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<mods:title id="4A64F421E86FAAF584E5FEDA42A7ADC3">A revision of the Chilean Brachyglutini - Part 6. Revision of Achilia Reitter, 1890: A. grandiceps, A. valdiviensis, and A. bicornis species groups (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="09D9B1FCF6F389F15B8ACEEBC12800BE">Sabella, Giorgio</mods:namePart>
<mods:affiliation id="F97E07C32F26D9815D213087D9CFC92B">Dipartimento di Scienze Biologiche, Geologiche ed Ambientali dell’Università - sezione Biologia Animale, via</mods:affiliation>
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<mods:namePart id="E5B20236162713888BB45AC128BE66C5">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="B7FA86A4895F80636425AF7F0A832101">Muséum d’histoire naturelle, CP 6434, CH- 1211 Genève 6, Switzerland.</mods:affiliation>
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<mods:namePart id="FE1A49C410C123D568FC0430DA3BF35C">Kurbatov, Sergey A.</mods:namePart>
<mods:affiliation id="3730785E6AFF75005F37A055200D0EFB">Museum of Entomology, All-Russian Plant Quarantine Center, Pogranichnaya 32, Bykovo 140150, Russia. E-mail:</mods:affiliation>
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<treatment id="03DE232DB045FFF3FC285181D6DBFEED" ID-DOI="http://doi.org/10.5281/zenodo.6979931" ID-GBIF-Taxon="198967443" ID-Zenodo-Dep="6979931" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED" httpUri="http://treatment.plazi.org/id/03DE232DB045FFF3FC285181D6DBFEED" lastPageId="19" lastPageNumber="146" pageId="12" pageNumber="140">
<subSubSection id="C36DC1B0B045FFECFC285181D224FE69" box="[921,1347,481,505]" pageId="12" pageNumber="140" type="nomenclature">
<paragraph id="8BC8923BB045FFECFC285181D224FE69" blockId="12.[921,1347,481,537]" box="[921,1347,481,505]" pageId="12" pageNumber="140">
<heading id="D0802557B045FFECFC285181D224FE69" bold="true" box="[921,1347,481,505]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="140" reason="2">
<taxonomicName id="4C77E9B8B045FFECFC285181D224FE69" ID-CoL="9792" authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[921,1347,481,505]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B045FFECFC285181D224FE69" bold="true" box="[921,1347,481,505]" pageId="12" pageNumber="140">
<emphasis id="B9034E29B045FFECFC285181D308FE69" bold="true" box="[921,1135,481,505]" italics="true" pageId="12" pageNumber="140">Achilia valdiviensis</emphasis>
(
<bibRefCitation id="EFE6EFCAB045FFECFBCC5182D25CFE69" author="Blanchard C. E." box="[1149,1339,482,505]" pageId="12" pageNumber="140" pagination="1 - 564" refId="ref21916" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos [pp. 561 - 564]. In: Gay C., Historia fisica y politica de Chile. Zoologia, 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</emphasis>
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<subSubSection id="C36DC1B0B045FFECFC2C5262D258FD89" box="[925,1343,514,537]" pageId="12" pageNumber="140" type="description">
<paragraph id="8BC8923BB045FFECFC2C5262D258FD89" blockId="12.[921,1347,481,537]" box="[925,1343,514,537]" pageId="12" pageNumber="140">
<figureCitation id="134C8EBEB045FFECFC2C5262D486FD89" box="[925,993,514,537]" captionStart="Figs 1-6" captionStartId="10.[151,192,2025,2046]" captionTargetBox="[175,1415,715,1976]" captionTargetId="figure-193@10.[151,1436,686,2025]" captionTargetPageId="10" captionText="Figs 1-6. Aedeagi of Achilia species. (1) A. grandiceps. (2) A. delamarei. (3) A. jeanneli n. sp. (4) A. franzi n. sp. (5) A. elguetai n. sp. (6) A. valdiviensis." figureDoi="http://doi.org/10.5281/zenodo.5743393" httpUri="https://zenodo.org/record/5743393/files/figure.png" pageId="12" pageNumber="140">Figs 6</figureCitation>
,
<figureCitation id="134C8EBEB045FFECFC5C5262D34AFD89" box="[1005,1069,514,537]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="12" pageNumber="140">18-19</figureCitation>
,
<figureCitation id="134C8EBEB045FFECFB895262D31AFD89" box="[1080,1149,514,537]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="12" pageNumber="140">28, 35</figureCitation>
,
<figureCitation id="134C8EBEB045FFECFB395262D3AAFD89" box="[1160,1229,514,537]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="12" pageNumber="140">39, 43</figureCitation>
,
<figureCitation id="134C8EBEB045FFECFB695262D27EFD89" box="[1240,1305,514,537]" captionStart="Figs 75-78" captionStartId="22.[151,192,2053,2074]" captionTargetBox="[153,1436,892,2015]" captionTargetId="figure-373@22.[153,1438,867,2015]" captionTargetPageId="22" captionText="Figs 75-78. Achilia valdiviensis. Male head in (75) dorsal, (76) lateral, (77) semilateral, and (78) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743423" httpUri="https://zenodo.org/record/5743423/files/figure.png" pageId="12" pageNumber="140">75-78</figureCitation>
,
<figureCitation id="134C8EBEB045FFECFA955262D258FD89" box="[1316,1343,514,537]" captionStart="Fig" captionStartId="25.[831,866,1909,1930]" captionTargetBox="[868,1366,257,1880]" captionTargetId="figure-92@25.[918,1366,254,1877]" captionText="Fig. 84. Distribution map of Achilia bicornis and valdiviensis group. (l red circles) A. bicornis. (green squares) A. valdiviensis." figureDoi="http://doi.org/10.5281/zenodo.6979915" httpUri="https://zenodo.org/record/6979915/files/figure.png" pageId="12" pageNumber="140">84</figureCitation>
</paragraph>
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<paragraph id="8BC8923BB045FFECFC8E5250D391FDD5" blockId="12.[831,1437,560,840]" box="[831,1270,560,581]" pageId="12" pageNumber="140">
<treatmentCitationGroup id="AB67B515B045FFECFC8E5250D391FDD5" box="[831,1270,560,581]" pageId="12" pageNumber="140">
<treatmentCitation id="0AD6B42AB045FFECFC8E5250D395FDD5" author="Blanchard C. E." box="[831,1266,560,581]" page="563" pageId="12" pageNumber="140" year="1851">
<taxonomicName id="4C77E9B8B045FFECFC8E5250D395FDD5" ID-CoL="4NFP5" authority="Blanchard, 1851: 563" authorityName="Blanchard" authorityPageNumber="563" authorityYear="1851" box="[831,1266,560,581]" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B045FFECFC8E5250D37EFDD5" box="[831,1049,560,581]" italics="true" pageId="12" pageNumber="140">Pselaphus valdiviensis</emphasis>
<bibRefCitation id="EFE6EFCAB045FFECFBAE5250D395FDD5" author="Blanchard C. E." box="[1055,1266,560,581]" pageId="12" pageNumber="140" pagination="1 - 564" refId="ref21916" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos [pp. 561 - 564]. In: Gay C., Historia fisica y politica de Chile. Zoologia, 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851: 563</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BC8923BB045FFECFC8E522CD255FD0B" blockId="12.[831,1437,560,840]" pageId="12" pageNumber="140">
<treatmentCitationGroup id="AB67B515B045FFECFC8E522CD255FD0B" pageId="12" pageNumber="140">
<treatmentCitation id="0AD6B42AB045FFECFC8E522CD389FDF1" author="Raffray A." box="[831,1262,588,609]" page="138" pageId="12" pageNumber="140" year="1904">
<taxonomicName id="4C77E9B8B045FFECFC8E522CD389FDF1" ID-CoL="9792" authority="Raffray, 1904: 138" authorityName="Blch., A. Raffray" authorityPageNumber="138" authorityYear="1904" box="[831,1262,588,609]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B045FFECFC8E522CD36EFDF1" box="[831,1033,588,609]" italics="true" pageId="12" pageNumber="140">Achilia valdiviensis</emphasis>
<bibRefCitation id="EFE6EFCAB045FFECFBAC522CD389FDF1" author="Raffray A." box="[1053,1262,588,609]" pageId="12" pageNumber="140" pagination="1 - 400" refId="ref22353" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray, 1904: 138</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<bibRefCitation id="EFE6EFCAB045FFECFAB4522CD2FDFDF1" author="Raffray A." box="[1285,1434,588,609]" pageId="12" pageNumber="140" refId="ref22377" refString="Raffray A. 1908. Coleoptera. Fam. Pselaphidae. In: Wytsmann P. (ed.), Genera Insectorum, fasc. 64. Rome, 487 pp., 9 pls." type="book" year="1908">Raffray, 1908</bibRefCitation>
: pl. 2, fig. 13 (habitus);
<treatmentCitation id="0AD6B42AB045FFECFBCE5209D258FDEE" author="Jeannel R." box="[1151,1343,617,638]" page="424" pageId="12" pageNumber="140" year="1962">
<bibRefCitation id="EFE6EFCAB045FFECFBCE5209D258FDEE" author="Jeannel R." box="[1151,1343,617,638]" pageId="12" pageNumber="140" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962: 424</bibRefCitation>
</treatmentCitation>
, fig. 189 (aedeagus); (nec
<taxonomicName id="4C77E9B8B045FFECFB9E52E6D240FD0B" ID-CoL="9792" authority="Reitter, 1885" authorityName="Reitter" authorityYear="1885" box="[1071,1319,646,667]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B045FFECFB9E52E6D3C7FD0B" box="[1071,1184,646,667]" italics="true" pageId="12" pageNumber="140">valdiviensis</emphasis>
Reitter, 1885
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).
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<paragraph id="8BC8923BB045FFECFC8E52C3D488FD45" blockId="12.[831,1437,560,840]" pageId="12" pageNumber="140">
<treatmentCitationGroup id="AB67B515B045FFECFC8E52C3D488FD45" pageId="12" pageNumber="140">
<treatmentCitation id="0AD6B42AB045FFECFC8E52C3D3C2FD28" author="Reitter E." box="[831,1189,675,696]" page="327" pageId="12" pageNumber="140" year="1885">
<taxonomicName id="4C77E9B8B045FFECFC8E52C3D3C2FD28" ID-CoL="8XW3H" authority="Reitter, 1885 a: 327" authorityName="Reitter" authorityPageNumber="327" authorityYear="1885" box="[831,1189,675,696]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B045FFECFC8E52C3D4B3FD28" box="[831,980,675,696]" italics="true" pageId="12" pageNumber="140">Bryaxis nasuta</emphasis>
<bibRefCitation id="EFE6EFCAB045FFECFC5152C3D3C2FD28" author="Reitter E." box="[992,1189,675,696]" pageId="12" pageNumber="140" pagination="321 - 332" refId="ref22459" refString="Reitter E. 1885 a. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia, Zweiter Theil. Deutsche Entomologische Zeitschrift 29: 321 - 332, pl. II." type="journal article" year="1885">Reitter, 1885a: 327</bibRefCitation>
</taxonomicName>
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, pl. 2, fig. 7 (head and antennae).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC8923BB045FFECFC8E52BCD4BEFC9E" blockId="12.[831,1437,560,840]" pageId="12" pageNumber="140">
<treatmentCitationGroup id="AB67B515B045FFECFC8E52BCD4BEFC9E" pageId="12" pageNumber="140">
<treatmentCitation id="0AD6B42AB045FFECFC8E52BCD3FCFD61" author="Reitter E." box="[831,1179,732,753]" page="317" pageId="12" pageNumber="140" year="1885">
<taxonomicName id="4C77E9B8B045FFECFC8E52BCD3FCFD61" ID-CoL="5X4S3" authority="Reitter, 1885 b: 317" authorityName="Reitter" authorityPageNumber="317" authorityYear="1885" box="[831,1179,732,753]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="anas">
<emphasis id="B9034E29B045FFECFC8E52BCD4A3FD61" box="[831,964,732,753]" italics="true" pageId="12" pageNumber="140">Bryaxis anas</emphasis>
<bibRefCitation id="EFE6EFCAB045FFECFC6352BCD3FCFD61" author="Reitter E." box="[978,1179,732,753]" pageId="12" pageNumber="140" pagination="313 - 317" refId="ref22489" refString="Reitter E. 1885 b. Coleopterologische Notizen XIV. Wiener Entomologische Zeitung 4 (10): 313 - 317." type="journal article" year="1885">Reitter, 1885b: 317</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(new name for
<taxonomicName id="4C77E9B8B045FFECFAE252BCD4ACFC9E" ID-CoL="8XW3H" authorityName="Reitter" authorityYear="1885" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B045FFECFAE252BCD4ACFC9E" italics="true" pageId="12" pageNumber="140">Bryaxis nasuta</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC8923BB045FFECFC8E5376D4BEFCD8" blockId="12.[831,1437,560,840]" pageId="12" pageNumber="140">
<treatmentCitationGroup id="AB67B515B045FFECFC8E5376D4BEFCD8" pageId="12" pageNumber="140">
<treatmentCitation id="0AD6B42AB045FFECFC8E5376D3F9FCBB" author="Reitter E." box="[831,1182,790,811]" page="261" pageId="12" pageNumber="140" year="1893">
<taxonomicName id="4C77E9B8B045FFECFC8E5376D3F9FCBB" ID-CoL="NGSR" authority="Reitter, 1893: 261" authorityName="Reitter" authorityPageNumber="261" authorityYear="1893" box="[831,1182,790,811]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="nasina">
<emphasis id="B9034E29B045FFECFC8E5376D4B1FCBB" box="[831,982,790,811]" italics="true" pageId="12" pageNumber="140">Bryaxis nasina</emphasis>
<bibRefCitation id="EFE6EFCAB045FFECFC525376D3F9FCBB" author="Reitter E." box="[995,1182,790,811]" pageId="12" pageNumber="140" pagination="260 - 261" refId="ref22511" refString="Reitter E. 1893. Coleopterologische Notizen XLVIII. Wiener Entomologische Zeitung 12 (7): 260 - 261." type="journal article" year="1893">Reitter, 1893: 261</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(new name for
<taxonomicName id="4C77E9B8B045FFECFAE25376D4ACFCD8" ID-CoL="8XW3H" authorityName="Reitter" authorityYear="1885" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B045FFECFAE25376D4ACFCD8" italics="true" pageId="12" pageNumber="140">Bryaxis nasuta</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<caption id="DF08C2B3B045FFECFF265787D6F9F789" ID-DOI="http://doi.org/10.5281/zenodo.5743397" ID-Zenodo-Dep="5743397" httpUri="https://zenodo.org/record/5743397/files/figure.png" pageId="12" pageNumber="140" startId="12.[151,192,2023,2044]" targetBox="[298,1321,898,1988]" targetPageId="12">
<paragraph id="8BC8923BB045FFECFF265787D6F9F789" blockId="12.[151,1434,2023,2073]" pageId="12" pageNumber="140">
Figs 12-16. Male antennae of
<taxonomicName id="4C77E9B8B045FFECFE715787D562F86C" ID-CoL="8XJ8V" authorityName="Reitter" authorityYear="1890" box="[448,517,2023,2044]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B045FFECFE715787D562F86C" box="[448,517,2023,2044]" italics="true" pageId="12" pageNumber="140">Achilia</emphasis>
</taxonomicName>
species. (12)
<emphasis id="B9034E29B045FFECFD3D5787D47EF86C" box="[652,793,2023,2044]" italics="true" pageId="12" pageNumber="140">
<taxonomicName id="4C77E9B8B045FFECFD3D5787D471F86C" ID-CoL="977V" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[652,790,2023,2044]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="grandiceps">A. grandiceps</taxonomicName>
.
</emphasis>
(13)
<emphasis id="B9034E29B045FFECFCFF5787D4B6F86C" box="[846,977,2023,2044]" italics="true" pageId="12" pageNumber="140">
<taxonomicName id="4C77E9B8B045FFECFCFF5787D4AAF86C" ID-CoL="977J" box="[846,973,2023,2044]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="delamarei">A. delamarei</taxonomicName>
.
</emphasis>
(14)
<taxonomicName id="4C77E9B8B045FFECFBB45787D33FF86C" ID-CoL="8XSTV" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[1029,1112,2023,2044]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="franzi" status="sp. nov.">
<emphasis id="B9034E29B045FFECFBB45787D33FF86C" box="[1029,1112,2023,2044]" italics="true" pageId="12" pageNumber="140">A. franzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B045FFECFBEE5787D3F3F86C" box="[1119,1172,2023,2044]" pageId="12" pageNumber="140" rank="species">n. sp.</taxonomicNameLabel>
(15)
<taxonomicName id="4C77E9B8B045FFECFB795787D256F86C" ID-CoL="8XSTT" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[1224,1329,2023,2044]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="elguetai" status="sp. nov.">
<emphasis id="B9034E29B045FFECFB795787D256F86C" box="[1224,1329,2023,2044]" italics="true" pageId="12" pageNumber="140">A. elguetai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B045FFECFA895787D20BF86C" box="[1336,1388,2023,2044]" pageId="12" pageNumber="140" rank="species">n. sp.</taxonomicNameLabel>
(16)
<taxonomicName id="4C77E9B8B045FFECFEA05864D6FFF789" box="[273,408,2052,2073]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="140" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B045FFECFEA05864D6FFF789" box="[273,408,2052,2073]" italics="true" pageId="12" pageNumber="140">A. denticornis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC8923BB044FFEDFF265162D5A7FEA4" blockId="13.[151,756,258,308]" pageId="13" pageNumber="141">
<treatmentCitationGroup id="AB67B515B044FFEDFF265162D5A7FEA4" pageId="13" pageNumber="141">
<treatmentCitation id="0AD6B42AB044FFEDFF265162D69AFE87" author="Jeannel R." box="[151,509,258,279]" page="424" pageId="13" pageNumber="141" year="1962">
<taxonomicName id="4C77E9B8B044FFEDFF265162D69AFE87" ID-CoL="977Z" authority="Jeannel, 1962: 424" authorityName="Jeannel" authorityPageNumber="424" authorityYear="1962" box="[151,509,258,279]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B044FFEDFF265162D652FE87" box="[151,309,258,279]" italics="true" pageId="13" pageNumber="141">Achilia kuscheli</emphasis>
<bibRefCitation id="EFE6EFCAB044FFEDFE8E5162D69AFE87" author="Jeannel R." box="[319,509,258,279]" pageId="13" pageNumber="141" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962: 424</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 425, figs 186 (habitus), 187 (head of female), 188 (aedeagus) (
<emphasis id="B9034E29B044FFEDFDE6517FD5CCFEA4" bold="true" box="[599,683,287,308]" pageId="13" pageNumber="141">syn. nov</emphasis>
.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B044FFEFFF26512CD5DCFEEC" lastPageId="15" lastPageNumber="143" pageId="13" pageNumber="141" type="materials_examined">
<paragraph id="8BC8923BB044FFEDFF26512CD340FE69" blockId="13.[151,756,332,771]" lastBlockId="13.[831,1437,258,505]" pageId="13" pageNumber="141">
<materialsCitation id="3B1F9866B044FFEDFF26512CD6E3FDF3" ID-GBIF-Occurrence="3886337414" collectionCode="MNHN" country="Chile" determinerName="Cuccodoro &amp; Kurbatov" location="Region Los Rios" pageId="13" pageNumber="141" specimenCount="15" specimenCount-male="1" stateProvince="Region Los Rios" typeStatus="lectotype">
<emphasis id="B9034E29B044FFEDFF26512CD6F8FEF3" bold="true" box="[151,415,332,355]" pageId="13" pageNumber="141">
Type material (
<specimenCount id="9D7159B2B044FFEDFEFE512CD6F0FEF3" box="[335,407,332,355]" pageId="13" pageNumber="141" type="generic">14 ex.</specimenCount>
)
</emphasis>
: SOUTHERN
<collectingCountry id="F360D2ABB044FFEDFDFD512CD5FCFEF3" box="[588,667,332,355]" name="Chile" pageId="13" pageNumber="141" value="Chile">CHILE</collectingCountry>
<collectingRegion id="49B35CD9B044FFEDFD17512CD662FE13" country="Chile" name="Los Rios" pageId="13" pageNumber="141">Región Los Rios</collectingRegion>
: Valdivia prov.:
<collectionCode id="ED660AFEB044FFEDFE7F510CD543FE13" box="[462,548,364,387]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="141">MNHN</collectionCode>
;
<specimenCount id="9D7159B2B044FFEDFD89510CD50FFE13" box="[568,616,364,387]" pageId="13" pageNumber="141" type="male">1 ♂</specimenCount>
(
<typeStatus id="54CC2C99B044FFEDFDCE510CD588FE13" box="[639,751,364,387]" pageId="13" pageNumber="141" type="lectotype">Lectotype</typeStatus>
, here designated); label verbatim: “Lectotype / Museum Paris, Chili, Gay 1849 / H 49 / Valdivia / Gen.
<taxonomicName id="4C77E9B8B044FFEDFD1651CCD7B3FE73" authority="Reitt." authorityName="Reitter" authorityYear="1890" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B044FFEDFD1651CCD593FE53" box="[679,756,428,451]" italics="true" pageId="13" pageNumber="141">Achilia</emphasis>
Reitt.
</taxonomicName>
=,
<taxonomicName id="4C77E9B8B044FFEDFEBC51ACD6D4FE73" authority="Raffr." authorityName="Raffr." box="[269,435,460,483]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B044FFEDFEBC51ACD607FE73" box="[269,352,460,483]" italics="true" pageId="13" pageNumber="141">Bryaxis</emphasis>
Raffr.
</taxonomicName>
, =
<taxonomicName id="4C77E9B8B044FFEDFE5B51ACD5BDFE73" authority="Blanchard" authorityName="Blanchard" box="[490,730,460,483]" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B044FFEDFE5B51ACD53EFE73" box="[490,601,460,483]" italics="true" pageId="13" pageNumber="141">Pselaphus</emphasis>
Blanchard
</taxonomicName>
/
<taxonomicName id="4C77E9B8B044FFEDFF26518CD61DFD93" authority=", Blanch." authorityName="Blanch." box="[151,378,492,515]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B044FFEDFF26518CD671FD93" box="[151,278,492,515]" italics="true" pageId="13" pageNumber="141">valdiviensis</emphasis>
, Blanch.
</taxonomicName>
=
<taxonomicName id="4C77E9B8B044FFEDFE2C518CD55DFD93" authority="Reitter" authorityName="Reitter" authorityYear="1893" box="[413,570,492,515]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="nasina">
<emphasis id="B9034E29B044FFEDFE2C518CD683FD93" box="[413,484,492,515]" italics="true" pageId="13" pageNumber="141">nasina</emphasis>
Reitter
</taxonomicName>
, =
<taxonomicName id="4C77E9B8B044FFEDFDD0518CD66BFDB3" authority="Reitt., A. Raffray" authorityName="Reitt., A. Raffray" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B044FFEDFDD0518CD5CFFD93" box="[609,680,492,515]" italics="true" pageId="13" pageNumber="141">nasuta</emphasis>
Reitt., A. Raffray
</taxonomicName>
det. 1904 /
<taxonomicName id="4C77E9B8B044FFEDFE27526CD559FDB3" authority="Bl." authorityName="Bl." box="[406,574,524,547]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B044FFEDFE27526CD573FDB3" box="[406,532,524,547]" italics="true" pageId="13" pageNumber="141">valdiviensis</emphasis>
Bl.
</taxonomicName>
(handwritten by Jeannel) /
<taxonomicName id="4C77E9B8B044FFEDFEA3524CD79AFDF3" authority="Sabella, Cuccodoro &amp; Kurbatov" authorityName="Sabella, Cuccodoro &amp; Kurbatov" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B044FFEDFEA3524CD68BFDD3" box="[274,492,556,579]" italics="true" pageId="13" pageNumber="141">Achilia valdiviensis</emphasis>
Sabella,
<determinerName id="7E147EFFB044FFEDFDED524CD79AFDF3" pageId="13" pageNumber="141">Cuccodoro &amp; Kurbatov</determinerName>
</taxonomicName>
det. 2019”
</materialsCitation>
.
<materialsCitation id="3B1F9866B044FFEDFE24522CD668FC93" ID-GBIF-Occurrence="3886337560" collectionCode="MNHN" country="Chile" determinerName="Cuccodoro &amp; Kurbatov" location="Sabella" pageId="13" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="Valdivia" typeStatus="paralectotype">
–
<collectionCode id="ED660AFEB044FFEDFE1E522CD563FDF3" box="[431,516,588,611]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="141">MNHN</collectionCode>
;
<specimenCount id="9D7159B2B044FFEDFDA7522CD523FDF3" box="[534,580,588,611]" pageId="13" pageNumber="141" type="male">1 ♂</specimenCount>
(
<typeStatus id="54CC2C99B044FFEDFDE8522CD597FDF3" box="[601,752,588,611]" pageId="13" pageNumber="141" type="paralectotype">Paralectotype</typeStatus>
, here designated); label verbatim: “Paralectotype / Museum Paris, Chili, Gay 1849 / H 49/ Valdivia /
<taxonomicName id="4C77E9B8B044FFEDFF2652CCD53EFD53" authority="Blch., A. Raffray" authorityName="Blch., A. Raffray" authorityYear="1904" box="[151,601,684,707]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B044FFEDFF2652CCD610FD53" box="[151,375,684,707]" italics="true" pageId="13" pageNumber="141">Achilia valdiviensis</emphasis>
Blch., A. Raffray
</taxonomicName>
det. 1904 /
<taxonomicName id="4C77E9B8B044FFEDFF2652ACD593FD73" authority="Sabella, Cuccodoro &amp; Kurbatov" authorityName="Sabella, Cuccodoro &amp; Kurbatov" box="[151,756,716,739]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B044FFEDFF2652ACD613FD73" box="[151,372,716,739]" italics="true" pageId="13" pageNumber="141">Achilia valdiviensis</emphasis>
<location id="8EA8C4E0B044FFEDFE3352ACD6B3FD73" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B044FFEDFE3352ACD6B3FD73" box="[386,468,716,739]" country="Chile" name="Sabella" pageId="13" pageNumber="141" stateProvince="Valdivia">Sabella</location>
,
<determinerName id="7E147EFFB044FFEDFE5752ACD593FD73" box="[486,756,716,739]" pageId="13" pageNumber="141">Cuccodoro &amp; Kurbatov</determinerName>
</taxonomicName>
det. 2019”
</materialsCitation>
.
