400 lines
58 KiB
XML
400 lines
58 KiB
XML
<document id="C81AD6EB8AE163790A6A3F774C2C11E8" ID-DOI="10.5194/fr-19-1-2016" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965366" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712925834647" checkinUser="felipe" docAuthor="Thomas, B. A. & Seyfullah, L. J." docDate="2016" docId="D65A87992440D306FFA1FD33FB6AF983" docLanguage="en" docName="FossRec.19.1.1-9.pdf" docOrigin="Fossil Record 19 (1)" docSource="http://dx.doi.org/10.5194/fr-19-1-2016" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.1:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="1" docTitle="Sigillariostrobus saltwellensis Thomas & Seyfullah 2016, sp. nov." docType="treatment" docVersion="5" lastPageNumber="4" masterDocId="2A63FFE12442D305FFDDFFDFFFB3FFA8" masterDocTitle="Two new species of megasporangiate Sigillariostrobus Schimper (Sigillariostrobaceae) fructifications from the British Coal Measures" masterLastPageNumber="9" masterPageNumber="1" pageNumber="3" updateTime="1713860397130" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6893839004218B1636F202289D5D460A">Two new species of megasporangiate Sigillariostrobus Schimper (Sigillariostrobaceae) fructifications from the British Coal Measures</mods:title>
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<mods:namePart id="66CB5D53AA37878EFC3BD5FA182DF00C">Thomas, B. A.</mods:namePart>
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<mods:namePart id="282F2FB55BCE871F755EA89DB755E41B">Seyfullah, L. J.</mods:namePart>
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<mods:typeOfResource id="35EB659AD948291E034CD14770886502">text</mods:typeOfResource>
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<mods:title id="10CC988821954A80BBFB5D950294E5FF">Fossil Record</mods:title>
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<mods:part id="AD4D2F972DE9784AF007CC1EE4F90B07">
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<mods:date id="9C331E7AA8E84A0BAB3E5408E5B915F5">2016</mods:date>
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<mods:number id="2C76BFC31C4F11C6244B31DDDED66DAE">2015-11-16</mods:number>
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<mods:url id="16EF0D9B3C47D699F71519206CE3C40B">http://dx.doi.org/10.5194/fr-19-1-2016</mods:url>
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<mods:identifier id="E73CE54A4C1362570A4927CF92F123CF" type="DOI">10.5194/fr-19-1-2016</mods:identifier>
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<mods:identifier id="B1B964B7A03B1E509BF9AD76DD503D22" type="ISSN">2193-0074</mods:identifier>
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<treatment id="D65A87992440D306FFA1FD33FB6AF983" ID-DOI="http://doi.org/10.5281/zenodo.11045648" ID-Zenodo-Dep="11045648" LSID="urn:lsid:plazi:treatment:D65A87992440D306FFA1FD33FB6AF983" httpUri="http://treatment.plazi.org/id/D65A87992440D306FFA1FD33FB6AF983" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
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<subSubSection id="16E965042440D307FFA1FD33FDAFFCAC" box="[124,540,748,772]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="5E4C368F2440D307FFA1FD33FDAFFCAC" blockId="2.[124,540,748,804]" box="[124,540,748,772]" pageId="2" pageNumber="3">
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<heading id="050481E32440D307FFA1FD33FDAFFCAC" box="[124,540,748,772]" fontSize="10" level="2" pageId="2" pageNumber="3" reason="5">
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<emphasis id="6C87EA9D2440D307FFA1FD33FDAFFCAC" bold="true" box="[124,540,748,772]" pageId="2" pageNumber="3">
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<taxonomicName id="99F34D0C2440D307FFA1FD33FE72FCAC" authority="Thomas & Seyfullah, 2016" authorityName="Thomas & Seyfullah" authorityYear="2016" box="[124,449,748,772]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="saltwellensis" status="sp. nov.">
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<emphasis id="6C87EA9D2440D307FFA1FD33FE72FCAC" bold="true" box="[124,449,748,772]" italics="true" pageId="2" pageNumber="3">Sigillariostrobus saltwellensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="77B457E62440D307FE1AFD33FDAFFCAC" box="[455,540,748,772]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="16E965042440D306FFA1FCD2FB6AF983" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
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<paragraph id="5E4C368F2440D307FFA1FCD2FEBFFC8C" blockId="2.[124,540,748,804]" box="[124,268,781,804]" pageId="2" pageNumber="3">
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(
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<figureCitation id="C6C82A0A2440D307FF5EFCD2FEB7FC8C" box="[131,260,781,804]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStartId-0="3.[124,191,1315,1337]" captionStartId-1="5.[124,191,1166,1188]" captionTargetBox-0="[136,1454,177,1284]" captionTargetBox-1="[137,1448,178,1135]" captionTargetId-0="figure-15@3.[132,1455,177,1284]" captionTargetId-1="figure-15@5.[132,1455,177,1135]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figure 2. Type specimen of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 53-C.203, Hunterian Museum, Glasgow. Alluvial coalbearing facies – upper Langsettian–Duckmantian. (a) Cone (scale bar 10 mm). (b) Detail of apical portion of the cone (scale bar 5 mm). (c) Detail of basal portion of the cone (scale bar 5 mm). (d) In situ glabrous megaspores (scale bar 2 mm).(e) Spore referable to Laevigatisporites glabratus macerated from the cone (scale bar 1000µm)." captionText-1="Figure 3. Second cone of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 707, Hunterian Museum, Glasgow. (a) The cone (scale bar 20 mm). (b) Detail of the upper part of the cone (scale bar 5 mm). (c) Detail of the mid-region of the cone, showing the megasporophyll impressions at the edge of the cone, often without the cuticle preserved (scale bar 5 mm). (d) Detail of (c), showing the megasporophylls with fissile compression and scattered megaspores (scale bar 5 mm). (e) Detail of (d), showing details of megaspores within the cone (scale bar 2 mm). (f) Megaspore referable to Laevigatisporites glabratus macerated from the spore mass shown in (e) (scale bar 2000 µm)." figureDoi-0="http://doi.org/10.5281/zenodo.10965370" figureDoi-1="http://doi.org/10.5281/zenodo.10965372" httpUri-0="https://zenodo.org/record/10965370/files/figure.png" httpUri-1="https://zenodo.org/record/10965372/files/figure.png" pageId="2" pageNumber="3">Figures 2–3</figureCitation>
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)
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</paragraph>
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<paragraph id="5E4C368F2440D307FFA1FC95FEF9FC28" blockId="2.[124,766,842,896]" pageId="2" pageNumber="3">
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<materialsCitation id="EE9B3CD22440D307FFA1FC95FEF9FC28" collectionCode="C" collectorName="Specimen Pb." location="Hunterian Museum" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Glasgow City" typeStatus="holotype">
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<typeStatus id="8148882D2440D307FFA1FC95FF50FCC9" box="[124,227,842,865]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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:
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<collectorName id="F30653592440D307FF2DFC95FE36FCC9" box="[240,389,842,865]" pageId="2" pageNumber="3">Specimen Pb.</collectorName>
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53-
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<collectionCode id="38E2AE4A2440D307FE6CFC95FE77FCC9" box="[433,452,842,865]" country="Denmark" name="University of Copenhagen" pageId="2" pageNumber="3" type="Herbarium">C</collectionCode>
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.203, stored in the
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<location id="5B2C60542440D307FD49FC95FF6FFC28" LSID="urn:lsid:plazi:treatment:D65A87992440D306FFA1FD33FB6AF983:5B2C60542440D307FD49FC95FF6FFC28" name="Hunterian Museum" pageId="2" pageNumber="3" stateProvince="Glasgow City">Hunterian Museum</location>
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,
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<collectingRegion id="9C37F86D2440D307FF3BFCB6FEF5FC28" box="[230,326,873,896]" country="United Kingdom" name="Glasgow City" pageId="2" pageNumber="3">Glasgow</collectingRegion>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="5E4C368F2440D307FFA1FC4AFEE3FC64" blockId="2.[124,767,917,1921]" pageId="2" pageNumber="3">
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Type locality: Saltwells Pit, near Cradley Heath,
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<collectingRegion id="9C37F86D2440D307FD42FC4AFF01FC64" country="United Kingdom" name="Staffordshire" pageId="2" pageNumber="3">Staffordshire</collectingRegion>
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,
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<collectingCountry id="26E4761F2440D307FF60FC6AFEFFFC64" box="[189,332,949,972]" name="United Kingdom" pageId="2" pageNumber="3">Great Britain</collectingCountry>
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.
