228 lines
32 KiB
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228 lines
32 KiB
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<mods:title id="070F50E09EF78C7ECCE8BF317E7867AB">On the systematic placement of the fossil crab Lathahypossia aculeata (Busulini, Tessier & Visentin, 1984) (Crustacea: Decapoda: Brachyura)</mods:title>
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<paragraph id="8B8C010AFFE8FFAE42E6DEE7AD685DD7" blockId="3.[151,884,1942,2005]" box="[151,884,1942,1969]" pageId="3" pageNumber="268">
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<heading id="D0C4B666FFE8FFAE42E6DEE7AD685DD7" bold="true" box="[151,884,1942,1969]" fontSize="11" level="1" pageId="3" pageNumber="268" reason="1">
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<taxonomicName id="4C337A89FFE8FFAE42E6DEE7AD685DD7" authority="(Busulini, Tessier & Visentin, 1984)" baseAuthorityName="Busulini, Tessier & Visentin" baseAuthorityYear="1984" box="[151,884,1942,1969]" class="Malacostraca" family="Hypothalassiidae" genus="Lathahypossia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="aculeata">
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<emphasis id="B947DD18FFE8FFAE42E6DEE7AD685DD7" bold="true" box="[151,884,1942,1969]" pageId="3" pageNumber="268">
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<emphasis id="B947DD18FFE8FFAE42E6DEE7AFAA5DD7" bold="true" box="[151,438,1942,1969]" italics="true" pageId="3" pageNumber="268">Lathahypossia aculeata</emphasis>
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(
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<bibRefCitation id="EFA27CFBFFE8FFAE43B5DEE6AD705DD7" author="Busulini, A. & Tessier, G. & Visentin, M." box="[452,876,1943,1969]" pageId="3" pageNumber="268" pagination="107 - 117" refId="ref5192" refString="Busulini, A., Tessier, G. & Visentin, M. (1984) Titanocarcinus aculeatus nuova specie di brachiuro nell'Eocene del Veneto (Crustacea, Decapoda). Lavori - Societa Veneziana di Scienze Naturali, 9 (1), 107 - 117." type="journal article" year="1984">Busulini, Tessier & Visentin, 1984</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3295281FFE8FFAE42E6DECAAF0E5DB3" box="[151,274,1979,2005]" pageId="3" pageNumber="268" type="description">
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<paragraph id="8B8C010AFFE8FFAE42E6DECAAF0E5DB3" blockId="3.[151,884,1942,2005]" box="[151,274,1979,2005]" pageId="3" pageNumber="268">
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(
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<figureCitation id="13081D8FFFE8FFAE42EEDECAAF155DB3" box="[159,265,1979,2005]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1765,1789]" captionStartId-1="4.[151,250,1724,1748]" captionStartId-2="5.[151,250,1366,1390]" captionTargetBox-0="[177,1411,450,1738]" captionTargetBox-1="[177,1410,322,1701]" captionTargetBox-2="[177,1410,181,1342]" captionTargetId-0="figure-153@3.[177,1411,450,1738]" captionTargetId-1="figure-121@4.[177,1410,322,1701]" captionTargetId-2="figure-17@5.