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<mods:title id="A335301FBC5256084BCF260301A10F54">A new classification of Callianassidae and related families (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support</mods:title>
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<paragraph id="8B9D0B4DF24F082CFC80B0C9A919FB4F" blockId="17.[817,1078,1231,1253]" box="[817,1078,1231,1253]" pageId="17" pageNumber="90">
<heading id="D0D5BC21F24F082CFC80B0C9A919FB4F" box="[817,1078,1231,1253]" fontSize="9" level="2" pageId="17" pageNumber="90" reason="5">
<taxonomicName id="4C2270CEF24F082CFC80B0C9AEFBFB4F" ID-CoL="BMK39" authorityName="Poore &amp; Dworschak &amp; Robles &amp; Mantelatto &amp; Felder" authorityYear="2019" box="[817,980,1231,1253]" class="Malacostraca" family="Callianassidae" genus="Aqaballianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="90" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B956D75FF24F082CFC80B0C9AEFBFB4F" bold="true" box="[817,980,1231,1253]" italics="true" pageId="17" pageNumber="90">Aqaballianassa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2656A24F24F082CFC6AB0D6A919FB4F" box="[987,1078,1232,1253]" pageId="17" pageNumber="90" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8B9D0B4DF24F082CFCE6B0FDA888FAA4" blockId="17.[817,1447,1275,1320]" box="[855,1447,1275,1294]" pageId="17" pageNumber="90">http://zoobank.org/urn:lsid:zoobank.org:act:27E2FC97-B393-</paragraph>
</subSubSection>
<subSubSection id="C33858C6F24F082FFC80B113ACA3FCC8" lastPageId="18" lastPageNumber="91" pageId="17" pageNumber="90" type="description">
<paragraph id="8B9D0B4DF24F082CFC80B113A915FA82" blockId="17.[817,1447,1275,1320]" box="[817,1082,1301,1320]" pageId="17" pageNumber="90">48F5-B69F-83AFD8FF49D7</paragraph>
<paragraph id="8B9D0B4DF24F082CFC80B146AED4FAD8" blockId="17.[817,1448,1343,1394]" pageId="17" pageNumber="90">
<emphasis id="B956D75FF24F082CFC80B146AE94FAFF" box="[817,955,1344,1365]" italics="true" pageId="17" pageNumber="90">
<typeStatus id="5499B5EFF24F082CFC80B146AE4FFAFF" box="[817,864,1344,1365]" pageId="17" pageNumber="90">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C2270CEF24F082CFC74B139A8AEFAFE" authority="Dworschak, 2003" authorityName="Dworschak" authorityYear="2003" box="[965,1409,1343,1365]" class="Malacostraca" family="Callianassidae" genus="Callianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="90" phylum="Arthropoda" rank="species" species="aqabaensis">
<emphasis id="B956D75FF24F082CFC74B139A992FAFE" box="[965,1213,1343,1364]" italics="true" pageId="17" pageNumber="90">Callianassa aqabaensis</emphasis>
Dworschak, 2003
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, by present designation.
</paragraph>
<paragraph id="8B9D0B4DF24F082FFC80B18FAC96FDA8" blockId="17.[816,1449,1416,1903]" lastBlockId="18.[140,772,229,514]" lastPageId="18" lastPageNumber="91" pageId="17" pageNumber="90">
<emphasis id="B956D75FF24F082CFC80B18FAEB8FA34" box="[817,919,1417,1438]" italics="true" pageId="17" pageNumber="90">Diagnosis</emphasis>
. Hermaphrodite.
<emphasis id="B956D75FF24F082CFBFBB18EA88EFA11" bold="true" italics="true" pageId="17" pageNumber="90">Triangular sclerite separate from branchiostegite at anterior end of linea thalassinica</emphasis>
. Rostrum obsolete or obtusely triangular, flat, not reaching cornea. Pleomere 1 tergite undivided,
<emphasis id="B956D75FF24F082CFB04B1E6A9E4FA5F" box="[1205,1227,1504,1525]" italics="true" pageId="17" pageNumber="90">or</emphasis>
with weak transverse ridge. Pleomere 6 with sublateral ventral sharp ridge, flared posteriorly. Antennular peduncle articles 2 and 3 with single lateral row of 610 well-spaced long setae along lower margin. Mandibular molar calcified, swollen projection without sharp edge; incisor with few teeth. Maxilliped 3 merus wider at ischium-merus suture than long. Male major cheliped merus with oblique spine about one third to half-way along lower margin; propodus distal margin with small lateral tooth; dactylus with dense setae along upper margin. Minor cheliped two-thirds width of major cheliped, both flattened. Pereopod 3 propodus rectangular, lower margin deeply convex, leading to broadly rounded free proximal lobe. Male pleopod 2 present or absent. Uropodal endopod ovoid, usually longer than wide, anterior margin straight or slightly convex, posterodistal margin evenly convex, with spiniform setae near anterior and distal margins,
<emphasis id="B956D75FF24C082FFE98B526AC10FE9F" box="[297,319,288,309]" italics="true" pageId="18" pageNumber="91">or</emphasis>
with facial spiniform setae on rib. Uropodal exopod 1.01.8 times as long as wide, distal margin clearly differentiated from anterior margin, anterodistal corner right-angled, posterodistal margin with row of 68 long blade-like setae proximal to long setae on distal margin.
