311 lines
35 KiB
XML
311 lines
35 KiB
XML
<document id="6C7F4389D829126B18383A19C09948C0" ID="10.11646/zootaxa.4169.2.2" ID-DOI="10.11646/zootaxa.4169.2.2" ID-GBIF-Dataset="5509767a-7e3a-4919-bb01-6b025bb293e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="260910" ID-ZooBank="53B27ED7-8CFA-4058-A47E-3A340ED322FC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474353234057" checkinUser="plazi" docAuthor="Márquez, Juan & Asiain, Julieta" docDate="2016" docId="0E256050FF87CC32FF34FBB4FCBAFDD0" docLanguage="en" docName="zootaxa.4169.2.2.pdf" docOrigin="Zootaxa 4169 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thyreocephalus rufipennis Sharp 1885" docType="treatment" docVersion="8" lastPageNumber="268" masterDocId="F21C1828FF88CC23FFA3FF81FFDBFFEC" masterDocTitle="Taxonomy of the Mexican species of Thyreocephalus Guérin-Méneville (Coleoptera: Staphylinidae, Xantholinini)" masterLastPageNumber="285" masterPageNumber="251" pageNumber="266" updateTime="1698386383488" updateUser="plazi">
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<mods:titleInfo id="6831FFC24FD2221359EA7389DE13E679">
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<mods:title id="166219F56C56BB3BD8F011C221FAC198">Taxonomy of the Mexican species of Thyreocephalus Guérin-Méneville (Coleoptera: Staphylinidae, Xantholinini)</mods:title>
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<mods:name id="833E5C6259D3BF0E1C5F065116797C15" type="personal">
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<mods:namePart id="0BC0D0F65D32D9A0D519E90A840A2C61">Márquez, Juan</mods:namePart>
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<mods:namePart id="E46285D80C6CAFC5FCD89E1502933C41">Asiain, Julieta</mods:namePart>
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<mods:typeOfResource id="00064AEF663B7811384E37E192B67E3A">text</mods:typeOfResource>
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<mods:title id="F230AF43379777159E11D674F06D3DC5">Zootaxa</mods:title>
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<mods:part id="8B038794E1D35737A348EB7BF65A9576">
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<mods:date id="A6D5FA9A360F44FD8AD4C4932C5990F1">2016</mods:date>
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<mods:number id="E1BDEF70E8FF6F035212851338F33123">4169</mods:number>
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<mods:detail id="630A85752994B31E60C3AC5B6E4A5041" type="issue">
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<mods:number id="9B6C726606AD102975F994F0D4A5ADC6">2</mods:number>
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<mods:identifier id="C492FC017350D952D5BDB08A3B096007" type="DOI">10.11646/zootaxa.4169.2.2</mods:identifier>
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<treatment id="0E256050FF87CC32FF34FBB4FCBAFDD0" ID-DOI="http://doi.org/10.5281/zenodo.5623717" ID-GBIF-Taxon="124552122" ID-Zenodo-Dep="5623717" LSID="urn:lsid:plazi:treatment:0E256050FF87CC32FF34FBB4FCBAFDD0" httpUri="http://treatment.plazi.org/id/0E256050FF87CC32FF34FBB4FCBAFDD0" lastPageId="17" lastPageNumber="268" pageId="15" pageNumber="266">
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<subSubSection id="CE9682CDFF87CC2CFF34FBB4FE07FB9C" pageId="15" pageNumber="266" type="nomenclature">
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<paragraph id="8633D146FF87CC2CFF34FBB4FDA0FBA3" blockId="15.[151,635,1077,1136]" box="[151,635,1077,1103]" pageId="15" pageNumber="266">
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<heading id="DD7B662AFF87CC2CFF34FBB4FDA0FBA3" bold="true" box="[151,635,1077,1103]" fontSize="11" level="1" pageId="15" pageNumber="266" reason="1">
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<taxonomicName id="418CAAC5FF87CC2CFF34FBB4FDA0FBA3" ID-CoL="56RYC" authority="Sharp, 1885" authorityName="Sharp" authorityYear="1885" box="[151,635,1077,1103]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="rufipennis">
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<emphasis id="B4F80D54FF87CC2CFF34FBB4FDA0FBA3" bold="true" box="[151,635,1077,1103]" pageId="15" pageNumber="266">
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<emphasis id="B4F80D54FF87CC2CFF34FBB4FE06FBA3" bold="true" box="[151,477,1077,1103]" italics="true" pageId="15" pageNumber="266">Thyreocephalus rufipennis</emphasis>
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<bibRefCitation id="E21DACB7FF87CC2CFE47FBB4FDA0FBA3" author="Sharp" box="[484,635,1077,1103]" pageId="15" pageNumber="266" refString="Sharp, D. (1885) Staphylinidae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. 1 (2). Taylor & Francis, London, pp. 393 - 536, pls. 10 - 13." type="journal article" year="1885">Sharp, 1885</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8633D146FF87CC2CFF34FBD6FE07FB9C" blockId="15.[151,635,1077,1136]" box="[151,476,1111,1136]" pageId="15" pageNumber="266">
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(