<materialsCitation id="3B1F9866B044FFEDFEAE528CD496FEE9" ID-GBIF-Occurrence="3886337359" collectingDate="1958-10-03" collectionCode="MHNS" collectorName="G. Kuschel" country="Chile" location="Region Los Lagos" pageId="13" pageNumber="141" specimenCount="1" specimenCount-male="1" stateProvince="Region Los Lagos" typeStatus="holotype">
<collectingRegion id="49B35CD9B044FFEDFEAE528CD694FC93" box="[287,499,748,771]" country="Chile" name="Los Lagos" pageId="13" pageNumber="141">Región Los Lagos</collectingRegion>
:
<collectingRegion id="49B35CD9B044FFEDFDB3528CD5EBFC93" box="[514,652,748,771]" pageId="13" pageNumber="141">Chiloé prov</collectingRegion>
.:
<collectionCode id="ED660AFEB044FFEDFD2E528CD597FC93" box="[671,752,748,771]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="13" pageNumber="141">MHNS</collectionCode>
;
<specimenCount id="9D7159B2B044FFEDFC8E5162D40AFE89" box="[831,877,258,281]" pageId="13" pageNumber="141" type="male">1 ♂</specimenCount>
(
<typeStatus id="54CC2C99B044FFEDFC315162D487FE89" box="[896,992,258,281]" pageId="13" pageNumber="141" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C77E9B8B044FFEDFBBF5162D3EEFE89" authorityName="Jeannel" authorityYear="1962" box="[1038,1161,258,281]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B044FFEDFBBF5162D3EEFE89" box="[1038,1161,258,281]" italics="true" pageId="13" pageNumber="141">A. kuscheli</emphasis>
</taxonomicName>
); labels verbatim “Type / Chepu,
<date id="FFC9B4FBB044FFEDFC1D5142D347FEA9" box="[940,1056,290,313]" pageId="13" pageNumber="141" value="1958-10-03">
<collectingDate id="EF8D4D13B044FFEDFC1D5142D347FEA9" box="[940,1056,290,313]" pageId="13" pageNumber="141" value="1958-10-03">03.X.1958</collectingDate>
</date>
,
<collectorName id="2682F7EDB044FFEDFB805142D3C8FEA9" box="[1073,1199,290,313]" pageId="13" pageNumber="141">G. Kuschel</collectorName>
/
<taxonomicName id="4C77E9B8B044FFEDFB7E5142D2EFFEA9" authorityName="Jeannel" authorityYear="1962" box="[1231,1416,290,313]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B044FFEDFB7E5142D2EFFEA9" box="[1231,1416,290,313]" italics="true" pageId="13" pageNumber="141">Achillia kuscheli</emphasis>
</taxonomicName>
/
<taxonomicName id="4C77E9B8B044FFEDFC8E5122D4F1FEC9" authorityName="Jeannel" authorityYear="1962" box="[831,918,322,345]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B044FFEDFC8E5122D4F1FEC9" box="[831,918,322,345]" italics="true" pageId="13" pageNumber="141">kuscheli</emphasis>
</taxonomicName>
(handwritten by Jeannel) / CHILE; M.N.H.N.; Typo, n. 1850”
</materialsCitation>
.
<materialsCitation id="3B1F9866B044FFEDFBB05102D356FE29" ID-GBIF-Occurrence="3886337545" collectingDate="1958-10-03" collectionCode="MNHN" collectorName="Kuschel" country="Chile" pageId="13" pageNumber="141" specimenCount="10" specimenCount-female="9" specimenCount-male="1" stateProvince="Chepu" typeStatus="paratype">
–
<collectionCode id="ED660AFEB044FFEDFBAA5102D317FEE9" box="[1051,1136,354,377]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="141">MNHN</collectionCode>
,
<specimenCount id="9D7159B2B044FFEDFB305102D3D7FEE9" box="[1153,1200,354,377]" pageId="13" pageNumber="141" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D7159B2B044FFEDFB405102D278FEE9" box="[1265,1311,354,377]" pageId="13" pageNumber="141" type="female">9 ♀</specimenCount>
(
<typeStatus id="54CC2C99B044FFEDFA825102D2FBFEE9" box="[1331,1436,354,377]" pageId="13" pageNumber="141" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C77E9B8B044FFEDFCD451E2D484FE09" authorityName="Jeannel" authorityYear="1962" box="[869,995,386,409]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B044FFEDFCD451E2D484FE09" box="[869,995,386,409]" italics="true" pageId="13" pageNumber="141">A. kuscheli</emphasis>
</taxonomicName>
); labels verbatim “Paratype / Chepu,
<date id="FFC9B4FBB044FFEDFC8E51C2D4D3FE29" box="[831,948,418,441]" pageId="13" pageNumber="141" value="1958-10-03">
<collectingDate id="EF8D4D13B044FFEDFC8E51C2D4D3FE29" box="[831,948,418,441]" pageId="13" pageNumber="141" value="1958-10-03">03.X.1958</collectingDate>
</date>
,
<collectorName id="2682F7EDB044FFEDFC7851C2D340FE29" box="[969,1063,418,441]" pageId="13" pageNumber="141">Kuschel</collectorName>
”
</materialsCitation>
.
<materialsCitation id="3B1F9866B044FFEDFBF451C2D344FE69" ID-GBIF-Occurrence="3886337538" collectingDate="1958-10-11" collectionCode="MNHN" collectorName="Kuschel" country="Chile" pageId="13" pageNumber="141" specimenCount="1" specimenCount-female="1" stateProvince="Chepu" typeStatus="paratype">
–
<collectionCode id="ED660AFEB044FFEDFBD251C2D3DEFE29" box="[1123,1209,418,441]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="141">MNHN</collectionCode>
;
<specimenCount id="9D7159B2B044FFEDFB7C51C2D398FE29" box="[1229,1279,418,441]" pageId="13" pageNumber="141" type="female">1 ♀</specimenCount>
(
<typeStatus id="54CC2C99B044FFEDFAA751C2D212FE29" box="[1302,1397,418,441]" pageId="13" pageNumber="141" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C77E9B8B044FFEDFC8E51A2D4A3FE49" authorityName="Jeannel" authorityYear="1962" box="[831,964,450,473]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B044FFEDFC8E51A2D4A3FE49" box="[831,964,450,473]" italics="true" pageId="13" pageNumber="141">A. kuscheli</emphasis>
</taxonomicName>
); labels verbatim “Paratype / Chepu,
<date id="FFC9B4FBB044FFEDFC8E5182D4D6FE69" box="[831,945,482,505]" pageId="13" pageNumber="141" value="1958-10-11">
<collectingDate id="EF8D4D13B044FFEDFC8E5182D4D6FE69" box="[831,945,482,505]" pageId="13" pageNumber="141" value="1958-10-11">11.X.1958</collectingDate>
</date>
,
<collectorName id="2682F7EDB044FFEDFC0D5182D37EFE69" box="[956,1049,482,505]" pageId="13" pageNumber="141">Kuschel</collectorName>
”
</materialsCitation>
.
</paragraph>
<paragraph id="8BC8923BB044FFEFFC8E5271D5DCFEEC" blockId="13.[831,1437,529,776]" lastBlockId="15.[151,756,258,380]" lastPageId="15" lastPageNumber="143" pageId="13" pageNumber="141">
<materialsCitation id="3B1F9866B044FFEDFC8E5271D38DFD78" ID-GBIF-Occurrence="3886337475" collectionCode="MNHN, MHNG" collectorName="H. Franz" country="Chile" location="Region Los Lagos" pageId="13" pageNumber="141" specimenCount="67" specimenCount-male="2" stateProvince="Chiloe prov" typeStatus="holotype">
<emphasis id="B9034E29B044FFEDFC8E5271D39BFDB8" bold="true" box="[831,1276,529,552]" pageId="13" pageNumber="141">
Additional material examined (
<specimenCount id="9D7159B2B044FFEDFB1C5271D393FDB8" box="[1197,1268,529,552]" pageId="13" pageNumber="141" type="generic">65 ex.</specimenCount>
)
</emphasis>
:
<collectionCode id="ED660AFEB044FFEDFABF5271D204FDB8" box="[1294,1379,529,552]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="141">MNHN</collectionCode>
;
<specimenCount id="9D7159B2B044FFEDFAC05271D2FBFDB8" box="[1393,1436,529,552]" pageId="13" pageNumber="141" type="male">1 ♂</specimenCount>
(probably
<typeStatus id="54CC2C99B044FFEDFC025251D376FDD8" box="[947,1041,561,584]" pageId="13" pageNumber="141" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C77E9B8B044FFEDFB8F5251D221FDD8" authority="Reitter, 1893" authorityName="Reitter" authorityYear="1893" box="[1086,1350,561,584]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="nasina">
<emphasis id="B9034E29B044FFEDFB8F5251D3CFFDD8" box="[1086,1192,561,584]" italics="true" pageId="13" pageNumber="141">A. nasina</emphasis>
<bibRefCitation id="EFE6EFCAB044FFEDFB035251D221FDD8" author="Reitter E." box="[1202,1350,561,584]" pageId="13" pageNumber="141" pagination="260 - 261" refId="ref22511" refString="Reitter E. 1893. Coleopterologische Notizen XLVIII. Wiener Entomologische Zeitung 12 (7): 260 - 261." type="journal article" year="1893">Reitter, 1893</bibRefCitation>
</taxonomicName>
); labels verbatim “Chili / Museum Paris, 1917, coll. Raffray / Type /
<emphasis id="B9034E29B044FFEDFC155211D365FD18" box="[932,1026,625,648]" italics="true" pageId="13" pageNumber="141">chilensis</emphasis>
Reitter /
<taxonomicName id="4C77E9B8B044FFEDFBC55211D2FBFD18" authority=", A. Raffray" authorityName="Blch., A. Raffray" authorityYear="1904" box="[1140,1436,625,648]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B044FFEDFBC55211D272FD18" box="[1140,1301,625,648]" italics="true" pageId="13" pageNumber="141">A. valdiviensis</emphasis>
, A. Raffray
</taxonomicName>
det. /
<taxonomicName id="4C77E9B8B044FFEDFC2652F1D32AFD38" authority="Bl." authorityName="Bl." box="[919,1101,657,680]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B044FFEDFC2652F1D37FFD38" box="[919,1048,657,680]" italics="true" pageId="13" pageNumber="141">valdiviensis</emphasis>
Bl.
</taxonomicName>
(handwritten by Jeannel)”. SOUTHERN
<collectingCountry id="F360D2ABB044FFEDFC5452D1D35EFD58" box="[997,1081,689,712]" name="Chile" pageId="13" pageNumber="141">CHILE</collectingCountry>
:
<location id="8EA8C4E0B044FFEDFBE352D1D25CFD58" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B044FFEDFBE352D1D25CFD58" box="[1106,1339,689,712]" country="Chile" name="Region Los Lagos" pageId="13" pageNumber="141" stateProvince="Chiloe prov">Región Los Lagos</location>
:
<collectingRegion id="49B35CD9B044FFEDFAE552D1D411FD78" pageId="13" pageNumber="141">Chiloé prov</collectingRegion>
.:
<collectionCode id="ED660AFEB044FFEDFC3952B1D4BAFD78" box="[904,989,721,744]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="13" pageNumber="141" type="Museum">MHNG</collectionCode>
;
<specimenCount id="9D7159B2B044FFEDFC5D52B1D37FFD78" box="[1004,1048,721,744]" pageId="13" pageNumber="141" type="male">1 ♂</specimenCount>
; Chiloe;
<collectorName id="2682F7EDB044FFEDFB3452B1D38DFD78" box="[1157,1258,721,744]" pageId="13" pageNumber="141">H. Franz</collectorName>
</materialsCitation>
.
<materialsCitation id="3B1F9866B044FFEEFB4952B1D5E5FE89" ID-GBIF-Occurrence="3886337472" collectingDate="1958-10-03" collectionCode="MHNS" collectorName="G. Kuschel" country="Chile" lastPageId="14" lastPageNumber="142" location="Chepu" pageId="13" pageNumber="141" specimenCount="7" specimenCount-female="7" stateProvince="Chepu" typeStatus="paratype">
–
<collectionCode id="ED660AFEB044FFEDFAA152B1D206FD78" box="[1296,1377,721,744]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="13" pageNumber="141">MHNS</collectionCode>
;
<specimenCount id="9D7159B2B044FFEDFAC152B1D2FBFD78" box="[1392,1436,721,744]" pageId="13" pageNumber="141" type="female">7 ♀</specimenCount>
(mislabelled as paratypes of
<taxonomicName id="4C77E9B8B044FFEDFB3B5291D261FC98" authorityName="Jeannel" authorityYear="1962" box="[1162,1286,753,776]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B044FFEDFB3B5291D261FC98" box="[1162,1286,753,776]" italics="true" pageId="13" pageNumber="141">A. kuscheli</emphasis>
</taxonomicName>
n. 1851 and n. 2072-77);
<location id="8EA8C4E0B047FFEEFE985162D615FE89" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B047FFEEFE985162D615FE89" box="[297,370,258,281]" country="Chile" name="Chepu" pageId="14" pageNumber="142" stateProvince="Chepu">Chepu</location>
;
<date id="FFC9B4FBB047FFEEFE315162D693FE89" box="[384,500,258,281]" pageId="14" pageNumber="142" value="1958-10-03">
<collectingDate id="EF8D4D13B047FFEEFE315162D693FE89" box="[384,500,258,281]" pageId="14" pageNumber="142" value="1958-10-03">03.X.1958</collectingDate>
</date>
;
<collectorName id="2682F7EDB047FFEEFDB25162D5E5FE89" box="[515,642,258,281]" pageId="14" pageNumber="142">G. Kuschel</collectorName>
</materialsCitation>
.