<taxonomicName id="4C2270CEF24C082FFD73B593AC24FE62" class="Copepoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="91" phylum="Arthropoda" rank="species" species="anterolateral">Telson anterolateral</taxonomicName>
lobe obsolete, undefined; truncate or slightly convex between posterolateral angles,
<emphasis id="B956D75FF24C082FFD9FB5D6AF6BFE4F" box="[558,580,464,485]" italics="true" pageId="18" pageNumber="91">or</emphasis>
slightly concave, sometimes with medial spine.
</paragraph>
<paragraph id="8B9D0B4DF24C082FFF3DB61FAF21FDE6" blockId="18.[140,771,537,588]" pageId="18" pageNumber="91">
<emphasis id="B956D75FF24C082FFF3DB61FAC2EFD84" box="[140,257,537,558]" italics="true" pageId="18" pageNumber="91">Etymology.</emphasis>
An alliteration of the name of the
<typeStatus id="5499B5EFF24C082FFDEDB61FAFA7FD84" box="[604,648,537,558]" pageId="18" pageNumber="91">type</typeStatus>
species and
<taxonomicName id="4C2270CEF24C082FFF3DB630AC29FDE1" authorityName="Leach" authorityYear="1814" box="[140,262,566,587]" class="Malacostraca" family="Callianassidae" genus="Callianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="B956D75FF24C082FFF3DB630AC29FDE1" box="[140,262,566,587]" italics="true" pageId="18" pageNumber="91">Callianassa</emphasis>
</taxonomicName>
,
<typeStatus id="5499B5EFF24C082FFEA2B631AC10FDE6" box="[275,319,567,588]" pageId="18" pageNumber="91">type</typeStatus>
genus of the family.
</paragraph>
<paragraph id="8B9D0B4DF24C082FFF3DB664ACA3FCC8" blockId="18.[138,771,610,866]" pageId="18" pageNumber="91">
<emphasis id="B956D75FF24C082FFF3DB664ADCAFDDD" box="[140,229,610,631]" italics="true" pageId="18" pageNumber="91">Remarks</emphasis>
. The genus is recognisable by the unique triangular sclerite separated by a clear suture or discontinuity from the branchiostegite at the anterior end of the linea thalassinica. All specimens of
<taxonomicName id="4C2270CEF24C082FFEA8B6BDAC86FD65" authorityName="Dworschak" authorityYear="2003" box="[281,425,698,720]" class="Malacostraca" family="Callianassidae" genus="Callianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="91" phylum="Arthropoda" rank="species" species="aqabaensis">
<emphasis id="B956D75FF24C082FFEA8B6BDAC86FD65" box="[281,425,698,720]" italics="true" pageId="18" pageNumber="91">A. aqabaensis</emphasis>
</taxonomicName>
have male and female gonopores, but pleopods 1 and 2 are sexually dimorphic (Dworschak, 2003;
<bibRefCitation id="EFB376BCF24C082FFF65B6F3AF17FCA1" author="Markham, J. C. &amp; Dworschak, P. C." box="[212,568,757,779]" pageId="18" pageNumber="91" pagination="413 - 419" refId="ref52882" refString="Markham, J. C., and Dworschak, P. C. 2005. A new species of Entophilus Richardson, 1903 (Isopoda: Bopyridae: Entophilinae) from the Gulf of Aqaba, Jordon. Journal of Crustacean Biology 25: 413 - 419. https: // doi. org / 10.1651 / C- 2566" type="journal article" year="2005">Markham and Dworschak, 2005</bibRefCitation>
). Two species are included besides the
<typeStatus id="5499B5EFF24C082FFEE9B714ACABFC8D" box="[344,388,786,807]" pageId="18" pageNumber="91">type</typeStatus>
species,
<taxonomicName id="4C2270CEF24C082FFE6AB714AF43FC8D" authorityName="Poore &amp; Dworschak &amp; Robles &amp; Mantelatto &amp; Felder" authorityYear="2019" baseAuthorityName="Sakai" baseAuthorityYear="2002" box="[475,620,786,807]" class="Malacostraca" family="Callianassidae" genus="Aqaballianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="91" phylum="Arthropoda" rank="species" species="brevirostris">
<emphasis id="B956D75FF24C082FFE6AB714AF43FC8D" box="[475,620,786,807]" italics="true" pageId="18" pageNumber="91">A. brevirostris</emphasis>
</taxonomicName>
from
<collectingCountry id="F3354BDDF24C082FFD16B714AE2DFC82" box="[679,770,786,808]" name="Thailand" pageId="18" pageNumber="91">Thailand</collectingCountry>
and
<taxonomicName id="4C2270CEF24C082FFF07B736AC01FCEE" baseAuthorityName="Ngoc-Ho" baseAuthorityYear="1994" box="[182,302,815,837]" class="Malacostraca" family="Callianassidae" genus="Aqaballianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="91" phylum="Arthropoda" rank="species" species="lewtonae">
<emphasis id="B956D75FF24C082FFF07B736AC01FCEE" box="[182,302,815,837]" italics="true" pageId="18" pageNumber="91">A. lewtonae</emphasis>
</taxonomicName>
from Queensland, plus an undescribed species from
<collectingCountry id="F3354BDDF24C082FFF72B74BACA7FCC8" box="[195,392,844,866]" name="Papua New Guinea" pageId="18" pageNumber="91">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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