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<document id="AEC2F1ED0060DF5707F578F0E8778E5D" ID-CLB-Dataset="117406" ID-DOI="10.35929/RSZ.0013" ID-GBIF-Dataset="8458d5d1-44ea-4c33-b453-b746ddaa2534" ID-ISSN="0035-418" ID-Zenodo-Dep="5743391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638275126813" checkinUser="carolina" docAuthor="Sabella, Giorgio, Cuccodoro, Giulio &amp; Kurbatov, Sergey A." docDate="2020" docId="03DE232DB041FFE8FEE95160D2FDFBC9" docLanguage="en" docName="RevueSuisseZool.127.1.129-156.pdf" docOrigin="Revue suisse de Zoologie 127 (1)" docSource="http://dx.doi.org/10.35929/rsz.0013" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Achilia franzi Sabella &amp; Cuccodoro &amp; Kurbatov 2020, n. sp." docType="treatment" docVersion="10" lastPageNumber="136" masterDocId="FFE75B55B049FFE0FFB15060D767FF90" masterDocTitle="A revision of the Chilean Brachyglutini - Part 6. Revision of Achilia Reitter, 1890: A. grandiceps, A. valdiviensis, and A. bicornis species groups (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="156" masterPageNumber="129" pageNumber="136" updateTime="1699471439041" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8D18486DB4EDDCB3F92E2BC74B83A75E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="625DE9E1DFA7EBBBF200DA14CED6077F">A revision of the Chilean Brachyglutini - Part 6. Revision of Achilia Reitter, 1890: A. grandiceps, A. valdiviensis, and A. bicornis species groups (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="DF2B2A9418296A6F55AE0C3567F95F8A">Sabella, Giorgio</mods:namePart>
<mods:affiliation id="06320DF73912E49CADB71EA0F6B3EA61">Dipartimento di Scienze Biologiche, Geologiche ed Ambientali dellUniversità - sezione Biologia Animale, via</mods:affiliation>
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<mods:namePart id="05DFC033956B29456710DC26C25EEEAB">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="486B4399C662BBF0FE8BB228ADC60096">Muséum dhistoire naturelle, CP 6434, CH- 1211 Genève 6, Switzerland.</mods:affiliation>
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<mods:namePart id="F0C8FE137F53AEDC56D088A75A48BC01">Kurbatov, Sergey A.</mods:namePart>
<mods:affiliation id="3999356C95EB0E97D47D2F152C48204F">Museum of Entomology, All-Russian Plant Quarantine Center, Pogranichnaya 32, Bykovo 140150, Russia. E-mail:</mods:affiliation>
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<treatment id="03DE232DB041FFE8FEE95160D2FDFBC9" ID-DOI="http://doi.org/10.5281/zenodo.6979925" ID-GBIF-Taxon="198967439" ID-Zenodo-Dep="6979925" LSID="urn:lsid:plazi:treatment:03DE232DB041FFE8FEE95160D2FDFBC9" httpUri="http://treatment.plazi.org/id/03DE232DB041FFE8FEE95160D2FDFBC9" lastPageNumber="136" pageId="8" pageNumber="136">
<subSubSection id="C36DC1B0B041FFE8FEE95160D554FE88" box="[344,563,256,280]" pageId="8" pageNumber="136" type="nomenclature">
<paragraph id="8BC8923BB041FFE8FEE95160D554FE88" blockId="8.[265,642,256,313]" box="[344,563,256,280]" pageId="8" pageNumber="136">
<heading id="D0802557B041FFE8FEE95160D554FE88" bold="true" box="[344,563,256,280]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="136" reason="2">
<emphasis id="B9034E29B041FFE8FEE95160D554FE88" bold="true" box="[344,563,256,280]" pageId="8" pageNumber="136">
<taxonomicName id="4C77E9B8B041FFE8FEE95160D697FE88" ID-CoL="8XSTV" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[344,496,256,280]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="136" phylum="Arthropoda" rank="species" species="franzi" status="sp. nov.">
<emphasis id="B9034E29B041FFE8FEE95160D697FE88" bold="true" box="[344,496,256,280]" italics="true" pageId="8" pageNumber="136">Achilia franzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B041FFE8FE465161D554FE88" box="[503,563,257,280]" pageId="8" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B041FFE8FEB85142D5E5FEA9" box="[265,642,290,313]" pageId="8" pageNumber="136" type="description">
<paragraph id="8BC8923BB041FFE8FEB85142D5E5FEA9" blockId="8.[265,642,256,313]" box="[265,642,290,313]" pageId="8" pageNumber="136">