<materialsCitation id="3B1F9866B047FFEEFD3E5162D38FFE89" ID-GBIF-Occurrence="3886337476" collectedFrom="leaf &amp; log litter" collectingDate="1997-01-14" collectingMethod="berlese" collectionCode="FMNH" collectorName="A. Newton &amp; M. Thayer" country="Chile" elevation="10" latitude="-42.666" location="Puente La Caldera" longLatPrecision="11" longitude="-74.011665" pageId="14" pageNumber="142" specimenCode="FMHD# 97-21" specimenCount="32" specimenCount-female="17" specimenCount-male="15" stateProvince="Cucao">
–
<collectionCode id="ED660AFEB047FFEEFD165162D593FE89" box="[679,756,258,281]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="14" pageNumber="142" type="Museum">FMNH</collectionCode>
(
<specimenCode id="DBD13A40B047FFEEFF2F5142D620FEA9" box="[158,327,290,313]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="14" pageNumber="142" type="Museum">FMHD# 97-21</specimenCode>
);
<specimenCount id="9D7159B2B047FFEEFEEE5142D6FFFEA9" box="[351,408,290,313]" pageId="14" pageNumber="142" type="male">15 ♂</specimenCount>
and
<specimenCount id="9D7159B2B047FFEEFE655142D56AFEA9" box="[468,525,290,313]" pageId="14" pageNumber="142" type="female">17 ♀</specimenCount>
;
<location id="8EA8C4E0B047FFEEFDAE5142D597FEA9" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B047FFEEFDAE5142D597FEA9" box="[543,752,290,313]" country="Chile" latitude="-42.666" longLatPrecision="11" longitude="-74.011665" name="Puente La Caldera" pageId="14" pageNumber="142" stateProvince="Cucao">Puente La Caldera</location>
,
<locationDeviation id="9A7A8B19B047FFEEFF265122D610FEC9" box="[151,375,322,345]" pageId="14" pageNumber="142">
<quantity id="4C8F3FDEB047FFEEFF265122D781FEC9" box="[151,230,322,345]" metricMagnitude="3" metricUnit="m" metricValue="9.8" pageId="14" pageNumber="142" unit="km" value="9.8">9.8 km</quantity>
E of Cucao
</locationDeviation>
;
<geoCoordinate id="EE43F4FCB047FFEEFE375122D56BFEC9" box="[390,524,322,345]" degrees="42" direction="south" minutes="39.96" orientation="latitude" pageId="14" pageNumber="142" precision="9" value="-42.666">42° 39.96’S</geoCoordinate>
<geoCoordinate id="EE43F4FCB047FFEEFDA65122D5C1FEC9" box="[535,678,322,345]" degrees="74" direction="west" minutes="00.70" orientation="longitude" pageId="14" pageNumber="142" precision="9" value="-74.011665">74° 00.70’W</geoCoordinate>
<quantity id="4C8F3FDEB047FFEEFD005122D597FEC9" box="[689,752,322,345]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="142" unit="m" value="10.0">
<elevation id="005A7508B047FFEEFD005122D597FEC9" box="[689,752,322,345]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="142" unit="m" value="10.0">10 m</elevation>
</quantity>
;
<date id="FFC9B4FBB047FFEEFF265102D665FEE9" box="[151,258,354,377]" pageId="14" pageNumber="142" value="1997-01-14">
<collectingDate id="EF8D4D13B047FFEEFF265102D665FEE9" box="[151,258,354,377]" pageId="14" pageNumber="142" value="1997-01-14">14.I.1997</collectingDate>
</date>
;
<location id="8EA8C4E0B047FFEEFEA25102D617FEE9" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B047FFEEFEA25102D617FEE9" box="[275,368,354,377]" country="Chile" latitude="-42.666" longLatPrecision="11" longitude="-74.011665" name="site 991" pageId="14" pageNumber="142" stateProvince="Cucao">site 991</location>
,
<location id="8EA8C4E0B047FFEEFE315102D53CFEE9" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B047FFEEFE315102D53CFEE9" box="[384,603,354,377]" country="Chile" latitude="-42.666" longLatPrecision="11" longitude="-74.011665" name="valdivian rainforest" pageId="14" pageNumber="142" stateProvince="Cucao">valdivian rainforest</location>
,
<collectingMethod id="5236EA2CB047FFEEFDDD5102D5DCFEE9" box="[620,699,354,377]" pageId="14" pageNumber="142">berlese</collectingMethod>
,
<collectedFrom id="386BD1C6B047FFEEFD7D5102D4A6FE89" box="[716,961,258,377]" pageId="14" pageNumber="142">leaf &amp; log litter</collectedFrom>
;
<collectorName id="2682F7EDB047FFEEFC7F5162D38FFE89" box="[974,1256,258,281]" pageId="14" pageNumber="142">A. Newton &amp; M. Thayer</collectorName>
</materialsCitation>
.
<materialsCitation id="3B1F9866B047FFEEFB445162D34EFEA9" ID-GBIF-Occurrence="3886337519" collectionCode="MHNG" country="Chile" pageId="14" pageNumber="142" specimenCount="13" specimenCount-female="6" specimenCount-male="7" stateProvince="Cucao">
–
<collectionCode id="ED660AFEB047FFEEFABC5162D205FE89" box="[1293,1378,258,281]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="142" type="Museum">MHNG</collectionCode>
;
<specimenCount id="9D7159B2B047FFEEFAC05162D2FBFE89" box="[1393,1436,258,281]" pageId="14" pageNumber="142" type="male">7 ♂</specimenCount>
and
<specimenCount id="9D7159B2B047FFEEFCC25142D4C6FEA9" box="[883,929,290,313]" pageId="14" pageNumber="142" type="female">6 ♀</specimenCount>
; same data
</materialsCitation>
.
<materialsCitation id="3B1F9866B047FFEEFB885142D408FEC9" ID-GBIF-Occurrence="3886337346" collectionCode="MHNS" country="Chile" pageId="14" pageNumber="142" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Cucao">
–
<collectionCode id="ED660AFEB047FFEEFBE25142D3C3FEA9" box="[1107,1188,290,313]" collectionName="Chile, Santiago, Museo Nacional de Historia Natural" pageId="14" pageNumber="142">MHNS</collectionCode>
;
<specimenCount id="9D7159B2B047FFEEFB045142D384FEA9" box="[1205,1251,290,313]" pageId="14" pageNumber="142" type="male">2 ♂</specimenCount>
and
<specimenCount id="9D7159B2B047FFEEFA925142D236FEA9" box="[1315,1361,290,313]" pageId="14" pageNumber="142" type="female">2 ♀</specimenCount>
; same data
</materialsCitation>
.
<materialsCitation id="3B1F9866B047FFEEFCCD5122D39AFEC9" ID-GBIF-Occurrence="3886337526" box="[892,1277,322,345]" country="Chile" pageId="14" pageNumber="142" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cucao">
– PCPH;
<specimenCount id="9D7159B2B047FFEEFC565122D375FEC9" box="[999,1042,322,345]" pageId="14" pageNumber="142" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D7159B2B047FFEEFBFC5122D31FFEC9" box="[1101,1144,322,345]" pageId="14" pageNumber="142" type="female">1 ♀</specimenCount>
; same data.
</materialsCitation>
<materialsCitation id="3B1F9866B047FFEFFAB75122D697FE89" ID-GBIF-Occurrence="3886337489" collectingDate="2013-12-06" collectingMethod="car net" collectionCode="MHNG" country="Chile" lastPageId="15" lastPageNumber="143" latitude="-40.5833" location="Corral" longitude="-73.333336" pageId="14" pageNumber="142" specimenCount="1" specimenCount-male="1" stateProvince="Region Los Rios">
–
<collectingRegion id="49B35CD9B047FFEEFAAC5122D413FEE9" country="Chile" name="Los Rios" pageId="14" pageNumber="142">Región Los Ríos</collectingRegion>
:
<collectingRegion id="49B35CD9B047FFEEFC345102D345FEE9" box="[901,1058,354,377]" pageId="14" pageNumber="142">Valdivia prov</collectingRegion>
.:
<collectionCode id="ED660AFEB047FFEEFB875102D3ECFEE9" box="[1078,1163,354,377]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="14" pageNumber="142" type="Museum">MHNG</collectionCode>
;
<specimenCount id="9D7159B2B047FFEEFB2D5102D3ADFEE9" box="[1180,1226,354,377]" pageId="14" pageNumber="142" type="male">1 ♂</specimenCount>
;
<location id="8EA8C4E0B047FFEEFB6E5102D240FEE9" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B047FFEEFB6E5102D240FEE9" box="[1247,1319,354,377]" country="Chile" longitude="-73.333336" name="Corral" pageId="14" pageNumber="142" stateProvince="Region Los Rios">Corral</location>
, 39° 95’S 73° 20’W;
<date id="FFC9B4FBB046FFEFFEA05162D6F0FE89" box="[273,407,258,281]" pageId="15" pageNumber="143" value="2013-12-06">
<collectingDate id="EF8D4D13B046FFEFFEA05162D6F0FE89" box="[273,407,258,281]" pageId="15" pageNumber="143" value="2013-12-06">06.XII.2013</collectingDate>
</date>
;
<collectingMethod id="5236EA2CB046FFEFFE125162D697FE89" box="[419,496,258,281]" pageId="15" pageNumber="143">car net</collectingMethod>
</materialsCitation>
.
<materialsCitation id="3B1F9866B046FFEFFE4D5162D5BFFECB" ID-GBIF-Occurrence="3886337493" collectedFrom="sifting litter" collectingDate="2008-02-27" collectionCode="PCPH" collectorName="W. D. Shepard" country="Chile" latitude="-39.976665" location="Reserva Costera Valdiviana" longLatPrecision="715" longitude="-73.583336" pageId="15" pageNumber="143" specimenCount="2" specimenCount-male="2" stateProvince="Region Los Rios">
–
<collectionCode id="ED660AFEB046FFEFFDA35162D53FFE89" box="[530,600,258,281]" pageId="15" pageNumber="143">PCPH</collectionCode>
;
<specimenCount id="9D7159B2B046FFEFFDD55162D5E9FE89" box="[612,654,258,281]" pageId="15" pageNumber="143" type="male">2 ♂</specimenCount>
;
<location id="8EA8C4E0B046FFEFFD2F5162D60CFEAA" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B046FFEFFD2F5162D60CFEAA" country="Chile" latitude="-39.976665" longLatPrecision="715" longitude="-73.583336" name="Reserva Costera Valdiviana" pageId="15" pageNumber="143" stateProvince="Region Los Rios">Reserva Costera Valdiviana</location>
,
<location id="8EA8C4E0B046FFEFFECA5143D686FEAA" LSID="urn:lsid:plazi:treatment:03DE232DB045FFF3FC285181D6DBFEED:8EA8C4E0B046FFEFFECA5143D686FEAA" box="[379,481,291,314]" country="Chile" latitude="-39.976665" longLatPrecision="715" longitude="-73.583336" municipality="Reserva Costera Valdiviana" name="Chaihuin" pageId="15" pageNumber="143" stateProvince="Region Los Rios">Chaihuín</location>
; WDS-T-207;
<geoCoordinate id="EE43F4FCB046FFEFFD365143D593FEAA" box="[647,756,291,314]" degrees="39" direction="south" minutes="58.6" orientation="latitude" pageId="15" pageNumber="143" precision="92" value="-39.976665">39°58.6’S</geoCoordinate>
<geoCoordinate id="EE43F4FCB046FFEFFF265124D660FECB" box="[151,263,324,347]" degrees="73" direction="west" minutes="35" orientation="longitude" pageId="15" pageNumber="143" precision="925" value="-73.583336">73° 35’W</geoCoordinate>
;
<date id="FFC9B4FBB046FFEFFEA65124D6EDFECB" box="[279,394,324,347]" pageId="15" pageNumber="143" value="2008-02-27">
<collectingDate id="EF8D4D13B046FFEFFEA65124D6EDFECB" box="[279,394,324,347]" pageId="15" pageNumber="143" value="2008-02-27">27.II.2008</collectingDate>
</date>
;
<collectedFrom id="386BD1C6B046FFEFFE285124D578FECB" box="[409,543,324,347]" pageId="15" pageNumber="143">sifting litter</collectedFrom>
;
<collectorName id="2682F7EDB046FFEFFD9F5124D5BFFECB" box="[558,728,324,347]" pageId="15" pageNumber="143">W. D. Shepard</collectorName>
</materialsCitation>
.