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<figureCitation id="1EB7CDC3FF87CC2CFF3CFBD6FF37FB9C" box="[159,236,1111,1136]" captionStart="FIGURE 3" captionStartId="16.[151,250,1867,1889]" captionTargetBox="[173,1421,206,1830]" captionTargetId="figure@16.[151,1436,193,1846]" captionTargetPageId="16" captionText="FIGURE 3. Thyreocephalus puncticeps: a) dorsal view of the head and pronotum; b) ventral view of the head (mandibles excluded). Thyreocephalus rufipennis: c) dorsal view of the head and pronotum; d) ventral view of the head (mandibles excluded). Scale = 2 mm." httpUri="https://zenodo.org/record/260912/files/figure.png" pageId="15" pageNumber="266">Figs. 3</figureCitation>
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c, d,
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, 9f,
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<quantity id="41747CA3FF87CC2CFED1FBD9FE45FB9C" box="[370,414,1112,1136]" metricMagnitude="-2" metricUnit="kg" metricValue="1.1" pageId="15" pageNumber="266" unit="g" value="11.0">11g</quantity>
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, 12b)
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</paragraph>
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</subSubSection>
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<subSubSection id="CE9682CDFF87CC2CFF34FB21FDAEFB38" pageId="15" pageNumber="266" type="reference_group">
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<paragraph id="8633D146FF87CC2CFF34FB21FD82FB5A" blockId="15.[151,629,1184,1236]" box="[151,601,1184,1206]" pageId="15" pageNumber="266">
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<treatmentCitationGroup id="A69CF668FF87CC2CFF34FB21FD82FB5A" box="[151,601,1184,1206]" pageId="15" pageNumber="266">
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<taxonomicName id="418CAAC5FF87CC2CFF34FB21FD82FB5A" ID-CoL="56RYC" authority="Sharp, 1885: 500" authorityName="Sharp" authorityPageNumber="500" authorityYear="1885" box="[151,601,1184,1206]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="rufipennis">
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<emphasis id="B4F80D54FF87CC2CFF34FB21FE7EFB5A" box="[151,421,1184,1206]" italics="true" pageId="15" pageNumber="266">Thyreocephalus rufipennis</emphasis>
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<treatmentCitation id="072DF757FF87CC2CFE08FB21FD82FB5A" author="Sharp" box="[427,601,1184,1206]" page="500" pageId="15" pageNumber="266" year="1885">
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<bibRefCitation id="E21DACB7FF87CC2CFE08FB21FDF2FB5A" author="Sharp" box="[427,553,1184,1206]" pageId="15" pageNumber="266" refString="Sharp, D. (1885) Staphylinidae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. 1 (2). Taylor & Francis, London, pp. 393 - 536, pls. 10 - 13." type="journal article" year="1885">Sharp, 1885</bibRefCitation>
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: 500
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8633D146FF87CC2CFF34FB3FFDAEFB38" blockId="15.[151,629,1184,1236]" box="[151,629,1214,1236]" pageId="15" pageNumber="266">
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<treatmentCitationGroup id="A69CF668FF87CC2CFF34FB3FFDAEFB38" box="[151,629,1214,1236]" pageId="15" pageNumber="266">
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<taxonomicName id="418CAAC5FF87CC2CFF34FB3FFDAEFB38" ID-CoL="56RYC" authority="Smetana, 1977: 349" authorityName="Smetana" authorityPageNumber="349" authorityYear="1977" box="[151,629,1214,1236]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="rufipennis">
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<emphasis id="B4F80D54FF87CC2CFF34FB3FFE7EFB38" box="[151,421,1214,1236]" italics="true" pageId="15" pageNumber="266">Thyreocephalus rufipennis</emphasis>
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<treatmentCitation id="072DF757FF87CC2CFE08FB3FFDAEFB38" author="Smetana" box="[427,629,1214,1236]" page="349" pageId="15" pageNumber="266" year="1977">
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<bibRefCitation id="E21DACB7FF87CC2CFE08FB3FFD9EFB38" author="Smetana" box="[427,581,1214,1236]" pageId="15" pageNumber="266" refString="Smetana, A. (1977) Lectotype designation and taxonomic remarks on some Xantholinine genera and species from Central and South America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 31 (4), 347 - 362." type="journal article" year="1977">Smetana, 1977</bibRefCitation>
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: 349
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="CE9682CDFF87CC2CFF34FA80FCFBF9D5" pageId="15" pageNumber="266" type="diagnosis">
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<paragraph id="8633D146FF87CC2CFF34FA80FCFBF9D5" blockId="15.[151,1437,1281,1954]" pageId="15" pageNumber="266">
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<emphasis id="B4F80D54FF87CC2CFF34FA80FECAFAF6" bold="true" box="[151,273,1281,1306]" pageId="15" pageNumber="266">Diagnosis.</emphasis>
