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<document id="559893A716333FF05262469D217A73B2" ID-DOI="10.26107/RBZ-2022-0018" ID-ISSN="2345-7600" ID-Zenodo-Dep="7174164" ID-ZooBank="E3A5AA23-BEAE-445E-981F-93A9E7A79417" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665290411274" checkinUser="felipe" docAuthor="Sang, Do Duc &amp; Son, Nguyen Thanh" docDate="2022" docId="03A68780FFC5FFA98F0C06F17B0F14B9" docLanguage="en" docName="RafflesBZool.70.364–375.pdf" docOrigin="Raffles Bulletin of Zoology 70" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Pincerna costulosus Bavay &amp; Dautzenberg 1912" docType="treatment" docVersion="3" lastPageNumber="369" masterDocId="FF9FFFF8FFC4FFAC8C6000767808166D" masterDocTitle="The land snail genus Pincerna Preston, 1907 (Gastropoda: Alycaeidae) from Vietnam and Laos, with description of a new species" masterLastPageNumber="375" masterPageNumber="364" pageNumber="365" updateTime="1699379731378" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="43782BE71AF963FE3F820591ED7B362E">The land snail genus Pincerna Preston, 1907 (Gastropoda: Alycaeidae) from Vietnam and Laos, with description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="38CC0CCD5B720D2AFFAE218E06E7071F" type="personal">
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<mods:namePart id="DA570152D9278A5BA647355F64FE91EC">Sang, Do Duc</mods:namePart>
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<mods:namePart id="20E86C9BE370FC8E0584B395FB2AB0C6">Son, Nguyen Thanh</mods:namePart>
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<mods:title id="C5A3F4B54C126328CAC5777B09D702C3">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="11358DA7C7EB22B6DE89A9C5AB109D60">
<mods:date id="E7AC5BACD496D4C56D717B86D34EEB94">2022</mods:date>
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<mods:number id="E0F4A5D696D22E02BF50A6356EDA0C36">2022-06-01</mods:number>
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<mods:detail id="7CF97D9DEC35C11F059C989C030729C5" type="volume">
<mods:number id="0BE956EFE8EBBFF4E2690E1C3440E34D">70</mods:number>
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<mods:identifier id="A15471A351840AE590BF345921CA33C6" type="DOI">10.26107/RBZ-2022-0018</mods:identifier>
<mods:identifier id="E34C35ECDED3AB6057620382774FC6F1" type="ISSN">2345-7600</mods:identifier>
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</mods:mods>
<treatment id="03A68780FFC5FFA98F0C06F17B0F14B9" LSID="urn:lsid:plazi:treatment:03A68780FFC5FFA98F0C06F17B0F14B9" httpUri="http://treatment.plazi.org/id/03A68780FFC5FFA98F0C06F17B0F14B9" lastPageId="5" lastPageNumber="369" pageId="1" pageNumber="365">
<subSubSection id="C315651DFFC5FFAD8F0C06F17D9110F2" box="[876,1433,1670,1695]" pageId="1" pageNumber="365" type="nomenclature">
<paragraph id="8BB03696FFC5FFAD8F0C06F17D9110F2" blockId="1.[876,1433,1670,1727]" box="[876,1433,1670,1695]" pageId="1" pageNumber="365">
<heading id="D0F881FAFFC5FFAD8F0C06F17D9110F2" bold="true" box="[876,1433,1670,1695]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="365" reason="2">
<taxonomicName id="4C0F4D15FFC5FFAD8F0C06F17D9110F2" ID-CoL="7PYC5" authority="Bavay &amp; Dautzenberg, 1912" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[876,1433,1670,1695]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="365" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFC5FFAD8F0C06F17D9110F2" bold="true" box="[876,1433,1670,1695]" pageId="1" pageNumber="365">
<emphasis id="B97BEA84FFC5FFAD8F0C06F17C4310F3" bold="true" box="[876,1099,1670,1695]" italics="true" pageId="1" pageNumber="365">Pincerna costulosus</emphasis>
<bibRefCitation id="EF9E4B67FFC5FFAD883406F17D9110F2" author="Bavay A &amp; Dautzenberg P" box="[1108,1433,1671,1695]" pageId="1" pageNumber="365" pagination="1 - 54" refId="ref5945" refString="Bavay A &amp; Dautzenberg P (1912) Description de Coquilles Nouvelles de l'Indo-Chine. Journal de Conchyliologie, 60: 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC5FFAD8FB106D17D3D10D2" box="[977,1333,1703,1727]" pageId="1" pageNumber="365" type="description">