<materialsCitation id="3B1F9866B046FFEFFD575124D5D0FEEC" ID-GBIF-Occurrence="3886337448" collectionCode="FMNH, MHNG" country="Chile" pageId="15" pageNumber="143" specimenCount="2" specimenCount-male="2" stateProvince="Region Los Rios">
–
<collectionCode id="ED660AFEB046FFEFFF265105D78FFEEC" box="[151,232,357,380]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="15" pageNumber="143" type="Museum">FMNH</collectionCode>
;
<specimenCount id="9D7159B2B046FFEFFF435105D67DFEEC" box="[242,282,357,380]" pageId="15" pageNumber="143" type="male">1 ♂</specimenCount>
; same data. –
<collectionCode id="ED660AFEB046FFEFFE055105D56EFEEC" box="[436,521,357,380]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="143" type="Museum">MHNG</collectionCode>
;
<specimenCount id="9D7159B2B046FFEFFDA25105D55CFEEC" box="[531,571,357,380]" pageId="15" pageNumber="143" type="male">1 ♂</specimenCount>
; same data
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF08C2B3B044FFEDFF2657CFD6F2F78A" ID-DOI="http://doi.org/10.5281/zenodo.5743401" ID-Zenodo-Dep="5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="13" pageNumber="141" startId="13.[151,192,1967,1988]" targetBox="[251,1307,833,1928]" targetPageId="13">
<paragraph id="8BC8923BB044FFEDFF2657CFD6F2F78A" blockId="13.[151,1435,1967,2074]" pageId="13" pageNumber="141">
Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of
<taxonomicName id="4C77E9B8B044FFEDFB8057CFD311F854" authorityName="Reitter" authorityYear="1890" box="[1073,1142,1967,1988]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B044FFEDFB8057CFD311F854" box="[1073,1142,1967,1988]" italics="true" pageId="13" pageNumber="141">Achilia</emphasis>
</taxonomicName>
. (17)
<taxonomicName id="4C77E9B8B044FFEDFB0857CFD242F854" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[1209,1317,1967,1988]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="jeanneli" status="sp. nov.">
<emphasis id="B9034E29B044FFEDFB0857CFD242F854" box="[1209,1317,1967,1988]" italics="true" pageId="13" pageNumber="141">A. jeanneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B044FFEDFA8157CFD20FF854" box="[1328,1384,1967,1988]" pageId="13" pageNumber="141" rank="species">n. sp.</taxonomicNameLabel>
(18)
<taxonomicName id="4C77E9B8B044FFEDFEA057ABD6C7F870" box="[273,416,1995,2016]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B044FFEDFEA057ABD6C7F870" box="[273,416,1995,2016]" italics="true" pageId="13" pageNumber="141">A. valdiviensis</emphasis>
</taxonomicName>
, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19)
<emphasis id="B9034E29B044FFEDFBD657ABD399F870" box="[1127,1278,1995,2016]" italics="true" pageId="13" pageNumber="141">
<taxonomicName id="4C77E9B8B044FFEDFBD657ABD39DF870" box="[1127,1274,1995,2016]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="valdiviensis">A. valdiviensis</taxonomicName>
,
</emphasis>
specimen from Chiloé, Puente La Caldera. (20)
<emphasis id="B9034E29B044FFEDFDFB5788D5DDF86D" box="[586,698,2024,2045]" italics="true" pageId="13" pageNumber="141">
<taxonomicName id="4C77E9B8B044FFEDFDFB5788D5D1F86D" authorityName="Jeannel" authorityYear="1962" box="[586,694,2024,2045]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="bicornis">A. bicornis</taxonomicName>
,
</emphasis>
specimen from Chiloé, Mocopulli. (21)
<emphasis id="B9034E29B044FFEDFBF05788D3D6F86D" box="[1089,1201,2024,2045]" italics="true" pageId="13" pageNumber="141">
<taxonomicName id="4C77E9B8B044FFEDFBF05788D3C9F86D" authorityName="Jeannel" authorityYear="1962" box="[1089,1198,2024,2045]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="141" phylum="Arthropoda" rank="species" species="bicornis">A. bicornis</taxonomicName>
,
</emphasis>
specimen from Osorno, Bahia Mansa.
</paragraph>
</caption>
<caption id="DF08C2B3B047FFEEFF2657ACD526F78B" ID-DOI="http://doi.org/10.5281/zenodo.5743405" ID-Zenodo-Dep="5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="14" pageNumber="142" startId="14.[151,192,1996,2017]" targetBox="[160,1427,435,1963]" targetPageId="14">
<paragraph id="8BC8923BB047FFEEFF2657ACD526F78B" blockId="14.[151,1437,1996,2075]" pageId="14" pageNumber="142">
Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of
<taxonomicName id="4C77E9B8B047FFEEFBAD57ACD306F871" authorityName="Reitter" authorityYear="1890" box="[1052,1121,1996,2017]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B047FFEEFBAD57ACD306F871" box="[1052,1121,1996,2017]" italics="true" pageId="14" pageNumber="142">Achilia</emphasis>
</taxonomicName>
. (22, 29)
<emphasis id="B9034E29B047FFEEFB0E57ACD22BF871" box="[1215,1356,1996,2017]" italics="true" pageId="14" pageNumber="142">
<taxonomicName id="4C77E9B8B047FFEEFB0E57ACD22FF871" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[1215,1352,1996,2017]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="species" species="grandiceps">A. grandiceps</taxonomicName>
.
</emphasis>
(23, 30)
<taxonomicName id="4C77E9B8B047FFEEFEA05789D6ECF86E" box="[273,395,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="species" species="delamarei">
<emphasis id="B9034E29B047FFEEFEA05789D6ECF86E" box="[273,395,2025,2046]" italics="true" pageId="14" pageNumber="142">A. delamarei</emphasis>
</taxonomicName>
. (24, 32)
<taxonomicName id="4C77E9B8B047FFEEFE585789D536F86E" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[489,593,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="species" species="jeanneli" status="sp. nov.">
<emphasis id="B9034E29B047FFEEFE585789D536F86E" box="[489,593,2025,2046]" italics="true" pageId="14" pageNumber="142">A. jeanneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B047FFEEFDE65789D5ECF86E" box="[599,651,2025,2046]" pageId="14" pageNumber="142" rank="species">n. sp.</taxonomicNameLabel>
(25, 33)
<taxonomicName id="4C77E9B8B047FFEEFD505789D453F86E" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[737,820,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="species" species="franzi" status="sp. nov.">
<emphasis id="B9034E29B047FFEEFD505789D453F86E" box="[737,820,2025,2046]" italics="true" pageId="14" pageNumber="142">A. franzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B047FFEEFC8B5789D409F86E" box="[826,878,2025,2046]" pageId="14" pageNumber="142" rank="species">n. sp.</taxonomicNameLabel>
(26-27, 34)
<taxonomicName id="4C77E9B8B047FFEEFC555789D32BF86E" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[996,1100,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="species" species="elguetai" status="sp. nov.">
<emphasis id="B9034E29B047FFEEFC555789D32BF86E" box="[996,1100,2025,2046]" italics="true" pageId="14" pageNumber="142">A. elguetai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B047FFEEFBE35789D3E2F86E" box="[1106,1157,2025,2046]" pageId="14" pageNumber="142" rank="species">n. sp.</taxonomicNameLabel>
(28, 35)
<taxonomicName id="4C77E9B8B047FFEEFB6C5789D20FF86E" box="[1245,1384,2025,2046]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B047FFEEFB6C5789D20FF86E" box="[1245,1384,2025,2046]" italics="true" pageId="14" pageNumber="142">A. valdiviensis</emphasis>
</taxonomicName>
. (31)
<taxonomicName id="4C77E9B8B047FFEEFEA05866D6FFF78B" box="[273,408,2054,2075]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B047FFEEFEA05866D6FFF78B" box="[273,408,2054,2075]" italics="true" pageId="14" pageNumber="142">A. denticornis</emphasis>
</taxonomicName>
. (36)
<taxonomicName id="4C77E9B8B047FFEEFE635866D55CF78B" authorityName="Jeannel" authorityYear="1962" box="[466,571,2054,2075]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="142" phylum="Arthropoda" rank="species" species="bicornis">
<emphasis id="B9034E29B047FFEEFE635866D55CF78B" box="[466,571,2054,2075]" italics="true" pageId="14" pageNumber="142">A. bicornis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C36DC1B0B046FFF0FF2651F5D5E0FCD8" lastPageId="16" lastPageNumber="144" pageId="15" pageNumber="143" type="description">
<paragraph id="8BC8923BB046FFEFFF2651F5D6C3FCCA" blockId="15.[151,757,405,1189]" pageId="15" pageNumber="143">
<emphasis id="B9034E29B046FFEFFF2651F5D641FE3C" bold="true" box="[151,294,405,428]" pageId="15" pageNumber="143">Description:</emphasis>
Body
<quantity id="4C8F3FDEB046FFEFFECC51F5D57AFE3C" box="[381,541,405,428]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="15" pageNumber="143" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.50-1.70 mm</quantity>
long; dark brown with reddish elytra, the latter generally darker at apex and along sutural stria, sometimes also at base; antennae and legs reddish; palpi yellowish. Head wider than long. Pronotum slightly wider than head and wider than long; disc moderately convex; median antebasal fovea slightly smaller than lateral ones; lateral margins with anterior portion distinctly convergent and sinuate anteriorly, and posterior portion slightly convergent and not sinuate. Elytra with discal stria extending to about elytral midlength. First abdominal tergite with slightly diverging basal striae extending to about one-third of paratergal length, and separated at base by more than one-third of tergal width.
</paragraph>
<paragraph id="8BC8923BB046FFF0FF265304D5B8FE69" blockId="15.[151,757,405,1189]" lastBlockId="16.[151,756,258,697]" lastPageId="16" lastPageNumber="144" pageId="15" pageNumber="143">
<emphasis id="B9034E29B046FFEFFF265304D7A9FCEB" box="[151,206,868,891]" italics="true" pageId="15" pageNumber="143">Male</emphasis>
: Head as in
<figureCitation id="134C8EBEB046FFEFFECA5304D69CFCEB" box="[379,507,868,891]" captionStart="Figs 75-78" captionStartId="22.[151,192,2053,2074]" captionTargetBox="[153,1436,892,2015]" captionTargetId="figure-373@22.[153,1438,867,2015]" captionTargetPageId="22" captionText="Figs 75-78. Achilia valdiviensis. Male head in (75) dorsal, (76) lateral, (77) semilateral, and (78) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743423" httpUri="https://zenodo.org/record/5743423/files/figure.png" pageId="15" pageNumber="143">Figs 75-78</figureCitation>
, sub-triangular; frons slightly raised, its surface with some sparse big punctures at middle; two very big and deep vertexal foveae at center of frons at point even with center of eyes, at same distance from each other as from nearest eye; frontal sulcus lacking; frontal lobe pointed in middle, with thick tuft of straight yellowish bristles extended forwards; clypeus strongly prolonged forward, duck’s beak-shaped. Eyes protruding, distinctly longer than short, with convex temples. Antennae (
<figureCitation id="134C8EBEB046FFEFFEC154EED68EFB35" box="[368,489,1166,1189]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="15" pageNumber="143">Figs 18-19</figureCitation>
) with scape longer than wide, more (
<figureCitation id="134C8EBEB046FFEFFC795162D371FE89" box="[968,1046,258,281]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="15" pageNumber="143">Fig. 19</figureCitation>
) or less (
<figureCitation id="134C8EBEB046FFEFFBCA5162D3AEFE89" box="[1147,1225,258,281]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="15" pageNumber="143">Fig. 18</figureCitation>
) elongated; pedicel misaligned, flattened and excavated on medial surface, as long as wide with mesal apical edge very pronounced (
<figureCitation id="134C8EBEB046FFEFFCF65103D4F2FEEA" box="[839,917,355,378]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="15" pageNumber="143">Fig. 18</figureCitation>
), or subrectangular in shape and distinctly longer than wide (
<figureCitation id="134C8EBEB046FFEFFC0651E4D361FE0B" box="[951,1030,388,411]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="15" pageNumber="143">Fig. 19</figureCitation>
); antennomere III slightly longer than wide; antennomere IV wider than long; antennomere V as long as wide; VI slightly wider than long; antennomere VII slightly longer than wide; antennomere VIII transverse; antennomere IX transverse with denticulate margins and protruding mesal margin; antennomere X wider than IX, wider than long with denticulate margin and protruding mesal margin; antennomere XI very elongate, also with denticulate margins, distinctly longer than VII-X combined, usually bearing long subbasal seta on mesal margin. Metaventrite with distal half covered by convergent long bristles, raised at middle for twothirds of its distal portion, this surface entirely divided by median sulcus. Protrochanters (
<figureCitation id="134C8EBEB046FFEFFB085349D26EFCD0" box="[1209,1289,809,832]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="15" pageNumber="143">Fig. 28</figureCitation>
) with ventral margin formed as spine bearing one long basal seta; mesotrochanters (
<figureCitation id="134C8EBEB046FFEFFBB1530AD335FC11" box="[1024,1106,874,897]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="15" pageNumber="143">Fig. 35</figureCitation>
) with ventral margin forming long spine at basal third; mesofemora (
<figureCitation id="134C8EBEB046FFEFFB6653EBD243FC32" box="[1239,1316,907,930]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="15" pageNumber="143">Fig. 35</figureCitation>
) with basal third of ventral margin covered by broad, short and thick setae, those setae longer than in
<taxonomicName id="4C77E9B8B046FFEFFB2B53ACD248FC73" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[1178,1327,972,995]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="B9034E29B046FFEFFB2B53ACD248FC73" box="[1178,1327,972,995]" italics="true" pageId="15" pageNumber="143">A. grandiceps</emphasis>
</taxonomicName>
. Protibiae (
<figureCitation id="134C8EBEB046FFEFFCF6538CD4FDFB93" box="[839,922,1004,1027]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="15" pageNumber="143">Fig. 39</figureCitation>
) enlarged for distal half, with medial margin forming small apical spine; mesotibiae (
<figureCitation id="134C8EBEB046FFEFFAB7546CD23EFBB3" box="[1286,1369,1036,1059]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="15" pageNumber="143">Fig. 43</figureCitation>
) with medial margin without recurved setae, apical margin forming two short spines. First abdominal sternite very long, projecting over second sternite which is thus almost entirely concealed. Aedeagus (
<figureCitation id="134C8EBEB046FFEFFB2E54EED380FB35" box="[1183,1255,1166,1189]" captionStart="Figs 1-6" captionStartId="10.[151,192,2025,2046]" captionTargetBox="[175,1415,715,1976]" captionTargetId="figure-193@10.[151,1436,686,2025]" captionTargetPageId="10" captionText="Figs 1-6. Aedeagi of Achilia species. (1) A. grandiceps. (2) A. delamarei. (3) A. jeanneli n. sp. (4) A. franzi n. sp. (5) A. elguetai n. sp. (6) A. valdiviensis." figureDoi="http://doi.org/10.5281/zenodo.5743393" httpUri="https://zenodo.org/record/5743393/files/figure.png" pageId="15" pageNumber="143">Fig. 6</figureCitation>
)
<quantity id="4C8F3FDEB046FFEFFB4F54EED2FBFB35" box="[1278,1436,1166,1189]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" metricValueMax="3.1" metricValueMin="2.9" pageId="15" pageNumber="143" unit="mm" value="0.3" valueMax="0.31" valueMin="0.29">0.29-0.31 mm</quantity>
long; with suboval dorsal plate, dorsal longitudinal struts slightly divergent. Parameres relatively wide with large and short seta on small outer lobe; tips rounded bearing large median seta. Copulatory pieces consisting of a pair of long wide medial sclerites basally recurved, sclerotized and apically rounded, with pair of thin lateral sclerites subequal in length, recurved, enlarged, sclerotized at base, with distal half thickly pitted near lateral margin.
</paragraph>
<caption id="DF08C2B3B046FFEFFF265714D4E1F789" ID-DOI="http://doi.org/10.5281/zenodo.5743407" ID-Zenodo-Dep="5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="15" pageNumber="143" startId="15.[151,192,1908,1929]" targetBox="[189,1416,1253,1860]" targetPageId="15">
<paragraph id="8BC8923BB046FFEFFF265714D4E1F789" blockId="15.[151,1437,1908,2073]" pageId="15" pageNumber="143">
Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of
<emphasis id="B9034E29B046FFEFFCB75714D435F819" box="[774,850,1908,1929]" italics="true" pageId="15" pageNumber="143">
<taxonomicName id="4C77E9B8B046FFEFFCB75714D429F819" authorityName="Reitter" authorityYear="1890" box="[774,846,1908,1929]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="genus">Achilia</taxonomicName>
.
</emphasis>
(37, 49)
<taxonomicName id="4C77E9B8B046FFEFFC035714D37AF819" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[946,1053,1908,1929]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="elguetai" status="sp. nov.">
<emphasis id="B9034E29B046FFEFFC035714D37AF819" box="[946,1053,1908,1929]" italics="true" pageId="15" pageNumber="143">A. elguetai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B046FFEFFB975714D33AF819" box="[1062,1117,1908,1929]" pageId="15" pageNumber="143" rank="species">n. sp.</taxonomicNameLabel>
(38, 44)
<taxonomicName id="4C77E9B8B046FFEFFB0F5714D220F819" box="[1214,1351,1908,1929]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="denticornis">
<emphasis id="B9034E29B046FFEFFB0F5714D220F819" box="[1214,1351,1908,1929]" italics="true" pageId="15" pageNumber="143">A. denticornis</emphasis>
</taxonomicName>
. (39)
<taxonomicName id="4C77E9B8B046FFEFFA395714D61BF836" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="vadiviensis">
<emphasis id="B9034E29B046FFEFFA395714D61BF836" italics="true" pageId="15" pageNumber="143">A. vadiviensis</emphasis>
</taxonomicName>
. (40-41)
<taxonomicName id="4C77E9B8B046FFEFFE6257F1D53FF836" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[467,600,1937,1958]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="B9034E29B046FFEFFE6257F1D53FF836" box="[467,600,1937,1958]" italics="true" pageId="15" pageNumber="143">A. grandiceps</emphasis>
</taxonomicName>
, specimen from: (40) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42)
<taxonomicName id="4C77E9B8B046FFEFFCB357CED41BF853" box="[770,892,1966,1987]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="delamarei">
<emphasis id="B9034E29B046FFEFFCB357CED41BF853" box="[770,892,1966,1987]" italics="true" pageId="15" pageNumber="143">A. delamarei</emphasis>
</taxonomicName>
. (43)
<taxonomicName id="4C77E9B8B046FFEFFC0557CED358F853" box="[948,1087,1966,1987]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B046FFEFFC0557CED358F853" box="[948,1087,1966,1987]" italics="true" pageId="15" pageNumber="143">A. valdiviensis</emphasis>
</taxonomicName>
. (45-46)
<taxonomicName id="4C77E9B8B046FFEFFB2957CED399F853" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[1176,1278,1966,1987]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="jeanneli" status="sp. nov.">
<emphasis id="B9034E29B046FFEFFB2957CED399F853" box="[1176,1278,1966,1987]" italics="true" pageId="15" pageNumber="143">A. jeanneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B046FFEFFAB557CED25EF853" box="[1284,1337,1966,1987]" pageId="15" pageNumber="143" rank="species">n. sp.</taxonomicNameLabel>
, specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48)
<taxonomicName id="4C77E9B8B046FFEFFD205787D581F86C" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[657,742,2023,2044]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="franzi" status="sp. nov.">
<emphasis id="B9034E29B046FFEFFD205787D581F86C" box="[657,742,2023,2044]" italics="true" pageId="15" pageNumber="143">A. franzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B046FFEFFD5C5787D442F86C" box="[749,805,2023,2044]" pageId="15" pageNumber="143" rank="species">n. sp.</taxonomicNameLabel>
, specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50)
<taxonomicName id="4C77E9B8B046FFEFFCA65864D4E7F789" authorityName="Jeannel" authorityYear="1962" box="[791,896,2052,2073]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="143" phylum="Arthropoda" rank="species" species="bicornis">
<emphasis id="B9034E29B046FFEFFCA65864D4E7F789" box="[791,896,2052,2073]" italics="true" pageId="15" pageNumber="143">A. bicornis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC8923BB059FFF0FF265262D610FD29" blockId="16.[151,756,258,697]" pageId="16" pageNumber="144">
<emphasis id="B9034E29B059FFF0FF265262D780FD89" box="[151,231,514,537]" italics="true" pageId="16" pageNumber="144">Female</emphasis>
: Similar to male except eyes smaller and less protruding, frontal lobe barely pointed at middle and without tuft of setae, clypeus just slightly elongated, antennae with pedicel barely misaligned, subrectangular and not excavated. Metaventrite, abdominal sternites, and legs unmodified.
</paragraph>
<paragraph id="8BC8923BB059FFF0FF2652B1D5E0FCD8" blockId="16.[151,757,721,840]" pageId="16" pageNumber="144">
<emphasis id="B9034E29B059FFF0FF2652B1D634FD78" bold="true" box="[151,339,721,744]" pageId="16" pageNumber="144">Collecting data:</emphasis>
Collected from October to February in valdivian rainforests, presumably at low elevations. All specimens came from sifted samples of leaf and log litter, except for
<specimenCount id="9D7159B2B059FFF0FEF95351D6CEFCD8" box="[328,425,817,840]" pageId="16" pageNumber="144" type="male">one male</specimenCount>
collected by car net.
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B059FFF0FC8E5162D4C8FEC9" pageId="16" pageNumber="144" type="distribution">
<paragraph id="8BC8923BB059FFF0FC8E5162D4C8FEC9" blockId="16.[831,1436,258,345]" pageId="16" pageNumber="144">
<emphasis id="B9034E29B059FFF0FC8E5162D4B3FE89" bold="true" box="[831,980,258,281]" pageId="16" pageNumber="144">Distribution:</emphasis>
The species is known only from southern Chile (
<figureCitation id="134C8EBEB059FFF0FC385142D4BDFEA9" box="[905,986,290,313]" captionStart="Fig" captionStartId="25.[831,866,1909,1930]" captionTargetBox="[868,1366,257,1880]" captionTargetId="figure-92@25.[918,1366,254,1877]" captionText="Fig. 84. Distribution map of Achilia bicornis and valdiviensis group. (l red circles) A. bicornis. (green squares) A. valdiviensis." figureDoi="http://doi.org/10.5281/zenodo.6979915" httpUri="https://zenodo.org/record/6979915/files/figure.png" pageId="16" pageNumber="144">Fig. 84</figureCitation>
: green squares) from Chiloé and Valdivia provinces.
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B059FFF3FC8E5111D6DBFEED" lastPageId="19" lastPageNumber="147" pageId="16" pageNumber="144" type="discussion">
<paragraph id="8BC8923BB059FFF0FC8E5111D4ACFDD8" blockId="16.[831,1437,369,840]" pageId="16" pageNumber="144">
<emphasis id="B9034E29B059FFF0FC8E5111D4A1FE18" bold="true" box="[831,966,369,392]" pageId="16" pageNumber="144">Comments:</emphasis>
Blanchard described
<taxonomicName id="4C77E9B8B059FFF0FB685111D375FE38" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="144" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B059FFF0FB685111D4EFFE38" italics="true" pageId="16" pageNumber="144">Pselaphus valdiviensis</emphasis>
(1851: 563)
</taxonomicName>
based on an unspecified number of specimens from Valdivia. The description, very concise, did not report clear diagnostic characters such that subsequent authors (
<bibRefCitation id="EFE6EFCAB059FFF0FB985191D3ABFD98" author="Reitter E." box="[1065,1228,497,520]" pageId="16" pageNumber="144" pagination="321 - 332" refId="ref22459" refString="Reitter E. 1885 a. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia, Zweiter Theil. Deutsche Entomologische Zeitschrift 29: 321 - 332, pl. II." type="journal article" year="1885">Reitter, 1885a</bibRefCitation>
;
<bibRefCitation id="EFE6EFCAB059FFF0FB6F5191D2FFFD98" author="Schaufuss L. W." box="[1246,1432,497,520]" pageId="16" pageNumber="144" pagination="241 - 296" refId="ref22694" refString="Schaufuss L. W. 1886. Beschreibung neuer Pselaphiden aus der Sammlung des Museums Ludwig Salvador. Ein Beitrag zur Fauna Brasiliens, der Kgl. niederlandischen Besitzungen in Indien und Neuholland. Tijdschrift voor Entomologie 29: 241 - 296." type="journal article" year="1886">Schaufuss, 1886</bibRefCitation>
; and
<bibRefCitation id="EFE6EFCAB059FFF0FCC15271D360FDB8" author="Raffray A." box="[880,1031,529,552]" pageId="16" pageNumber="144" pagination="391 - 400" refId="ref22316" refString="Raffray A. 1895. Revision du genre Tyropsis Saulcy (Aplodea Reitter) et description de deux genres nouveaux du meme groupe. Annales de la Societe Entomologique de France 64: 391 - 400." type="journal article" year="1895">Raffray, 1895</bibRefCitation>
) thought this species was a member of the Tyrini.
</paragraph>
<paragraph id="8BC8923BB059FFF1FC8E5231D63DFE29" blockId="16.[831,1437,369,840]" lastBlockId="17.[151,756,258,793]" lastPageId="17" lastPageNumber="145" pageId="16" pageNumber="144">
<bibRefCitation id="EFE6EFCAB059FFF0FC8E5231D346FDF8" author="Reitter E." box="[831,1057,593,616]" pageId="16" pageNumber="144" pagination="47 - 54" refId="ref22433" refString="Reitter E. 1883. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia. Deutsche Entomologische Zeitschrift 27: 47 - 54, pl. I." type="journal article" year="1883">Reitter (1883: 50)</bibRefCitation>
described
<taxonomicName id="4C77E9B8B059FFF0FB045231D2FBFDF8" authorityName="Reitter" authorityYear="1883" box="[1205,1436,593,616]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="144" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B059FFF0FB045231D2FBFDF8" box="[1205,1436,593,616]" italics="true" pageId="16" pageNumber="144">Bryaxis valdiviensis</emphasis>
</taxonomicName>
(technically
<emphasis id="B9034E29B059FFF0FC765211D322FD18" box="[967,1093,625,648]" italics="true" pageId="16" pageNumber="144">valvidiensis</emphasis>
in the original description due to a
<emphasis id="B9034E29B059FFF0FCCA52F1D372FD38" box="[891,1045,657,680]" italics="true" pageId="16" pageNumber="144">lapsus calami</emphasis>
) based on an unspecified number of specimens from Valdivia, without any mention of
<taxonomicName id="4C77E9B8B059FFF0FC8E52B1D354FD78" authorityName="Blanchard" authorityYear="1851" box="[831,1075,721,744]" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="144" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B059FFF0FC8E52B1D354FD78" box="[831,1075,721,744]" italics="true" pageId="16" pageNumber="144">Pselaphus valdiviensis</emphasis>
</taxonomicName>
, and two years later he described (
<bibRefCitation id="EFE6EFCAB059FFF0FCF65291D371FC98" author="Reitter E." box="[839,1046,753,776]" pageId="16" pageNumber="144" pagination="321 - 332" refId="ref22459" refString="Reitter E. 1885 a. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia, Zweiter Theil. Deutsche Entomologische Zeitschrift 29: 321 - 332, pl. II." type="journal article" year="1885">Reitter 1885a: 325</bibRefCitation>
and 327)
<taxonomicName id="4C77E9B8B059FFF0FB3C5291D257FC98" box="[1165,1328,753,776]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="144" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B059FFF0FB3C5291D257FC98" box="[1165,1328,753,776]" italics="true" pageId="16" pageNumber="144">Bryaxis nasuta</emphasis>
</taxonomicName>
based on a single male specimen from Valdivia that had been collected by the cousins Elsbeth and Elfride Kindermann. In the same year
<bibRefCitation id="EFE6EFCAB058FFF1FEE25162D555FE89" author="Reitter E." box="[339,562,258,281]" pageId="17" pageNumber="145" pagination="313 - 317" refId="ref22489" refString="Reitter E. 1885 b. Coleopterologische Notizen XIV. Wiener Entomologische Zeitung 4 (10): 313 - 317." type="journal article" year="1885">Reitter (1885b: 317)</bibRefCitation>
, realizing that the name
<taxonomicName id="4C77E9B8B058FFF1FF505142D6EDFEA9" box="[225,394,290,313]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B058FFF1FF505142D6EDFEA9" box="[225,394,290,313]" italics="true" pageId="17" pageNumber="145">Bryaxis nasuta</emphasis>
</taxonomicName>
was preoccupied, proposed to change the species name to
<emphasis id="B9034E29B058FFF1FE7C5122D506FEC9" box="[461,609,322,345]" italics="true" pageId="17" pageNumber="145">
<taxonomicName id="4C77E9B8B058FFF1FE7C5122D539FEC9" authorityName="Reitter" authorityYear="1885" box="[461,606,322,345]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="anas">Bryaxis anas</taxonomicName>
.
</emphasis>
Later
<bibRefCitation id="EFE6EFCAB058FFF1FD1A5122D67CFEE9" author="Reitter E." pageId="17" pageNumber="145" pagination="260 - 261" refId="ref22511" refString="Reitter E. 1893. Coleopterologische Notizen XLVIII. Wiener Entomologische Zeitung 12 (7): 260 - 261." type="journal article" year="1893">Reitter (1893: 261)</bibRefCitation>
, apparently forgetting he already done so, proposed a second name for
<taxonomicName id="4C77E9B8B058FFF1FE7451E2D504FE09" box="[453,611,386,409]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B058FFF1FE7451E2D504FE09" box="[453,611,386,409]" italics="true" pageId="17" pageNumber="145">Bryaxis nasuta</emphasis>
</taxonomicName>
, but this time as
<taxonomicName id="4C77E9B8B058FFF1FF0551C2D634FE29" authorityName="Reitter" authorityYear="1893" box="[180,339,418,441]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="nasina">
<emphasis id="B9034E29B058FFF1FF0551C2D634FE29" box="[180,339,418,441]" italics="true" pageId="17" pageNumber="145">Bryaxis nasina</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF08C2B3B059FFF0FF275867D2E9F78C" ID-DOI="http://doi.org/10.5281/zenodo.5743409" ID-Zenodo-Dep="5743409" box="[150,1422,2055,2076]" httpUri="https://zenodo.org/record/5743409/files/figure.png" pageId="16" pageNumber="144" startId="16.[150,191,2055,2076]" targetBox="[151,1435,901,2016]" targetPageId="16">
<paragraph id="8BC8923BB059FFF0FF275867D2E9F78C" blockId="16.[150,1422,2055,2076]" box="[150,1422,2055,2076]" pageId="16" pageNumber="144">
Figs 51-54.
<taxonomicName id="4C77E9B8B059FFF0FEA05867D6A0F78C" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[273,455,2055,2076]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="144" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="B9034E29B059FFF0FEA05867D6A0F78C" box="[273,455,2055,2076]" italics="true" pageId="16" pageNumber="144">Achilia grandiceps</emphasis>
</taxonomicName>
. Male head in (51) dorsal, (52) lateral, (53) semilateral, and (54) frontal views. Scale bars (200 μm).
</paragraph>
</caption>
<paragraph id="8BC8923BB058FFF1FF2651A2D7BAFD69" blockId="17.[151,756,258,793]" pageId="17" pageNumber="145">
<bibRefCitation id="EFE6EFCAB058FFF1FF2651A2D60DFE49" author="Raffray A." box="[151,362,450,473]" pageId="17" pageNumber="145" pagination="1 - 400" refId="ref22353" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray (1904: 138)</bibRefCitation>
, having studied the
<typeStatus id="54CC2C99B058FFF1FD8851A2D500FE49" box="[569,615,450,473]" pageId="17" pageNumber="145">type</typeStatus>
of
<taxonomicName id="4C77E9B8B058FFF1FD3751A2D6A8FE69" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B058FFF1FD3751A2D672FE69" italics="true" pageId="17" pageNumber="145">Pselaphus valdiviensis</emphasis>
<bibRefCitation id="EFE6EFCAB058FFF1FEAD5182D6A8FE69" author="Blanchard C. E." box="[284,463,482,505]" pageId="17" pageNumber="145" pagination="1 - 564" refId="ref21916" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos [pp. 561 - 564]. In: Gay C., Historia fisica y politica de Chile. Zoologia, 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
, which he believed to be in the MNHN collections, established that this species was not a member of the Tyrini, but should be transferred to
<taxonomicName id="4C77E9B8B058FFF1FF265222D783FDC9" authorityName="Reitter" authorityYear="1890" box="[151,228,578,601]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B058FFF1FF265222D783FDC9" box="[151,228,578,601]" italics="true" pageId="17" pageNumber="145">Achilia</emphasis>
</taxonomicName>
in the Brachyglutini, and therefore proposed the substitute name
<taxonomicName id="4C77E9B8B058FFF1FEE35202D5A0FDE9" authority="Raffray, 1904" authorityName="Raffray" authorityYear="1904" box="[338,711,610,633]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="blanchardi">
<emphasis id="B9034E29B058FFF1FEE35202D546FDE9" box="[338,545,610,633]" italics="true" pageId="17" pageNumber="145">Achilia blanchardi</emphasis>
<bibRefCitation id="EFE6EFCAB058FFF1FD9F5202D5A0FDE9" author="Raffray A." box="[558,711,610,633]" pageId="17" pageNumber="145" pagination="1 - 400" refId="ref22353" refString="Raffray A. 1904. Genera et Catalogue des Pselaphides. Annales de la Societe Entomologique de France 73: 1 - 400." type="journal article" year="1904">Raffray, 1904</bibRefCitation>
</taxonomicName>
for
<taxonomicName id="4C77E9B8B058FFF1FF2652E2D562FD09" authority="Reitter, 1883" authorityName="Reitter" authorityYear="1883" box="[151,517,642,665]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B058FFF1FF2652E2D608FD09" box="[151,367,642,665]" italics="true" pageId="17" pageNumber="145">Bryaxis valdiviensis</emphasis>
<bibRefCitation id="EFE6EFCAB058FFF1FEC652E2D562FD09" author="Reitter E." box="[375,517,642,665]" pageId="17" pageNumber="145" pagination="47 - 54" refId="ref22433" refString="Reitter E. 1883. Beitrag zur Kenntniss der Pselaphiden-Fauna von Valdivia. Deutsche Entomologische Zeitschrift 27: 47 - 54, pl. I." type="journal article" year="1883">Reitter, 1883</bibRefCitation>
</taxonomicName>
, and pointing out that
<taxonomicName id="4C77E9B8B058FFF1FF2652C2D6A8FD29" authority="Reitter, 1885" authorityName="Reitter" authorityYear="1885" box="[151,463,674,697]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B058FFF1FF2652C2D65EFD29" box="[151,313,674,697]" italics="true" pageId="17" pageNumber="145">Bryaxis nasuta</emphasis>
Reitter, 1885
</taxonomicName>
(=
<taxonomicName id="4C77E9B8B058FFF1FE4B52C2D7A9FD49" authority="Reitter, 1893" authorityName="Reitter" authorityYear="1893" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="nasina">
<emphasis id="B9034E29B058FFF1FE4B52C2D5FBFD29" box="[506,668,674,697]" italics="true" pageId="17" pageNumber="145">Bryaxis nasina</emphasis>
<bibRefCitation id="EFE6EFCAB058FFF1FD1752C2D7A9FD49" author="Reitter E." pageId="17" pageNumber="145" pagination="260 - 261" refId="ref22511" refString="Reitter E. 1893. Coleopterologische Notizen XLVIII. Wiener Entomologische Zeitung 12 (7): 260 - 261." type="journal article" year="1893">Reitter, 1893</bibRefCitation>
</taxonomicName>
) was identical to
<taxonomicName id="4C77E9B8B058FFF1FE2952A2D7BFFD69" authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B058FFF1FE2952A2D50AFD49" box="[408,621,706,729]" italics="true" pageId="17" pageNumber="145">Achilia valdiviensis</emphasis>
(
<bibRefCitation id="EFE6EFCAB058FFF1FDCF52A2D7B7FD69" author="Blanchard C. E." pageId="17" pageNumber="145" pagination="1 - 564" refId="ref21916" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos [pp. 561 - 564]. In: Gay C., Historia fisica y politica de Chile. Zoologia, 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BC8923BB058FFF1FF265362D4B2FE49" blockId="17.[151,756,258,793]" lastBlockId="17.[831,1437,258,793]" pageId="17" pageNumber="145">
Then
<bibRefCitation id="EFE6EFCAB058FFF1FF675362D6B9FC89" author="Jeannel R." box="[214,478,770,793]" pageId="17" pageNumber="145" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel (1962: 424-425)</bibRefCitation>
claimed that the
<typeStatus id="54CC2C99B058FFF1FD2C5362D5B2FC89" box="[669,725,770,793]" pageId="17" pageNumber="145">types</typeStatus>
of
<taxonomicName id="4C77E9B8B058FFF1FC8E5162D3E4FE89" authority="Reitter, 1885" authorityName="Reitter" authorityYear="1885" box="[831,1155,258,281]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B058FFF1FC8E5162D482FE89" box="[831,997,258,281]" italics="true" pageId="17" pageNumber="145">Bryaxis nasuta</emphasis>
Reitter, 1885
</taxonomicName>
and
<taxonomicName id="4C77E9B8B058FFF1FB755162D369FEA9" authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B058FFF1FB755162D2FBFE89" box="[1220,1436,258,281]" italics="true" pageId="17" pageNumber="145">Achilia valdiviensis</emphasis>
(
<bibRefCitation id="EFE6EFCAB058FFF1FCF65142D362FEA9" author="Blanchard C. E." box="[839,1029,290,313]" pageId="17" pageNumber="145" pagination="1 - 564" refId="ref21916" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos [pp. 561 - 564]. In: Gay C., Historia fisica y politica de Chile. Zoologia, 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</taxonomicName>
, both collected in Valdivia, were present in the MNHN collections, and among the material examined, he mentions only
<specimenCount id="9D7159B2B058FFF1FB385102D259FEE9" box="[1161,1342,354,377]" pageId="17" pageNumber="145" type="generic">three specimens</specimenCount>
: Central Chile: Valdivia prov.: Env. of Valdivia (39 ° 50 ‘lat. S) male and female (Cl. Gay), and another male (E. and E. Kindermann).
</paragraph>
<paragraph id="8BC8923BB058FFF2FC8E5182D5CBFDC9" blockId="17.[831,1437,258,793]" lastBlockId="18.[151,756,258,697]" lastPageId="18" lastPageNumber="146" pageId="17" pageNumber="145">
However in MNHN we could find only
<specimenCount id="9D7159B2B058FFF1FB485182D22AFE69" box="[1273,1357,482,505]" pageId="17" pageNumber="145" type="male">3 males</specimenCount>
of this species:
<specimenCount id="9D7159B2B058FFF1FC2C5262D48FFD89" box="[925,1000,514,537]" pageId="17" pageNumber="145" type="male">1 male</specimenCount>
in the Raffray collection labeled “Chili / Museum Paris, 1917, coll. Raffray /
<typeStatus id="54CC2C99B058FFF1FB5C5242D245FDA9" box="[1261,1314,546,569]" pageId="17" pageNumber="145">Type</typeStatus>
/
<emphasis id="B9034E29B058FFF1FA8E5242D2FBFDA9" box="[1343,1436,546,569]" italics="true" pageId="17" pageNumber="145">chilensis</emphasis>
Reitter /
<taxonomicName id="4C77E9B8B058FFF1FC175222D3AAFDC9" authority=", A. Raffray" authorityName="Blch., A. Raffray" authorityYear="1904" box="[934,1229,578,601]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B058FFF1FC175222D320FDC9" box="[934,1095,578,601]" italics="true" pageId="17" pageNumber="145">A. valdiviensis</emphasis>
, A. Raffray
</taxonomicName>
det. /
<taxonomicName id="4C77E9B8B058FFF1FAAF5222D407FDE9" authority="Bl." authorityName="Bl." class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B058FFF1FAAF5222D2FBFDC9" box="[1310,1436,578,601]" italics="true" pageId="17" pageNumber="145">valdiviensis</emphasis>
Bl.
</taxonomicName>
(handwritten by Jeannel)”, and
<specimenCount id="9D7159B2B058FFF1FB585202D222FDE9" box="[1257,1349,610,633]" pageId="17" pageNumber="145" type="male">2 males</specimenCount>
in the general Chile collection – one labeled: “Museum Paris, Chili, Gay 1849 / H 49 / Valdivia / Gen.
<taxonomicName id="4C77E9B8B058FFF1FABA52C2D2FAFD29" authority="Reitt." authorityName="Reitter" authorityYear="1890" box="[1291,1437,674,697]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B058FFF1FABA52C2D23FFD29" box="[1291,1368,674,697]" italics="true" pageId="17" pageNumber="145">Achilia</emphasis>
Reitt.
</taxonomicName>
=,
<taxonomicName id="4C77E9B8B058FFF1FCEF52A2D49FFD49" authority="Raffr." authorityName="Raffr." box="[862,1016,706,729]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B058FFF1FCEF52A2D4C8FD49" box="[862,943,706,729]" italics="true" pageId="17" pageNumber="145">Bryaxis</emphasis>
Raffr.
</taxonomicName>
, =
<taxonomicName id="4C77E9B8B058FFF1FBAA52A2D398FD49" authority="Blanchard" authorityName="Blanchard" box="[1051,1279,706,729]" class="Insecta" family="Staphylinidae" genus="Pselaphus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B9034E29B058FFF1FBAA52A2D3EEFD49" box="[1051,1161,706,729]" italics="true" pageId="17" pageNumber="145">Pselaphus</emphasis>
Blanchard
</taxonomicName>
/
<taxonomicName id="4C77E9B8B058FFF1FAA652A2D2F2FD49" box="[1303,1429,706,729]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B058FFF1FAA652A2D2F2FD49" box="[1303,1429,706,729]" italics="true" pageId="17" pageNumber="145">valdiviensis</emphasis>
</taxonomicName>
; Blanch. =
<taxonomicName id="4C77E9B8B058FFF1FC1E5282D320FD69" authority="Reitter" authorityName="Reitter" authorityYear="1893" box="[943,1095,738,761]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="nasina">
<emphasis id="B9034E29B058FFF1FC1E5282D491FD69" box="[943,1014,738,761]" italics="true" pageId="17" pageNumber="145">nasina</emphasis>
Reitter
</taxonomicName>
, =
<taxonomicName id="4C77E9B8B058FFF1FBD95282D209FD69" authority="Reitt., A. Raffray" authorityName="Reitt., A. Raffray" box="[1128,1390,738,761]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B058FFF1FBD95282D3C8FD69" box="[1128,1199,738,761]" italics="true" pageId="17" pageNumber="145">nasuta</emphasis>
Reitt., A. Raffray
</taxonomicName>
det. 1904 /
<taxonomicName id="4C77E9B8B058FFF1FC235362D35BFC89" authority="Bl." authorityName="Bl." box="[914,1084,770,793]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B058FFF1FC235362D377FC89" box="[914,1040,770,793]" italics="true" pageId="17" pageNumber="145">valdiviensis</emphasis>
Bl.
</taxonomicName>
(handwritten by Jeannel)”, and the second labeled “Museum Paris, Chili, Gay 1849 / H 49/ Valdivia /
<taxonomicName id="4C77E9B8B05BFFF2FEE75142D593FEA9" authority="Blch., A. Raffray" authorityName="Blch., A. Raffray" authorityYear="1904" box="[342,756,290,313]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B05BFFF2FEE75142D54BFEA9" box="[342,556,290,313]" italics="true" pageId="18" pageNumber="146">Achilia valdiviensis</emphasis>
Blch., A. Raffray
</taxonomicName>
det. 1904”. We think that these
<specimenCount id="9D7159B2B05BFFF2FE465122D501FEC9" box="[503,614,322,345]" pageId="18" pageNumber="146" type="male">two males</specimenCount>
collected by Gay and studied by Raffray are part of the typical series of
<taxonomicName id="4C77E9B8B05BFFF2FF0751E2D53FFE09" authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[182,600,386,409]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B05BFFF2FF0751E2D6EEFE09" box="[182,393,386,409]" italics="true" pageId="18" pageNumber="146">Achilia valdiviensis</emphasis>
(
<bibRefCitation id="EFE6EFCAB05BFFF2FE2851E2D528FE09" author="Blanchard C. E." box="[409,591,386,409]" pageId="18" pageNumber="146" pagination="1 - 564" refId="ref21916" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos [pp. 561 - 564]. In: Gay C., Historia fisica y politica de Chile. Zoologia, 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</taxonomicName>
, and therefore designate them here as
<typeStatus id="54CC2C99B05BFFF2FE1251C2D562FE29" box="[419,517,418,441]" pageId="18" pageNumber="146" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54CC2C99B05BFFF2FDF251C2D5B4FE29" box="[579,723,418,441]" pageId="18" pageNumber="146" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C77E9B8B05BFFF2FF2651A2D55BFE49" authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[151,572,450,473]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B05BFFF2FF2651A2D60BFE49" box="[151,364,450,473]" italics="true" pageId="18" pageNumber="146">Achilia valdiviensis</emphasis>
(
<bibRefCitation id="EFE6EFCAB05BFFF2FECD51A2D553FE49" author="Blanchard C. E." box="[380,564,450,473]" pageId="18" pageNumber="146" pagination="1 - 564" refId="ref21916" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos [pp. 561 - 564]. In: Gay C., Historia fisica y politica de Chile. Zoologia, 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</taxonomicName>
. It is very likely that the male in the Raffray collection, which is labeled as the
<typeStatus id="54CC2C99B05BFFF2FF575262D673FD89" box="[230,276,514,537]" pageId="18" pageNumber="146">type</typeStatus>
of
<taxonomicName id="4C77E9B8B05BFFF2FEF25262D683FD89" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[323,484,514,537]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B05BFFF2FEF25262D683FD89" box="[323,484,514,537]" italics="true" pageId="18" pageNumber="146">A. valdiviensis</emphasis>
</taxonomicName>
despite the handwritten label
<emphasis id="B9034E29B05BFFF2FF685242D651FDA9" box="[217,310,546,569]" italics="true" pageId="18" pageNumber="146">chilensis</emphasis>
by Reitter, is the
<typeStatus id="54CC2C99B05BFFF2FDA65242D513FDA9" box="[535,628,546,569]" pageId="18" pageNumber="146" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C77E9B8B05BFFF2FD165242D6E6FDC9" authority="(Reitter, 1885)" baseAuthorityName="Reitter" baseAuthorityYear="1885" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="nasuta">
<emphasis id="B9034E29B05BFFF2FD165242D7B9FDC9" italics="true" pageId="18" pageNumber="146">Achilia nasuta</emphasis>
(Reitter, 1885)
</taxonomicName>
(=
<taxonomicName id="4C77E9B8B05BFFF2FE175222D5F8FDC9" authority="Reitter, 1893" authorityName="Reitter" authorityYear="1893" box="[422,671,578,601]" class="Insecta" family="Staphylinidae" genus="Bryaxis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="nasina">
<emphasis id="B9034E29B05BFFF2FE175222D56CFDC9" box="[422,523,578,601]" italics="true" pageId="18" pageNumber="146">A. nasina</emphasis>
<bibRefCitation id="EFE6EFCAB05BFFF2FDA35222D5F8FDC9" author="Reitter E." box="[530,671,578,601]" pageId="18" pageNumber="146" pagination="260 - 261" refId="ref22511" refString="Reitter E. 1893. Coleopterologische Notizen XLVIII. Wiener Entomologische Zeitung 12 (7): 260 - 261." type="journal article" year="1893">Reitter, 1893</bibRefCitation>
</taxonomicName>
).
</paragraph>
<caption id="DF08C2B3B058FFF1FF265866D21DF78B" ID-DOI="http://doi.org/10.5281/zenodo.5743413" ID-Zenodo-Dep="5743413" box="[151,1402,2054,2075]" httpUri="https://zenodo.org/record/5743413/files/figure.png" pageId="17" pageNumber="145" startId="17.[151,192,2054,2075]" targetBox="[151,1435,871,2000]" targetPageId="17">
<paragraph id="8BC8923BB058FFF1FF265866D21DF78B" blockId="17.[151,1402,2054,2075]" box="[151,1402,2054,2075]" pageId="17" pageNumber="145">
Figs 55-58.
<taxonomicName id="4C77E9B8B058FFF1FEA05866D6DBF78B" box="[273,444,2054,2075]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="145" phylum="Arthropoda" rank="species" species="delamarei">
<emphasis id="B9034E29B058FFF1FEA05866D6DBF78B" box="[273,444,2054,2075]" italics="true" pageId="17" pageNumber="145">Achilia delamarei</emphasis>
</taxonomicName>
. Male head in (55) dorsal, (56) lateral, (57) semilateral, and (58) frontal views. Scale bar (200 μm).
</paragraph>
</caption>
<paragraph id="8BC8923BB05BFFF2FF265202D26EFDA9" blockId="18.[151,756,258,697]" lastBlockId="18.[831,1437,258,697]" pageId="18" pageNumber="146">
We have also compared the
<typeStatus id="54CC2C99B05BFFF2FE045202D68AFDE9" box="[437,493,610,633]" pageId="18" pageNumber="146">types</typeStatus>
and other supplementary material of
<taxonomicName id="4C77E9B8B05BFFF2FEA552E2D541FD09" authority="Jeannel, 1962" authorityName="Jeannel" authorityYear="1962" box="[276,550,642,665]" class="Insecta" family="Staphylinidae" genus="Achillia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B05BFFF2FEA552E2D6EDFD09" box="[276,394,642,665]" italics="true" pageId="18" pageNumber="146">A. kuscheli</emphasis>
<bibRefCitation id="EFE6EFCAB05BFFF2FE2352E2D541FD09" author="Jeannel R." box="[402,550,642,665]" pageId="18" pageNumber="146" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962</bibRefCitation>
</taxonomicName>
with the
<typeStatus id="54CC2C99B05BFFF2FD3C52E2D5A2FD09" box="[653,709,642,665]" pageId="18" pageNumber="146">types</typeStatus>
and other supplementary material of
<taxonomicName id="4C77E9B8B05BFFF2FE5E52C2D5EDFD29" baseAuthorityName="Blanchard" baseAuthorityYear="1851" box="[495,650,674,697]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B05BFFF2FE5E52C2D5EDFD29" box="[495,650,674,697]" italics="true" pageId="18" pageNumber="146">A. valdiviensis</emphasis>
</taxonomicName>
. The only difference we could find between these two potential taxa is the morphology of the first two antennomeres (see
<figureCitation id="134C8EBEB05BFFF2FC8E5122D48EFEC9" box="[831,1001,322,345]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="18" pageNumber="146">Figs 18 and 19</figureCitation>
), but for all other characters, including the aedeagus, they are identical. Our opinion is that the different morphology of the first two antennomeres (scape and pedicel), although it has some geographical bearing, pertains to infraspecific variability and, we consequently decided that
<taxonomicName id="4C77E9B8B05BFFF2FBCC5182D413FD89" authority="Jeannel, 1962" authorityName="Jeannel" authorityYear="1962" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="kuscheli">
<emphasis id="B9034E29B05BFFF2FBCC5182D254FE69" box="[1149,1331,482,505]" italics="true" pageId="18" pageNumber="146">Achilia kuscheli</emphasis>
<bibRefCitation id="EFE6EFCAB05BFFF2FAF75182D413FD89" author="Jeannel R." pageId="18" pageNumber="146" pagination="295 - 479" refId="ref22040" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D., E. Rapoport (eds), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962</bibRefCitation>
</taxonomicName>
must be considered a junior synonym of
<taxonomicName id="4C77E9B8B05BFFF2FAFE5262D3E1FDA9" authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="146" phylum="Arthropoda" rank="species" species="valdiviensis">
<emphasis id="B9034E29B05BFFF2FAFE5262D4DAFDA9" italics="true" pageId="18" pageNumber="146">Achilia valdiviensis</emphasis>
(
<bibRefCitation id="EFE6EFCAB05BFFF2FC7A5242D319FDA9" author="Blanchard C. E." box="[971,1150,546,569]" pageId="18" pageNumber="146" pagination="1 - 564" refId="ref21916" refString="Blanchard C. E. 1851. Fauna Chilena, Insectos, Coleopteros. LVI. Pselafianos [pp. 561 - 564]. In: Gay C., Historia fisica y politica de Chile. Zoologia, 5: 1 - 564 + ii, pls 1 - 32." type="journal article" year="1851">Blanchard, 1851</bibRefCitation>
)
</taxonomicName>
(
<emphasis id="B9034E29B05BFFF2FB275242D3DAFDA9" bold="true" box="[1174,1213,546,569]" pageId="18" pageNumber="146">syn</emphasis>
.
<emphasis id="B9034E29B05BFFF2FB785242D395FDA9" bold="true" box="[1225,1266,546,569]" pageId="18" pageNumber="146">nov</emphasis>
.).
</paragraph>
<paragraph id="8BC8923BB05BFFF3FC8E5222D6DBFEED" blockId="18.[831,1437,258,697]" lastBlockId="19.[151,757,258,381]" lastPageId="19" lastPageNumber="147" pageId="18" pageNumber="146">
The males of this species are easily distinguished from their congeners by the peculiar morphology of the head (
<figureCitation id="134C8EBEB05BFFF2FCF652E2D4A5FD09" box="[839,962,642,665]" captionStart="Figs 75-78" captionStartId="22.[151,192,2053,2074]" captionTargetBox="[153,1436,892,2015]" captionTargetId="figure-373@22.[153,1438,867,2015]" captionTargetPageId="22" captionText="Figs 75-78. Achilia valdiviensis. Male head in (75) dorsal, (76) lateral, (77) semilateral, and (78) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743423" httpUri="https://zenodo.org/record/5743423/files/figure.png" pageId="18" pageNumber="146">Figs 75-78</figureCitation>
) and antennae (
<figureCitation id="134C8EBEB05BFFF2FBCC52E2D39CFD09" box="[1149,1275,642,665]" captionStart="Figs 17-21" captionStartId="13.[151,192,1967,1988]" captionTargetBox="[251,1307,833,1928]" captionTargetId="figure-342@13.[218,1374,814,1938]" captionTargetPageId="13" captionText="Figs 17-21. Male antennae (17-18, 20), and variability of the base of antennae (19-21) of Achilia. (17) A. jeanneli n. sp. (18) A. valdiviensis, specimen from Valdivia, Reserva Costera Valdiviana, Chaihuín. (19) A. valdiviensis, specimen from Chiloé, Puente La Caldera. (20) A. bicornis, specimen from Chiloé, Mocopulli. (21) A. bicornis, specimen from Osorno, Bahia Mansa." figureDoi="http://doi.org/10.5281/zenodo.5743401" httpUri="https://zenodo.org/record/5743401/files/figure.png" pageId="18" pageNumber="146">Figs 18-19</figureCitation>
). The females (note here that we could examine only female specimens from Chiloe) are characterized by the subtriangular head with a prolonged clypeus and especially by the misaligned antennal pedicel, which is subrectangular and distinctly longer than wide.
</paragraph>
</subSubSection>
</treatment>
</document>
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