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<taxonomicName id="418CAAC5FF87CC2CFEBAFA80FD9FFAF6" box="[281,580,1281,1306]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="rufipennis">
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<emphasis id="B4F80D54FF87CC2CFEBAFA80FD9FFAF6" box="[281,580,1281,1306]" italics="true" pageId="15" pageNumber="266">Thyreocephalus rufipennis</emphasis>
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</taxonomicName>
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is distinguished from
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<taxonomicName id="418CAAC5FF87CC2CFCE1FA83FC3CFAF6" box="[834,999,1281,1306]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="cribripennis">
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<emphasis id="B4F80D54FF87CC2CFCE1FA83FC3CFAF6" box="[834,999,1281,1306]" italics="true" pageId="15" pageNumber="266">T. cribripennis</emphasis>
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</taxonomicName>
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by denser umbilicate punctures on the dorsal and ventral surface of the head (
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<figureCitation id="1EB7CDC3FF87CC2CFDF4FAA4FD7CFAD2" box="[599,679,1317,1342]" captionStart="FIGURE 3" captionStartId="16.[151,250,1867,1889]" captionTargetBox="[173,1421,206,1830]" captionTargetId="figure@16.[151,1436,193,1846]" captionTargetPageId="16" captionText="FIGURE 3. Thyreocephalus puncticeps: a) dorsal view of the head and pronotum; b) ventral view of the head (mandibles excluded). Thyreocephalus rufipennis: c) dorsal view of the head and pronotum; d) ventral view of the head (mandibles excluded). Scale = 2 mm." httpUri="https://zenodo.org/record/260912/files/figure.png" pageId="15" pageNumber="266">Figs. 3</figureCitation>
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c, d), head almost as long as wide (proportion = 1.02 times), neck with two cavities connected by an anterior fine suture (
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<figureCitation id="1EB7CDC3FF87CC2CFCA0FAC8FC9EFA8E" box="[771,837,1353,1378]" captionStart="FIGURE 3" captionStartId="16.[151,250,1867,1889]" captionTargetBox="[173,1421,206,1830]" captionTargetId="figure@16.[151,1436,193,1846]" captionTargetPageId="16" captionText="FIGURE 3. Thyreocephalus puncticeps: a) dorsal view of the head and pronotum; b) ventral view of the head (mandibles excluded). Thyreocephalus rufipennis: c) dorsal view of the head and pronotum; d) ventral view of the head (mandibles excluded). Scale = 2 mm." httpUri="https://zenodo.org/record/260912/files/figure.png" pageId="15" pageNumber="266">Fig. 3</figureCitation>
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c) and aedeagus of median size, with their parameres slender and acute apically (
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<figureCitation id="1EB7CDC3FF87CC2CFE6DFAECFDC7FA6A" box="[462,540,1389,1414]" captionStart="FIGURE 9" captionStartId="24.[151,250,1913,1935]" captionTargetBox="[157,1412,211,1885]" captionTargetId="figure@24.[151,1436,193,1892]" captionTargetPageId="24" captionText="FIGURE 9. Ventral view of the aedeagus of: a) Thyreocephalus unicolor, b) T. sp. aff. subtilis, c) T. jalapaensis, d) T. scutellaris, e) T. cribripennis, f) T. rufipennis, g) T. halffteri. Scale = 0.5 mm." httpUri="https://zenodo.org/record/260917/files/figure.png" pageId="15" pageNumber="266">Figs. 9</figureCitation>
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f,
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<quantity id="41747CA3FF87CC2CFD96FAEFFDBAFA6A" box="[565,609,1390,1414]" metricMagnitude="-2" metricUnit="kg" metricValue="1.1" pageId="15" pageNumber="266" unit="g" value="11.0">11g</quantity>
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); whereas
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<taxonomicName id="418CAAC5FF87CC2CFD78FAEFFC5AFA6A" box="[731,897,1389,1414]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="cribripennis">
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<emphasis id="B4F80D54FF87CC2CFD78FAEFFC5AFA6A" box="[731,897,1389,1414]" italics="true" pageId="15" pageNumber="266">T. cribripennis</emphasis>
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</taxonomicName>
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has a lower density of umbilicate punctures on the dorsal and ventral surfaces of the head, with some of these punctures forming two close central rows, head 1.12 times as long as wide, neck with only one large cavity resulting from the clear connection of the two lateral cavities, and a large aedeagus, with wide parameres, with apex slightly acute. Also, it is distinguished easily from the remaining three species of the group by the lower density of umbilicate punctures on the dorsal and ventral surface of the head, and because the head is more convex.
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</paragraph>
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</subSubSection>
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<subSubSection id="CE9682CDFF87CC32FF64F9C4FAF4FF38" lastPageId="17" lastPageNumber="268" pageId="15" pageNumber="266" type="discussion">
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<paragraph id="8633D146FF87CC2CFF64F9C4FD70F84E" blockId="15.[151,1437,1281,1954]" pageId="15" pageNumber="266">
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<emphasis id="B4F80D54FF87CC2CFF64F9C4FE97F9B2" bold="true" box="[199,332,1605,1630]" pageId="15" pageNumber="266">Discussion.</emphasis>
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We agree with the observation by
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<bibRefCitation id="E21DACB7FF87CC2CFD79F9C4FCB5F9B2" author="Sharp" box="[730,878,1605,1630]" pageId="15" pageNumber="266" refString="Sharp, D. (1885) Staphylinidae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. 1 (2). Taylor & Francis, London, pp. 393 - 536, pls. 10 - 13." type="journal article" year="1885">Sharp (1885)</bibRefCitation>
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in respect to
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<taxonomicName id="418CAAC5FF87CC2CFBA8F9C7FB42F9B2" box="[1035,1177,1606,1630]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="rufipennis">
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<emphasis id="B4F80D54FF87CC2CFBA8F9C7FB42F9B2" box="[1035,1177,1606,1630]" italics="true" pageId="15" pageNumber="266">T. rufipennis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="418CAAC5FF87CC2CFB0BF9C7FA95F9B2" box="[1192,1358,1605,1630]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="cribripennis">
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<emphasis id="B4F80D54FF87CC2CFB0BF9C7FA95F9B2" box="[1192,1358,1605,1630]" italics="true" pageId="15" pageNumber="266">T. cribripennis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="418CAAC5FF87CC2CFA24F9C7FED4F96D" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="puncticeps">
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<emphasis id="B4F80D54FF87CC2CFA24F9C7FED4F96D" italics="true" pageId="15" pageNumber="266">T. puncticeps</emphasis>
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</taxonomicName>
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: "This and the following species are readily distinguished from
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<taxonomicName id="418CAAC5FF87CC2CFC49F9EBFB89F96E" box="[1002,1106,1641,1666]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="salvini">
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<emphasis id="B4F80D54FF87CC2CFC49F9EBFB89F96E" box="[1002,1106,1641,1666]" italics="true" pageId="15" pageNumber="266">T. salvini</emphasis>
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</taxonomicName>
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and its allies by the fact that the prosternum is acutely carinate behind, and the punctation more evenly distributed over the upper surface of the head". Thanks to the designation of the lectotype made by
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<bibRefCitation id="E21DACB7FF87CC2CFCF7F933FBD7F926" author="Smetana" box="[852,1036,1714,1738]" pageId="15" pageNumber="266" refString="Smetana, A. (1977) Lectotype designation and taxonomic remarks on some Xantholinine genera and species from Central and South America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 31 (4), 347 - 362." type="journal article" year="1977">Smetana (1977)</bibRefCitation>
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, the drawing of the aedeagus, the distinctive characteristics he provided, and the advantage that we reviewed this specimen, the recognition of the species was facilitated. A pending aspect is the only specimen in the type series mentioned by
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<bibRefCitation id="E21DACB7FF87CC2CFB69F978FA85F8FE" author="Sharp" box="[1226,1374,1785,1810]" pageId="15" pageNumber="266" refString="Sharp, D. (1885) Staphylinidae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. 1 (2). Taylor & Francis, London, pp. 393 - 536, pls. 10 - 13." type="journal article" year="1885">Sharp (1885)</bibRefCitation>
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from Guatemala (San Geronimo), which was not studied by
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<bibRefCitation id="E21DACB7FF87CC2CFCA3F89FFC69F8DA" author="Smetana" box="[768,946,1822,1846]" pageId="15" pageNumber="266" refString="Smetana, A. (1977) Lectotype designation and taxonomic remarks on some Xantholinine genera and species from Central and South America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 31 (4), 347 - 362." type="journal article" year="1977">Smetana (1977)</bibRefCitation>
|
|
nor by us in this work to corroborate that it belongs to the same species, in which case it can be considered as paralectotype. It is interesting that with exception of the unique specimen from Guatemala, the remaining known specimens of this species are from few localities of the Sierra Madre del Sur in the state of Oaxaca.
|
|
</paragraph>
|
|
<caption id="D2F381CEFF98CC33FF34F8CAFE41F872" httpUri="https://zenodo.org/record/260912/files/figure.png" pageId="16" pageNumber="267" targetBox="[173,1421,206,1830]" targetPageId="16">
|
|
<paragraph id="8633D146FF98CC33FF34F8CAFE41F872" blockId="16.[151,1436,1867,1950]" pageId="16" pageNumber="267">
|
|
<emphasis id="B4F80D54FF98CC33FF34F8CAFECEF88C" bold="true" box="[151,277,1867,1889]" pageId="16" pageNumber="267">FIGURE 3.</emphasis>
|
|
<taxonomicName id="418CAAC5FF98CC33FEBCF8CAFDEEF88D" box="[287,565,1867,1889]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="puncticeps">
|
|
<emphasis id="B4F80D54FF98CC33FEBCF8CAFDEEF88D" box="[287,565,1867,1889]" italics="true" pageId="16" pageNumber="267">Thyreocephalus puncticeps</emphasis>
|
|
</taxonomicName>
|
|
: a) dorsal view of the head and pronotum; b) ventral view of the head (mandibles excluded).