[177,1410,181,1342]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. A, B, Lathahypossia aculeata (Busulini, Tessier & Visentin, 1984), holotype, MSNVE-25083, from the Middle Eocene of the Cava Main, Arzignano (northeast Italy). A, dorsal view; B, frontal view. Scale bar = 10 mm (pictures by G. Tessier)." captionText-1="FIGURE 2. A–F, Lathahypossia aculeata (Busulini, Tessier & Visentin, 1984), from the Middle Eocene of the Cava Main, Arzignano (northeast Italy). A–E, male, MSNVE-27378: A, dorsal view; B, ventral view; C, frontal view, D, posterior view; E, right lateral view. F, male, MSNVE-27377, ventral view. Abbreviations, s: pleonal somites; st: thoracic sternites; t: telson. Scale bar = 10 mm." captionText-2="FIGURE 3. A–G, Lathahypossia aculeata (Busulini, Tessier & Visentin, 1984), from the Middle Eocene of the Cava Main, Arzignano (NE Italy). A, B, D–F, male, MSNVE-27378: A, view of sub-cephalic region; B, view of endostomial region; D, detail of pleon; E–G, detail and close-up of ‘press-button’, male pleonal locking mechanism. C, male, MSNVE-27377, detail of anterior part of sternum. Abbreviations, aa: basal antennular article; ab: basal antennal article; en: endostome; ep; epistome; er: endostomial ridge; ex: exopod of third maxilliped; is: ischium of third maxilliped; mxp3: third maxilliped; P1: first pereopod; pb: ‘press-button’; s: pleonal somites; st: thoracic sternites. Scale bar = 10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8391501" figureDoi-1="http://doi.org/10.5281/zenodo.8391503" figureDoi-2="http://doi.org/10.5281/zenodo.8391507" httpUri-0="https://zenodo.org/record/8391501/files/figure.png" httpUri-1="https://zenodo.org/record/8391503/files/figure.png" httpUri-2="https://zenodo.org/record/8391507/files/figure.png" pageId="3" pageNumber="268">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3295281FFEFFFA942E6D9E6AB805B6C" pageId="4" pageNumber="269" type="reference_group">
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<paragraph id="8B8C010AFFEFFFA942E6D9E6AD375AC9" blockId="4.[151,1436,151,297]" box="[151,811,151,175]" pageId="4" pageNumber="269">
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<taxonomicName id="4C337A89FFEFFFA942E6D9E6AFCB5AC9" authorityName="A. Milne-Edwards" authorityYear="1864" box="[151,471,151,175]" class="Malacostraca" family="Hypothalassiidae" genus="Titanocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species">
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<emphasis id="B947DD18FFEFFFA942E6D9E6AF325AC8" box="[151,302,151,174]" italics="true" pageId="4" pageNumber="269">Titanocarcinus</emphasis>
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aff.
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<emphasis id="B947DD18FFEFFFA9432CD9E6AFCB5AC9" box="[349,471,151,175]" italics="true" pageId="4" pageNumber="269">Raulinianus</emphasis>
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</taxonomicName>
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<emphasis id="B947DD18FFEFFFA943A6D9E6AFE85AC9" box="[471,500,151,175]" italics="true" pageId="4" pageNumber="269">.—</emphasis>
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<bibRefCitation id="EFA27CFBFFEFFFA94385D9E6ACF95AC9" author="Busulini, A. & Tessier, G. & Visentin, M." box="[500,741,151,175]" pageId="4" pageNumber="269" pagination="75 - 84" refId="ref5142" refString="Busulini, A., Tessier, G. & Visentin, M. (1982) Brachyura della cava Main (Arzignano) - Lessini orientali (Vicenza) (Crustacea, Decapoda). Lavori - Societa Veneziana di Scienze Naturali, 7, 75 - 84." type="journal article" year="1982">
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Busulini
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<emphasis id="B947DD18FFEFFFA94021D9E6AC615AC9" box="[592,637,151,175]" italics="true" pageId="4" pageNumber="269">et al</emphasis>
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. 1982: 82
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</bibRefCitation>
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, fig. 3.
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</paragraph>
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<paragraph id="8B8C010AFFEFFFA942E6D9C7AB805B6C" blockId="4.[151,1436,151,297]" pageId="4" pageNumber="269">
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<taxonomicName id="4C337A89FFEFFFA942E6D9C7AD465AAB" ID-CoL="e20179c5-49cd-47f9-a607-036d151954b6" authority="Busulini, Tessier & Visentin, 1984: 107 - 114" authorityName="Busulini, Tessier & Visentin" authorityPageNumber="107 - 114" authorityYear="1984" box="[151,858,181,205]" class="Malacostraca" family="Xanthidae" genus="Titanocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="aculeatus">
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<emphasis id="B947DD18FFEFFFA942E6D9C7AF895AAB" box="[151,405,181,205]" italics="true" pageId="4" pageNumber="269">Titanocarcinus aculeatus</emphasis>
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<bibRefCitation id="EFA27CFBFFEFFFA943EBD9C4AD465AAB" author="Busulini, A. & Tessier, G. & Visentin, M." box="[410,858,181,205]" pageId="4" pageNumber="269" pagination="107 - 117" refId="ref5192" refString="Busulini, A., Tessier, G. & Visentin, M. (1984) Titanocarcinus aculeatus nuova specie di brachiuro nell'Eocene del Veneto (Crustacea, Decapoda). Lavori - Societa Veneziana di Scienze Naturali, 9 (1), 107 - 117." type="journal article" year="1984">Busulini, Tessier & Visentin, 1984: 107–114</bibRefCitation>
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||
</taxonomicName>
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||
, pl. 1 figs. 1–3, pl. 2 figs, 1, 2, pl. 3 figs. 1, 2.—
|
||
<bibRefCitation id="EFA27CFBFFEFFFA94745D9C4AB805AAB" author="De Angeli, A. & Beschin, C." box="[1332,1436,181,205]" pageId="4" pageNumber="269" pagination="5 - 54" refId="ref5532" refString="De Angeli, A. & Beschin, C. (2001) I Crostacei fossili del territorio Vicentino. Natura Vicentina, 5, 5 - 54." type="journal article" year="2001">De Angeli</bibRefCitation>
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& Beschin 2001, fig. 36.—
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<bibRefCitation id="EFA27CFBFFEFFFA943A9D9A5ACBC5A8A" author="Beschin, C. & Busulini, A. & De Angeli, A. & Tessier, G." box="[472,672,212,236]" pageId="4" pageNumber="269" pagination="7 - 28" refId="ref4853" refString="Beschin, C., Busulini, A., De Angeli, A. & Tessier, G. (2002) Aggiornamento ai crostacei eocenici di cava " Main " di Arzignano (Vicenza, Italia settentrionale) (Crustacea, Decapoda). Studi e Ricerche - Associazione Amici del Museo - Museo Civico " G. Zannato ", Monlccchio Maggiore (Viccnza), 15 Dicembre 2002, 7 - 28." type="book chapter" year="2002">
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Beschin
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<emphasis id="B947DD18FFEFFFA94043D9A4AC435A8A" box="[562,607,212,236]" italics="true" pageId="4" pageNumber="269">et al</emphasis>
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. 2002
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</bibRefCitation>
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||
, text-fig. 15, pl. 4 fig. 3.—
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<bibRefCitation id="EFA27CFBFFEFFFA941DCD9A5AB145A8A" author="Karasawa, H. & Schweitzer, C. E." box="[941,1288,212,236]" pageId="4" pageNumber="269" pagination="23 - 73" refId="ref6142" refString="Karasawa, H. & Schweitzer, C. E. (2006) A new classification of the Xanthoidea sensu lato (Crustacea: Decapoda: Brachyura) based on phylogenetic analysis and traditional systematics and evaluation of all fossil Xanthoidea sensu lato. Contributions to Zoology, 75 (1 / 2), 23 - 73. https: // doi. org / 10.1163 / 18759866 - 0750102002" type="journal article" year="2006">Karasawa & Schweitzer 2006: 53</bibRefCitation>
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||
.
|
||
<taxonomicName id="4C337A89FFEFFFA942E6D982AF965B6D" baseAuthorityName="Busulini, Tessier & Visentin" baseAuthorityYear="1984" box="[151,394,243,267]" class="Malacostraca" family="Hypothalassiidae" genus="Lathahypossia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="aculeata">
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<emphasis id="B947DD18FFEFFFA942E6D982AF965B6D" box="[151,394,243,267]" italics="true" pageId="4" pageNumber="269">Lathahypossia aculeata</emphasis>
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</taxonomicName>
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.—
|
||
<bibRefCitation id="EFA27CFBFFEFFFA943D9D982ACD55B6D" author="Schweitzer C. E. & Artal P. & Van Bakel B. & Jagt J. W. M. & Karasawa, H." box="[424,713,243,267]" pageId="4" pageNumber="269" pagination="278 - 295" refId="ref8010" refString="Schweitzer C. E., Artal P., Van Bakel B., Jagt J. W. M., Karasawa, H. (2007) Revision of the genus Titanocarcinus (Decapoda: Brachyura: Xanthoidea) with two new genera and one new species. Journal of Crustacean Biology, 27 (2), 278 - 295. https: // doi. org / 10.1651 / S- 2713.1" type="journal article" year="2007">
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Schweitzer
|
||
<emphasis id="B947DD18FFEFFFA94052D982AC4D5B6D" box="[547,593,243,267]" italics="true" pageId="4" pageNumber="269">et al</emphasis>
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||
. 2007: 292
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</bibRefCitation>
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, 293.—Beschin
|
||
<emphasis id="B947DD18FFEFFFA94104D982ADBF5B6D" box="[885,931,243,267]" italics="true" pageId="4" pageNumber="269">et al</emphasis>
|
||
. 2012: 72, text-fig. 64, pl. 10 fig. 5.—
|
||
<bibRefCitation id="EFA27CFBFFEFFFA94745D983AB805B6C" author="Jagt, J. W. M. & Van Bakel, B. W. M. & Guinot, D. & Fraaije, R. H. B. & Artal, P." box="[1332,1436,242,267]" pageId="4" pageNumber="269" pagination="847 - 920" refId="ref6474" refString="Jagt, J. W. M., Van Bakel, B. W. M., Guinot, D., Fraaije, R. H. B. & Artal, P. (2015) Fossil Brachyura. In: Castro, P., Davie, P. J. F., Guinot, D., Schram, F. R. & von Vaupel Klein, J. C. (Eds.), The Crustacea, Treatise on Zoology - Anatomy, Taxonomy, Biology. Vol. 9. Part C - II. Decapoda: Brachyura (Part 2): Fossil Brachyura. Brill, Leiden, pp. 847 - 920. https: // doi. org / 10.1163 / 9789004190832 _ 018" type="book chapter" year="2015">
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Jagt
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<emphasis id="B947DD18FFEFFFA94719D982AB8A5B6D" box="[1384,1430,243,267]" italics="true" pageId="4" pageNumber="269">et al</emphasis>
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||
.
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||
</bibRefCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="C3295281FFEFFFA942B6D860AC345D93" pageId="4" pageNumber="269" type="materials_examined">
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<paragraph id="8B8C010AFFEFFFA942B6D860AA255B4F" blockId="4.[151,1436,151,297]" box="[199,1081,273,297]" pageId="4" pageNumber="269">
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||
2015: 898, fig. 71D.—
|
||
<bibRefCitation id="EFA27CFBFFEFFFA943DDD860ACA65B4F" author="De Angeli, A. & Garassino, A. & Pasini, G." box="[428,698,273,297]" pageId="4" pageNumber="269" pagination="1 - 87" refId="ref5561" refString="De Angeli, A., Garassino, A. & Pasini, G. (2019) Catalog and bibliography of fossil Stomatopoda and Decapoda from Italy (2007 - 2018). Memorie della Societa Italiana di Scienze Naturali e del Museo Di Storia Naturale Di Milano, 45, 1 - 87." type="journal article" year="2019">
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De Angeli
|
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<emphasis id="B947DD18FFEFFFA9406DD863AC545B4F" box="[540,584,273,297]" italics="true" pageId="4" pageNumber="269">et al</emphasis>
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||
. 2019: 898
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||
</bibRefCitation>
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.—
|
||
<bibRefCitation id="EFA27CFBFFEFFFA940A6D860ADC45B4F" author="Schweitzer, C. E. & Feldmann, R. M. & Karasawa, H." box="[727,984,273,297]" pageId="4" pageNumber="269" pagination="1 - 8" refId="ref8086" refString="Schweitzer, C. E., Feldmann, R. M. & Karasawa, H. (2020) Part R, Revised, Volume 1, Chapter 8 T 10: Systematic descriptions: Superfamily Eriphioidea. Treatise Online, 132, 1 - 8. https: // doi. org / 10.17161 / to. vi. 13577" type="journal article" year="2020">
|
||
Schweitzer
|
||
<emphasis id="B947DD18FFEFFFA94120D863AD625B4F" box="[849,894,273,297]" italics="true" pageId="4" pageNumber="269">et al</emphasis>
|
||
. 2020: 2
|
||
</bibRefCitation>
|
||
, fig. 1, 4.
|
||
</paragraph>
|
||
<caption id="DF4C5182FFEFFFA942E6DFCDAF3D5D59" ID-DOI="http://doi.org/10.5281/zenodo.8391503" ID-Zenodo-Dep="8391503" httpUri="https://zenodo.org/record/8391503/files/figure.png" pageId="4" pageNumber="269" startId="4.[151,250,1724,1748]" targetBox="[177,1410,322,1701]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8B8C010AFFEFFFA942E6DFCDAF3D5D59" blockId="4.[151,1436,1724,1856]" pageId="4" pageNumber="269">
|
||
<emphasis id="B947DD18FFEFFFA942E6DFCDAF095CB2" bold="true" box="[151,277,1724,1748]" pageId="4" pageNumber="269">FIGURE 2.</emphasis>
|
||
A–F,
|
||
<taxonomicName id="4C337A89FFEFFFA94327DFCDADD75CB2" authority="(Busulini, Tessier & Visentin, 1984)" baseAuthorityName="Busulini, Tessier & Visentin" baseAuthorityYear="1984" box="[342,971,1724,1748]" class="Malacostraca" family="Hypothalassiidae" genus="Lathahypossia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="aculeata">
|
||
<emphasis id="B947DD18FFEFFFA94327DFCDAC565CB2" box="[342,586,1724,1748]" italics="true" pageId="4" pageNumber="269">Lathahypossia aculeata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA27CFBFFEFFFA9402BDFCDADD85CB2" author="Busulini, A. & Tessier, G. & Visentin, M." box="[602,964,1724,1748]" pageId="4" pageNumber="269" pagination="107 - 117" refId="ref5192" refString="Busulini, A., Tessier, G. & Visentin, M. (1984) Titanocarcinus aculeatus nuova specie di brachiuro nell'Eocene del Veneto (Crustacea, Decapoda). Lavori - Societa Veneziana di Scienze Naturali, 9 (1), 107 - 117." type="journal article" year="1984">Busulini, Tessier & Visentin, 1984</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, from the Middle Eocene of the Cava Main, Arzignano (northeast Italy). A–E, male, MSNVE-27378: A, dorsal view; B, ventral view; C, frontal view, D, posterior view; E, right lateral view. F, male, MSNVE-27377, ventral view. Abbreviations,
|
||
<emphasis id="B947DD18FFEFFFA94101DE75AD665D7A" bold="true" box="[880,890,1796,1820]" pageId="4" pageNumber="269">s</emphasis>
|
||
: pleonal somites;
|
||
<emphasis id="B947DD18FFEFFFA94640DE75AA5F5D7A" bold="true" box="[1073,1091,1796,1820]" pageId="4" pageNumber="269">st</emphasis>
|
||
: thoracic sternites;
|
||
<emphasis id="B947DD18FFEFFFA94779DE75AB0C5D7A" bold="true" box="[1288,1296,1796,1820]" pageId="4" pageNumber="269">t</emphasis>
|
||
: telson. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C010AFFEFFFA942E6DEE3AC345D93" blockId="4.[151,1436,1938,2037]" pageId="4" pageNumber="269">
|
||
<emphasis id="B947DD18FFEFFFA942E6DEE3AF9C5DCA" bold="true" box="[151,384,1938,1964]" pageId="4" pageNumber="269">Material examined.</emphasis>
|
||
One complete carapace without pereopods: MSNVE-27378, L = 40.5; W = 54.5; FOW = 36.0; FW = 16.0/19.0;
|
||
<collectionCode id="ED2299CFFFEFFFA94325DEC6AF605DB7" box="[340,380,1975,2001]" country="USA" name="Pratt Museum" pageId="4" pageNumber="269">PM</collectionCode>
|
||
= 23.0. One specimen preserving partial sternopleonal features and chelipeds: MSNVE-27377, width from carpus to carpus = 67.5.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4C5182FFEEFFA842E6DC27ADA35C44" ID-DOI="http://doi.org/10.5281/zenodo.8391507" ID-Zenodo-Dep="8391507" httpUri="https://zenodo.org/record/8391507/files/figure.png" pageId="5" pageNumber="270" startId="5.[151,250,1366,1390]" targetBox="[177,1410,181,1342]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8B8C010AFFEEFFA842E6DC27ADA35C44" blockId="5.[151,1437,1366,1570]" pageId="5" pageNumber="270">
|
||
<emphasis id="B947DD18FFEEFFA842E6DC27AF095F08" bold="true" box="[151,277,1366,1390]" pageId="5" pageNumber="270">FIGURE 3.</emphasis>
|
||
A–G,
|
||
<taxonomicName id="4C337A89FFEEFFA8432DDC27ADD25F08" authority="(Busulini, Tessier & Visentin, 1984)" baseAuthorityName="Busulini, Tessier & Visentin" baseAuthorityYear="1984" box="[348,974,1366,1390]" class="Malacostraca" family="Hypothalassiidae" genus="Lathahypossia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="aculeata">
|
||
<emphasis id="B947DD18FFEEFFA8432DDC27AC535F08" box="[348,591,1366,1390]" italics="true" pageId="5" pageNumber="270">Lathahypossia aculeata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA27CFBFFEEFFA8402FDC27ADDB5F08" author="Busulini, A. & Tessier, G. & Visentin, M." box="[606,967,1366,1390]" pageId="5" pageNumber="270" pagination="107 - 117" refId="ref5192" refString="Busulini, A., Tessier, G. & Visentin, M. (1984) Titanocarcinus aculeatus nuova specie di brachiuro nell'Eocene del Veneto (Crustacea, Decapoda). Lavori - Societa Veneziana di Scienze Naturali, 9 (1), 107 - 117." type="journal article" year="1984">Busulini, Tessier & Visentin, 1984</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, from the Middle Eocene of the Cava Main, Arzignano (NE Italy). A, B, D–F, male, MSNVE-27378: A, view of sub-cephalic region; B, view of endostomial region; D, detail of pleon; E–G, detail and close-up of ‘press-button’, male pleonal locking mechanism. C, male, MSNVE-27377, detail of anterior part of sternum. Abbreviations,
|
||
<emphasis id="B947DD18FFEEFFA8405ADCB3AC585FBC" bold="true" box="[555,580,1474,1498]" pageId="5" pageNumber="270">aa</emphasis>
|
||
: basal antennular article;
|
||
<emphasis id="B947DD18FFEEFFA84136DCB3AD7E5FBC" bold="true" box="[839,866,1474,1498]" pageId="5" pageNumber="270">ab</emphasis>
|
||
: basal antennal article;
|
||
<emphasis id="B947DD18FFEEFFA84621DCB3AA755FBC" bold="true" box="[1104,1129,1474,1498]" pageId="5" pageNumber="270">en</emphasis>
|
||
: endostome;
|
||
<emphasis id="B947DD18FFEEFFA84680DCB3AB165FBC" bold="true" box="[1265,1290,1474,1498]" pageId="5" pageNumber="270">ep</emphasis>
|
||
; epistome;
|
||
<emphasis id="B947DD18FFEEFFA8470EDCB3AB895FBC" bold="true" box="[1407,1429,1474,1498]" pageId="5" pageNumber="270">er</emphasis>
|
||
: endostomial ridge;
|
||
<emphasis id="B947DD18FFEEFFA8432DDC97AF685F98" bold="true" box="[348,372,1510,1534]" pageId="5" pageNumber="270">ex</emphasis>
|
||
: exopod of third maxilliped;
|
||
<emphasis id="B947DD18FFEEFFA840D1DC97ACAD5F98" bold="true" box="[672,689,1510,1534]" pageId="5" pageNumber="270">is</emphasis>
|
||
: ischium of third maxilliped;
|
||
<emphasis id="B947DD18FFEEFFA84193DC97AA035F98" bold="true" box="[994,1055,1510,1534]" pageId="5" pageNumber="270">mxp3</emphasis>
|
||
: third maxilliped;
|
||
<emphasis id="B947DD18FFEEFFA846ACDC97AAE55F98" bold="true" box="[1245,1273,1510,1534]" pageId="5" pageNumber="270">P1</emphasis>
|
||
: first pereopod;
|
||
<emphasis id="B947DD18FFEEFFA842E6DF7BAEAF5C44" bold="true" box="[151,179,1546,1570]" pageId="5" pageNumber="270">pb</emphasis>
|
||
: ‘press-button’;
|
||
<emphasis id="B947DD18FFEEFFA8432DDF7BAF7A5C44" bold="true" box="[348,358,1546,1570]" pageId="5" pageNumber="270">s</emphasis>
|
||
: pleonal somites;
|
||
<emphasis id="B947DD18FFEEFFA8406EDF7BAC2D5C44" bold="true" box="[543,561,1546,1570]" pageId="5" pageNumber="270">st</emphasis>
|
||
: thoracic sternites. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3295281FFEEFFA842B6DF05AF2B5DEC" pageId="5" pageNumber="270" type="diagnosis">
|
||
<paragraph id="8B8C010AFFEEFFA842B6DF05AF2B5DEC" blockId="5.[151,1436,1652,2039]" pageId="5" pageNumber="270">
|
||
<emphasis id="B947DD18FFEEFFA842B6DF05AFDB5CE8" bold="true" box="[199,455,1652,1678]" pageId="5" pageNumber="270">Diagnosis (emended).</emphasis>
|
||
Carapace transversely ovate, wider than long, L/W ratio about 0.75; surface granulate, fairly vaulted longitudinally, regions well defined, spiny anterior half of outline; front bilobed, armed with acute spines; orbits ovate, complete, supraorbital margin spiny, with one closed medial fissure; anterolateral margins with 4 spiny lobes; posterior margin convex, posteriorly convergent; posterior margin straight, rimmed. Male thoracic sternum relatively broad, strongly vaulted longitudinally; sternopleonal cavity deep, exceeding midpoint of P1 coxa, reaching 2/3 of sternite 4, press-button on anterior margin of sternite 5. Pleon subtrapezoidal, somite 3 widest, somites 3–5 fused with subtle demarcations visible. Cheliped strongly spinous, heterochelous. Ambulatory legs slender, spiny.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3295281FFEEFFAB42B6DEE5ACE05FA7" lastPageId="6" lastPageNumber="271" pageId="5" pageNumber="270" type="description">
|
||
<paragraph id="8B8C010AFFEEFFAB42B6DEE5AB0C595F" blockId="5.[151,1436,1652,2039]" lastBlockId="6.[151,1437,151,1509]" lastPageId="6" lastPageNumber="271" pageId="5" pageNumber="270">
|
||
<emphasis id="B947DD18FFEEFFA842B6DEE5AF455DC8" bold="true" box="[199,345,1940,1966]" pageId="5" pageNumber="270">Description.</emphasis>
|
||
Carapace transversely ovate, wider than long, L/W ratio about 0.75, maximum width half carapace length, fairly vaulted longitudinally, slightly vaulted transversely; surface covered with scattered granules and setal pits. Regions swollen, well-defined by shallow, smooth grooves. Front bilobed, notched medially; each lobe, sinuous, with 3 acute spines internally and 1 acute spine at outer corner. Orbits transversely ovate, complete, slightly obliquely placed; supraorbital margin raised, spiny, with closed medial fissure; deep notch separating inner orbital spine from front; infraorbital margin spiny, dorsally visible; ocular peduncle short. Anterolateral margins gently convex, with 4 lobes or teeth: first lobe smaller, last lobe (epibranchial) largest, all armed with long, acute spines. Posterolateral margin slightly convex, with rounded edges, posteriorly convergent. Posterior margin straight in dorsal view, medially slightly concave in posterior view, with fine rim. Frontal region depressed, directed downwards. Epigastric lobes fairly swollen, stepped distally. Protogastric lobes slightly swollen, adjacent to epigastric lobes. Mesogastric region very slender anteriorly reaching middle of epigastric lobes, broadly subpentagonal posteriorly. Metagastric region weakly differentiated from mesogastric region. Urogastric region bounded laterally by branchiocardiac grooves, separated from meso-, metagastric lobes by cervical groove; adjacent and undifferentiated from cardiac regions, with 2 regions forming pentagon with the apex pointed downwards. Intestinal region short, smooth. Hepatic lobes swollen, demarcated by cervical and gastrohepatic grooves. Epibranchial region defined by 3 swollen lobes, separated by smooth grooves. Metabranchial region swollen, weakly defined. Metabranchial regions demarcated by 2 transverse swellings surrounding intestinal region. Suborbital region defined by smooth fringe between row of spines parallel to lateral and infraorbital margins. Subhepatic lobes swollen; with oblique shallow smooth grooves on ventral carapace surface. Pterygostomial regions subtrapezoidal, concave in upper and lower margins, finely granulated. Endostome broad, endostomial ridge weakly marked in lower part. Epistome widely subtriangular, edges ornate, with granules. Proepistome prominent, granulated. Basal antennular article subtrapezoidal, with granulated oblique ridge, folding transversely in antennular fossae. Basal antennal article spiny, obliquely placed.
|
||
</paragraph>
|
||
<paragraph id="8B8C010AFFEDFFAB42B6DA32AFED59C3" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="271">Ischium of third maxilliped with convex inner margin, with 2 longitudinal sulci, 1 medial and second parallel to inner margin, lined by setal pits, inner margin subtruncate, lined by setal pits; merus subtrapezoidal; exopod almost straight, half width of ischium.</paragraph>
|
||
<paragraph id="8B8C010AFFEDFFAB42B6DADEADEB5EA3" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="271">Male thoracic sternum relatively broad, strongly vaulted longitudinally. Sternites 1 and 2 form a triangular structure with margins lined by fine granules; sternite 3 surface medially concave, distal margin granulated; sternite 4 subtrapezoidal in shape, surface medially concave, divided by sternopleonal cavity, broadly vaulted laterally, lateral edges granulated. Sternites 5–7 subtrapezoidal, sternite 5 longer, sternite 7 shorter; sternite 8 not visible when pleon closed. Episternites 4–7 posteriorly directed, sutures not visible. Suture 1/2 distinct; suture 2/3 complete; suture 3/4 only visible laterally; sutures 4/5, 5/6 and 6/7, laterally complete. Sternopleonal cavity deep, narrow, reaching beyond gynglyme for P1, reaching two–thirds of sternite 4 to become deeper, margins ornate, lined with granules. Press button on anterior margin of sternite 5, adjacent to suture 4/5.</paragraph>
|
||
<paragraph id="8B8C010AFFEDFFAB42B6DDBEADFB5F33" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="271">Male pleon relatively narrow, slightly vaulted ventrally; somites subrectangular, transversely elongate; somites 1 and 2 shorter; somites 3–5 subequal in length, fused, with demarcations shallow but visible, somite 3 widest; somite 6 slightly longer than somite 5, with sockets in inner distal corner to receive tubercle of press-button of male pleonal-locking mechanism; telson subtriangular elongate with rounded tip.</paragraph>
|
||
<paragraph id="8B8C010AFFEDFFAB42B6DC2EADB55FFB" blockId="6.[151,1437,151,1509]" pageId="6" pageNumber="271">Chelipeds strongly spinous, heterochelous, palm robust, armed with rows of acute spines on outer and upper margins; dactylus about one–third length of propodus, with black tips.</paragraph>
|
||
<paragraph id="8B8C010AFFEDFFAB42B6DCD6ACE05FA7" blockId="6.[151,1437,151,1509]" box="[199,764,1447,1473]" pageId="6" pageNumber="271">Ambulatory legs slender, margins weakly spinose.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3295281FFEDFFAB42B6DCBAAC1C5F83" box="[199,512,1483,1509]" pageId="6" pageNumber="271" type="discussion">
|
||
<paragraph id="8B8C010AFFEDFFAB42B6DCBAAC1C5F83" blockId="6.[151,1437,151,1509]" box="[199,512,1483,1509]" pageId="6" pageNumber="271">
|
||
<emphasis id="B947DD18FFEDFFAB42B6DCBAAF275F83" bold="true" box="[199,315,1483,1509]" pageId="6" pageNumber="271">Remarks.</emphasis>
|
||
As for the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |