treatments-xml/data/A1/76/B5/A176B5C93EBC568B9539914B640AA181.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="A176B5C93EBC568B9539914B640AA181" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana taurina Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="85E80E52-59E3-47E8-BF42-7DFAD573998B" docUuidSource="ZooBank" docVersion="1" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1674309819629" updateUser="pensoft">
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<mods:title>Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart>Cavichioli, Rodney Ramiro</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-01-20</mods:number>
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<taxonomicName LSID="https://zoobank.org/85E80E52-59E3-47E8-BF42-7DFAD573998B" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Coarctana taurina" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="taurina" status="sp. nov.">Coarctana taurina</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">Figs 28</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">, 32I, J</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="diagnosis">
<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
Pygofer (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28G</figureCitation>
) with apex slightly tapered, subacute. Subgenital plate (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28H</figureCitation>
) with small group of filiform setae on apex. Style, in lateral view (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28K</figureCitation>
), blade with ventral margin not serrated; apex enlarged, forming a rounded dorsal process. Aedeagus (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28L, M</figureCitation>
) apodemal processes narrow basally, curved in right angle dorsally; distal portion, in lateral view, expanded medially, external surface, in posterior view, bearing acute process curved dorsally; shaft apex with pair of minute processes, directed ventrally.
</paragraph>
<caption doi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79" start="Figure 28" startId="F28">
<paragraph pageId="0" pageNumber="79">
<emphasis bold="true" pageId="0" pageNumber="79">Figure 28.</emphasis>
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana taurina" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="taurina">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana taurina</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male. 
<emphasis bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing. 
<emphasis bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view. 
<emphasis bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view. 
<emphasis bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view. 
<emphasis bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: holotype male 7.8 mm; paratypes, males (n = 5) 7.7-8.0 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head, in frontal view (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28B</figureCitation>
), frons 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately the width of scape; clypeus 1.2 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins slightly convergent apically; apex straight. Head, in lateral view (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28C</figureCitation>
), crown-face transition thin, with 4 transverse carinae; clypeus not inflated. Profemur AV row with 4-5 setae; PV row with 1-2 setae. Protibia PD row with 3 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 21-23, 12 and 14-16 macrosetae respectively. Metatarsomere I 4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 5 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
Head and thorax (Fig. 
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32I, J</figureCitation>
) reddish-brown. Head (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28A</figureCitation>
) with pair of small circular black maculae near posterior margin behind ocelli. Face (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28B</figureCitation>
) yellowish without maculae. Pronotum (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28A</figureCitation>
) with black punctures and pair of circular black spots near anterior margin behind eye; proepimeron (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28C</figureCitation>
), with narrow black band below pronotal carina; lateral carina yellow. Forewing (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28D</figureCitation>
) with black maculae on apex of anal veins, cross veins of inner discal and apical cells; transverse brown stripe over subapical cells. Metatibia (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
Sternite VIII (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28E</figureCitation>
) 1.6 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; lateral margins weakly convergent posterad; posterior margin rounded. Valve (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28F</figureCitation>
) 3.5 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; dorsal margin slightly excavated; posterior margin straight. Pygofer, in lateral view (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28G</figureCitation>
), 2 
<normalizedToken originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process thin and short, extending ventrally for 1/4 height of lateral lobe; anteroventral, posterodorsal and posteroventral margins rounded; apex slightly tapered, subacute. Subgenital plate, in lateral view (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28G</figureCitation>
), short, not reaching pygofer apex; in ventral view (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28H</figureCitation>
), elongated, 4.5 
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, maximum width near basal third; outer margin with minute setae, apex with small group of filiform setae; inner margin slightly rounded; external margin straight; apex rounded. Connective (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28I</figureCitation>
) 2 
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; anterior margin excavated; dorsal keel and stem reduced; stem wide. Style, in dorsal view (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28J</figureCitation>
), with outer lobe reduced and rounded; in lateral view (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28K</figureCitation>
), blade weakly curved dorsally, wider near base and tapered toward apex; ventral margin not serrated; apex enlarged, forming rounded dorsal process. Aedeagus (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28L, M</figureCitation>
) preatrium weakly developed; dorsal apodeme with dorsal margin straight, lateral margins slightly produced laterally; apodemal processes narrow basally, curved in right angle dorsally at basal third; distal portion, in lateral view, expanded medially, external surface, in posterior view, bearing acute process curved dorsally; apex rounded, with subapical constriction; shaft symmetrical, strongly curved dorsally near base, with basolateral expansion that surrounds the previous portion of shaft, middle portion cylindrical; apex weakly curved ventrally, with pair of minute processes, directed ventrally. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female">
<paragraph pageId="0" pageNumber="79">Female.</paragraph>
<paragraph pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="1986-10-17" collectorName="C. Elias" country="Brazil" location="Rondonia" specimenCount="1" stateProvince="Rondonia" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="79">
<typeStatus>Holotype</typeStatus>
male: 
<collectingCountry name="Brazil">Brazil</collectingCountry>
: 
<normalizedToken originalValue="Rondônia">
<collectingRegion country="Brazil" name="Rondonia">Rondonia</collectingRegion>
</normalizedToken>
</emphasis>
: &quot;Vilhena, RO\ 
<collectingDate value="1986-10-17">17/X/1986</collectingDate>
\ 
<collectorName>C. Elias</collectorName>
, leg.\ Polonoroeste&quot; (DZUP)
</materialsCitation>
. 
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="C. Elias" country="Brazil" location="Paratypes" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Paratypes">
<typeStatus>Paratypes</typeStatus>
</collectingRegion>
</emphasis>
: 
<specimenCount type="male">1♂</specimenCount>
, same data as holotype (DZUP)
</materialsCitation>
; 
<materialsCitation collectingDate="1986-10-15" collectorName="C. Elias" country="Brazil" location="Rondonia" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, same data as holotype except 
<collectingDate value="1986-10-15">15/X/1986</collectingDate>
(DZUP)
</materialsCitation>
. - 
<materialsCitation collectingDate="2006-03-13" collectingMethod="light trap" collectorName="AR Calor" country="Brazil" county="Rio Azul" location="Brasil" municipality="Serra do Divisor" specimenCount="1" specimenCount-male="1" stateProvince="Acre" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Acre">Acre</collectingRegion>
</emphasis>
: 
<specimenCount type="male">1♂</specimenCount>
, &quot; 
<location LSID="urn:lsid:plazi:treatment:A176B5C93EBC568B9539914B640AA181:09A93ED097758EAE5676B7B5DB104B4C" country="Brazil" county="Rio Azul" municipality="Serra do Divisor" name="Brasil" stateProvince="Acre">Brasil</location>
, 
<location LSID="urn:lsid:plazi:treatment:A176B5C93EBC568B9539914B640AA181:E6154C65A80998BCE18B5E5612037A28" country="Brazil" county="Rio Azul" municipality="Serra do Divisor" name="Acre" stateProvince="Acre">Acre</location>
, 
<location LSID="urn:lsid:plazi:treatment:A176B5C93EBC568B9539914B640AA181:0BC5B90DEAFD214C15966A464175E4C7" country="Brazil" county="Rio Azul" municipality="Serra do Divisor" name="Mancio" stateProvince="Acre">Mancio</location>
\ 
<location LSID="urn:lsid:plazi:treatment:A176B5C93EBC568B9539914B640AA181:FAF4E2DE5475277272DABBE894610DA8" country="Brazil" county="Rio Azul" municipality="Serra do Divisor" name="Lima" stateProvince="Acre">Lima</location>
, 
<location LSID="urn:lsid:plazi:treatment:A176B5C93EBC568B9539914B640AA181:635C7DA9AA22A3F4EF4105773F901BE8" country="Brazil" county="Rio Azul" municipality="Serra do Divisor" name="Parque Nacional" stateProvince="Acre">Parque Nacional</location>
\ 
<collectingMunicipality>Serra do Divisor</collectingMunicipality>
,\ 
<collectingCounty>Rio Azul</collectingCounty>
<collectingDate value="2006-03-13">13.iii.2006</collectingDate>
\ 
<collectingMethod>light trap</collectingMethod>
\ 
<collectorName>AR Calor</collectorName>
leg&quot; (MZSP). - 
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingRegion country="Brazil" name="Mato Grosso">Mato Grosso</collectingRegion>
</emphasis>
</materialsCitation>
: 
<materialsCitation collectingDate="1966-10-01" collectingDateMax="1966-10-12" collectingDateMin="1966-10-01" collectorName="Lenko, Pereira" country="Brazil" location="Utiariti" municipality="Rio Papagaio" specimenCount="1" specimenCount-male="1" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCount type="male">1♂</specimenCount>
, &quot; 
<location LSID="urn:lsid:plazi:treatment:A176B5C93EBC568B9539914B640AA181:53463091B9DEBE71844B4EC07C5F5C65" country="Brazil" municipality="Rio Papagaio" name="Utiariti" stateProvince="Mato Grosso">Utiariti</location>
\ 
<collectingMunicipality>Rio Papagaio</collectingMunicipality>
, MT\ 
<collectingDate value="1966-10-01" valueMax="1966-10-12" valueMin="1966-10-01">1-12.X.1966</collectingDate>
\ 
<collectorName>Lenko</collectorName>
&amp; 
<collectorName>Pereira</collectorName>
&quot; (MZSP). -
</materialsCitation>
<materialsCitation collectingDate="2013-07-01" collectingDateMax="2013-07-15" collectingDateMin="2013-07-01" collectorName="J. A. Rafael, F. Limeira-de-Oliveira, T. T. A. Silva, S. Onkone Gare Camp Reserva Etnica, T. L. Erwin" country="Ecuador" county="Riacho Cancela" elevation="225" latitude="-7.111722" location="Brasil" longLatPrecision="1" longitude="-47.29911" municipality="Carolina" specimenCount="1" specimenCount-male="1" stateProvince="Maranhao" typeStatus="paratype">
