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<document id="AB6D5DFE5FD7539AEE3A0EC08FB390AE" ID-DOI="10.11646/zootaxa.1745.1.3" ID-ISSN="1175­5334" ID-Zenodo-Dep="5108342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626382332205" checkinUser="felipe" docAuthor="Sabu, Thomas K., Abhitha, Prabhakar &amp; Zhao, Danyang" docDate="2008" docId="03B97607FFF9647F47D62C8FFCF2F9C5" docLanguage="en" docName="zt01745p046.pdf" docOrigin="Zootaxa 1745 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Helluodes taprobanae Westwood 1847" docType="treatment" docVersion="4" lastPageNumber="36" masterDocId="FF800E7FFFFF647947412D70FF96FF81" masterDocTitle="A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae)" masterLastPageNumber="46" masterPageNumber="30" pageNumber="36" updateTime="1699011263822" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="00D0DB0FBD125F4DE20A3C73F30FA4E2">
<mods:title id="06C624DF2FD2A71CEA361C49CD037CD7">A new Helluodes Westwood species from Western Ghats with notes on habitat, distribution and phylogenetic relationships of genera Helluodes Westwood and Physocrotaphus Parry of the tribe Physocrotaphini Chaudoir (Insecta: Coleoptera: Carabidae: Anthiinae)</mods:title>
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<mods:namePart id="6F0F2BBD3F094BB9E7C652A52F5BF863">Sabu, Thomas K.</mods:namePart>
<mods:affiliation id="850D22713EB12302BCB22FE165A35E75">Litter Entomology Research Unit, P. G. &amp; Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources &amp; Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="8C51A8BD3A8E84CC8B49BEBF66212268" type="email">sabukthomas@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="08D947B5CC42B794531AFF048AB10AA6">Author</mods:roleTerm>
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<mods:namePart id="164391F11F87CCA98A0C53B14DD342EB">Abhitha, Prabhakar</mods:namePart>
<mods:affiliation id="632ED45A6D16D0D0299C375CCD7D6237">Litter Entomology Research Unit, P. G. &amp; Research Department of Zoology, St. Joseph's College, Devagiri, Calicut, Kerala, 673008, India Department of Entomology, College of Natural Resources &amp; Environment, South China Agricultural University, Wushan, Guangzhou, 510640, China Corresponding author. E-mail: sabukthomas @ gmail. com</mods:affiliation>
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<mods:namePart id="35A0884179680F5C5613C15FAFCD4547">Zhao, Danyang</mods:namePart>
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<mods:typeOfResource id="8FC3631FBC938A609BADC3D5FA8EFC39">text</mods:typeOfResource>
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<mods:title id="D9AFB9D7B3F34EA3416B9A84CA6A6969">Zootaxa</mods:title>
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<mods:date id="540D0FF278052921B7C0022737922E09">2008</mods:date>
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<mods:number id="D48A03B795793C86BD2EE5BB9EC2F9A6">2008-04-09</mods:number>
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<mods:detail id="BB2F08D7018E626E9C977ADE2D8659E3" type="volume">
<mods:number id="0A1F69FD241E51412BC2A8B6CB922E4F">1745</mods:number>
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<mods:url id="67AA65B90875A4ABBEFDC6DB78D54A30">https://biotaxa.org/Zootaxa/article/view/zootaxa.1745.1.3</mods:url>
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<mods:identifier id="3BF5CF31DF47043F053FF764AA788F12" type="DOI">10.11646/zootaxa.1745.1.3</mods:identifier>
<mods:identifier id="E55D573D772D971CFA481C299C180744" type="ISSN">1175­5334</mods:identifier>
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<treatment id="03B97607FFF9647F47D62C8FFCF2F9C5" ID-DOI="http://doi.org/10.5281/zenodo.5108824" ID-Zenodo-Dep="5108824" LSID="urn:lsid:plazi:treatment:03B97607FFF9647F47D62C8FFCF2F9C5" httpUri="http://treatment.plazi.org/id/03B97607FFF9647F47D62C8FFCF2F9C5" lastPageNumber="36" pageId="6" pageNumber="36">
<subSubSection id="C30A949AFFF9647F47D62C8FFDE7FD98" box="[151,625,511,537]" pageId="6" pageNumber="36" type="nomenclature">
<paragraph id="8BAFC711FFF9647F47D62C8FFDE7FD98" blockId="6.[151,625,511,572]" box="[151,625,511,537]" pageId="6" pageNumber="36">
<heading id="D0E7707DFFF9647F47D62C8FFDE7FD98" bold="true" box="[151,625,511,537]" fontSize="11" level="1" pageId="6" pageNumber="36" reason="1">
<taxonomicName id="4C10BC92FFF9647F47D62C8FFDE7FD98" ID-CoL="3KFT5" authority="Westwood, 1847" authorityName="Westwood" authorityYear="1847" box="[151,625,511,537]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="species" species="taprobanae">
<emphasis id="B9641B03FFF9647F47D62C8FFDE7FD98" bold="true" box="[151,625,511,537]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F47D62C8FFE36FD98" bold="true" box="[151,416,511,537]" italics="true" pageId="6" pageNumber="36">Helluodes taprobanae</emphasis>
<bibRefCitation id="EF81BAE0FFF9647F46E92C8FFDE7FD98" author="Westwood, J. O." box="[424,625,511,537]" pageId="6" pageNumber="36" pagination="278 - 280" refId="ref10481" refString="Westwood, J. O. (1847) Descriptions of two new genera of carabideous insects. Transactions of the Entomological Society of London, Series I, vol IV, 278 - 280." type="journal article" year="1847">Westwood, 1847</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A949AFFF9647F47D62F52FEB3FDBD" box="[151,293,546,572]" pageId="6" pageNumber="36" type="description">
<paragraph id="8BAFC711FFF9647F47D62F52FEB3FDBD" blockId="6.[151,625,511,572]" box="[151,293,546,572]" pageId="6" pageNumber="36">
(
<figureCitation id="132BDB94FFF9647F47DE2F52FE8AFDBD" box="[159,284,546,572]" captionStart="FIGURES 6–14" captionStartId="4.[151,269,1435,1459]" captionTargetBox="[175,1403,493,1408]" captionTargetId="figure-287@4.[151,1436,474,1411]" captionTargetPageId="4" captionText="FIGURES 6–14. Labrum, maxillary palpi and pronotum: 6–8 Helluodes devagiriensis n.sp.; 9–11 Helluodes taprobanae; 12–14 Helluodes westwoodi. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.5108350" httpUri="https://zenodo.org/record/5108350/files/figure.png" pageId="6" pageNumber="36">Figs. 9–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A949AFFF9647F47D62F02FEC8FCCD" pageId="6" pageNumber="36" type="reference_group">
<paragraph id="8BAFC711FFF9647F47D62F02FCF4FD08" blockId="6.[151,866,626,649]" box="[151,866,626,649]" pageId="6" pageNumber="36">
<treatmentCitationGroup id="AB00E03FFFF9647F47D62F02FCF4FD08" box="[151,866,626,649]" pageId="6" pageNumber="36">
<treatmentCitation id="0AB1E100FFF9647F47D62F02FDE3FD08" author="Westwood, J. O." box="[151,629,626,649]" page="279" pageId="6" pageNumber="36" year="1847">
<taxonomicName id="4C10BC92FFF9647F47D62F02FDE3FD08" ID-CoL="3KFT5" authority="Westwood, 1847: 279" authorityName="Westwood" authorityPageNumber="279" authorityYear="1847" box="[151,629,626,649]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="species" species="taprobanae">
<emphasis id="B9641B03FFF9647F47D62F02FE12FD08" box="[151,388,626,649]" italics="true" pageId="6" pageNumber="36">Helluodes taprobanae</emphasis>
<bibRefCitation id="EF81BAE0FFF9647F46CB2F02FDE3FD08" author="Westwood, J. O." box="[394,629,626,649]" pageId="6" pageNumber="36" pagination="278 - 280" refId="ref10481" refString="Westwood, J. O. (1847) Descriptions of two new genera of carabideous insects. Transactions of the Entomological Society of London, Series I, vol IV, 278 - 280." type="journal article" year="1847">Westwood, 1847: 279</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AB1E100FFF9647F45C12F02FCC8FD08" author="Chaudoir, M." box="[640,862,626,649]" page="302" pageId="6" pageNumber="36" year="1862">
<bibRefCitation id="EF81BAE0FFF9647F45C12F02FCC8FD08" author="Chaudoir, M." box="[640,862,626,649]" pageId="6" pageNumber="36" pagination="275 - 320" refId="ref9253" refString="Chaudoir, M. (1862) Mat e riaux pour servir l'etude des Cicind eletes et Carabiques 3 - e partie. Extraits du Bulletin de la Societe Imperiale des Naturalist de Moscou, 35 (4), 275 - 320." type="journal article" year="1862">Chaudoir, 1862: 302</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BAFC711FFF9647F47D62FCAFEC8FCCD" blockId="6.[151,1437,698,1604]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F47D62FCAFF77FD55" bold="true" box="[151,225,698,724]" pageId="6" pageNumber="36">Head.</emphasis>
Labrum large, as wide as long, anterior angles rounded, apex between angles slightly emarginate. Maxillary palpomere 4 roundly dilate from middle to apex, inflexed medially and medial surface with distinct, deep puncture, roundly narrowed at apex. Ligula elongate, narrow and tapering towards apex with two elongate apical setae.
</paragraph>
</subSubSection>
<subSubSection id="C30A949AFFF9647F47842E2AFDB2FB5D" pageId="6" pageNumber="36" type="description">
<paragraph id="8BAFC711FFF9647F47842E2AFB37FC6D" blockId="6.[151,1437,698,1604]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F47842E2AFEDDFCF5" bold="true" box="[197,331,858,884]" pageId="6" pageNumber="36">Pronotum.</emphasis>
Apical angles projected and rounded; surface densely punctate and pubescent; anterior and posterior transverse sulci shallow; lateral margin explanate and reflexed; posterior angles weakly reflexed and blunt at apex; basal foveae deep and wide, extended forward as deep lateral furrows; lateral margins weakly sinuate anterior to posterior angles; base distinctly oblique laterally, narrower than apex.
</paragraph>
<paragraph id="8BAFC711FFF9647F47842E8AFE77FBBD" blockId="6.[151,1437,698,1604]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F47842E8AFE8AFB95" bold="true" box="[197,284,1018,1044]" pageId="6" pageNumber="36">Elytra.</emphasis>
Apices obliquely truncate with wide membranous margins; intervals densely pubescent and punctuated; lateral edges narrow.
</paragraph>
<paragraph id="8BAFC711FFF9647F4784293AFC51FBE5" blockId="6.[151,1437,698,1604]" box="[197,967,1098,1124]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F4784293AFEB6FBE5" bold="true" box="[197,288,1098,1124]" pageId="6" pageNumber="36">Venter.</emphasis>
Sternites
<quantity id="4CE86AF4FFF9647F46D2293AFE64FBE5" box="[403,498,1098,1124]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="6" pageNumber="36" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3 to 4 in</quantity>
male with one pair of paramedian setae
</paragraph>
<paragraph id="8BAFC711FFF9647F47842902FADDFB0D" blockId="6.[151,1437,698,1604]" box="[197,1355,1138,1164]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F47842902FD80FB0D" bold="true" box="[197,534,1138,1164]" pageId="6" pageNumber="36">Measurements. Male (n=1)</emphasis>
, TLA = 24.5; TLB = 19.5; TW = 7.0; PL = 3.2; PW = 5.5; EL = 12.0.
</paragraph>
<paragraph id="8BAFC711FFF9647F478429EAFDB2FB5D" blockId="6.[151,1437,698,1604]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F478429EAFD8AFB35" bold="true" box="[197,540,1178,1204]" pageId="6" pageNumber="36">Specimens examined (n=1).</emphasis>
<materialsCitation id="3B78CD4CFFF9647F456529EAFDB2FB5D" collectionCode="MNHN" country="Sri Lanka" county="Nietner" location="Ex-Musaeo Chaudoir" municipality="Taprobanae Westwoodi" pageId="6" pageNumber="36" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54AB79B3FFF9647F456529EAFD0FFB35" box="[548,665,1178,1204]" pageId="6" pageNumber="36" type="lectotype">Lectotype</typeStatus>
male, labeled “
<location id="8ECF91CAFFF9647F441129EAFBC4FB35" LSID="urn:lsid:plazi:treatment:03B97607FFF9647F47D62C8FFCF2F9C5:8ECF91CAFFF9647F441129EAFBC4FB35" box="[848,1106,1178,1204]" country="Sri Lanka" county="Nietner" municipality="Taprobanae Westwoodi" name="Ex-Musaeo Chaudoir" pageId="6" pageNumber="36">Ex-Musaeo Chaudoir</location>
” / “
<collectingMunicipality id="6BCB5D6BFFF9647F433F29EAFA0EFB35" box="[1150,1432,1178,1204]" pageId="6" pageNumber="36">Taprobanae Westwoodi</collectingMunicipality>
,
<collectingCountry id="F3078781FFF9647F47D629B2FF7BFB5D" box="[151,237,1218,1244]" name="Sri Lanka" pageId="6" pageNumber="36">Ceylan</collectingCountry>
,
<collectingCounty id="62CEBF9DFFF9647F47B929B2FEC5FB5D" box="[248,339,1218,1244]" pageId="6" pageNumber="36">Nietner</collectingCounty>
” at
<collectionCode id="ED015FD4FFF9647F46C029B2FE4DFB5D" box="[385,475,1218,1244]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="36">MNHN</collectionCode>
Paris.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A949AFFF9647F4784299AFE1AFAAD" pageId="6" pageNumber="36" type="distribution">
<paragraph id="8BAFC711FFF9647F4784299AFE1AFAAD" blockId="6.[151,1437,698,1604]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F4784299AFEF0FA85" bold="true" box="[197,358,1258,1284]" pageId="6" pageNumber="36">Distribution.</emphasis>
<collectingCountry id="F3078781FFF9647F4631299AFE73FA85" box="[368,485,1258,1284]" name="Sri Lanka" pageId="6" pageNumber="36">Sri Lanka</collectingCountry>
(Kandy, Kitugalle, Kotte, Matale, Galgamuwa, Peradeniya-Sri Lanka Lowland rainforest ecoregion)
</paragraph>
</subSubSection>
<subSubSection id="C30A949AFFF9647F4784284AFCF2F9C5" pageId="6" pageNumber="36" type="discussion">
<paragraph id="8BAFC711FFF9647F4784284AFCF2F9C5" blockId="6.[151,1437,698,1604]" pageId="6" pageNumber="36">
<emphasis id="B9641B03FFF9647F4784284AFEA8FAD5" bold="true" box="[197,318,1338,1364]" pageId="6" pageNumber="36">Remarks.</emphasis>
With slightly emarginate margin between anterior angles of labrum, rounded anterior angles of pronotum and slender ligula,
<taxonomicName id="4C10BC92FFF9647F46B12813FD0CFAFD" box="[496,666,1379,1404]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="species" species="taprobanae">
<emphasis id="B9641B03FFF9647F46B12813FD0CFAFD" box="[496,666,1379,1404]" italics="true" pageId="6" pageNumber="36">H. taprobanae</emphasis>
</taxonomicName>
is more similar to
<taxonomicName id="4C10BC92FFF9647F44362813FBAEFAFD" authority="Sabu &amp; Abhitha &amp; Zhao, 2008" authorityName="Sabu &amp; Abhitha &amp; Zhao" authorityYear="2008" box="[887,1080,1379,1404]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="species" species="devagiriensis" status="sp. nov.">
<emphasis id="B9641B03FFF9647F44362813FBAEFAFD" box="[887,1080,1379,1404]" italics="true" pageId="6" pageNumber="36">H. devagiriensis</emphasis>
</taxonomicName>
<emphasis id="B9641B03FFF9647F437E2812FBECFAFD" bold="true" box="[1087,1146,1378,1404]" pageId="6" pageNumber="36">
<taxonomicNameLabel id="A257A678FFF9647F437E2812FBECFAFD" box="[1087,1146,1378,1404]" pageId="6" pageNumber="36" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
, but differs from the latter in the absence of dentation on the anterior margin of labrum, maxillary palpomere 4 roundly dilated from middle to apex and roundly narrowed at apex, and in more densely punctuate pronotal disc. Although females of
<taxonomicName id="4C10BC92FFF9647F47F928ABFEF2FA75" box="[184,356,1499,1524]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="species" species="taprobanae">
<emphasis id="B9641B03FFF9647F47F928ABFEF2FA75" box="[184,356,1499,1524]" italics="true" pageId="6" pageNumber="36">H. taprobanae</emphasis>
</taxonomicName>
are not known, we predict that the venter of the female will not have paramedial setae on segments 3 and 4 and that this will be a sexually dimorphic character similar to that found in all species of the genus
<taxonomicName id="4C10BC92FFF9647F47A22B5BFECCF9C5" box="[227,346,1579,1604]" class="Insecta" family="Carabidae" genus="Helluodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B9641B03FFF9647F47A22B5BFECCF9C5" box="[227,346,1579,1604]" italics="true" pageId="6" pageNumber="36">Helluodes</emphasis>
</taxonomicName>
and tribe
<taxonomicName id="4C10BC92FFF9647F468E2B5AFD1BF9C5" authorityName="Chaudoir" authorityYear="1863" box="[463,653,1578,1604]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="36" phylum="Arthropoda" rank="tribe" tribe="Physocrotaphini">Physocrotaphini</taxonomicName>
examined to date.
</paragraph>
</subSubSection>
</treatment>
</document>
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