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</paragraph>
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<paragraph id="5E4C368F2440D307FFA1FC3FFE2FFBBF" blockId="2.[124,767,917,1921]" pageId="2" pageNumber="3">Type stratum: Alluvial coal-bearing facies – upper Langsettian–Duckmantian.</paragraph>
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<paragraph id="5E4C368F2440D307FFA1FBF4FE35FBCA" blockId="2.[124,767,917,1921]" pageId="2" pageNumber="3">
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Etymology: Named after the locality of the specimen that we designate as the
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<typeStatus id="8148882D2440D307FE8DFB94FE32FBCA" box="[336,385,1099,1122]" pageId="2" pageNumber="3">type</typeStatus>
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.
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</paragraph>
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<paragraph id="5E4C368F2440D307FFA1FBA9FD34FA24" blockId="2.[124,767,917,1921]" pageId="2" pageNumber="3">
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Diagnosis: Cone at least
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<quantity id="990B9B6A2440D307FE7EFBA9FE58FB25" box="[419,491,1142,1165]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="2" pageNumber="3" unit="mm" value="86.0">86 mm</quantity>
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long, ca.
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<quantity id="990B9B6A2440D307FDB6FBA9FD68FB25" box="[619,731,1142,1165]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="2" pageNumber="3" unit="mm" value="18.5" valueMax="20.0" valueMin="17.0">17–20 mm</quantity>
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||
in diameter, axis 1.2 mm, pedicel ca.
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||
<quantity id="990B9B6A2440D307FDD1FB49FDF4FB05" box="[524,583,1174,1197]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="3" unit="mm" value="8.0">8 mm</quantity>
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long, sporangia
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<quantity id="990B9B6A2440D307FFA1FB69FF04FB65" box="[124,183,1206,1229]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="3" unit="mm" value="7.0">7 mm</quantity>
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long, sporophyll laminae 12 broad at base, 6.3 mm long spreading outwards from the cone. Sporangia
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<quantity id="990B9B6A2440D307FD1EFB09FD4DFB45" box="[707,766,1238,1261]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="3" unit="mm" value="7.0">7 mm</quantity>
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||
long, 2.5 mm high. Megaspores trilete, probably originally plano-convex, average diameter 2000 µm in their equatorial planes, smooth walls with a thickened area between the arms of the trilete mark. Lesurae approximately half the spore radius, ca. 15 µm broad at the centre of the spore.
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</paragraph>
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<paragraph id="5E4C368F2440D307FFA1FA7FFF11F89E" blockId="2.[124,767,917,1921]" pageId="2" pageNumber="3">
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Description: Both cones are preserved as dark compression (
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<figureCitation id="C6C82A0A2440D307FF59FA1FFF4EFA7F" box="[132,253,1472,1495]" captionStart="Figure 2" captionStartId="3.[124,191,1315,1337]" captionTargetBox="[136,1454,177,1284]" captionTargetId="figure-15@3.[132,1455,177,1284]" captionTargetPageId="3" captionText="Figure 2. Type specimen of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 53-C.203, Hunterian Museum, Glasgow. Alluvial coalbearing facies – upper Langsettian–Duckmantian. (a) Cone (scale bar 10 mm). (b) Detail of apical portion of the cone (scale bar 5 mm). (c) Detail of basal portion of the cone (scale bar 5 mm). (d) In situ glabrous megaspores (scale bar 2 mm).(e) Spore referable to Laevigatisporites glabratus macerated from the cone (scale bar 1000µm)." figureDoi="http://doi.org/10.5281/zenodo.10965370" httpUri="https://zenodo.org/record/10965370/files/figure.png" pageId="2" pageNumber="3">Figs. 2a–b</figureCitation>
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,
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<figureCitation id="C6C82A0A2440D307FED3FA1FFEF6FA7F" box="[270,325,1472,1495]" captionStart="Figure 3" captionStartId="5.[124,191,1166,1188]" captionTargetBox="[137,1448,178,1135]" captionTargetId="figure-15@5.[132,1455,177,1135]" captionTargetPageId="5" captionText="Figure 3. Second cone of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 707, Hunterian Museum, Glasgow. (a) The cone (scale bar 20 mm). (b) Detail of the upper part of the cone (scale bar 5 mm). (c) Detail of the mid-region of the cone, showing the megasporophyll impressions at the edge of the cone, often without the cuticle preserved (scale bar 5 mm). (d) Detail of (c), showing the megasporophylls with fissile compression and scattered megaspores (scale bar 5 mm). (e) Detail of (d), showing details of megaspores within the cone (scale bar 2 mm). (f) Megaspore referable to Laevigatisporites glabratus macerated from the spore mass shown in (e) (scale bar 2000 µm)." figureDoi="http://doi.org/10.5281/zenodo.10965372" httpUri="https://zenodo.org/record/10965372/files/figure.png" pageId="2" pageNumber="3">3a–b</figureCitation>
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), although they are broken somewhere above their bases (
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<figureCitation id="C6C82A0A2440D307FE88FA3FFE60FA5F" box="[341,467,1504,1527]" captionStart="Figure 2" captionStartId="3.[124,191,1315,1337]" captionTargetBox="[136,1454,177,1284]" captionTargetId="figure-15@3.[132,1455,177,1284]" captionTargetPageId="3" captionText="Figure 2. Type specimen of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 53-C.203, Hunterian Museum, Glasgow. Alluvial coalbearing facies – upper Langsettian–Duckmantian. (a) Cone (scale bar 10 mm). (b) Detail of apical portion of the cone (scale bar 5 mm). (c) Detail of basal portion of the cone (scale bar 5 mm). (d) In situ glabrous megaspores (scale bar 2 mm).(e) Spore referable to Laevigatisporites glabratus macerated from the cone (scale bar 1000µm)." figureDoi="http://doi.org/10.5281/zenodo.10965370" httpUri="https://zenodo.org/record/10965370/files/figure.png" pageId="2" pageNumber="3">Figs. 2a, c</figureCitation>
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,
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<figureCitation id="C6C82A0A2440D307FE39FA3FFDACFA5F" box="[484,543,1504,1527]" captionStart="Figure 3" captionStartId="5.[124,191,1166,1188]" captionTargetBox="[137,1448,178,1135]" captionTargetId="figure-15@5.[132,1455,177,1135]" captionTargetPageId="5" captionText="Figure 3. Second cone of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 707, Hunterian Museum, Glasgow. (a) The cone (scale bar 20 mm). (b) Detail of the upper part of the cone (scale bar 5 mm). (c) Detail of the mid-region of the cone, showing the megasporophyll impressions at the edge of the cone, often without the cuticle preserved (scale bar 5 mm). (d) Detail of (c), showing the megasporophylls with fissile compression and scattered megaspores (scale bar 5 mm). (e) Detail of (d), showing details of megaspores within the cone (scale bar 2 mm). (f) Megaspore referable to Laevigatisporites glabratus macerated from the spore mass shown in (e) (scale bar 2000 µm)." figureDoi="http://doi.org/10.5281/zenodo.10965372" httpUri="https://zenodo.org/record/10965372/files/figure.png" pageId="2" pageNumber="3">3a, c</figureCitation>
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), so their complete length is unknown. The megasporophylls have free tips approximately 6.3 mm long (
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<figureCitation id="C6C82A0A2440D307FE73F9FFFE4FF99F" box="[430,508,1568,1591]" captionStart="Figure 3" captionStartId="5.[124,191,1166,1188]" captionTargetBox="[137,1448,178,1135]" captionTargetId="figure-15@5.[132,1455,177,1135]" captionTargetPageId="5" captionText="Figure 3. Second cone of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 707, Hunterian Museum, Glasgow. (a) The cone (scale bar 20 mm). (b) Detail of the upper part of the cone (scale bar 5 mm). (c) Detail of the mid-region of the cone, showing the megasporophyll impressions at the edge of the cone, often without the cuticle preserved (scale bar 5 mm). (d) Detail of (c), showing the megasporophylls with fissile compression and scattered megaspores (scale bar 5 mm). (e) Detail of (d), showing details of megaspores within the cone (scale bar 2 mm). (f) Megaspore referable to Laevigatisporites glabratus macerated from the spore mass shown in (e) (scale bar 2000 µm)." figureDoi="http://doi.org/10.5281/zenodo.10965372" httpUri="https://zenodo.org/record/10965372/files/figure.png" pageId="2" pageNumber="3">Fig. 3d</figureCitation>
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), only slightly diverging from the sides of the cone. The megasporophylls are not well preserved (
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<figureCitation id="C6C82A0A2440D307FF29F9BFFEDAF9DF" box="[244,361,1632,1655]" captionStart="Figure 2" captionStartId="3.[124,191,1315,1337]" captionTargetBox="[136,1454,177,1284]" captionTargetId="figure-15@3.[132,1455,177,1284]" captionTargetPageId="3" captionText="Figure 2. Type specimen of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 53-C.203, Hunterian Museum, Glasgow. Alluvial coalbearing facies – upper Langsettian–Duckmantian. (a) Cone (scale bar 10 mm). (b) Detail of apical portion of the cone (scale bar 5 mm). (c) Detail of basal portion of the cone (scale bar 5 mm). (d) In situ glabrous megaspores (scale bar 2 mm).(e) Spore referable to Laevigatisporites glabratus macerated from the cone (scale bar 1000µm)." figureDoi="http://doi.org/10.5281/zenodo.10965370" httpUri="https://zenodo.org/record/10965370/files/figure.png" pageId="2" pageNumber="3">Figs. 2b–c</figureCitation>
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,
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<figureCitation id="C6C82A0A2440D307FEABF9BFFE21F9DF" box="[374,402,1632,1655]" captionStart="Figure 3" captionStartId="5.[124,191,1166,1188]" captionTargetBox="[137,1448,178,1135]" captionTargetId="figure-15@5.[132,1455,177,1135]" captionTargetPageId="5" captionText="Figure 3. Second cone of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 707, Hunterian Museum, Glasgow. (a) The cone (scale bar 20 mm). (b) Detail of the upper part of the cone (scale bar 5 mm). (c) Detail of the mid-region of the cone, showing the megasporophyll impressions at the edge of the cone, often without the cuticle preserved (scale bar 5 mm). (d) Detail of (c), showing the megasporophylls with fissile compression and scattered megaspores (scale bar 5 mm). (e) Detail of (d), showing details of megaspores within the cone (scale bar 2 mm). (f) Megaspore referable to Laevigatisporites glabratus macerated from the spore mass shown in (e) (scale bar 2000 µm)." figureDoi="http://doi.org/10.5281/zenodo.10965372" httpUri="https://zenodo.org/record/10965372/files/figure.png" pageId="2" pageNumber="3">3d</figureCitation>
|
||
) and the compression material is very brittle, so the cuticle could not be prepared. Numerous obvious large red-brown megaspores are visible to the naked eye (
|
||
<figureCitation id="C6C82A0A2440D307FF72F960FEBAF97E" box="[175,265,1727,1750]" captionStart="Figure 2" captionStartId="3.[124,191,1315,1337]" captionTargetBox="[136,1454,177,1284]" captionTargetId="figure-15@3.[132,1455,177,1284]" captionTargetPageId="3" captionText="Figure 2. Type specimen of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 53-C.203, Hunterian Museum, Glasgow. Alluvial coalbearing facies – upper Langsettian–Duckmantian. (a) Cone (scale bar 10 mm). (b) Detail of apical portion of the cone (scale bar 5 mm). (c) Detail of basal portion of the cone (scale bar 5 mm). (d) In situ glabrous megaspores (scale bar 2 mm).(e) Spore referable to Laevigatisporites glabratus macerated from the cone (scale bar 1000µm)." figureDoi="http://doi.org/10.5281/zenodo.10965370" httpUri="https://zenodo.org/record/10965370/files/figure.png" pageId="2" pageNumber="3">Figs. 2d</figureCitation>
|
||
,
|
||
<figureCitation id="C6C82A0A2440D307FEC9F960FE83F97E" box="[276,304,1727,1750]" captionStart="Figure 3" captionStartId="5.[124,191,1166,1188]" captionTargetBox="[137,1448,178,1135]" captionTargetId="figure-15@5.[132,1455,177,1135]" captionTargetPageId="5" captionText="Figure 3. Second cone of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 707, Hunterian Museum, Glasgow. (a) The cone (scale bar 20 mm). (b) Detail of the upper part of the cone (scale bar 5 mm). (c) Detail of the mid-region of the cone, showing the megasporophyll impressions at the edge of the cone, often without the cuticle preserved (scale bar 5 mm). (d) Detail of (c), showing the megasporophylls with fissile compression and scattered megaspores (scale bar 5 mm). (e) Detail of (d), showing details of megaspores within the cone (scale bar 2 mm). (f) Megaspore referable to Laevigatisporites glabratus macerated from the spore mass shown in (e) (scale bar 2000 µm)." figureDoi="http://doi.org/10.5281/zenodo.10965372" httpUri="https://zenodo.org/record/10965372/files/figure.png" pageId="2" pageNumber="3">3e</figureCitation>
|
||
). They all measure approximately 2000 µm in diameter in their equatorial plane, with lesurae 50 µm long. The entire surfaces of the spores are smooth (
|
||
<figureCitation id="C6C82A0A2440D307FD7FF920FD49F8BE" box="[674,762,1791,1814]" captionStart="Figure 2" captionStartId="3.[124,191,1315,1337]" captionTargetBox="[136,1454,177,1284]" captionTargetId="figure-15@3.[132,1455,177,1284]" captionTargetPageId="3" captionText="Figure 2. Type specimen of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 53-C.203, Hunterian Museum, Glasgow. Alluvial coalbearing facies – upper Langsettian–Duckmantian. (a) Cone (scale bar 10 mm). (b) Detail of apical portion of the cone (scale bar 5 mm). (c) Detail of basal portion of the cone (scale bar 5 mm). (d) In situ glabrous megaspores (scale bar 2 mm).(e) Spore referable to Laevigatisporites glabratus macerated from the cone (scale bar 1000µm)." figureDoi="http://doi.org/10.5281/zenodo.10965370" httpUri="https://zenodo.org/record/10965370/files/figure.png" pageId="2" pageNumber="3">Figs. 2e</figureCitation>
|
||
,
|
||
<figureCitation id="C6C82A0A2440D307FFA1F8C0FF26F89E" box="[124,149,1823,1846]" captionStart="Figure 3" captionStartId="5.[124,191,1166,1188]" captionTargetBox="[137,1448,178,1135]" captionTargetId="figure-15@5.[132,1455,177,1135]" captionTargetPageId="5" captionText="Figure 3. Second cone of Sigillariostrobus saltwellensis sp. nov. Specimen Pb. 707, Hunterian Museum, Glasgow. (a) The cone (scale bar 20 mm). (b) Detail of the upper part of the cone (scale bar 5 mm). (c) Detail of the mid-region of the cone, showing the megasporophyll impressions at the edge of the cone, often without the cuticle preserved (scale bar 5 mm). (d) Detail of (c), showing the megasporophylls with fissile compression and scattered megaspores (scale bar 5 mm). (e) Detail of (d), showing details of megaspores within the cone (scale bar 2 mm). (f) Megaspore referable to Laevigatisporites glabratus macerated from the spore mass shown in (e) (scale bar 2000 µm)." figureDoi="http://doi.org/10.5281/zenodo.10965372" httpUri="https://zenodo.org/record/10965372/files/figure.png" pageId="2" pageNumber="3">3f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5E4C368F2440D307FFA1F895FC30FF4F" blockId="2.[124,767,917,1921]" lastBlockId="2.[819,1463,176,1900]" pageId="2" pageNumber="3">
|
||
Comments:
|
||
<collectionCode id="38E2AE4A2440D307FF22F895FEA1F8C9" box="[255,274,1866,1889]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="2" pageNumber="3" type="Herbarium">A</collectionCode>
|
||
second specimen, Pb. 707, also stored in the Hunterian Museum,
|
||
<collectingRegion id="9C37F86D2440D307FE84F8B5FE0BF829" box="[345,440,1898,1921]" country="United Kingdom" name="Glasgow City" pageId="2" pageNumber="3">Glasgow</collectingRegion>
|
||
, is referred to this species, but the locality and horizon information for this specimen is not known.
|
||
</paragraph>
|
||
<paragraph id="5E4C368F2440D307FCEEFF24FBD4FD79" blockId="2.[819,1463,176,1900]" pageId="2" pageNumber="3">
|
||
Discussion: The megaspores clearly fit within the parameters of the dispersed spore genus
|
||
<taxonomicName id="99F34D0C2440D307FB17FEC4FA31FE9B" box="[1226,1410,283,307]" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="6C87EA9D2440D307FB17FEC4FA31FE9B" box="[1226,1410,283,307]" italics="true" pageId="2" pageNumber="3">Laevigatisporites</emphasis>
|
||
</taxonomicName>
|
||
and are closest to the species
|
||
<taxonomicName id="99F34D0C2440D307FBBBFEE4FC44FEDA" authority="(Zerndt) Potonie and Kremp, 1956" authorityName="Potonie and Kremp" authorityYear="1956" baseAuthorityName="Zerndt" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2440D307FBBBFEE4FB5CFEFB" box="[1126,1263,315,339]" italics="true" pageId="2" pageNumber="3">L. glabratus</emphasis>
|
||
(Zerndt) Potonié and Kremp, 1956
|
||
</taxonomicName>
|
||
. We agree with
|
||
<bibRefCitation id="3A624B7E2440D307FB6CFE84FA37FEDA" author="Karczewska, J." box="[1201,1412,347,370]" pageId="2" pageNumber="3" pagination="267 - 345" refId="ref5662" refString="Karczewska, J.: Carboniferous spores from the Chelm I boring (Eastern Poland), Acta Palaeontol. Pol., 12, 267 - 345, 12 Taf, 1967." type="journal article" year="1967">Karczewska (1967)</bibRefCitation>
|
||
that the spores described by
|
||
<bibRefCitation id="3A624B7E2440D307FB99FEA4FB6EFE3A" author="Zerndt, J." box="[1092,1245,379,402]" pageId="2" pageNumber="3" pagination="39 - 70" refId="ref6481" refString="Zerndt, J.: Megasporen aus einem Floz in Libiaz (Stephanien). Akademie Polonaise des Sciences Bulletin International, ser. BI, 39 - 70, 1930." type="book chapter" year="1930">Zerndt (1930)</bibRefCitation>
|
||
as
|
||
<typeStatus id="8148882D2440D307FAD5FEA4FAF4FE3A" box="[1288,1351,379,402]" pageId="2" pageNumber="3">Types</typeStatus>
|
||
9 and 10 should be referred to
|
||
<taxonomicName id="99F34D0C2440D307FBC1FE45FAF2FE1A" box="[1052,1345,410,434]" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2440D307FBC1FE45FAF2FE1A" box="[1052,1345,410,434]" italics="true" pageId="2" pageNumber="3">Laevigatisporites glabratus</emphasis>
|
||
</taxonomicName>
|
||
as a result of
|
||
<bibRefCitation id="3A624B7E2440D307FC8BFE64FB98FE7A" author="Bochenski, T. A." box="[854,1067,443,466]" pageId="2" pageNumber="3" pagination="193 - 240" refId="ref5312" refString="Bochenski, T. A.: Uber sporophyllstande [Blutem] einige Lepidophyten aus dem produktiven Karbon Polens, Jargang XII (1936) der Polischen Geologischen Gesellschaft, 193 - 240, 1936." type="journal article" year="1936">Bochenski’s (1936)</bibRefCitation>
|
||
description of the spores from his
|
||
<taxonomicName id="99F34D0C2440D307FCEEFE05FBEAFE5A" authorityName="Bochenski" authorityYear="1936" box="[819,1113,474,498]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="czarnockii">
|
||
<emphasis id="6C87EA9D2440D307FCEEFE05FBEAFE5A" box="[819,1113,474,498]" italics="true" pageId="2" pageNumber="3">Sigillariostrobus czarnockii</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="3A624B7E2440D307FBBBFE04FA90FE5A" author="Bochenski, T. A." box="[1126,1315,475,498]" pageId="2" pageNumber="3" pagination="193 - 240" refId="ref5312" refString="Bochenski, T. A.: Uber sporophyllstande [Blutem] einige Lepidophyten aus dem produktiven Karbon Polens, Jargang XII (1936) der Polischen Geologischen Gesellschaft, 193 - 240, 1936." type="journal article" year="1936">Bochenski (1936)</bibRefCitation>
|
||
described the spores as 440 to 2700 µm in diameter, and underdeveloped spores which were triangular, thicker than the normal ones and with strongly marked lesurae.
|
||
<bibRefCitation id="3A624B7E2440D307FB1BFDE5FAD1FDF9" author="Zerndt, J." box="[1222,1378,570,593]" pageId="2" pageNumber="3" pagination="10 - 16" refId="ref6513" refString="Zerndt, J.: Megasporen aus dem Zwikauer und Lugau-Olsnitzer Karbon, Jahrbuch Berg- und Huttenwesen in Sachsen 1932, 10 - 16, 1932." type="journal article" year="1932">Zerndt (1932)</bibRefCitation>
|
||
agreed that Bochenski’s finding of his megaspores of
|
||
<typeStatus id="8148882D2440D307FA86FD85FA29FDD9" box="[1371,1434,602,625]" pageId="2" pageNumber="3">Types</typeStatus>
|
||
9 and 10 (
|
||
<bibRefCitation id="3A624B7E2440D307FC4CFDA5FB93FD39" author="Zerndt, J." box="[913,1056,634,657]" pageId="2" pageNumber="3" pagination="39 - 70" refId="ref6481" refString="Zerndt, J.: Megasporen aus einem Floz in Libiaz (Stephanien). Akademie Polonaise des Sciences Bulletin International, ser. BI, 39 - 70, 1930." type="book chapter" year="1930">Zerndt, 1930</bibRefCitation>
|
||
) in the same cone was considerably important to the understanding of
|
||
<taxonomicName id="99F34D0C2440D307FB69FD46FAC7FD19" box="[1204,1396,665,689]" genus="Triletes" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2440D307FB69FD46FAC7FD19" box="[1204,1396,665,689]" italics="true" pageId="2" pageNumber="3">Triletes glabratus</emphasis>
|
||
</taxonomicName>
|
||
(now
|
||
<taxonomicName id="99F34D0C2440D307FCEEFD66FBE4FD79" box="[819,1111,697,721]" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2440D307FCEEFD66FBE4FD79" box="[819,1111,697,721]" italics="true" pageId="2" pageNumber="3">Laevigatisporites glabratus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5E4C368F2440D307FC93FD05FB1FFB67" blockId="2.[819,1463,176,1900]" pageId="2" pageNumber="3">
|
||
We also agree that there are no reliable grounds to distinguish such species as
|
||
<taxonomicName id="99F34D0C2440D307FB93FD26FA05FCB9" authority="Wicher" authorityName="Wicher" box="[1102,1462,761,785]" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="primus">
|
||
<emphasis id="6C87EA9D2440D307FB93FD26FAEEFCB9" box="[1102,1373,761,785]" italics="true" pageId="2" pageNumber="3">Laevigatisporites primus</emphasis>
|
||
Wicher
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="99F34D0C2440D307FCBBFCC6FB51FC98" authority="Ibrahim" authorityName="Ibrahim" box="[870,1250,793,817]" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="reinschi">
|
||
<emphasis id="6C87EA9D2440D307FCBBFCC6FBCDFC99" box="[870,1150,793,817]" italics="true" pageId="2" pageNumber="3">Laevigatisporites reinschi</emphasis>
|
||
Ibrahim
|
||
</taxonomicName>
|
||
, as has been done by
|
||
<bibRefCitation id="3A624B7E2440D307FC88FCE6FBC4FCF8" author="Potonie, R. & Kremp, G." box="[853,1143,825,848]" pageId="2" pageNumber="3" pagination="1 - 136" refId="ref5877" refString="Potonie, R. and Kremp, G.: Die Sporae dispersae des Ruhrkarbons, ihre morphographie und stratigraphie mit ausblicken auf arten anderer gebiete und zeitabschnitte, Palaeontogr. Abt. B., 98, 1 - 136, 1955." type="journal article" year="1955">Potonié and Kremp (1955)</bibRefCitation>
|
||
. Erecting the species
|
||
<taxonomicName id="99F34D0C2440D307FA82FCE6FB9EFCD8" authority=", Wicher (1934)" authorityName=", Wicher" authorityYear="1934" class="Sphagnopsida" family="Sphagnaceae" genus="Sporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="primus">
|
||
<emphasis id="6C87EA9D2440D307FA82FCE6FCCEFCD9" italics="true" pageId="2" pageNumber="3">Sporites primus</emphasis>
|
||
,
|
||
<bibRefCitation id="3A624B7E2440D307FC53FC86FB9EFCD8" author="Wicher, C. A." box="[910,1069,857,880]" pageId="2" pageNumber="3" pagination="165 - 212" refId="ref6311" refString="Wicher, C. A.: Sporenformen der Flammkohle des Ruhrgebietes, Arbeit. Instit. Palaobot. Petrograph. Brennsteine, 4, 165 - 212, 1934." type="journal article" year="1934">Wicher (1934)</bibRefCitation>
|
||
</taxonomicName>
|
||
assigned only the specific name to the spores, described by
|
||
<bibRefCitation id="3A624B7E2440D307FB8CFCA6FA30FC38" author="Bennie, J. & Kidston, R." box="[1105,1411,889,912]" pageId="2" pageNumber="3" pagination="82 - 117" refId="ref5118" refString="Bennie, J. and Kidston, R.: On the occurrence of spores in the Carboniferous formation of Scotland, Proceedings of the Royal Physical Society of Edinburgh, 9, 82 - 117, 1886." type="journal article" year="1886">Bennie and Kidston (1886)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="3A624B7E2440D307FCEEFC46FC7AFC18" author="Zerndt, J." box="[819,969,921,944]" pageId="2" pageNumber="3" pagination="39 - 70" refId="ref6481" refString="Zerndt, J.: Megasporen aus einem Floz in Libiaz (Stephanien). Akademie Polonaise des Sciences Bulletin International, ser. BI, 39 - 70, 1930." type="book chapter" year="1930">Zerndt (1930)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="99F34D0C2440D307FC33FC47FB8DFC18" authorityName="P.F.Reinsch" authorityYear="1881" box="[1006,1086,920,944]" genus="Triletes" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6C87EA9D2440D307FC33FC47FB8DFC18" box="[1006,1086,920,944]" italics="true" pageId="2" pageNumber="3">Triletes</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="8148882D2440D307FB9BFC46FBC9FC18" box="[1094,1146,921,944]" pageId="2" pageNumber="3">Type</typeStatus>
|
||
<collectionCode id="38E2AE4A2440D307FB5FFC46FB38FC18" box="[1154,1163,921,944]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&quot;Alexandru Ioan Cuza&quot; University" pageId="2" pageNumber="3" type="Herbarium">I</collectionCode>
|
||
Kidston.
|
||
<bibRefCitation id="3A624B7E2440D307FB28FC46FA22FC18" author="Wicher, C. A." box="[1269,1425,921,944]" pageId="2" pageNumber="3" pagination="165 - 212" refId="ref6311" refString="Wicher, C. A.: Sporenformen der Flammkohle des Ruhrgebietes, Arbeit. Instit. Palaobot. Petrograph. Brennsteine, 4, 165 - 212, 1934." type="journal article" year="1934">Wicher (1934)</bibRefCitation>
|
||
interpreted the species
|
||
<taxonomicName id="99F34D0C2440D307FBC8FC67FA8FFC78" authority="Ibrahim" authorityName="Ibrahim" box="[1045,1340,952,976]" genus="Sporonites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" pageId="2" pageNumber="3" rank="species" species="reinschi">
|
||
<emphasis id="6C87EA9D2440D307FBC8FC67FB53FC78" box="[1045,1248,952,976]" italics="true" pageId="2" pageNumber="3">Sporonites reinschi</emphasis>
|
||
Ibrahim
|
||
</taxonomicName>
|
||
to be identical to
|
||
<taxonomicName id="99F34D0C2440D307FC5EFC07FB98FC58" authorityName=", Wicher" authorityYear="1934" box="[899,1067,984,1008]" class="Sphagnopsida" family="Sphagnaceae" genus="Sporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="primus">
|
||
<emphasis id="6C87EA9D2440D307FC5EFC07FB98FC58" box="[899,1067,984,1008]" italics="true" pageId="2" pageNumber="3">Sporites primus</emphasis>
|
||
</taxonomicName>
|
||
; he erected himself and – contrary to the law of priority – included
|
||
<taxonomicName id="99F34D0C2440D307FBAEFC27FA29FBB8" authority="Ibrahim" authorityName="Ibrahim" box="[1139,1434,1016,1040]" genus="Sporonites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" pageId="2" pageNumber="3" rank="species" species="reinschi">
|
||
<emphasis id="6C87EA9D2440D307FBAEFC27FA8CFBB8" box="[1139,1343,1016,1040]" italics="true" pageId="2" pageNumber="3">Sporonites reinschi</emphasis>
|
||
Ibrahim
|
||
</taxonomicName>
|
||
in the synonymy of
|
||
<taxonomicName id="99F34D0C2440D307FC2DFBC7FB02FB87" authority="Wicher." authorityName=", Wicher" authorityYear="1934" box="[1008,1201,1048,1072]" class="Sphagnopsida" family="Sphagnaceae" genus="Sporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="primus">
|
||
<emphasis id="6C87EA9D2440D307FC2DFBC7FBE5FB98" box="[1008,1110,1048,1072]" italics="true" pageId="2" pageNumber="3">S. primus</emphasis>
|
||
Wicher.
|
||
</taxonomicName>
|
||
After the publication of Bochenski’s work, it became clear that the spores of
|
||
<taxonomicName id="99F34D0C2440D307FABBFBE7FA05FBF8" authorityName="P.F.Reinsch" authorityYear="1881" box="[1382,1462,1080,1104]" genus="Triletes" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6C87EA9D2440D307FABBFBE7FA05FBF8" box="[1382,1462,1080,1104]" italics="true" pageId="2" pageNumber="3">Triletes</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="8148882D2440D307FCEEFB87FCD4FBC7" box="[819,871,1112,1135]" pageId="2" pageNumber="3">Type</typeStatus>
|
||
<collectionCode id="38E2AE4A2440D307FCAFFB87FCC8FBC7" box="[882,891,1112,1135]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&quot;Alexandru Ioan Cuza&quot; University" pageId="2" pageNumber="3" type="Herbarium">I</collectionCode>
|
||
Kidston (=
|
||
<taxonomicName id="99F34D0C2440D307FBD6FB87FB6AFBD8" box="[1035,1241,1112,1136]" genus="Sporonites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" pageId="2" pageNumber="3" rank="species" species="reinschi">
|
||
<emphasis id="6C87EA9D2440D307FBD6FB87FB6AFBD8" box="[1035,1241,1112,1136]" italics="true" pageId="2" pageNumber="3">Sporonites reinschi</emphasis>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName id="99F34D0C2440D307FADFFB87FA1EFBD8" authorityName=", Wicher" authorityYear="1934" box="[1282,1453,1112,1136]" class="Sphagnopsida" family="Sphagnaceae" genus="Sporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="primus">
|
||
<emphasis id="6C87EA9D2440D307FADFFB87FA1EFBD8" box="[1282,1453,1112,1136]" italics="true" pageId="2" pageNumber="3">Sporites primus</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="99F34D0C2440D307FCB8FBA7FBC4FB27" authority="Zerndt" authorityName="Zerndt" box="[869,1143,1144,1168]" genus="Triletes" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2440D307FCB8FBA7FB96FB38" box="[869,1061,1144,1168]" italics="true" pageId="2" pageNumber="3">Triletes glabratus</emphasis>
|
||
Zerndt
|
||
</taxonomicName>
|
||
belong to the same species.
|
||
<taxonomicName id="99F34D0C2440D307FCEEFB47FB3AFB07" authority="Zerndt, 1930" authorityName="Zerndt" authorityYear="1930" box="[819,1161,1176,1200]" genus="Triletes" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2440D307FCEEFB47FC42FB18" box="[819,1009,1176,1200]" italics="true" pageId="2" pageNumber="3">Triletes glabratus</emphasis>
|
||
<bibRefCitation id="3A624B7E2440D307FC26FB47FB3AFB07" author="Zerndt, J." box="[1019,1161,1176,1199]" pageId="2" pageNumber="3" pagination="39 - 70" refId="ref6481" refString="Zerndt, J.: Megasporen aus einem Floz in Libiaz (Stephanien). Akademie Polonaise des Sciences Bulletin International, ser. BI, 39 - 70, 1930." type="book chapter" year="1930">Zerndt, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
, is the oldest specific name and, therefore, has specific priority.
|
||
</paragraph>
|
||
<paragraph id="5E4C368F2440D307FC93FB07FAB8FA66" blockId="2.[819,1463,176,1900]" pageId="2" pageNumber="3">
|
||
In situ spores identified as being identical to
|
||
<taxonomicName id="99F34D0C2440D307FAAFFB08FAC4FAA7" authority="(Zerndt) Potonie and Kremp" authorityName="Potonie and Kremp" baseAuthorityName="Zerndt" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2440D307FAAFFB08FB97FAA7" italics="true" pageId="2" pageNumber="3">Laevigatisporites glabratus</emphasis>
|
||
(Zerndt) Potonié and Kremp
|
||
</taxonomicName>
|
||
have been also described from four species of
|
||
<taxonomicName id="99F34D0C2440D307FB23FAC8FA1CFA87" authorityName="Feistmantel" authorityYear="1876" baseAuthorityName="Schimper" baseAuthorityYear="1870" box="[1278,1455,1303,1327]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6C87EA9D2440D307FB23FAC8FA1CFA87" box="[1278,1455,1303,1327]" italics="true" pageId="2" pageNumber="3">Sigillariostrobus</emphasis>
|
||
</taxonomicName>
|
||
. They are
|
||
<taxonomicName id="99F34D0C2440D307FC7EFAE8FB04FAE6" authority="Zeiller, 1884" authorityName="Zeiller" authorityYear="1884" box="[931,1207,1335,1359]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="tieghemi">
|
||
<emphasis id="6C87EA9D2440D307FC7EFAE8FBAEFAE7" box="[931,1053,1335,1359]" italics="true" pageId="2" pageNumber="3">S. tieghemi</emphasis>
|
||
<bibRefCitation id="3A624B7E2440D307FBF7FAE8FB04FAE6" author="Zeiller, R." box="[1066,1207,1335,1358]" pageId="2" pageNumber="3" pagination="256 - 280" refId="ref6413" refString="Zeiller, R.: Cones de fructifications de Sigillaires, Ann. Sci. nat. Bot., 19, 256 - 280, pls 11 - 12, 1884." type="journal article" year="1884">Zeiller, 1884</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="3A624B7E2440D307FB16FAE8FAECFAE6" author="Schenk, A." box="[1227,1375,1335,1358]" pageId="2" pageNumber="3" pagination="127 - 131" refId="ref5921" refString="Schenk, A.: Uber Sigillariostrobus. Abdruck a. d. Berichte d. math. - phys. Classeb d. Konigl, Sachsischen Gesellschaft. d. Wissenschaften zu Leipzig. Ber. 37, S 127 - 131, 1885." type="journal article" year="1885">Schenk 1885</bibRefCitation>
|
||
, Zeiller 1888),
|
||
<taxonomicName id="99F34D0C2440D307FCA2FA88FAD3FAC6" authority="(Lesquereux) White, 1903" authorityName="White" authorityYear="1903" baseAuthorityName="Lesquereux" box="[895,1376,1367,1391]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="quadrangularis">
|
||
<emphasis id="6C87EA9D2440D307FCA2FA88FBF3FAC7" box="[895,1088,1367,1391]" italics="true" pageId="2" pageNumber="3">S. quadrangularis</emphasis>
|
||
(Lesquereux)
|
||
<bibRefCitation id="3A624B7E2440D307FB01FA88FAD3FAC6" author="White, D." box="[1244,1376,1367,1390]" pageId="2" pageNumber="3" pagination="1 - 106" refId="ref6262" refString="White, D.: Stratigraphy and paleontology of the Upper Carboniferous rocks of the Kansas Section, Department of the Interior, U. S. Geological Survey Bulletin 211, Ser. C, Systematic Geology: Paleontology, 6, 1 - 106, 1903." type="journal article" year="1903">White, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="3A624B7E2440D307FAB2FA88FCD8FA26" author="Wood, J. M." pageId="2" pageNumber="3" pagination="141 - 154" refId="ref6377" refString="Wood, J. M.: The morphology and relationships of sigillarian fructifications from the Lower Pennsylvanian of Indiana, Am. Mid. Nat., 58, 141 - 154, 1957." type="journal article" year="1957">Wood, 1957</bibRefCitation>
|
||
),
|
||
<taxonomicName id="99F34D0C2440D307FCA0FAA8FB0DFA26" authority="Bochenski, 1936" authorityName="Bochenski" authorityYear="1936" box="[893,1214,1399,1423]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="czarnockii">
|
||
<emphasis id="6C87EA9D2440D307FCA0FAA8FBB6FA27" box="[893,1029,1399,1423]" italics="true" pageId="2" pageNumber="3">S. czarnockii</emphasis>
|
||
<bibRefCitation id="3A624B7E2440D307FBD7FAA8FB0DFA26" author="Bochenski, T. A." box="[1034,1214,1399,1422]" pageId="2" pageNumber="3" pagination="193 - 240" refId="ref5312" refString="Bochenski, T. A.: Uber sporophyllstande [Blutem] einige Lepidophyten aus dem produktiven Karbon Polens, Jargang XII (1936) der Polischen Geologischen Gesellschaft, 193 - 240, 1936." type="journal article" year="1936">Bochenski, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="99F34D0C2440D307FB2EFAA8FC12FA06" authority="Abbott, 1963" authorityName="Abbott" authorityYear="1963" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="leiosporous">
|
||
<emphasis id="6C87EA9D2440D307FB2EFAA8FA34FA27" box="[1267,1415,1399,1423]" italics="true" pageId="2" pageNumber="3">S. leiosporous</emphasis>
|
||
Abbott, 1963
|
||
</taxonomicName>
|
||
. The features of these cones together with those of the British cones are summarised in
|
||
<tableCitation id="137103342440D307FB6BFA68FAB6FA66" box="[1206,1285,1463,1486]" captionStart="Table 1" captionStartId="4.[124,180,177,199]" captionTargetPageId="4" captionText="Table 1. Morphological comparison of Sigillariostrobus cones with in situ spores referable to Laevigatisporites glabratus." httpUri="http://table.plazi.org/id/0A8C66072446D301FFA1FF6EFABFFF6F" pageId="2" pageNumber="3" tableUuid="0A8C66072446D301FFA1FF6EFABFFF6F">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5E4C368F2440D306FC93FA08FE16F9A3" blockId="2.[819,1463,176,1900]" lastBlockId="3.[124,767,1492,1898]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
Both
|
||
<taxonomicName id="99F34D0C2440D307FC5AFA09FBBDFA46" authorityName="Bochenski" authorityYear="1936" box="[903,1038,1494,1518]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="czarnockii">
|
||
<emphasis id="6C87EA9D2440D307FC5AFA09FBBDFA46" box="[903,1038,1494,1518]" italics="true" pageId="2" pageNumber="3">S. czarnockii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="99F34D0C2440D307FBE2FA09FB60FA46" authorityName="Abbott" authorityYear="1963" box="[1087,1235,1494,1518]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="leiosporous">
|
||
<emphasis id="6C87EA9D2440D307FBE2FA09FB60FA46" box="[1087,1235,1494,1518]" italics="true" pageId="2" pageNumber="3">S. leiosporous</emphasis>
|
||
</taxonomicName>
|
||
have sporophyll laminae with ciliate edges, which is a feature not observed in the cones from localities in Britain.
|
||
<taxonomicName id="99F34D0C2440D307FB73F9C9FADEF986" authorityName="White" authorityYear="1903" baseAuthorityName="Lesquereux" box="[1198,1389,1558,1582]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="quadrangularis">
|
||
<emphasis id="6C87EA9D2440D307FB73F9C9FADEF986" box="[1198,1389,1558,1582]" italics="true" pageId="2" pageNumber="3">S. quadrangularis</emphasis>
|
||
</taxonomicName>
|
||
has its sporophylls in whorls, which is a most unusual character for
|
||
<taxonomicName id="99F34D0C2440D307FCEEF989FC57F9C6" authorityName="Feistmantel" authorityYear="1876" baseAuthorityName="Schimper" baseAuthorityYear="1870" box="[819,996,1622,1646]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6C87EA9D2440D307FCEEF989FC57F9C6" box="[819,996,1622,1646]" italics="true" pageId="2" pageNumber="3">Sigillariostrobus</emphasis>
|
||
</taxonomicName>
|
||
. This again is different from the helical arrangement in the cones from localities in Britain.
|
||
<taxonomicName id="99F34D0C2440D307FA9CF9A9FA06F926" authorityName="Zeiller" authorityYear="1884" box="[1345,1461,1654,1678]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="tieghemi">
|
||
<emphasis id="6C87EA9D2440D307FA9CF9A9FA06F926" box="[1345,1461,1654,1678]" italics="true" pageId="2" pageNumber="3">S. tieghemi</emphasis>
|
||
</taxonomicName>
|
||
is described as having downward-projecting heels at the distal end of the pedicel, a feature which is again not shown by the cones from localities in Britain. Although all these cones contain megaspores that would be included in the same dispersed spore species,
|
||
<taxonomicName id="99F34D0C2440D307FBFCF8CAFAF4F885" box="[1057,1351,1813,1837]" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="3" phylum="Bryophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2440D307FBFCF8CAFAF4F885" box="[1057,1351,1813,1837]" italics="true" pageId="2" pageNumber="3">Laevigatisporites glabratus</emphasis>
|
||
</taxonomicName>
|
||
, there are characters of the cones which lead us to the conclusion that they are all different in some respects. For that reason we accept them all as distinct species and therefore refer the British specimens to a new species.
|
||
</paragraph>
|
||
<caption id="0A8C66072441D306FFA1FAFCFD27FA39" ID-DOI="http://doi.org/10.5281/zenodo.10965370" ID-Zenodo-Dep="10965370" httpUri="https://zenodo.org/record/10965370/files/figure.png" pageId="3" pageNumber="4" startId="3.[124,191,1315,1337]" targetBox="[136,1454,177,1284]" targetPageId="3" targetType="figure">
|
||
<paragraph id="5E4C368F2441D306FFA1FAFCFD27FA39" blockId="3.[124,1462,1315,1425]" pageId="3" pageNumber="4">
|
||
<emphasis id="6C87EA9D2441D306FFA1FAFCFF6AFA91" bold="true" box="[124,217,1315,1337]" pageId="3" pageNumber="4">Figure 2.</emphasis>
|
||
Type specimen of
|
||
<taxonomicName id="99F34D0C2441D306FE4BFAFBFD04FA91" authority="Thomas & Seyfullah, 2016" authorityName="Thomas & Seyfullah" authorityYear="2016" box="[406,695,1316,1337]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="saltwellensis" status="sp. nov.">
|
||
<emphasis id="6C87EA9D2441D306FE4BFAFBFD04FA91" box="[406,695,1316,1337]" italics="true" pageId="3" pageNumber="4">Sigillariostrobus saltwellensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="77B457E62441D306FD62FAFBFCB8FA91" box="[703,779,1316,1337]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Specimen Pb. 53-C.203, Hunterian Museum, Glasgow. Alluvial coalbearing facies – upper Langsettian–Duckmantian.
|
||
<emphasis id="6C87EA9D2441D306FDBDFA9FFDC8FAFE" bold="true" box="[608,635,1344,1366]" pageId="3" pageNumber="4">(a)</emphasis>
|
||
Cone (scale bar 10 mm).
|
||
<emphasis id="6C87EA9D2441D306FCA9FA9FFC23FAFE" bold="true" box="[884,912,1344,1366]" pageId="3" pageNumber="4">(b)</emphasis>
|
||
Detail of apical portion of the cone (scale bar 5 mm).
|
||
<emphasis id="6C87EA9D2441D306FA46FA9FFA05FAFE" bold="true" box="[1435,1462,1344,1366]" pageId="3" pageNumber="4">(c)</emphasis>
|
||
Detail of basal portion of the cone (scale bar 5 mm).
|
||
<emphasis id="6C87EA9D2441D306FDB6FA81FD3BFADC" bold="true" box="[619,648,1374,1396]" pageId="3" pageNumber="4">(d)</emphasis>
|
||
In situ glabrous megaspores (scale bar 2 mm).
|
||
<emphasis id="6C87EA9D2441D306FB9FFA81FBEEFADC" bold="true" box="[1090,1117,1374,1396]" pageId="3" pageNumber="4">(e)</emphasis>
|
||
Spore referable to
|
||
<taxonomicName id="99F34D0C2441D306FACDFA81FF64FA38" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2441D306FACDFA81FF64FA38" italics="true" pageId="3" pageNumber="4">Laevigatisporites glabratus</emphasis>
|
||
</taxonomicName>
|
||
macerated from the cone (scale bar 1000 µm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5E4C368F2441D306FF4AF9CBFB6AF983" blockId="3.[124,767,1492,1898]" lastBlockId="3.[819,1462,1492,1579]" pageId="3" pageNumber="4">
|
||
The anatomically preserved monosporangiate
|
||
<taxonomicName id="99F34D0C2441D306FD45F9CBFEE4F9E3" authority="Benson, 1918" authorityName="Benson" authorityYear="1918" class="Lycopodiopsida" family="Sigillariaceae" genus="Mazocarpon" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6C87EA9D2441D306FD45F9CBFF17F9E3" italics="true" pageId="3" pageNumber="4">Mazocarpon</emphasis>
|
||
<bibRefCitation id="3A624B7E2441D306FF65F9EBFEE4F9E3" author="Benson, M." box="[184,343,1588,1611]" pageId="3" pageNumber="4" pagination="569 - 589" refId="ref5157" refString="Benson, M.: Mazocarpon or the structure of Sigillariostrobus, Ann. Bot., 32, 569 - 589, 1918." type="journal article" year="1918">Benson, 1918</bibRefCitation>
|
||
</taxonomicName>
|
||
includes two species that contain spores referable to
|
||
<taxonomicName id="99F34D0C2441D306FE86F98CFD35F9C3" box="[347,646,1619,1643]" class="Sphagnopsida" family="Sphagnaceae" genus="Laevigatisporites" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Sphagnales" pageId="3" pageNumber="4" phylum="Bryophyta" rank="species" species="glabratus">
|
||
<emphasis id="6C87EA9D2441D306FE86F98CFD35F9C3" box="[347,646,1619,1643]" italics="true" pageId="3" pageNumber="4">Laevigatisporites glabratus</emphasis>
|
||
</taxonomicName>
|
||
. They are
|
||
<taxonomicName id="99F34D0C2441D306FFA1F9ACFECBF923" authority="Schopf" authorityName="Schopf" box="[124,376,1651,1675]" class="Lycopodiopsida" family="Sigillariaceae" genus="Mazocarpon" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="oedipterum">
|
||
<emphasis id="6C87EA9D2441D306FFA1F9ACFEADF923" box="[124,286,1651,1675]" italics="true" pageId="3" pageNumber="4">M. oedipterum</emphasis>
|
||
Schopf
|
||
</taxonomicName>
|
||
, from the McLeansboro Group, in
|
||
<collectingRegion id="9C37F86D2441D306FFA1F94BFF74F903" box="[124,199,1684,1707]" country="United States of America" name="Illinois" pageId="3" pageNumber="4">Illinois</collectingRegion>
|
||
(
|
||
<bibRefCitation id="3A624B7E2441D306FF0AF94BFED8F903" author="Schopf, J. M." box="[215,363,1684,1707]" pageId="3" pageNumber="4" pagination="1 - 53" refId="ref6001" refString="Schopf, J. M.: Contributions to Pennsylvanian paleobotany Mazocarpon oedipterum, sp. nov, and sigillarian relationships, Ill. State Geo. Surv. Rep. Invest., 75, 1 - 53, 1941." type="journal article" year="1941">Schopf, 1941</bibRefCitation>
|
||
, late Pennsylvanian (Stephanian
|
||
<collectionCode id="38E2AE4A2441D306FD0EF94BFD5BF903" box="[723,744,1684,1707]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="4" type="Herbarium">A</collectionCode>
|
||
)), and
|
||
<taxonomicName id="99F34D0C2441D306FF6CF96CFECBF962" authority="Pigg" authorityName="Pigg" box="[177,376,1715,1739]" class="Lycopodiopsida" family="Sigillariaceae" genus="Mazocarpon" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="bensonii">
|
||
<emphasis id="6C87EA9D2441D306FF6CF96CFE84F963" box="[177,311,1715,1739]" italics="true" pageId="3" pageNumber="4">M. bensonii</emphasis>
|
||
Pigg
|
||
</taxonomicName>
|
||
, from the Duquesne Coal, Conemaugh Group,
|
||
<collectingRegion id="9C37F86D2441D306FEFAF90CFEEFF942" box="[295,348,1747,1770]" country="United States of America" name="Ohio" pageId="3" pageNumber="4">Ohio</collectingRegion>
|
||
(late Pennsylvanian (Stephanian
|
||
<collectionCode id="38E2AE4A2441D306FD09F90CFD54F942" box="[724,743,1747,1770]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="3" pageNumber="4" type="Herbarium">B</collectionCode>
|
||
)).
|
||
<bibRefCitation id="3A624B7E2441D306FFA1F92CFF48F8A2" author="Pigg, K. B." box="[124,251,1779,1802]" pageId="3" pageNumber="4" pagination="600 - 613" refId="ref5813" refString="Pigg, K. B.: The morphology and reproductive biology of the Sigillarian cone Mazocarpon, Bot. Gaz., 144, 600 - 613, 1983." type="journal article" year="1983">Pigg (1983)</bibRefCitation>
|
||
showed the two species to have sufficiently different anatomical features to be separated. Although it is almost certain that
|
||
<taxonomicName id="99F34D0C2441D306FEEDF8ECFE52F8E3" authorityName="Feistmantel" authorityYear="1876" baseAuthorityName="Schimper" baseAuthorityYear="1870" box="[304,481,1843,1867]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6C87EA9D2441D306FEEDF8ECFE52F8E3" box="[304,481,1843,1867]" italics="true" pageId="3" pageNumber="4">Sigillariostrobus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="99F34D0C2441D306FDCFF8ECFD2BF8E3" authorityName="Benson" authorityYear="1918" box="[530,664,1843,1867]" class="Lycopodiopsida" family="Sigillariaceae" genus="Mazocarpon" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="6C87EA9D2441D306FDCFF8ECFD2BF8E3" box="[530,664,1843,1867]" italics="true" pageId="3" pageNumber="4">Mazocarpon</emphasis>
|
||
</taxonomicName>
|
||
represent the same
|
||
<typeStatus id="8148882D2441D306FF3AF88CFEA6F8C2" box="[231,277,1875,1898]" pageId="3" pageNumber="4">type</typeStatus>
|
||
of sigillarian cone but are merely preserved differently, we hesitate to include our
|
||
<taxonomicName id="99F34D0C2441D306FB15FA0BFAD8FA44" authorityName="Thomas & Seyfullah" authorityYear="2016" box="[1224,1387,1492,1516]" class="Lycopodiopsida" family="Sigillariaceae" genus="Sigillariostrobus" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="saltwellensis">
|
||
<emphasis id="6C87EA9D2441D306FB15FA0BFAD8FA44" box="[1224,1387,1492,1516]" italics="true" pageId="3" pageNumber="4">S. saltwellensis</emphasis>
|
||
</taxonomicName>
|
||
within either of these two species because of the lack of anatomical evidence and their rather different ages.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |