100AC" authorityName="Assing" authorityYear="2014" box="[690,836,438,460]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="474" phylum="Arthropoda" rank="species" species="granulosus">
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<emphasis id="A295EAEDFFF2BD581D9CA8C3936100AC" box="[690,836,438,460]" italics="true" pageId="13" pageNumber="474">O. granulosus</emphasis>
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</taxonomicName>
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; sternite VIII (
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<figureCitation id="08DA2A7AFFF2BD581CC6A8C2941000AC" box="[1000,1077,438,460]" captionStart="Figs 44-48" captionStartId="15.[178,218,779,799]" captionTargetBox="[196,1048,218,767]" captionTargetId="figure-2@15.[178,1091,212,768]" captionTargetPageId="15" captionText="Figs 44-48: Orsunius cuneatus nov.sp. (44-46) and O. heissi nov.sp. (47-48): (44) male sternite VII; (45, 47) male sternite VIII; (46, 48) posterior portion of male sternite VIII. Scale bars: 44-45, 47: 0.2 mm; 46, 48: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305073" httpUri="https://zenodo.org/record/5305073/files/figure.png" pageId="13" pageNumber="474">Fig. 45</figureCitation>
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) distinctly oblong and with distinct posterior excision of nearly semi-circular shape (
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<figureCitation id="08DA2A7AFFF2BD581B3AA8A090EA0367" captionStart="Figs 44-48" captionStartId="15.[178,218,779,799]" captionTargetBox="[196,1048,218,767]" captionTargetId="figure-2@15.[178,1091,212,768]" captionTargetPageId="15" captionText="Figs 44-48: Orsunius cuneatus nov.sp. (44-46) and O. heissi nov.sp. (47-48): (44) male sternite VII; (45, 47) male sternite VIII; (46, 48) posterior portion of male sternite VIII. Scale bars: 44-45, 47: 0.2 mm; 46, 48: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305073" httpUri="https://zenodo.org/record/5305073/files/figure.png" pageId="13" pageNumber="474">Fig. 46</figureCitation>
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); aedeagus (
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<figureCitation id="08DA2A7AFFF2BD581E77A88591F50367" box="[345,464,497,519]" captionStart="Figs 36-43" captionStartId="14.[178,218,1484,1504]" captionTargetBox="[192,1078,212,1464]" captionTargetId="figure-2@14.[178,1091,212,1474]" captionTargetPageId="14" captionText="Figs 36-43: Orsunius cuneatus nov.sp. (36-39) and O. heissi nov.sp. (40-43): (36, 40) habitus; (37, 41) forebody; (38, 42) aedeagus in lateral view; (39, 43) aedeagus in ventral view. Scale bars: 36- 37, 40-41: 1.0 mm; 38-39, 42-43: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5305071" httpUri="https://zenodo.org/record/5305071/files/figure.png" pageId="13" pageNumber="474">Figs 38-39</figureCitation>
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) small, approximately
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<quantity id="57199B1AFFF2BD581DE3A885930B0367" box="[717,814,497,519]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="474" unit="mm" value="0.35">0.35 mm</quantity>
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long; ventral process of acutely triangular shape in ventral view; internal sac with a pair of distinctly sclerotized apical structures and with a pair of short, weakly sclerotized basal structures.
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</paragraph>
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</subSubSection>
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<subSubSection id="D8FB6574FFF2BD581F9CAB2592C60399" pageId="13" pageNumber="474" type="discussion">
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<paragraph id="905E36FFFFF2BD581F9CAB2592C60399" blockId="13.[178,1086,593,761]" pageId="13" pageNumber="474">
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Comparative notes: As can be inferred from the practically identical external characters, including the similarly derived punctation of the forebody, as well as from the similarly derived morphology of the aedeagus,
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<taxonomicName id="57E14D7CFFF2BD581DE4ABF8936303C1" authorityName="Assing" authorityYear="2014" box="[714,838,652,673]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="474" phylum="Arthropoda" rank="species" species="cuneatus">
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<emphasis id="A295EAEDFFF2BD581DE4ABF8936303C1" box="[714,838,652,673]" italics="true" pageId="13" pageNumber="474">O. cuneatus</emphasis>
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</taxonomicName>
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is closely related to
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<taxonomicName id="57E14D7CFFF2BD581B0AABF8910703DF" authorityName="Assing" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="474" phylum="Arthropoda" rank="species" species="granulosus">
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<emphasis id="A295EAEDFFF2BD581B0AABF8910703DF" italics="true" pageId="13" pageNumber="474">O. granulosus</emphasis>
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</taxonomicName>
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, from which it differs by the shape of the posterior excision of the male sternite VIII, by the acutely triangular shape of the ventral process of the aedeagus, and by the shapes of the internal structures of the aedeagus.
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</paragraph>
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</subSubSection>
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<subSubSection id="D8FB6574FFF2BD581F9CAA7C933D025C" pageId="13" pageNumber="474" type="distribution">
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<paragraph id="905E36FFFFF2BD581F9CAA7C933D025C" blockId="13.[178,1085,776,828]" pageId="13" pageNumber="474">
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Distribution and natural history: The
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<typeStatus id="4F5A885DFFF2BD581C19AA7D9346027E" box="[823,867,777,798]" pageId="13" pageNumber="474">type</typeStatus>
|
|
locality is situated in Central
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|
<collectingCountry id="E8F6766FFFF2BD581E2DAA539111025C" box="[259,308,807,828]" name="Laos" pageId="13" pageNumber="474">Laos</collectingCountry>
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(
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<figureCitation id="08DA2A7AFFF2BD581E6FAA5291A0025C" box="[321,389,806,828]" captionStart="Map 2" captionStartId="2.[178,223,1075,1095]" captionTargetBox="[178,1092,212,1064]" captionTargetId="figure-2@2.[178,1092,212,1064]" captionTargetPageId="2" captionText="Map 2: Distributions of Orsunius gladiator (white and black circles), O. latexcisus (black circle), O. flavovirgatus (black circle), O. heissi (black star), O. stimulans (black and grey triangles), O. granulosus (white and grey triangles), O. cuneatus (white diamond), and O. spathulatus (black square)." figureDoi="http://doi.org/10.5281/zenodo.5305061" httpUri="https://zenodo.org/record/5305061/files/figure.png" pageId="13" pageNumber="474">Map 2</figureCitation>
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) at an altitude of approximately
|
|
<specimenCount id="86E7FD76FFF2BD581DFFAA52933D025C" box="[721,792,806,828]" pageId="13" pageNumber="474" type="generic">
|
|
<quantity id="57199B1AFFF2BD581DFFAA529336025C" box="[721,787,806,828]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="474" unit="m" value="200.0">200 m</quantity>
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.
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|
</specimenCount>
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</paragraph>
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|
</subSubSection>
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</treatment>
|
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</document> |