<emphasis bold="true" pageId="0" pageNumber="79">
<normalizedToken originalValue="Maranhão">
<collectingRegion country="Brazil" name="Maranhao">Maranhao</collectingRegion>
</normalizedToken>
</emphasis>
: 
<specimenCount type="male">1♂</specimenCount>
, &quot; 
<location LSID="urn:lsid:plazi:treatment:A176B5C93EBC568B9539914B640AA181:09631857FD89D28C29215D1E99505A2E" country="Ecuador" county="Riacho Cancela" latitude="-7.111722" longLatPrecision="1" longitude="-47.29911" municipality="Carolina" name="Brasil" stateProvince="Maranhao">Brasil</location>
(MA), 
<collectingMunicipality>Carolina</collectingMunicipality>
\ PARNA Chapada das Mesas,\ 
<collectingCounty>Riacho Cancela</collectingCounty>
, 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.25" unit="m" value="225.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.25" unit="m" value="225.0">225m</elevation>
</quantity>
\ 
<geoCoordinate degrees="7" direction="south" minutes="06" orientation="latitude" precision="1" seconds="42.2" value="-7.111722">7°06′42.2S</geoCoordinate>
<geoCoordinate degrees="47" direction="west" minutes="17" orientation="longitude" precision="1" seconds="56.8" value="-47.29911">47°17′56.8″W</geoCoordinate>
, &quot;Armadilha de Malaise\ 
<collectingDate value="2013-07-01" valueMax="2013-07-15" valueMin="2013-07-01">01-15.vii.2013</collectingDate>
; 
<collectorName>J.A. Rafael</collectorName>
\ 
<collectorName>F. Limeira-de-Oliveira</collectorName>
&amp;\ 
<collectorName>T.T.A. Silva</collectorName>
, cols.&quot; (CZMA). - 
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
</emphasis>
: &quot;Ecuador Orellana Transect Ent. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="km" value="1.0">1 km</quantity>
<collectorName>S. Onkone Gare Camp Reserva Etnica</collectorName>
Waorani Onkone Gare Camp 
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.2" unit="m" value="220.0">220m</elevation>
</quantity>
03/jul/95 
<geoCoordinate degrees="00" direction="south" minutes="39" orientation="latitude" precision="15" seconds="10" value="-0.65277773">00 39′ 10&quot; S</geoCoordinate>
<geoCoordinate degrees="076" direction="west" minutes="26" orientation="longitude" precision="15" seconds="00" value="-76.433334">076 26′ 00&quot; W</geoCoordinate>
<collectorName>T.L.Erwin</collectorName>
et al Fogging terre firme forest&quot; (EPNC - on indefinite loan to the United States National Museum, Washington, USA)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="etymology">
<paragraph pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="79">The new species name refers to the aedeagus with apodemal processes bearing projections at the mid-length of external surface resembling bull horns.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" genus="Coarctana" kingdom="Animalia" lsidName="Coarctana taurina" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="taurina">
<emphasis italics="true" pageId="0" pageNumber="79">Coarctana taurina</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
can be easily distinguished from 
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. oricula" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="oricula">
<emphasis italics="true" pageId="0" pageNumber="79">C. oricula</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
by the style (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28K</figureCitation>
) with an apex enlarged, forming a rounded dorsal process, the apodemal processes of aedeagus (Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Coarctana taurina sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure28" httpUri="https://binary.pensoft.net/fig/798181" pageId="0" pageNumber="79">28L, M</figureCitation>
) with an acute process curved dorsally on external surface, and the shaft with pair of minute apical processes, directed ventrally.
</paragraph>
</subSubSection>
</treatment>
</document>