<figureCitation id="134C8EBEB041FFE8FEB85142D62AFEA9" box="[265,333,290,313]" captionStart="Figs 1-6" captionStartId="10.[151,192,2025,2046]" captionTargetBox="[175,1415,715,1976]" captionTargetId="figure-193@10.[151,1436,686,2025]" captionTargetPageId="10" captionText="Figs 1-6. Aedeagi of Achilia species. (1) A. grandiceps. (2) A. delamarei. (3) A. jeanneli n. sp. (4) A. franzi n. sp. (5) A. elguetai n. sp. (6) A. valdiviensis." figureDoi="http://doi.org/10.5281/zenodo.5743393" httpUri="https://zenodo.org/record/5743393/files/figure.png" pageId="8" pageNumber="136">Figs 4</figureCitation>
,
<figureCitation id="134C8EBEB041FFE8FEE95142D612FEA9" box="[344,373,290,313]" captionStart="Figs 12-16" captionStartId="12.[151,192,2023,2044]" captionTargetBox="[298,1321,898,1988]" captionTargetId="figure-288@12.[237,1392,882,2013]" captionTargetPageId="12" captionText="Figs 12-16. Male antennae of Achilia species. (12) A. grandiceps. (13) A. delamarei. (14) A. franzi n. sp. (15) A. elguetai n. sp. (16) A. denticornis." figureDoi="http://doi.org/10.5281/zenodo.5743397" httpUri="https://zenodo.org/record/5743397/files/figure.png" pageId="8" pageNumber="136">14</figureCitation>
,
<figureCitation id="134C8EBEB041FFE8FE315142D6A2FEA9" box="[384,453,290,313]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="8" pageNumber="136">25, 33</figureCitation>
,
<figureCitation id="134C8EBEB041FFE8FE615142D576FEA9" box="[464,529,290,313]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="8" pageNumber="136">47-48</figureCitation>
,
<figureCitation id="134C8EBEB041FFE8FDAD5142D53BFEA9" box="[540,604,290,313]" captionStart="Figs 63-66" captionStartId="19.[151,192,2053,2074]" captionTargetBox="[151,1435,763,2004]" captionTargetId="figure-199@19.[151,1436,753,2004]" captionTargetPageId="19" captionText="Figs 63-66. Achilia franzi n. sp. Male head in (63) dorsal, (64) lateral, (65) semilateral, and (66) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743417" httpUri="https://zenodo.org/record/5743417/files/figure.png" pageId="8" pageNumber="136">63-66</figureCitation>
,
<figureCitation id="134C8EBEB041FFE8FDD65142D5E5FEA9" box="[615,642,290,313]" captionStart="Fig" captionStartId="25.[151,186,1909,1930]" captionTargetBox="[236,716,257,1880]" captionTargetId="figure-90@25.[236,687,254,1880]" captionTargetPageId="25" captionText="Fig. 83. Distribution map of Achilia grandiceps group. (u green diamonds) A. grandiceps. (red squares) A. delamarei. (p blue triangles) A. denticornis. (« fuchsia stars) A. jeanneli n. sp. (¤ squares edged in fuchsia) A. franzi n. sp. (q red inverted triangles) A. elguetai n. sp." figureDoi="http://doi.org/10.5281/zenodo.5743427" httpUri="https://zenodo.org/record/5743427/files/figure.png" pageId="8" pageNumber="136">83</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B041FFE8FF265131D78DFCEE" pageId="8" pageNumber="136" type="materials_examined">
<paragraph id="8BC8923BB041FFE8FF265131D5BBFD9B" blockId="8.[151,756,337,523]" pageId="8" pageNumber="136">
<materialsCitation id="3B1F9866B041FFE8FF265131D5BFFD9B" ID-GBIF-Occurrence="3886337341" collectedFrom="moss on tree trunks and forest floor" collectingDate="1993-01-05" collectingDateMax="1993-01-06" collectingDateMin="1993-01-05" collectingMethod="sifting" collectionCode="MHNG" collectorName="D. Burckhardt" country="Chile" county="Llanquihue prov." elevation="550" latitude="-41.666668" location="Laguna Triangulo" longLatPrecision="1155" longitude="-72.583336" municipality="Alerce Andino National Park" pageId="8" pageNumber="136" specimenCount="1" specimenCount-male="1" stateProvince="Region Los Lagos" typeStatus="holotype">
<emphasis id="B9034E29B041FFE8FF265131D66DFEF8" bold="true" box="[151,266,337,360]" pageId="8" pageNumber="136">
<typeStatus id="54CC2C99B041FFE8FF265131D663FEF8" box="[151,260,337,360]" pageId="8" pageNumber="136" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED660AFEB041FFE8FEAA5131D60AFEF8" box="[283,365,337,360]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="136" type="Museum">MHNG</collectionCode>
(# MHNG-ENTO-81580);
<specimenCount id="9D7159B2B041FFE8FD085131D58BFEF8" box="[697,748,337,360]" pageId="8" pageNumber="136" type="male">1 ♂</specimenCount>
; SOUTHERN
<collectingCountry id="F360D2ABB041FFE8FE825112D6E1FE19" box="[307,390,370,393]" name="Chile" pageId="8" pageNumber="136" value="Chile">CHILE</collectingCountry>
:
<collectingRegion id="49B35CD9B041FFE8FE245112D50DFE19" box="[405,618,370,393]" country="Chile" name="Los Lagos" pageId="8" pageNumber="136">Región Los Lagos</collectingRegion>
:
<collectingCounty id="62A9EAB7B041FFE8FDC85112D7B5FE39" pageId="8" pageNumber="136">Llanquihue prov.</collectingCounty>
:
<collectingMunicipality id="6BAC0841B041FFE8FF6D51F2D579FE39" box="[220,542,402,425]" pageId="8" pageNumber="136">Alerce Andino National Park</collectingMunicipality>
,
<location id="8EA8C4E0B041FFE8FD9A51F2D597FE39" LSID="urn:lsid:plazi:treatment:03DE232DB041FFE8FEE95160D2FDFBC9:8EA8C4E0B041FFE8FD9A51F2D597FE39" box="[555,752,402,425]" country="Chile" county="Llanquihue prov." latitude="-41.666668" longLatPrecision="1155" longitude="-72.583336" municipality="Alerce Andino National Park" name="Laguna Triangulo" pageId="8" pageNumber="136" stateProvince="Region Los Lagos">Laguna Triángulo</location>
;
<geoCoordinate id="EE43F4FCB041FFE8FF2651D3D79CFE5A" box="[151,251,435,458]" degrees="41" direction="south" minutes="40" orientation="latitude" pageId="8" pageNumber="136" precision="925" value="-41.666668">41° 40S</geoCoordinate>
<geoCoordinate id="EE43F4FCB041FFE8FEB451D3D612FE5A" box="[261,373,435,458]" degrees="72" direction="west" minutes="35" orientation="longitude" pageId="8" pageNumber="136" precision="925" value="-72.583336">72° 35W</geoCoordinate>
;
<quantity id="4C8F3FDEB041FFE8FE3451D3D6B7FE5A" box="[389,464,435,458]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="136" unit="m" value="550.0">
<elevation id="005A7508B041FFE8FE3451D3D6B7FE5A" box="[389,464,435,458]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="136" unit="m" value="550.0">550 m</elevation>
</quantity>
;
<date id="FFC9B4FBB041FFE8FE6E51D3D50AFE5A" box="[479,621,435,458]" pageId="8" pageNumber="136" value="1993-01-05" valueMax="1993-01-06" valueMin="1993-01-05">
<collectingDate id="EF8D4D13B041FFE8FE6E51D3D50AFE5A" box="[479,621,435,458]" pageId="8" pageNumber="136" value="1993-01-05" valueMax="1993-01-06" valueMin="1993-01-05">05-06.I.1993</collectingDate>
</date>
; sclerophyll rainforest, site 38b,
<collectingMethod id="5236EA2CB041FFE8FEC951B3D6DAFE7A" box="[376,445,467,490]" pageId="8" pageNumber="136">sifting</collectingMethod>
of
<collectedFrom id="386BD1C6B041FFE8FE5651B3D669FD9B" pageId="8" pageNumber="136">moss on tree trunks and forest floor</collectedFrom>
, and of vegetational debris;
<collectorName id="2682F7EDB041FFE8FD8D5194D5BFFD9B" box="[572,728,500,523]" pageId="8" pageNumber="136">D. Burckhardt</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC8923BB041FFE8FF265243D78DFCEE" blockId="8.[151,757,547,894]" pageId="8" pageNumber="136">
<materialsCitation id="3B1F9866B041FFE8FF265243D79FFD0B" ID-GBIF-Occurrence="3886337536" collectionCode="MHNG, MHNG-" country="Chile" location="Region Los Lagos" pageId="8" pageNumber="136" specimenCount="187" specimenCount-female="40" specimenCount-male="51" stateProvince="Region Los Lagos" typeStatus="paratype">
<emphasis id="B9034E29B041FFE8FF265243D66CFDAA" bold="true" box="[151,267,547,570]" pageId="8" pageNumber="136">
<typeStatus id="54CC2C99B041FFE8FF265243D66CFDAA" box="[151,267,547,570]" pageId="8" pageNumber="136" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D7159B2B041FFE8FE965243D614FDAA" box="[295,371,547,570]" pageId="8" pageNumber="136" type="generic">
<emphasis id="B9034E29B041FFE8FE965243D60AFDAA" bold="true" box="[295,365,547,570]" pageId="8" pageNumber="136">96 ex</emphasis>
.
</specimenCount>
): SOUTHERN
<collectingCountry id="F360D2ABB041FFE8FD8D5243D5F7FDAA" box="[572,656,547,570]" name="Chile" pageId="8" pageNumber="136" value="Chile">CHILE</collectingCountry>
:
<collectingRegion id="49B35CD9B041FFE8FD175243D67FFDCB" country="Chile" name="Los Lagos" pageId="8" pageNumber="136">Región Los Lagos</collectingRegion>
:
<collectingRegion id="49B35CD9B041FFE8FE9F5224D695FDCB" box="[302,498,580,603]" pageId="8" pageNumber="136">Llanquihue prov</collectingRegion>
.:
<collectionCode id="ED660AFEB041FFE8FDBD5224D539FDCB" box="[524,606,580,603]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="136" type="Museum">MHNG</collectionCode>
(#
<collectionCode id="ED660AFEB041FFE8FD285224D593FDCB" box="[665,756,580,603]" pageId="8" pageNumber="136">MHNG-</collectionCode>
ENTO-81581-672);
<specimenCount id="9D7159B2B041FFE8FECC5204D6DBFDEB" box="[381,444,612,635]" pageId="8" pageNumber="136" type="male">51 ♂</specimenCount>
and
<specimenCount id="9D7159B2B041FFE8FDB25204D525FDEB" box="[515,578,612,635]" pageId="8" pageNumber="136" type="female">40 ♀</specimenCount>
; same data as holotype
</materialsCitation>
.