<paragraph id="8BB03696FFC5FFAD8FB106D17D3D10D2" blockId="1.[876,1433,1670,1727]" box="[977,1333,1703,1727]" pageId="1" pageNumber="365">
(
<figureCitation id="13342A13FFC5FFAD8FB906D17C2C10D2" box="[985,1060,1703,1727]" captionStart="Fig" captionStartId="2.[113,148,1941,1963]" captionTargetBox="[113,1452,311,1926]" captionTargetId="figure-11@2.[113,1452,311,1926]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Pincerna species in Vietnam and Laos. Triangle: Pincerna anceyi; Grey circle: Pincerna costulosus; Black circles: Pincerna mouhoti; Squares: Pincerna vanbuensis; Star: Pincerna clausa, new species. The localities are detailed in Table 1." figureDoi="http://doi.org/10.5281/zenodo.7174166" httpUri="https://zenodo.org/record/7174166/files/figure.png" pageId="1" pageNumber="365">Figs. 1</figureCitation>
,
<figureCitation id="13342A13FFC5FFAD885306D17C7E10D3" box="[1075,1142,1703,1727]" captionStart="Fig" captionStartId="3.[136,171,2006,2028]" captionTargetBox="[168,1442,189,1991]" captionTargetId="figure-8@3.[168,1442,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Shells (apertural view, dorsal view, ventral view, and side view) of Pincerna species. A, Pincerna anceyi (Mabille, 1887), syntype MNHN-IM-2000-31797; B, C, Pincerna mouhoti (L. Pfeiffer, 1862): B, specimen from MƯờng La, SƠn La, Vietnam, ZMHU; C, syntype NHMUK 20170120; D, E, Pincerna costulosus Bavay &amp; Dautzenberg, 1912: D, syntype MNHN-IM-2000-31786; E, specimen from Phong Thổ, Lai Châu, Vietnam, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174168" httpUri="https://zenodo.org/record/7174168/files/figure.png" pageId="1" pageNumber="365">2D, E</figureCitation>
,
<figureCitation id="13342A13FFC5FFAD88E306D17CAD10D2" box="[1155,1189,1703,1727]" captionStart="Fig" captionStartId="8.[113,149,1150,1172]" captionTargetBox="[113,1451,188,1135]" captionTargetId="figure-34@8.[113,1451,188,1135]" captionTargetPageId="8" captionText="Fig. 6. Synoptic view of Vietnamese and Laos Pincerna species. A, MNHN-IM-2000-31797; B, NHMUK 20170120; C, MNHN- IM-2000-31786; D, VNMN-IZ 002.304; E, MNHN-IM-2000-31798." figureDoi="http://doi.org/10.5281/zenodo.7174176" httpUri="https://zenodo.org/record/7174176/files/figure.png" pageId="1" pageNumber="365">6C</figureCitation>
;
<tableCitation id="C68D032DFFC5FFAD88D206D17D0710D2" box="[1202,1295,1703,1727]" captionStart="Table 1" captionStartId="9.[136,191,189,211]" captionText="Table 1. Shell size variation of Pincerna spp. specimen collections in this study" pageId="1" pageNumber="365">Tables 1</tableCitation>
,
<tableCitation id="C68D032DFFC5FFAD897F06D17D2510D2" box="[1311,1325,1703,1727]" captionStart="Table 2" captionStartId="9.[136,191,1135,1157]" captionText="Table 2. Comparison of diagnostic conchological characters among Pincerna species in this study." pageId="1" pageNumber="365">2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC5FFAD8F2306937CF911BF" pageId="1" pageNumber="365" type="reference_group">
<paragraph id="8BB03696FFC5FFAD8F2306937D601175" blockId="1.[835,1475,1765,1874]" pageId="1" pageNumber="365">
<treatmentCitationGroup id="AB1F11B8FFC5FFAD8F2306937D601175" pageId="1" pageNumber="365">
<treatmentCitation id="0AAE1087FFC5FFAD8F2306937D5A1096" author="Bavay A &amp; Dautzenberg P" box="[835,1362,1765,1787]" page="49" pageId="1" pageNumber="365" year="1912">
<taxonomicName id="4C0F4D15FFC5FFAD8F2306937D5A1096" ID-CoL="7PYC5" authority="Bavay &amp; Dautzenberg, 1912: 49" authorityName="Bavay &amp; Dautzenberg" authorityPageNumber="49" authorityYear="1912" box="[835,1362,1765,1787]" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="365" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFC5FFAD8F2306937C0E1097" box="[835,1030,1765,1786]" italics="true" pageId="1" pageNumber="365">Alycaeus costulosus</emphasis>
<materialsCitation id="3B673CCBFFC5FFAD886F06907D5A1096" box="[1039,1362,1765,1787]" country="Vietnam" location="Phong Th" pageId="1" pageNumber="365" specimenCount="1" stateProvince="Lai Chau">