|
|
<taxonomicName id="418CAAC5FF98CC33FEAFF8EBFDC5F86C" box="[268,542,1898,1920]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="rufipennis">
|
|
<emphasis id="B4F80D54FF98CC33FEAFF8EBFDC5F86C" box="[268,542,1898,1920]" italics="true" pageId="16" pageNumber="267">Thyreocephalus rufipennis</emphasis>
|
|
</taxonomicName>
|
|
: c) dorsal view of the head and pronotum; d) ventral view of the head (mandibles excluded). Scale = 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8633D146FF99CC32FF64FF16FAF4FF38" blockId="17.[151,1436,151,573]" pageId="17" pageNumber="268">
|
|
<emphasis id="B4F80D54FF99CC32FF64FF16FE5EFF5C" bold="true" box="[199,389,151,176]" pageId="17" pageNumber="268">Natural history.</emphasis>
|
|
It is possible that this species has the same life habits as remaining species of this group, as it has been collected on rotten "nopales" (
|
|
<taxonomicName id="418CAAC5FF99CC32FDF0FF3CFD74FF38" box="[595,687,189,212]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="17" pageNumber="268" phylum="Tracheophyta" rank="genus">
|
|
<emphasis id="B4F80D54FF99CC32FDF0FF3CFD74FF38" box="[595,687,189,212]" italics="true" pageId="17" pageNumber="268">Opuntia</emphasis>
|
|
</taxonomicName>
|
|
sp.), rotten mandarins, and with flight interception traps.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CE9682CDFF99CC32FF64FF5EFEA5FEF0" pageId="17" pageNumber="268" type="distribution">
|
|
<paragraph id="8633D146FF99CC32FF64FF5EFEA5FEF0" blockId="17.[151,1436,151,573]" pageId="17" pageNumber="268">
|
|
<emphasis id="B4F80D54FF99CC32FF64FF5EFE82FF14" bold="true" box="[199,345,223,248]" pageId="17" pageNumber="268">Distribution</emphasis>
|
|
(
|
|
<figureCitation id="1EB7CDC3FF99CC32FECAFF5EFE67FF14" box="[361,444,223,248]" captionStart="FIGURE 12" captionStartId="27.[151,250,1969,1991]" captionTargetBox="[201,1373,206,1910]" captionTargetId="figure@27.[181,1397,193,1932]" captionTargetPageId="27" captionText="FIGURE 12. Geographic distribution and individual track of: a) T. puncticeps (black squares) and T. unicolor (black circles); b) T. arizonicus (gray squares), T. jalapaensis (black circles) and T. rufipennis (white circles)." httpUri="https://zenodo.org/record/260920/files/figure.png" pageId="17" pageNumber="268">Fig. 12</figureCitation>
|
|
b).
|
|
<collectingRegion id="44481FA4FF99CC32FE42FF5EFDEDFF14" box="[481,566,223,248]" country="Mexico" name="Mexico" pageId="17" pageNumber="268">
|
|
<collectingCountry id="FE9B91D6FF99CC32FE42FF5EFDEDFF14" box="[481,566,223,248]" name="Mexico" pageId="17" pageNumber="268">Mexico</collectingCountry>
|
|
</collectingRegion>
|
|
(
|
|
<collectingRegion id="44481FA4FF99CC32FDE5FF61FD46FF14" box="[582,669,224,248]" country="Mexico" name="Oaxaca" pageId="17" pageNumber="268">Oaxaca</collectingRegion>
|
|
) and
|
|
<collectingRegion id="44481FA4FF99CC32FD7EFF5EFC8DFF14" box="[733,854,223,248]" country="Guatemala" name="Guatemala" pageId="17" pageNumber="268">
|
|
<collectingCountry id="FE9B91D6FF99CC32FD7EFF5EFC8DFF14" box="[733,854,223,248]" name="Guatemala" pageId="17" pageNumber="268">Guatemala</collectingCountry>
|
|
</collectingRegion>
|
|
(San Geronimo) (
|
|
<tableCitation id="CB0EE4FDFF99CC32FB87FF5EFBA0FF14" box="[1060,1147,223,248]" captionStart="TABLE 2" captionStartId="7.[151,239,616,639]" captionTargetBox="[159,1401,698,1307]" captionTargetPageId="7" captionText="TABLE 2. Mexican species of Thyreocephalus with country and state records (Herman 2001, Navarrete-Heredia et al. 2002, Asenjo 2004, Newton et al. 2005, Asenjo et al. 2013). * First Mexican record, ** first Mexican state record." httpUri="http://table.plazi.org/id/D2F381CEFF8FCC24FF34FDE9FA8EFD73" pageId="17" pageNumber="268" tableUuid="D2F381CEFF8FCC24FF34FDE9FA8EFD73">Table 2</tableCitation>
|
|
) (
|
|
<bibRefCitation id="E21DACB7FF99CC32FB31FF5EFAC3FF14" author="Sharp" box="[1170,1304,223,248]" pageId="17" pageNumber="268" refString="Sharp, D. (1885) Staphylinidae. In: Godman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. 1 (2). Taylor & Francis, London, pp. 393 - 536, pls. 10 - 13." type="journal article" year="1885">Sharp 1885</bibRefCitation>
|
|
, Navarrete-
|
|
<collectingRegion id="44481FA4FF99CC32FF34FE85FF2BFEF1" box="[151,240,260,285]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="268">Heredia</collectingRegion>
|
|
<emphasis id="B4F80D54FF99CC32FF55FE84FEF3FEF1" box="[246,296,260,285]" italics="true" pageId="17" pageNumber="268">et al</emphasis>
|
|
. 2002).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CE9682CDFF99CC32FF64FEA6FCBAFDD0" pageId="17" pageNumber="268" type="materials_examined">
|
|
<paragraph id="8633D146FF99CC32FF64FEA6FE5FFE40" blockId="17.[151,1436,151,573]" pageId="17" pageNumber="268">
|
|
<materialsCitation id="36E4DB1BFF99CC32FF64FEA6FA88FEAC" ID-GBIF-Occurrence="1313797560" box="[199,1363,295,320]" collectorName="Type D. S." country="Mexico" location="Oaxaca" pageId="17" pageNumber="268" specimenCount="3" stateProvince="Oaxaca" typeStatus="lectotype">
|
|
<emphasis id="B4F80D54FF99CC32FF64FEA6FE29FEAC" bold="true" box="[199,498,295,320]" pageId="17" pageNumber="268">
|
|
<typeStatus id="59376FE4FF99CC32FF64FEA6FED9FEAC" box="[199,258,295,320]" pageId="17" pageNumber="268">Type</typeStatus>
|
|
material examined.