<materialsCitation id="3B1F9866B041FFE8FEB552E4D5E0FD0B" ID-GBIF-Occurrence="3886337351" box="[260,647,644,667]" collectionCode="MNHS" country="Chile" pageId="8" pageNumber="136" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Region Los Lagos">
<collectionCode id="ED660AFEB041FFE8FEAB52E4D60DFD0B" box="[282,362,644,667]" pageId="8" pageNumber="136">MNHS</collectionCode>
;
<specimenCount id="9D7159B2B041FFE8FEC952E4D6C5FD0B" box="[376,418,644,667]" pageId="8" pageNumber="136" type="male">1 ♂</specimenCount>
and
<specimenCount id="9D7159B2B041FFE8FE6B52E4D563FD0B" box="[474,516,644,667]" pageId="8" pageNumber="136" type="female">1 ♀</specimenCount>
, same data.
</materialsCitation>
<materialsCitation id="3B1F9866B041FFE8FD3E52E4D596FD6D" ID-GBIF-Occurrence="3886337312" collectingDate="1982-12-18" collectingDateMax="1982-12-24" collectingDateMin="1982-12-18" collectingMethod="window trap" collectionCode="UNHC" collectorName="A. Newton &amp; M. Thayer" country="Chile" elevation="690" location="Puyehue National Park" pageId="8" pageNumber="136" specimenCount="1" specimenCount-male="1" stateProvince="Osorno prov.">
<collectingRegion id="49B35CD9B041FFE8FD1452E4D7B5FD2C" pageId="8" pageNumber="136">Osorno prov.</collectingRegion>
:
<collectionCode id="ED660AFEB041FFE8FF6E52C5D649FD2C" box="[223,302,677,700]" pageId="8" pageNumber="136">UNHC</collectionCode>
;
<specimenCount id="9D7159B2B041FFE8FE8D52C5D600FD2C" box="[316,359,677,700]" pageId="8" pageNumber="136" type="male">1 ♂</specimenCount>
;
<location id="8EA8C4E0B041FFE8FEC952C5D51FFD2C" LSID="urn:lsid:plazi:treatment:03DE232DB041FFE8FEE95160D2FDFBC9:8EA8C4E0B041FFE8FEC952C5D51FFD2C" box="[376,632,677,700]" country="Chile" name="Puyehue National Park" pageId="8" pageNumber="136" stateProvince="Osorno prov.">Puyehue National Park</location>
,
<location id="8EA8C4E0B041FFE8FD3452C5D7A8FD4C" LSID="urn:lsid:plazi:treatment:03DE232DB041FFE8FEE95160D2FDFBC9:8EA8C4E0B041FFE8FD3452C5D7A8FD4C" country="Chile" name="Antillanca road;" pageId="8" pageNumber="136" stateProvince="Osorno prov.">Antillanca road;</location>
<quantity id="4C8F3FDEB041FFE8FF5352A5D651FD4C" box="[226,310,709,732]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="8" pageNumber="136" unit="m" value="690.0">
<elevation id="005A7508B041FFE8FF5352A5D651FD4C" box="[226,310,709,732]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="8" pageNumber="136" unit="m" value="690.0">690 m</elevation>
</quantity>
;
<date id="FFC9B4FBB041FFE8FEFF52A5D69CFD4C" box="[334,507,709,732]" pageId="8" pageNumber="136" value="1982-12-18" valueMax="1982-12-24" valueMin="1982-12-18">
<collectingDate id="EF8D4D13B041FFE8FEFF52A5D69CFD4C" box="[334,507,709,732]" pageId="8" pageNumber="136" value="1982-12-18" valueMax="1982-12-24" valueMin="1982-12-18">18-24.XII.1982</collectingDate>
</date>
; site 661,
<location id="8EA8C4E0B041FFE8FD3E52A5D665FD6D" LSID="urn:lsid:plazi:treatment:03DE232DB041FFE8FEE95160D2FDFBC9:8EA8C4E0B041FFE8FD3E52A5D665FD6D" country="Chile" name="valdivian rainforest" pageId="8" pageNumber="136" stateProvince="Osorno prov.">valdivian rainforest</location>
,
<collectingMethod id="5236EA2CB041FFE8FEA75286D6CCFD6D" box="[278,427,742,765]" pageId="8" pageNumber="136">window trap</collectingMethod>
;
<collectorName id="2682F7EDB041FFE8FE0F5286D596FD6D" box="[446,753,742,765]" pageId="8" pageNumber="136">A. Newton &amp; M. Thayer</collectorName>
</materialsCitation>
.
<materialsCitation id="3B1F9866B041FFE8FF265366D6B9FC8D" ID-GBIF-Occurrence="3886337540" box="[151,478,774,797]" collectionCode="FMNH" country="Chile" pageId="8" pageNumber="136" specimenCount="1" specimenCount-male="1" stateProvince="Osorno">
<collectionCode id="ED660AFEB041FFE8FF045366D661FC8D" box="[181,262,774,797]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="136" type="Museum">FMNH</collectionCode>
;
<specimenCount id="9D7159B2B041FFE8FEAA5366D62AFC8D" box="[283,333,774,797]" pageId="8" pageNumber="136" type="male">1 ♂</specimenCount>
; same data
</materialsCitation>
.
<materialsCitation id="3B1F9866B041FFE8FE405366D781FCEE" ID-GBIF-Occurrence="3886337410" collectingDate="1984-12-18" collectingDateMax="1984-12-20" collectingDateMin="1984-12-18" collectingMethod="car netting" collectionCode="MHNG, MHNG-" collectorName="S. &amp; J. Peck" country="Chile" elevation="750" location="Puyehue National Park" pageId="8" pageNumber="136" specimenCount="1" specimenCount-male="1" stateProvince="Puyehue National Park">
<collectionCode id="ED660AFEB041FFE8FDBE5366D506FC8D" box="[527,609,774,797]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="136" type="Museum">MHNG</collectionCode>
(#
<collectionCode id="ED660AFEB041FFE8FD285366D593FC8D" box="[665,756,774,797]" pageId="8" pageNumber="136">MHNG-</collectionCode>
ENTO-81673);
<specimenCount id="9D7159B2B041FFE8FEF35347D60AFCAE" box="[322,365,807,830]" pageId="8" pageNumber="136" type="male">1 ♂</specimenCount>
;
<location id="8EA8C4E0B041FFE8FECC5347D51DFCAE" LSID="urn:lsid:plazi:treatment:03DE232DB041FFE8FEE95160D2FDFBC9:8EA8C4E0B041FFE8FECC5347D51DFCAE" box="[381,634,807,830]" country="Chile" name="Puyehue National Park" pageId="8" pageNumber="136" stateProvince="Puyehue National Park">Puyehue National Park</location>
,
<location id="8EA8C4E0B041FFE8FD345347D7A9FCCE" LSID="urn:lsid:plazi:treatment:03DE232DB041FFE8FEE95160D2FDFBC9:8EA8C4E0B041FFE8FD345347D7A9FCCE" country="Chile" name="Antillanca road;" pageId="8" pageNumber="136" stateProvince="Puyehue National Park">Antillanca road;</location>
<quantity id="4C8F3FDEB041FFE8FF6A5327D601FCCE" box="[219,358,839,862]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="136" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">
<elevation id="005A7508B041FFE8FF6A5327D601FCCE" box="[219,358,839,862]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="8" pageNumber="136" unit="m" value="750.0" valueMax="1000.0" valueMin="500.0">500-1000 m</elevation>
</quantity>
;
<date id="FFC9B4FBB041FFE8FEC75327D546FCCE" box="[374,545,839,862]" pageId="8" pageNumber="136" value="1984-12-18" valueMax="1984-12-20" valueMin="1984-12-18">
<collectingDate id="EF8D4D13B041FFE8FEC75327D546FCCE" box="[374,545,839,862]" pageId="8" pageNumber="136" value="1984-12-18" valueMax="1984-12-20" valueMin="1984-12-18">18-20.XII.1984</collectingDate>
</date>
;
<collectingMethod id="5236EA2CB041FFE8FD835327D5CAFCCE" box="[562,685,839,862]" pageId="8" pageNumber="136">car netting</collectingMethod>
;
<collectorName id="2682F7EDB041FFE8FD0C5327D781FCEE" pageId="8" pageNumber="136">S. &amp; J. Peck</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B041FFE8FF2653F7D3E9FD1A" pageId="8" pageNumber="136" type="description">
<paragraph id="8BC8923BB041FFE8FF2653F7D5E0FBE1" blockId="8.[151,756,919,2077]" pageId="8" pageNumber="136">
<emphasis id="B9034E29B041FFE8FF2653F7D641FC3E" bold="true" box="[151,294,919,942]" pageId="8" pageNumber="136">Description:</emphasis>
Body
<quantity id="4C8F3FDEB041FFE8FECC53F7D57AFC3E" box="[381,541,919,942]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.6" metricValueMin="1.45" pageId="8" pageNumber="136" unit="mm" value="1.525" valueMax="1.6" valueMin="1.45">1.45-1.60 mm</quantity>
long, dark brown with reddish elytra; antennae and legs reddish, palpi yellowish. Pronotum with disc moderately convex; posterior portion of lateral margins subparallel. First abdominal tergite with diverging basal striae extending to about one-fourth of paratergal length, and separated at base by more than one-third of tergal width.
</paragraph>
<paragraph id="8BC8923BB041FFE8FF26541AD35FFEC9" blockId="8.[151,756,919,2077]" lastBlockId="8.[831,1437,258,474]" pageId="8" pageNumber="136">
<emphasis id="B9034E29B041FFE8FF26541AD7A9FB01" box="[151,206,1146,1169]" italics="true" pageId="8" pageNumber="136">Male</emphasis>
: Head as in
<figureCitation id="134C8EBEB041FFE8FEC4541AD694FB01" box="[373,499,1146,1169]" captionStart="Figs 63-66" captionStartId="19.[151,192,2053,2074]" captionTargetBox="[151,1435,763,2004]" captionTargetId="figure-199@19.[151,1436,753,2004]" captionTargetPageId="19" captionText="Figs 63-66. Achilia franzi n. sp. Male head in (63) dorsal, (64) lateral, (65) semilateral, and (66) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743417" httpUri="https://zenodo.org/record/5743417/files/figure.png" pageId="8" pageNumber="136">Figs 63-66</figureCitation>
, with occipital region and basal half of frons raised with V-shaped median notch; lateral arms of notch densely pubescent apically; anterior portion of frons deeply excavated; median apophysis curved and directed backwards, originating from very large median sub-basal clypeal hump, tip in dorsal view fusiform; sub-basal clypeal hump sparsely pubescent. Antennal tubercles very prominent. Antennae (
<figureCitation id="134C8EBEB041FFE8FF2E551DD797FA04" box="[159,240,1405,1428]" captionStart="Figs 12-16" captionStartId="12.[151,192,2023,2044]" captionTargetBox="[298,1321,898,1988]" captionTargetId="figure-288@12.[237,1392,882,2013]" captionTargetPageId="12" captionText="Figs 12-16. Male antennae of Achilia species. (12) A. grandiceps. (13) A. delamarei. (14) A. franzi n. sp. (15) A. elguetai n. sp. (16) A. denticornis." figureDoi="http://doi.org/10.5281/zenodo.5743397" httpUri="https://zenodo.org/record/5743397/files/figure.png" pageId="8" pageNumber="136">Fig. 14</figureCitation>
) with scape and pedicel distinctly longer than wide; antennomeres III and V distinctly longer than wide; antennomeres IV and VI-VIII wider than long; antennomere IX wider than long with protruding mesal margin pointed at distal angle; antennomere X wider than long, with protruding mesal margin; antennomere XI elongate, slightly longer than VII-X combined, with denticulate margins. Metaventrite with distal half pubescent, raised at middle for two thirds of its distal portion, this surface entirely divided by median sulcus. Protrochanters (
<figureCitation id="134C8EBEB041FFE8FE2856A2D68CF949" box="[409,491,1730,1753]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="8" pageNumber="136">Fig. 25</figureCitation>
) bearing one long seta; profemora slightly enlarged at middle and slightly hollowed medially near ventral margin; mesotrochanters (
<figureCitation id="134C8EBEB041FFE8FF2E5743D78BF8AA" box="[159,236,1827,1850]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="8" pageNumber="136">Fig. 33</figureCitation>
) with ventral margin forming spine at basal third; mesofemora (
<figureCitation id="134C8EBEB041FFE8FE9A5724D61DF8CB" box="[299,378,1860,1883]" captionStart="Figs 22-36" captionStartId="14.[151,192,1996,2017]" captionTargetBox="[160,1427,435,1963]" captionTargetId="figure-153@14.[152,1437,416,1963]" captionTargetPageId="14" captionText="Figs 22-36. Male protrochanters (22-28), mesotrochanters and base of mesofemur (29-36) of Achilia. (22, 29) A. grandiceps. (23, 30) A. delamarei. (24, 32) A. jeanneli n. sp. (25, 33) A. franzi n. sp. (26-27, 34) A. elguetai n. sp. (28, 35) A. valdiviensis. (31) A. denticornis. (36) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743405" httpUri="https://zenodo.org/record/5743405/files/figure.png" pageId="8" pageNumber="136">Fig. 33</figureCitation>
) with basal third of ventral margin covered by broad, short and thick setae, very similar to mesofemora of
<taxonomicName id="4C77E9B8B041FFE8FE8C57E4D6D7F80B" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[317,432,1924,1947]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="136" phylum="Arthropoda" rank="species" species="jeanneli" status="sp. nov.">
<emphasis id="B9034E29B041FFE8FE8C57E4D6D7F80B" box="[317,432,1924,1947]" italics="true" pageId="8" pageNumber="136">A. jeanneli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B041FFE8FE0757E4D696F80B" box="[438,497,1924,1947]" pageId="8" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
; mesotibiae with medial margin bearing recurved setae (
<figureCitation id="134C8EBEB041FFE8FE5A57C5D55BF82C" box="[491,572,1957,1980]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="8" pageNumber="136">Fig. 48</figureCitation>
), or not recurved (
<figureCitation id="134C8EBEB041FFE8FF2E57A5D794F84C" box="[159,243,1989,2012]" captionStart="Figs 37-50" captionStartId="15.[151,192,1908,1929]" captionTargetBox="[189,1416,1253,1860]" captionTargetId="figure-589@15.[151,1436,1249,1871]" captionTargetPageId="15" captionText="Figs 37-50. Male protibiae (37-39) and mesotibiae (40-50) of Achilia. (37, 49) A. elguetai n. sp. (38, 44) A. denticornis. (39) A. vadiviensis. (40-41) A. grandiceps, specimen from: (40) Llanquihue,AlerceAndino National Park, Laguna Triàngulo; (41) Malleco, Purén, Contulmo Natural Monument. (42) A. delamarei. (43) A. valdiviensis. (45-46) A. jeanneli n. sp., specimen from: (45) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 31b; (46) Malleco, Nahuelbuta National Park, Piedra del Aquila, station 10a. (47-48) A. franzi n. sp., specimen from: (47) Llanquihue, Alerce Andino National Park, Laguna Triàngulo; (48) Osorno, Antillanca road. (50) A. bicornis." figureDoi="http://doi.org/10.5281/zenodo.5743407" httpUri="https://zenodo.org/record/5743407/files/figure.png" pageId="8" pageNumber="136">Fig. 47</figureCitation>
) and forming very short and rounded apical spine; apical margin denticulate. Aedeagus (
<figureCitation id="134C8EBEB041FFE8FDDF5786D5C9F86D" box="[622,686,2022,2045]" captionStart="Figs 1-6" captionStartId="10.[151,192,2025,2046]" captionTargetBox="[175,1415,715,1976]" captionTargetId="figure-193@10.[151,1436,686,2025]" captionTargetPageId="10" captionText="Figs 1-6. Aedeagi of Achilia species. (1) A. grandiceps. (2) A. delamarei. (3) A. jeanneli n. sp. (4) A. franzi n. sp. (5) A. elguetai n. sp. (6) A. valdiviensis." figureDoi="http://doi.org/10.5281/zenodo.5743393" httpUri="https://zenodo.org/record/5743393/files/figure.png" pageId="8" pageNumber="136">Fig. 4</figureCitation>
)
<quantity id="4C8F3FDEB041FFE8FD0D5786D79EF78D" metricMagnitude="-4" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="8" pageNumber="136" unit="mm" value="0.28" valueMax="0.29" valueMin="0.27">0.27- 0.29 mm</quantity>
long; similar to that of
<taxonomicName id="4C77E9B8B041FFE8FDBD5866D5C3F78D" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[524,676,2054,2077]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="136" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="B9034E29B041FFE8FDBD5866D5C3F78D" box="[524,676,2054,2077]" italics="true" pageId="8" pageNumber="136">A. grandiceps</emphasis>
</taxonomicName>
except parameres wider with larger outer lobe, and copulatory pieces with lateral sclerites wider, longer, pointed and apically bent outwards.
</paragraph>
<paragraph id="8BC8923BB041FFE8FC8E5102D4DFFE4A" blockId="8.[831,1437,258,474]" pageId="8" pageNumber="136">
<emphasis id="B9034E29B041FFE8FC8E5102D4E8FEE9" box="[831,911,354,377]" italics="true" pageId="8" pageNumber="136">Female</emphasis>
: Similar to male except: head not modified, but frons flattened behind frontal sulcus, which is well impressed, anterior margin of the frontal lobe pointed in the middle.
</paragraph>
<paragraph id="8BC8923BB041FFE8FC8E5192D3E9FD1A" blockId="8.[831,1437,498,650]" pageId="8" pageNumber="136">
<emphasis id="B9034E29B041FFE8FC8E5192D49EFD99" bold="true" box="[831,1017,498,521]" pageId="8" pageNumber="136">Collecting data:</emphasis>
Collected from December to January in sclerophyll or valdivian rainforest, at elevations ranging from
<quantity id="4C8F3FDEB041FFE8FC6A5252D343FDD9" box="[987,1060,562,585]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="136" unit="m" value="550.0">550 m</quantity>
to
<quantity id="4C8F3FDEB041FFE8FBFF5252D3C1FDD9" box="[1102,1190,562,585]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="136" unit="m" value="1000.0">1000 m</quantity>
. The specimens came from sifted samples of moss and vegetational debris, windows traps, and car netting.
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B041FFE8FC8E52C2D2EAFD69" pageId="8" pageNumber="136" type="distribution">
<paragraph id="8BC8923BB041FFE8FC8E52C2D2EAFD69" blockId="8.[831,1436,674,761]" pageId="8" pageNumber="136">
<emphasis id="B9034E29B041FFE8FC8E52C2D4B3FD29" bold="true" box="[831,980,674,697]" pageId="8" pageNumber="136">Distribution:</emphasis>
<taxonomicName id="4C77E9B8B041FFE8FC6C52C2D315FD29" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[989,1138,674,697]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="136" phylum="Arthropoda" rank="species" species="franzi" status="sp. nov.">
<emphasis id="B9034E29B041FFE8FC6C52C2D315FD29" box="[989,1138,674,697]" italics="true" pageId="8" pageNumber="136">Achilia franzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B041FFE8FBCD52C2D3DFFD29" box="[1148,1208,674,697]" pageId="8" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
is known only from southern Chile (
<figureCitation id="134C8EBEB041FFE8FC4752A2D32CFD49" box="[1014,1099,706,729]" captionStart="Fig" captionStartId="25.[151,186,1909,1930]" captionTargetBox="[236,716,257,1880]" captionTargetId="figure-90@25.[236,687,254,1880]" captionTargetPageId="25" captionText="Fig. 83. Distribution map of Achilia grandiceps group. (u green diamonds) A. grandiceps. (red squares) A. delamarei. (p blue triangles) A. denticornis. (« fuchsia stars) A. jeanneli n. sp. (¤ squares edged in fuchsia) A. franzi n. sp. (q red inverted triangles) A. elguetai n. sp." figureDoi="http://doi.org/10.5281/zenodo.5743427" httpUri="https://zenodo.org/record/5743427/files/figure.png" pageId="8" pageNumber="136">Fig. 83</figureCitation>
: squares edged in fuchsia) in Llanquihue and Osorno provinces (Región Los Lagos).
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B041FFE8FC8E5372D351FCD9" pageId="8" pageNumber="136" type="etymology">
<paragraph id="8BC8923BB041FFE8FC8E5372D351FCD9" blockId="8.[831,1437,786,841]" pageId="8" pageNumber="136">
<emphasis id="B9034E29B041FFE8FC8E5372D4A3FCB9" bold="true" box="[831,964,786,809]" pageId="8" pageNumber="136">Etymology:</emphasis>
This species is dedicated to the Austrian entomologist H. Franz.
</paragraph>
</subSubSection>
<subSubSection id="C36DC1B0B041FFE8FC8E5301D2FDFBC9" pageId="8" pageNumber="136" type="discussion">
<paragraph id="8BC8923BB041FFE8FC8E5301D2FDFBC9" blockId="8.[831,1437,865,1113]" pageId="8" pageNumber="136">
<emphasis id="B9034E29B041FFE8FC8E5301D4A3FCE8" bold="true" box="[831,964,865,888]" pageId="8" pageNumber="136">Comments:</emphasis>
Within the
<taxonomicName id="4C77E9B8B041FFE8FBF65301D3B9FCE8" authorityName="Jeannel" authorityYear="1962" baseAuthorityName="grandiceps Jeannel" baseAuthorityYear="1962" box="[1095,1246,865,888]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="136" phylum="Arthropoda" rank="species" species="grandiceps">
<emphasis id="B9034E29B041FFE8FBF65301D3B9FCE8" box="[1095,1246,865,888]" italics="true" pageId="8" pageNumber="136">A. grandiceps</emphasis>
</taxonomicName>
group, the males of
<taxonomicName id="4C77E9B8B041FFE8FCEC53E1D4DEFC08" authority="Sabella &amp; Cuccodoro &amp; Kurbatov, 2020" authorityName="Sabella &amp; Cuccodoro &amp; Kurbatov" authorityYear="2020" box="[861,953,897,920]" class="Insecta" family="Staphylinidae" genus="Achilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="136" phylum="Arthropoda" rank="species" species="franzi" status="sp. nov.">
<emphasis id="B9034E29B041FFE8FCEC53E1D4DEFC08" box="[861,953,897,920]" italics="true" pageId="8" pageNumber="136">A. franzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A230F352B041FFE8FC7153E1D49DFC08" box="[960,1018,897,920]" pageId="8" pageNumber="136" rank="species">n. sp.</taxonomicNameLabel>
are easily distinguished from the other species by the peculiar shape of their head (
<figureCitation id="134C8EBEB041FFE8FAAF53C1D2F3FC28" box="[1310,1428,929,952]" captionStart="Figs 63-66" captionStartId="19.[151,192,2053,2074]" captionTargetBox="[151,1435,763,2004]" captionTargetId="figure-199@19.[151,1436,753,2004]" captionTargetPageId="19" captionText="Figs 63-66. Achilia franzi n. sp. Male head in (63) dorsal, (64) lateral, (65) semilateral, and (66) frontal views. Scale bar (200 μm)." figureDoi="http://doi.org/10.5281/zenodo.5743417" httpUri="https://zenodo.org/record/5743417/files/figure.png" pageId="8" pageNumber="136">Figs 63-66</figureCitation>
) and antennae (
<figureCitation id="134C8EBEB041FFE8FC6E53A2D355FC49" box="[991,1074,962,985]" captionStart="Figs 12-16" captionStartId="12.[151,192,2023,2044]" captionTargetBox="[298,1321,898,1988]" captionTargetId="figure-288@12.[237,1392,882,2013]" captionTargetPageId="12" captionText="Figs 12-16. Male antennae of Achilia species. (12) A. grandiceps. (13) A. delamarei. (14) A. franzi n. sp. (15) A. elguetai n. sp. (16) A. denticornis." figureDoi="http://doi.org/10.5281/zenodo.5743397" httpUri="https://zenodo.org/record/5743397/files/figure.png" pageId="8" pageNumber="136">Fig. 14</figureCitation>
). The females of this species are characterized by antennomeres wider than long, or at most as wide as long, the frontal sulcus well-impressed, the anterior margin of frontal lobe distinctly pointed in middle, and the frons flattened behind the frontal sulcus.
</paragraph>
</subSubSection>
</treatment>
</document>