<bibRefCitation id="EF9E4B67FFC5FFAD886F06907D5A1096" author="Bavay A &amp; Dautzenberg P" box="[1039,1362,1765,1787]" pageId="1" pageNumber="365" pagination="1 - 54" refId="ref5945" refString="Bavay A &amp; Dautzenberg P (1912) Description de Coquilles Nouvelles de l'Indo-Chine. Journal de Conchyliologie, 60: 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912: 49</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, pl. 4, figs. 1–4 (
<typeStatus id="54B48834FFC5FFAD8FFB07747BCE117A" box="[923,966,1794,1815]" pageId="1" pageNumber="365">type</typeStatus>
locality:
<location id="8ED0604DFFC5FFAD884407747C8F1177" LSID="urn:lsid:plazi:treatment:03A68780FFC5FFA98F0C06F17B0F14B9:8ED0604DFFC5FFAD884407747C8F1177" box="[1060,1159,1794,1818]" country="Vietnam" name="Phong Th" pageId="1" pageNumber="365" stateProvince="Lai Chau">Phong Th</location>
ổ,
<collectingRegion id="49CBF874FFC5FFAD88FE07747CF31175" box="[1182,1275,1794,1816]" country="Vietnam" name="Lai Chau" pageId="1" pageNumber="365">Lai Châu</collectingRegion>
,
<collectingCountry id="F3187606FFC5FFAD896607747D551175" box="[1286,1373,1794,1816]" name="Vietnam" pageId="1" pageNumber="365">Vietnam</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB03696FFC5FFAD8F2307567D19113C" blockId="1.[835,1475,1765,1874]" pageId="1" pageNumber="365">
<treatmentCitationGroup id="AB1F11B8FFC5FFAD8F2307567D19113C" pageId="1" pageNumber="365">
<taxonomicName id="4C0F4D15FFC5FFAD8F2307567BF61159" ID-CoL="7SQ3L" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" box="[835,1022,1823,1845]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="1" pageNumber="365" phylum="Mollusca" rank="species" species="costulosa">
<emphasis id="B97BEA84FFC5FFAD8F2307567BF61159" box="[835,1022,1823,1845]" italics="true" pageId="1" pageNumber="365">Pincerna costulosa</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AAE1087FFC5FFAD887907697D391159" author="Pall-Gergely B &amp; Hunyadi A &amp; Do SD &amp; Naggs F &amp; Asami T" box="[1049,1329,1823,1845]" page="10" pageId="1" pageNumber="365" year="2017">
<bibRefCitation id="EF9E4B67FFC5FFAD887907697D391159" author="Pall-Gergely B &amp; Hunyadi A &amp; Do SD &amp; Naggs F &amp; Asami T" box="[1049,1329,1823,1845]" pageId="1" pageNumber="365" pagination="1 - 124" refId="ref6902" refString="Pall-Gergely B, Hunyadi A, Do SD, Naggs F &amp; Asami T (2017) Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus. Zootaxa, 4331: 1 - 124." type="journal article" year="2017">Páll-Gergely et al., 2017: 10</bibRefCitation>
</treatmentCitation>
, fig. 3F; 2020: 172;
<treatmentCitation id="0AAE1087FFC5FFAD8FFC074A7C71113F" author="Chen ZY &amp; Wu M" box="[924,1145,1852,1874]" page="45" pageId="1" pageNumber="365" year="2020">
<bibRefCitation id="EF9E4B67FFC5FFAD8FFC074A7C71113F" author="Chen ZY &amp; Wu M" box="[924,1145,1852,1874]" pageId="1" pageNumber="365" pagination="37 - 46" refId="ref5993" refString="Chen ZY &amp; Wu M (2020) Two new cyclophoroid species from Hubei, China, with proposal of a new genus (Gastropoda, Caenogastropoda, Diplommatinidae and Alycaeidae). ZooKeys, 935: 37 - 46." type="journal article" year="2020">Chen &amp; Wu, 2020: 45</bibRefCitation>
</treatmentCitation>
, figs. 1, 4C, D.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB03696FFC5FFAD8F23070C7CF911BF" blockId="1.[835,1475,1914,2003]" pageId="1" pageNumber="365">
<emphasis id="B97BEA84FFC5FFAD8F23070C7C1B11FF" bold="true" box="[835,1043,1914,1938]" pageId="1" pageNumber="365">Types examined.</emphasis>
<materialsCitation id="3B673CCBFFC5FFAD887F070C7CE611BF" collectionCode="RBINS" collectorName="Messager &amp; Messager, No." country="Vietnam" location="Tonkin" pageId="1" pageNumber="365" specimenCode="RBINS 525426" specimenCount="1" typeStatus="syntype">
Phong Tho,
<collectingCountry id="F3187606FFC5FFAD88D3070C7D0011FF" box="[1203,1288,1914,1938]" name="Vietnam" pageId="1" pageNumber="365">Tonkin</collectingCountry>
, col.
<collectorName id="26FA5340FFC5FFAD892E070D7DB511FE" box="[1358,1469,1915,1939]" pageId="1" pageNumber="365">Messager</collectorName>
, MNHN-IM-2000-31786 (
<typeStatus id="54B48834FFC5FFAD883A07EC7CB211DF" box="[1114,1210,1946,1970]" pageId="1" pageNumber="365" type="syntype">syntypes</typeStatus>
) (
<figureCitation id="13342A13FFC5FFAD88B407EC7D2611DF" box="[1236,1326,1946,1970]" captionStart="Fig" captionStartId="3.[136,171,2006,2028]" captionTargetBox="[168,1442,189,1991]" captionTargetId="figure-8@3.[168,1442,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Shells (apertural view, dorsal view, ventral view, and side view) of Pincerna species. A, Pincerna anceyi (Mabille, 1887), syntype MNHN-IM-2000-31797; B, C, Pincerna mouhoti (L. Pfeiffer, 1862): B, specimen from MƯờng La, SƠn La, Vietnam, ZMHU; C, syntype NHMUK 20170120; D, E, Pincerna costulosus Bavay &amp; Dautzenberg, 1912: D, syntype MNHN-IM-2000-31786; E, specimen from Phong Thổ, Lai Châu, Vietnam, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174168" httpUri="https://zenodo.org/record/7174168/files/figure.png" pageId="1" pageNumber="365">Fig. 2D</figureCitation>
); Phong-Tho, col.
<collectorName id="26FA5340FFC5FFAD8F1307CD7C1911BF" box="[883,1041,1979,2003]" pageId="1" pageNumber="365">Messager, No.</collectorName>
29,
<specimenCode id="DBA99EEDFFC5FFAD882307CC7CE611BF" box="[1091,1262,1978,2002]" collectionCode="RBINS" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="1" pageNumber="365" type="Museum">RBINS 525426</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC5FFA98F23078C78F21779" lastPageId="5" lastPageNumber="369" pageId="1" pageNumber="365" type="materials_examined">
<paragraph id="8BB03696FFC5FFAD8F23078C7DCB1E5F" blockId="1.[835,1475,2042,2101]" pageId="1" pageNumber="365">
<emphasis id="B97BEA84FFC5FFAD8F23078C7CCB1E7F" bold="true" box="[835,1219,2042,2066]" pageId="1" pageNumber="365">Additional material examined.</emphasis>
<materialsCitation id="3B673CCBFFC5FFAD88AF078C7DCB1E5F" collectionCode="ZMHU" country="Vietnam" county="Phong Th" location="Kh" municipality="District" pageId="1" pageNumber="365" specimenCode="ZMHU/ 4" specimenCount="1" stateProvince="Lai Chau">
<specimenCode id="DBA99EEDFFC5FFAD88AF078C7D4A1E7F" box="[1231,1346,2042,2066]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="1" pageNumber="365">ZMHU/ 4</specimenCode>
(
<figureCitation id="13342A13FFC5FFAD8937078C7DBB1E7F" box="[1367,1459,2042,2066]" captionStart="Fig" captionStartId="3.[136,171,2006,2028]" captionTargetBox="[168,1442,189,1991]" captionTargetId="figure-8@3.[168,1442,189,1991]" captionTargetPageId="3" captionText="Fig. 2. Shells (apertural view, dorsal view, ventral view, and side view) of Pincerna species. A, Pincerna anceyi (Mabille, 1887), syntype MNHN-IM-2000-31797; B, C, Pincerna mouhoti (L. Pfeiffer, 1862): B, specimen from MƯờng La, SƠn La, Vietnam, ZMHU; C, syntype NHMUK 20170120; D, E, Pincerna costulosus Bavay &amp; Dautzenberg, 1912: D, syntype MNHN-IM-2000-31786; E, specimen from Phong Thổ, Lai Châu, Vietnam, ZMHU." figureDoi="http://doi.org/10.5281/zenodo.7174168" httpUri="https://zenodo.org/record/7174168/files/figure.png" pageId="1" pageNumber="365">Fig. 2E</figureCitation>
),
<collectingCountry id="F3187606FFC5FFAD8F24086C7BAA1E5F" box="[836,930,2074,2098]" name="Vietnam" pageId="1" pageNumber="365">Vietnam</collectingCountry>
,
<collectingRegion id="49CBF874FFC5FFAD8FCB086C7C791E5F" box="[939,1137,2074,2098]" country="Vietnam" name="Lai Chau" pageId="1" pageNumber="365">Lai Châu Province</collectingRegion>
,
<collectingCounty id="62D14E1AFFC5FFAD881A086C7CE81E58" box="[1146,1248,2074,2101]" pageId="1" pageNumber="365">Phong Th</collectingCounty>
ổ
<collectingMunicipality id="6BD4ACECFFC5FFAD8892086C7D4C1E5F" box="[1266,1348,2074,2098]" pageId="1" pageNumber="365">District</collectingMunicipality>
,
<location id="8ED0604DFFC5FFAD892D086D7D661E58" LSID="urn:lsid:plazi:treatment:03A68780FFC5FFA98F0C06F17B0F14B9:8ED0604DFFC5FFAD892D086D7D661E58" box="[1357,1390,2075,2101]" country="Vietnam" county="Phong Th" municipality="District" name="Kh" pageId="1" pageNumber="365" stateProvince="Lai Chau">Kh</location>
ổng
<location id="8ED0604DFFC5FFAD89FA086C7DCB1E5F" LSID="urn:lsid:plazi:treatment:03A68780FFC5FFA98F0C06F17B0F14B9:8ED0604DFFC5FFAD89FA086C7DCB1E5F" box="[1434,1475,2074,2098]" country="Vietnam" county="Phong Th" municipality="District" name="Lao" pageId="1" pageNumber="365" stateProvince="Lai Chau">Lào</location>
</materialsCitation>
</paragraph>
<caption id="DF70661EFFC6FFAE8C1107E37D3C11AA" ID-DOI="http://doi.org/10.5281/zenodo.7174166" ID-Zenodo-Dep="7174166" httpUri="https://zenodo.org/record/7174166/files/figure.png" pageId="2" pageNumber="366" startId="2.[113,148,1941,1963]" targetBox="[113,1452,311,1926]" targetPageId="2">
<paragraph id="8BB03696FFC6FFAE8C1107E37D3C11AA" blockId="2.[113,1453,1941,1992]" pageId="2" pageNumber="366">
Fig. 1. Distribution of
<taxonomicName id="4C0F4D15FFC6FFAE8D2D07E079F911C6" box="[333,497,1941,1963]" pageId="2" pageNumber="366">
<emphasis id="B97BEA84FFC6FFAE8D2D07E079AD11C6" box="[333,421,1942,1963]" italics="true" pageId="2" pageNumber="366">Pincerna</emphasis>
species
</taxonomicName>
in Vietnam and Laos. Triangle:
<taxonomicName id="4C0F4D15FFC6FFAE8F5107E07BC711C6" baseAuthorityName="Mabille" baseAuthorityYear="1887" box="[817,975,1942,1963]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="366" phylum="Mollusca" rank="species" species="anceyi">
<emphasis id="B97BEA84FFC6FFAE8F5107E07BC711C6" box="[817,975,1942,1963]" italics="true" pageId="2" pageNumber="366">Pincerna anceyi</emphasis>
</taxonomicName>
; Grey circle:
<taxonomicName id="4C0F4D15FFC6FFAE883607E07D1111C7" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[1110,1305,1941,1963]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="366" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFC6FFAE883607E07D1111C7" box="[1110,1305,1941,1963]" italics="true" pageId="2" pageNumber="366">Pincerna costulosus</emphasis>
</taxonomicName>
; Black circles:
<taxonomicName id="4C0F4D15FFC6FFAE8C1107C5791711AA" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[113,287,1970,1992]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="366" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC6FFAE8C1107C5791711AA" box="[113,287,1970,1992]" italics="true" pageId="2" pageNumber="366">Pincerna mouhoti</emphasis>
</taxonomicName>
; Squares:
<taxonomicName id="4C0F4D15FFC6FFAE8DE807C57A5B11AA" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[392,595,1970,1992]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="366" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFC6FFAE8DE807C57A5B11AA" box="[392,595,1970,1992]" italics="true" pageId="2" pageNumber="366">Pincerna vanbuensis</emphasis>
</taxonomicName>
; Star:
<taxonomicName id="4C0F4D15FFC6FFAE8EF607C57B3D11AA" authority="Sang &amp; Son, 2022" authorityName="Sang &amp; Son" authorityYear="2022" box="[662,821,1970,1992]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="2" pageNumber="366" phylum="Mollusca" rank="species" species="clausa" status="sp. nov.">
<emphasis id="B97BEA84FFC6FFAE8EF607C57B3D11AA" box="[662,821,1970,1992]" italics="true" pageId="2" pageNumber="366">Pincerna clausa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24857FFFFC6FFAE8F2207C57BB211A5" box="[834,954,1970,1992]" pageId="2" pageNumber="366" rank="species">new species</taxonomicNameLabel>
. The localities are detailed in Table 1.
</paragraph>
</caption>
<caption id="DF70661EFFC7FFAF8CE807A07A101E2F" ID-DOI="http://doi.org/10.5281/zenodo.7174168" ID-Zenodo-Dep="7174168" httpUri="https://zenodo.org/record/7174168/files/figure.png" pageId="3" pageNumber="367" startId="3.[136,171,2006,2028]" targetBox="[168,1442,189,1991]" targetPageId="3">
<paragraph id="8BB03696FFC7FFAF8CE807A07A101E2F" blockId="3.[136,1474,2006,2116]" pageId="3" pageNumber="367">
Fig. 2. Shells (apertural view, dorsal view, ventral view, and side view) of
<taxonomicName id="4C0F4D15FFC7FFAF8F0007A17C2C1186" box="[864,1060,2006,2028]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="367" phylum="Mollusca" rank="species" species="undefined-a">
<emphasis id="B97BEA84FFC7FFAF8F0007A17BB01181" box="[864,952,2007,2028]" italics="true" pageId="3" pageNumber="367">Pincerna</emphasis>
species. A
</taxonomicName>
,
<taxonomicName id="4C0F4D15FFC7FFAF884F07A17D651186" authority="(Mabille, 1887)" baseAuthorityName="Mabille" baseAuthorityYear="1887" box="[1071,1389,2006,2028]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="367" phylum="Mollusca" rank="species" species="anceyi">
<emphasis id="B97BEA84FFC7FFAF884F07A17CC51181" box="[1071,1229,2007,2028]" italics="true" pageId="3" pageNumber="367">Pincerna anceyi</emphasis>
(
<bibRefCitation id="EF9E4B67FFC7FFAF88BA07A07D6E1186" author="Mabille J" box="[1242,1382,2006,2028]" pageId="3" pageNumber="367" pagination="73 - 164" refId="ref6829" refString="Mabille J (1887) On some molluscs from Northern Vietnam. Bulletin of the Malacological Society of France, 4: 73 - 164." type="journal article" year="1887">Mabille, 1887</bibRefCitation>
)
</taxonomicName>
, syntype MNHN-IM-2000-31797; B, C,
<taxonomicName id="4C0F4D15FFC7FFAF8DD907827B2B1E64" authority="(L. Pfeiffer, 1862)" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" box="[441,803,2035,2057]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="367" phylum="Mollusca" rank="species" species="mouhoti">
<emphasis id="B97BEA84FFC7FFAF8DD907827A6F1E65" box="[441,615,2035,2057]" italics="true" pageId="3" pageNumber="367">Pincerna mouhoti</emphasis>
(L. Pfeiffer, 1862)
</taxonomicName>
: B, specimen from MƯờng La, SƠn La, Vietnam, ZMHU; C, syntype NHMUK 20170120; D, E,
<taxonomicName id="4C0F4D15FFC7FFAF8DFD08667B981E48" authority="Bavay &amp; Dautzenberg, 1912" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[413,912,2064,2086]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="3" pageNumber="367" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFC7FFAF8DFD08667A6C1E48" box="[413,612,2064,2085]" italics="true" pageId="3" pageNumber="367">Pincerna costulosus</emphasis>
<bibRefCitation id="EF9E4B67FFC7FFAF8E0D08667B981E48" author="Bavay A &amp; Dautzenberg P" box="[621,912,2064,2086]" pageId="3" pageNumber="367" pagination="1 - 54" refId="ref5945" refString="Bavay A &amp; Dautzenberg P (1912) Description de Coquilles Nouvelles de l'Indo-Chine. Journal de Conchyliologie, 60: 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
</taxonomicName>
: D, syntype MNHN-IM-2000-31786; E, specimen from Phong Thổ, Lai Châu, Vietnam, ZMHU.
</paragraph>
</caption>
<caption id="DF70661EFFC0FFA88C1107857D941E2F" ID-DOI="http://doi.org/10.5281/zenodo.7174170" ID-Zenodo-Dep="7174170" httpUri="https://zenodo.org/record/7174170/files/figure.png" pageId="4" pageNumber="368" startId="4.[113,148,2035,2057]" targetBox="[134,1429,188,2020]" targetPageId="4">
<paragraph id="8BB03696FFC0FFA88C1107857D941E2F" blockId="4.[113,1451,2035,2115]" pageId="4" pageNumber="368">
Fig. 3. Shells (apertural view, dorsal view, ventral view, and side view) and operculum (outer surface, inner surface, and side view) of
<taxonomicName id="4C0F4D15FFC0FFA88C1108667A711E48" authority="(Bavay &amp; Dautzenberg, 1900)" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1900" box="[113,633,2064,2086]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="4" pageNumber="368" phylum="Mollusca" rank="species" species="vanbuensis">
<emphasis id="B97BEA84FFC0FFA88C11086679371E48" box="[113,319,2064,2085]" italics="true" pageId="4" pageNumber="368">Pincerna vanbuensis</emphasis>
(Bavay &amp; Dautzenberg, 1900)
</taxonomicName>
. A, syntype MNHN-IM-2000-31798; B, E, specimen from Thuận Châu, SƠn La, Vietnam, ZMHU; C, specimen from Khoune, Xieng Khouang, Laos, ZMHU; D, specimen from Xiengkho, Houaphanh, Laos, ZMHU.
</paragraph>
</caption>
<paragraph id="8BB03696FFC1FFA98CE800CB78F21779" blockId="5.[136,775,187,277]" pageId="5" pageNumber="369">
<materialsCitation id="3B673CCBFFC1FFA98CE800CB78FE1779" collectingDate="2015-07-14" collectionCode="V, T" collectorName="D. S. Do &amp; Sin" elevation="889" latitude="22.58889" longLatPrecision="21" longitude="103.39777" pageId="5" pageNumber="369" specimenCount="1">
Commune, limestone outcrops,
<geoCoordinate id="EE3B5051FFC1FFA98D8600CD7A6016B9" box="[486,616,187,212]" degrees="22" direction="north" minutes="35" orientation="latitude" pageId="5" pageNumber="369" precision="15" seconds="20" value="22.58889">22°35′20″N</geoCoordinate>
,
<geoCoordinate id="EE3B5051FFC1FFA98E1600CD7B0B16B9" box="[630,771,187,212]" degrees="103" direction="east" minutes="23" orientation="longitude" pageId="5" pageNumber="369" precision="15" seconds="52" value="103.39777">103°23′52″E</geoCoordinate>
,
<elevation id="0022D1A5FFC1FFA98CE800AB79721698" box="[136,378,221,245]" metricMagnitude="2" metricUnit="m" metricValue="8.89" pageId="5" pageNumber="369" unit="m" value="889.0">
<quantity id="4CF79B73FFC1FFA98CE800AB78C51699" box="[136,205,221,245]" metricMagnitude="2" metricUnit="m" metricValue="8.89" pageId="5" pageNumber="369" unit="m" value="889.0">889 m</quantity>
above sea level
</elevation>
(a.s.l.), col.
<collectorName id="26FA5340FFC1FFA98E6400AB7A541699" box="[516,604,221,245]" pageId="5" pageNumber="369">D.S. Do</collectorName>
&amp;
<collectionCode id="ED1EAE53FFC1FFA98EE100AB7A901699" box="[641,664,221,244]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="369" type="Museum">V</collectionCode>
.
<collectionCode id="ED1EAE53FFC1FFA98EFB00AB7AA61699" box="[667,686,221,244]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="369">T</collectionCode>
.
<collectorName id="26FA5340FFC1FFA98ED900AB7AD71698" box="[697,735,221,245]" pageId="5" pageNumber="369">Sin</collectorName>
,
<date id="FFB11056FFC1FFA98E8C00AB78FE1779" pageId="5" pageNumber="369" value="2015-07-14">
<collectingDate id="EFF5E9BEFFC1FFA98E8C00AB78FE1779" pageId="5" pageNumber="369" value="2015-07-14">14 July 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC1FFA98CE8014A7A1917F8" pageId="5" pageNumber="369" type="diagnosis">
<paragraph id="8BB03696FFC1FFA98CE8014A7A1917F8" blockId="5.[136,775,316,405]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98CE8014A78F41739" bold="true" box="[136,252,316,340]" pageId="5" pageNumber="369">Diagnosis.</emphasis>
<taxonomicName id="4C0F4D15FFC1FFA98D64014B79D61738" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[260,478,317,341]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFC1FFA98D64014B79D61738" box="[260,478,317,341]" italics="true" pageId="5" pageNumber="369">Pincerna costulosus</emphasis>
</taxonomicName>
is characterised by a small sized, translucent, high conical shell with strongly, regularly ribs on R1, and R2 extremely short.
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC1FFA98CE801CA79D31478" pageId="5" pageNumber="369" type="distribution">
<paragraph id="8BB03696FFC1FFA98CE801CA79D31478" blockId="5.[135,775,444,533]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98CE801CA791C17B9" bold="true" box="[136,276,444,468]" pageId="5" pageNumber="369">Distribution.</emphasis>
<taxonomicName id="4C0F4D15FFC1FFA98D7801CB79EE17B8" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[280,486,445,469]" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFC1FFA98D7801CB79EE17B8" box="[280,486,445,469]" italics="true" pageId="5" pageNumber="369">Pincerna costulosus</emphasis>
</taxonomicName>
is known from northwestern
<collectingCountry id="F3187606FFC1FFA98CE701AB78EB1798" box="[135,227,477,501]" name="Vietnam" pageId="5" pageNumber="369">Vietnam</collectingCountry>
(
<collectingRegion id="49CBF874FFC1FFA98C9001AB79BF1798" box="[240,439,477,501]" country="Vietnam" name="Lai Chau" pageId="5" pageNumber="369">Lai Châu Province</collectingRegion>
) and Southwest
<collectingCountry id="F3187606FFC1FFA98E0401AB7AAC1798" box="[612,676,477,501]" name="China" pageId="5" pageNumber="369">China</collectingCountry>
(
<collectingRegion id="49CBF874FFC1FFA98ED001AB78E11478" country="China" name="Yunnan" pageId="5" pageNumber="369">Yunnan Province</collectingRegion>
) (
<bibRefCitation id="EF9E4B67FFC1FFA98C9F018B79C71478" author="Chen ZY &amp; Wu M" box="[255,463,509,533]" pageId="5" pageNumber="369" pagination="37 - 46" refId="ref5993" refString="Chen ZY &amp; Wu M (2020) Two new cyclophoroid species from Hubei, China, with proposal of a new genus (Gastropoda, Caenogastropoda, Diplommatinidae and Alycaeidae). ZooKeys, 935: 37 - 46." type="journal article" year="2020">Chen &amp; Wu, 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C315651DFFC1FFA98CE8024B7B0F14B9" pageId="5" pageNumber="369" type="discussion">
<paragraph id="8BB03696FFC1FFA98CE8024B7B0F14B9" blockId="5.[136,775,573,725]" pageId="5" pageNumber="369">
<emphasis id="B97BEA84FFC1FFA98CE8024B78F31438" bold="true" box="[136,251,573,597]" pageId="5" pageNumber="369">Remarks.</emphasis>
The species is quite stable in terms of shell characters, although only a few shells are known.
<taxonomicName id="4C0F4D15FFC1FFA98EC5022B78FF14F8" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" class="Gastropoda" family="Alycaeidae" genus="Pincerna" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="369" phylum="Mollusca" rank="species" species="costulosus">
<emphasis id="B97BEA84FFC1FFA98EC5022B78FF14F8" italics="true" pageId="5" pageNumber="369">Pincerna costulosus</emphasis>
</taxonomicName>
has been recently recorded from Xishuangbanna,
<collectingRegion id="49CBF874FFC1FFA98CE802EB794F14D8" box="[136,327,669,693]" country="China" name="Yunnan" pageId="5" pageNumber="369">Yunnan Province</collectingRegion>
,
<collectingCountry id="F3187606FFC1FFA98D3302EB799B14D8" box="[339,403,669,693]" name="China" pageId="5" pageNumber="369">China</collectingCountry>
(
<bibRefCitation id="EF9E4B67FFC1FFA98DC302EB7A7B14D8" author="Chen ZY &amp; Wu M" box="[419,627,669,693]" pageId="5" pageNumber="369" pagination="37 - 46" refId="ref5993" refString="Chen ZY &amp; Wu M (2020) Two new cyclophoroid species from Hubei, China, with proposal of a new genus (Gastropoda, Caenogastropoda, Diplommatinidae and Alycaeidae). ZooKeys, 935: 37 - 46." type="journal article" year="2020">Chen &amp; Wu, 2020</bibRefCitation>
). The present record extends the distributed range circa
<quantity id="4CF79B73FFC1FFA98E5C02CB7A8314B8" box="[572,651,701,725]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="369" unit="km" value="400.0">400 km</quantity>
to the west.
</paragraph>
</subSubSection>
</treatment>
</document>
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