|
|
</emphasis>
|
|
<typeStatus id="59376FE4FF99CC32FE59FEA9FDB1FEAC" box="[506,618,296,320]" pageId="17" pageNumber="268" type="lectotype">Lectotype</typeStatus>
|
|
(male): “
|
|
<taxonomicName id="418CAAC5FF99CC32FD7AFEA6FBDFFEAC" box="[729,1028,295,320]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="rufipennis">
|
|
<emphasis id="B4F80D54FF99CC32FD7AFEA6FBDFFEAC" box="[729,1028,295,320]" italics="true" pageId="17" pageNumber="268">Thyreocephalus rufipennis</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
<collectorName id="2B79B490FF99CC32FBB7FEA9FB55FEAC" box="[1044,1166,296,320]" pageId="17" pageNumber="268">
|
|
<typeStatus id="59376FE4FF99CC32FBB7FEA9FB97FEAC" box="[1044,1100,296,320]" pageId="17" pageNumber="268">Type</typeStatus>
|
|
D. S.
|
|
</collectorName>
|
|
<collectingRegion id="44481FA4FF99CC32FB34FEA9FB35FEAC" box="[1175,1262,296,320]" country="Mexico" name="Oaxaca" pageId="17" pageNumber="268">Oaxaca</collectingRegion>
|
|
,
|
|
<collectingCountry id="FE9B91D6FF99CC32FB58FEA6FA88FEAC" box="[1275,1363,295,320]" name="Mexico" pageId="17" pageNumber="268">Mexico</collectingCountry>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="36E4DB1BFF99CC32FAF5FEA6FE5FFE40" ID-GBIF-Occurrence="1313797559" collectingDate="I. 2." collectionCode="BMNH" collectorName="B. C. A. Col." country="Mexico" location="Oaxaca" pageId="17" pageNumber="268" specimenCount="2" stateProvince="Oaxaca" typeStatus="lectotype">
|
|
Höge/ aedeagus on the plate/ genital segment on the plate/
|
|
<typeStatus id="59376FE4FF99CC32FC8AFECDFCBBFE88" box="[809,864,332,356]" pageId="17" pageNumber="268">Type</typeStatus>
|
|
/
|
|
<collectingRegion id="44481FA4FF99CC32FCD4FECDFC15FE88" box="[887,974,332,356]" country="Mexico" name="Oaxaca" pageId="17" pageNumber="268">Oaxaca</collectingRegion>
|
|
,
|
|
<collectingCountry id="FE9B91D6FF99CC32FC42FECDFBE2FE89" box="[993,1081,332,357]" name="Mexico" pageId="17" pageNumber="268">Mexico</collectingCountry>
|
|
, Hoege/
|
|
<collectorName id="2B79B490FF99CC32FB0EFECCFA82FE89" box="[1197,1369,332,357]" pageId="17" pageNumber="268">B. C. A. Col.</collectorName>
|
|
<collectingDate id="E2760E6EFF99CC32FACBFECCFA47FE88" box="[1384,1436,332,356]" pageId="17" pageNumber="268">
|
|
<date id="F232F786FF99CC32FACBFECCFA47FE88" box="[1384,1436,332,356]" pageId="17" pageNumber="268">I. 2.</date>
|
|
</collectingDate>
|
|
<taxonomicName id="418CAAC5FF99CC32FF34FEEEFDD2FE64" authority="Sharp" authorityName="Sharp" box="[151,521,367,392]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="rufipennis">
|
|
<emphasis id="B4F80D54FF99CC32FF34FEEEFE19FE64" box="[151,450,367,392]" italics="true" pageId="17" pageNumber="268">Thyreocephalus rufipennis</emphasis>
|
|
Sharp
|
|
</taxonomicName>
|
|
/ Sharp Coll. 1905,-313/
|
|
<typeStatus id="59376FE4FF99CC32FC82FEF1FC4AFE64" box="[801,913,368,392]" pageId="17" pageNumber="268" type="lectotype">Lectotype</typeStatus>
|
|
<taxonomicName id="418CAAC5FF99CC32FC3BFEEEFA47FE64" authority="Sharp. A. Smetana" authorityName="Sharp. A. Smetana" box="[920,1436,367,392]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="268" phylum="Arthropoda" rank="species" species="rufipennis">
|
|
<emphasis id="B4F80D54FF99CC32FC3BFEEEFB18FE64" box="[920,1219,367,392]" italics="true" pageId="17" pageNumber="268">Thyreocephalus rufipennis</emphasis>
|
|
Sharp. A. Smetana
|
|
</taxonomicName>
|
|
des. 1977” (
|
|
<collectionCode id="E09D4983FF99CC32FEBEFE15FEACFE40" box="[285,375,404,428]" pageId="17" pageNumber="268">BMNH</collectionCode>
|
|
).
|
|
</materialsCitation>
|
|
</paragraph>
|
|
<paragraph id="8633D146FF99CC32FF64FE36FCBAFDD0" blockId="17.[151,1436,151,573]" pageId="17" pageNumber="268">
|
|
<materialsCitation id="36E4DB1BFF99CC32FF64FE36FF3CFDF4" ID-GBIF-Occurrence="1313797562" collectingDate="2013-06-09" collectingDateMax="2013-07-09" collectingDateMin="2013-06-09" collectionCode="IEXA" collectorName="El Puerto & L. Delgado & E. Mora" country="Mexico" county="El Puerto" elevation="1250" location="San Miguel Chimalapa" municipality="Benito Juarez" pageId="17" pageNumber="268" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Oaxaca">
|
|
<emphasis id="B4F80D54FF99CC32FF64FE36FDECFE3C" bold="true" box="[199,567,439,464]" pageId="17" pageNumber="268">Additional material examined.</emphasis>
|
|
“
|
|
<collectingCountry id="FE9B91D6FF99CC32FDEAFE36FD7EFE3C" box="[585,677,439,464]" name="Mexico" pageId="17" pageNumber="268">Mexico</collectingCountry>
|
|
:
|
|
<collectingRegion id="44481FA4FF99CC32FD12FE36FCD0FE3C" box="[689,779,439,464]" country="Mexico" name="Oaxaca" pageId="17" pageNumber="268">
|
|
<emphasis id="B4F80D54FF99CC32FD12FE36FCD0FE3C" bold="true" box="[689,779,439,464]" pageId="17" pageNumber="268">Oaxaca</emphasis>
|
|
</collectingRegion>
|
|
,
|
|
<location id="8353879DFF99CC32FCB8FE39FBF8FE3C" LSID="urn:lsid:plazi:treatment:0E256050FF87CC32FF34FBB4FCBAFDD0:8353879DFF99CC32FCB8FE39FBF8FE3C" box="[795,1059,439,464]" country="Mexico" county="El Puerto" municipality="Benito Juarez" name="San Miguel Chimalapa" pageId="17" pageNumber="268" stateProvince="Oaxaca">San Miguel Chimalapa</location>
|
|
,
|
|
<collectingMunicipality id="66574B3CFF99CC32FB93FE36FB16FE3C" box="[1072,1229,439,464]" pageId="17" pageNumber="268">Benito Juárez</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="6F52A9CAFF99CC32FB79FE36FA93FE23" box="[1242,1352,439,464]" pageId="17" pageNumber="268">
|
|
<collectorName id="2B79B490FF99CC32FB79FE36FA93FE23" box="[1242,1352,439,464]" pageId="17" pageNumber="268">El Puerto</collectorName>
|
|
</collectingCounty>
|
|
, altitud
|
|
<elevation id="0DA13675FF99CC32FF34FE5DFF25FE18" box="[151,254,476,501]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="268" unit="m" value="1250.0">
|
|
<quantity id="41747CA3FF99CC32FF34FE5DFF25FE18" box="[151,254,476,501]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="268" unit="m" value="1250.0">1,250 m</quantity>
|
|
</elevation>
|
|
, trampa de intercepción de vuelo,
|
|
<collectingDate id="E2760E6EFF99CC32FD1BFE5DFC46FE18" box="[696,925,476,500]" pageId="17" pageNumber="268" value="2013-06-09" valueMax="2013-07-09" valueMin="2013-06-09">
|
|
<date id="F232F786FF99CC32FD1BFE5DFC46FE18" box="[696,925,476,500]" pageId="17" pageNumber="268" value="2013-06-09" valueMax="2013-07-09" valueMin="2013-06-09">9-VI a 9-VII-2013</date>
|
|
</collectingDate>
|
|
,
|
|
<collectorName id="2B79B490FF99CC32FC12FE5CFBE2FE19" box="[945,1081,476,501]" pageId="17" pageNumber="268">L. Delgado</collectorName>
|
|
y
|
|
<collectorName id="2B79B490FF99CC32FBCBFE5CFB16FE18" box="[1128,1229,477,500]" pageId="17" pageNumber="268">E. Mora</collectorName>
|
|
cols.” (
|
|
<specimenCount id="908A1ACFFF99CC32FA99FE5DFA80FE18" box="[1338,1371,476,500]" pageId="17" pageNumber="268" type="female">1♀</specimenCount>
|
|
,
|
|
<specimenCount id="908A1ACFFF99CC32FAD1FE5DFA43FE18" box="[1394,1432,476,500]" pageId="17" pageNumber="268" type="male">2♂</specimenCount>
|
|
,
|
|
<collectionCode id="E09D4983FF99CC32FF34FE7EFF04FDF4" box="[151,223,511,536]" pageId="17" pageNumber="268">IEXA</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="36E4DB1BFF99CC32FF49FE7EFC85FDD0" ID-GBIF-Occurrence="1313797567" collectingDate="1999-11-16" collectionCode="MZFC" collectorName="J. Marquez" country="Mexico" elevation="1436" latitude="20.31215" location="San Juan Chicomesuchitl" longLatPrecision="1" longitude="-96.51" municipality="Rio Grande" pageId="17" pageNumber="268" specimenCount="1" specimenCount-female="1" stateProvince="Oaxaca">
|
|
“
|
|
<collectingCountry id="FE9B91D6FF99CC32FF56FE7EFE8AFDF4" box="[245,337,511,536]" name="Mexico" pageId="17" pageNumber="268">Mexico</collectingCountry>
|
|
:
|
|
<collectingRegion id="44481FA4FF99CC32FEF8FD81FE69FDF4" box="[347,434,512,536]" country="Mexico" name="Oaxaca" pageId="17" pageNumber="268">Oaxaca</collectingRegion>
|
|
,
|
|
<location id="8353879DFF99CC32FE1FFD81FD01FDF4" LSID="urn:lsid:plazi:treatment:0E256050FF87CC32FF34FBB4FCBAFDD0:8353879DFF99CC32FE1FFD81FD01FDF4" box="[444,730,511,536]" country="Mexico" latitude="20.31215" longLatPrecision="1" longitude="-96.51" municipality="Rio Grande" name="San Juan Chicomesuchitl" pageId="17" pageNumber="268" stateProvince="Oaxaca">San Juan Chicomesuchitl</location>
|
|
,
|
|
<collectingMunicipality id="66574B3CFF99CC32FD47FD81FCB2FDF4" box="[740,873,511,536]" pageId="17" pageNumber="268">Río Grande</collectingMunicipality>
|
|
,
|
|
<geoCoordinate id="E3B8B781FF99CC32FCD7FD81FBD7FDF4" box="[884,1036,512,536]" direction="north" orientation="latitude" pageId="17" pageNumber="268" precision="1" value="20.31215">N20°18.729”</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="E3B8B781FF99CC32FBB5FD81FB60FDF4" box="[1046,1211,511,536]" direction="west" orientation="longitude" pageId="17" pageNumber="268" precision="1" value="-96.51">W 96°30.600”</geoCoordinate>
|
|
,
|
|
<elevation id="0DA13675FF99CC32FB65FE7EFAC7FDFB" box="[1222,1308,511,536]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="17" pageNumber="268" unit="m" value="1436.0">
|
|
<quantity id="41747CA3FF99CC32FB65FE7EFAC7FDFB" box="[1222,1308,511,536]" metricMagnitude="3" metricUnit="m" metricValue="1.436" pageId="17" pageNumber="268" unit="m" value="1436.0">1436 m</quantity>
|
|
</elevation>
|
|
, en nopal y mandarina podrida,
|
|
<collectingDate id="E2760E6EFF99CC32FED4FDA5FE25FDD1" box="[375,510,548,573]" pageId="17" pageNumber="268" value="1999-11-16">
|
|
<date id="F232F786FF99CC32FED4FDA5FE25FDD1" box="[375,510,548,573]" pageId="17" pageNumber="268" value="1999-11-16">16-XI-1999</date>
|
|
</collectingDate>
|
|
,
|
|
<collectorName id="2B79B490FF99CC32FDA9FDA4FD5CFDD0" box="[522,647,548,572]" pageId="17" pageNumber="268">J. Márquez</collectorName>
|
|
col.” (
|
|
<specimenCount id="908A1ACFFF99CC32FD75FDA5FD2CFDD0" box="[726,759,548,572]" pageId="17" pageNumber="268" type="female">1♀</specimenCount>
|
|
,
|
|
<collectionCode id="E09D4983FF99CC32FCA7FDA5FC8DFDD0" box="[772,854,548,572]" country="Mexico" httpUri="http://grbio.org/cool/fg24-x4zf" name="Museo de Zoologia &quot;Alfonso L. Herrera&quot;" pageId="17" pageNumber="268">MZFC</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |