403 lines
60 KiB
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403 lines
60 KiB
XML
<document id="6FC2DB966145E05E96A904619CC2B10A" ID-DOI="10.11646/zootaxa.4103.5.4" ID-GBIF-Dataset="b9b4b63c-d1af-47a8-8e0a-3c63a6cbcc91" ID-ISSN="1175-5326" ID-Zenodo-Dep="263213" ID-ZooBank="D31571A3-8049-423F-AF40-37E4D5FA18D2" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460701761610" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2016" docId="039A87B6C03BFFB1A7B1011443C4FD21" docLanguage="en" docName="zootaxa.4103.5.4.pdf.imf" docOrigin="Zootaxa 4103 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus" docType="treatment" docVersion="7" lastPageNumber="470" masterDocId="FFA3FFCEC03FFFB6A72603304003FF86" masterDocTitle="Taxonomy of ' Euconnus complex'. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepaloconnus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="472" masterPageNumber="463" pageNumber="467" updateTime="1698354660124" updateUser="plazi">
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<mods:titleInfo id="2C6E9955AC3D3C0519877A3FEAF1837E">
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<mods:title id="64D1A7BC0C9258C82DCFBC42C7BE4308">Taxonomy of ' Euconnus complex'. Part VIII. Subgenera Napoconnus, Himaloconnus and Nepaloconnus removed from Euconnus (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A99FDF8BAF1D428AE3AF005743588265" type="personal">
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<mods:namePart id="28517DF5E8FFF64D700163AC3EF468CB">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="C520F815941B6544C4C07FF19044F9B4">text</mods:typeOfResource>
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<mods:title id="73BB68AC71B60E16831E47150C915F7C">Zootaxa</mods:title>
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<mods:part id="9621F2885C768E145248FB40F30B68DC">
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<mods:date id="43D12010F05428EA85970CACBBFFA8F7">2016</mods:date>
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<mods:detail id="35CAC13318F51BD14CA309EB6E7DE196" type="volume">
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<mods:number id="84F772937840D96EF194B54504A9B96A">4103</mods:number>
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<mods:detail id="0282CEB14EA5CE752408A7E2D83EED28" type="issue">
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<mods:number id="9C13F5C16DAE184CC24357C850245CAB">5</mods:number>
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<mods:classification id="98861C760543099975C16C26EC9893C4">journal article</mods:classification>
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<mods:identifier id="C94FA594824F08264B091F2A69F156D6" type="DOI">10.11646/zootaxa.4103.5.4</mods:identifier>
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</mods:mods>
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<subSection id="E2BC2F4BC03BFFB1A7B1011443C4FD21" lastPageId="7" lastPageNumber="470" pageId="4" pageNumber="467" type="multiple">
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<treatment id="039A87B6C03BFFB1A7B1011443C4FD21" ID-DOI="http://doi.org/10.5281/zenodo.6073096" ID-GBIF-Taxon="127711644" ID-Zenodo-Dep="6073096" LSID="urn:lsid:plazi:treatment:039A87B6C03BFFB1A7B1011443C4FD21" httpUri="http://treatment.plazi.org/id/039A87B6C03BFFB1A7B1011443C4FD21" lastPageId="7" lastPageNumber="470" pageId="4" pageNumber="467">
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<subSubSection id="C329652BC03BFFB2A7B1011444C6FDB8" box="[151,1221,547,574]" pageId="4" pageNumber="467" type="nomenclature">
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<paragraph id="8B8C36A0C03BFFB2A7B1011444C6FDB8" blockId="4.[151,1221,547,574]" box="[151,1221,547,574]" pageId="4" pageNumber="467">
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<heading id="D0C481CCC03BFFB2A7B1011444C6FDB8" bold="true" box="[151,1221,547,574]" fontSize="11" level="1" pageId="4" pageNumber="467" reason="1">
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<keyStep id="30C72E05C03BFFB2A7B1011444C6FDB8" box="[151,1221,547,574]" pageId="4" pageNumber="467">
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<key id="FFA26191C03BFFB2A7B1011444C6FDB8" box="[151,1221,547,574]" pageId="4" pageNumber="467">
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<keyLead id="30C29595C03BFFB2A7B1011444C6FDB8" box="[151,1221,547,574]" pageId="4" pageNumber="467">
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<emphasis id="B947EAB2C03BFFB2A7B1011444C6FDB8" bold="true" box="[151,1221,547,574]" pageId="4" pageNumber="467">
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2. Morphological structures and taxonomic status of
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<taxonomicName id="4C334D23C03BFFB2A40F011343A6FDBB" box="[809,933,547,573]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03BFFB2A40F011343A6FDBB" bold="true" box="[809,933,547,573]" italics="true" pageId="4" pageNumber="467">Euconnus</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="4C334D23C03BFFB2A490011344BDFDB8" authority="Franz" authorityName="Franz" box="[950,1214,547,574]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03BFFB2A4900113446AFDBB" bold="true" box="[950,1129,547,573]" italics="true" pageId="4" pageNumber="467">Himaloconnus</emphasis>
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Franz
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</taxonomicName>
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)
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</emphasis>
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</keyLead>
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</key>
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</keyStep>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C329652BC03BFFB2A7B1015A4299FD20" pageId="4" pageNumber="467" type="discussion">
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<paragraph id="8B8C36A0C03BFFB2A7B1015A4299FD20" blockId="4.[151,1436,618,1939]" pageId="4" pageNumber="467">
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The general body shape of
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<taxonomicName id="4C334D23C03BFFB2A6FD015C4380FD05" box="[475,899,618,643]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
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<emphasis id="B947EAB2C03BFFB2A6FD015C424AFD05" box="[475,585,620,643]" italics="true" pageId="4" pageNumber="467">Euconnus</emphasis>
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(
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<emphasis id="B947EAB2C03BFFB2A57B015A4303FD05" box="[605,768,618,643]" italics="true" pageId="4" pageNumber="467">Himaloconnus</emphasis>
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)
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<emphasis id="B947EAB2C03BFFB2A433015A4380FD05" box="[789,899,618,643]" italics="true" pageId="4" pageNumber="467">shutjensis</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13082A25C03BFFB2A4B0015A43DDFD05" box="[918,990,618,643]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="4" pageNumber="467">Fig. 2</figureCitation>
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) is similar to that of many species of
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<taxonomicName id="4C334D23C03BFFB2A7B101BF4106FD20" box="[151,261,655,678]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03BFFB2A7B101BF4106FD20" box="[151,261,655,678]" italics="true" pageId="4" pageNumber="467">Euconnus</emphasis>
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</taxonomicName>
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, especially the subgenus
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<taxonomicName id="4C334D23C03BFFB2A50501BE4290FD21" box="[547,659,654,679]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03BFFB2A50501BE4290FD21" box="[547,659,654,679]" italics="true" pageId="4" pageNumber="467">Napochus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C329652BC03BFFB0A7E10182434BF822" lastPageId="6" lastPageNumber="469" pageId="4" pageNumber="467" type="description">
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<paragraph id="8B8C36A0C03BFFB2A7E1018243E1FA1D" blockId="4.[151,1436,618,1939]" pageId="4" pageNumber="467">
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The head capsule (
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<figureCitation id="13082A25C03BFFB2A6BF018241E0FD4D" box="[409,483,690,715]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="4" pageNumber="467">Figs 2</figureCitation>
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,
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<figureCitation id="13082A25C03BFFB2A6C90183420FFD4D" box="[495,524,691,715]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">13</figureCitation>
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) is divided into the posterior part ('neck' region) and anterior, exposed part by an occipital constriction broader than half width of head. The anterior part of the head is strongly elongate and slightly flattened; vertex distinctly demarcated from the 'neck' region and not bulging posterodorsally; frons subtrapezoidal; antennal insertions moderately broadly separated; compound eyes located anteriorly and the tempora are much longer than eyes. The tempora and genae are covered with thick bristles (
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<figureCitation id="13082A25C03BFFB2A4F000724459FCDD" box="[982,1114,834,859]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Figs 13, 14</figureCitation>
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). The posterior tentorial pits (
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<figureCitation id="13082A25C03BFFB2A7B9005640F1FCF9" box="[159,242,870,895]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 14</figureCitation>
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;
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<emphasis id="B947EAB2C03BFFB2A7D800574121FCF8" box="[254,290,871,894]" italics="true" pageId="4" pageNumber="467">ptp</emphasis>
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) are small, circular and located slightly in front of the transverse impression demarcating ventrally the anterior and posterior parts of the head capsule; the gular plate (
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<figureCitation id="13082A25C03BFFB2A44A00BA43C3FC25" box="[876,960,906,931]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 14</figureCitation>
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;
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<emphasis id="B947EAB2C03BFFB2A4EA00BC43EBFC25" box="[972,1000,908,931]" italics="true" pageId="4" pageNumber="467">gp</emphasis>
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) is broad and subtrapezoidal, but with its anterior portion adjacent to the tentorial pits narrowed and elongate. The submentum (
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<figureCitation id="13082A25C03BFFB2A394009E450AFC41" box="[1202,1289,942,967]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 13</figureCitation>
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;
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<emphasis id="B947EAB2C03BFFB2A23F009F4545FC40" box="[1305,1350,943,966]" italics="true" pageId="4" pageNumber="467">smn</emphasis>
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) is not demarcated laterally by sutures, bearing shallow and broad impression with presumably glandular openings (
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<figureCitation id="13082A25C03BFFB2A24900E240B6FB89" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 15</figureCitation>
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), and hypostomal ridges (
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<figureCitation id="13082A25C03BFFB2A6C600C64235FB89" box="[480,566,1014,1039]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 14</figureCitation>
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;
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<emphasis id="B947EAB2C03BFFB2A56200C6425FFB89" box="[580,604,1014,1039]" italics="true" pageId="4" pageNumber="467">hr</emphasis>
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) are long and posteriorly extending up to the level of posterior tentorial pits, but the ends of ridges are located laterally to the pits. Most of these characters can be found in
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<taxonomicName id="4C334D23C03BFFB2A200072C4597FBB5" box="[1318,1428,1052,1075]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
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<emphasis id="B947EAB2C03BFFB2A200072C4597FBB5" box="[1318,1428,1052,1075]" italics="true" pageId="4" pageNumber="467">Euconnus</emphasis>
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</taxonomicName>
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, except for the hypostomal ridges running laterad the posterior tentorial pits (in
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<taxonomicName id="4C334D23C03BFFB2A373070F44C0FBD0" box="[1109,1219,1087,1110]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B947EAB2C03BFFB2A373070F44C0FBD0" box="[1109,1219,1087,1110]" italics="true" pageId="4" pageNumber="467">Euconnus</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="A27457C9C03BFFB2A3F6070F4510FBD0" box="[1232,1299,1087,1110]" pageId="4" pageNumber="467" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C334D23C03BFFB2A203070E4596FBD1" box="[1317,1429,1086,1111]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A203070E4596FBD1" box="[1317,1429,1086,1111]" italics="true" pageId="4" pageNumber="467">Napochus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D23C03BFFB2A7B107524132FBFD" box="[151,305,1122,1147]" class="Insecta" family="Staphylinidae" genus="Filonapochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A7B107524132FBFD" box="[151,305,1122,1147]" italics="true" pageId="4" pageNumber="467">Filonapochus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D23C03BFFB2A666075241BAFBFD" box="[320,441,1122,1147]" class="Insecta" family="Staphylinidae" genus="Tetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A666075241BAFBFD" box="[320,441,1122,1147]" italics="true" pageId="4" pageNumber="467">Tetramelus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D23C03BFFB2A6ED0752427AFBFD" box="[459,633,1122,1147]" class="Insecta" family="Staphylinidae" genus="Paratetramelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A6ED0752427AFBFD" box="[459,633,1122,1147]" italics="true" pageId="4" pageNumber="467">Paratetramelus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C334D23C03BFFB2A5AF07544326FBFD" box="[649,805,1124,1147]" class="Insecta" family="Staphylinidae" genus="Heteroconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A5AF07544326FBFD" box="[649,805,1124,1147]" italics="true" pageId="4" pageNumber="467">Heteroconnus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B947EAB2C03BFFB2A478075243FBFBFD" box="[862,1016,1122,1147]" italics="true" pageId="4" pageNumber="467">Glabriconnus</emphasis>
|
||
the posterior ends of both ridges are very close to posterior tentorial pits, typically in front of each pit; in
|
||
<taxonomicName id="4C334D23C03BFFB2A49507B64463FB19" box="[947,1120,1158,1183]" class="Insecta" family="Staphylinidae" genus="Rhomboconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A49507B64463FB19" box="[947,1120,1158,1183]" italics="true" pageId="4" pageNumber="467">Rhomboconnus</emphasis>
|
||
</taxonomicName>
|
||
the ridges are connected at middle in front of the pits); also the presumably glandular impression of the submentum is unique for
|
||
<taxonomicName id="4C334D23C03BFFB2A7B107FE4139FB61" box="[151,314,1230,1255]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A7B107FE4139FB61" box="[151,314,1230,1255]" italics="true" pageId="4" pageNumber="467">Himaloconnus</emphasis>
|
||
</taxonomicName>
|
||
. A pair of enigmatic structures (
|
||
<figureCitation id="13082A25C03BFFB2A59207FE4309FB61" box="[692,778,1230,1255]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 14</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03BFFB2A43E07FF4325FB60" box="[792,806,1231,1254]" italics="true" pageId="4" pageNumber="467">?</emphasis>
|
||
) can be seen at each side of the prementum; these are nearly circular flat lobes which can represent modified (i.e., enlarged and lacking microtrichia) lateral lobes of the hypopharynx. However, in the studied specimens the mandibles are tightly closed and any attempts to open them could easily result in damaging the mouthparts. Therefore, it was not possible to see where these lobes are inserted; additional material available for disarticulation is required to clarify the nature of these unusual structures, which are not known in any species of
|
||
<taxonomicName id="4C334D23C03BFFB2A52606B242A0FA1D" box="[512,675,1410,1435]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
|
||
studied so far by the author.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C03BFFB2A7E10696423FF9AD" blockId="4.[151,1436,618,1939]" pageId="4" pageNumber="467">
|
||
The antennae (
|
||
<figureCitation id="13082A25C03BFFB2A654069641BAFA39" box="[370,441,1446,1471]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="4" pageNumber="467">Fig. 4</figureCitation>
|
||
) of
|
||
<taxonomicName id="4C334D23C03BFFB2A6CA06974390FA39" box="[492,915,1446,1471]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C03BFFB2A6CA06974259FA38" box="[492,602,1447,1470]" italics="true" pageId="4" pageNumber="467">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03BFFB2A54806964311FA39" box="[622,786,1446,1471]" italics="true" pageId="4" pageNumber="467">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03BFFB2A40306964390FA39" box="[805,915,1446,1471]" italics="true" pageId="4" pageNumber="467">shutjensis</emphasis>
|
||
</taxonomicName>
|
||
are moderately long, with scape and pedicel elongate and proximal flagellomeres subcylindrical and compactly assembled; the club is composed of three antennomeres subequal in width and loosely assembled. The antennae resemble those of
|
||
<taxonomicName id="4C334D23C03BFFB2A38006DE4515F981" box="[1190,1302,1518,1543]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A38006DE4515F981" box="[1190,1302,1518,1543]" italics="true" pageId="4" pageNumber="467">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, except for being three-, and not four-segmented.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C03BFFB2A7E1050642B8F93D" blockId="4.[151,1436,618,1939]" pageId="4" pageNumber="467">
|
||
The prothorax of
|
||
<taxonomicName id="4C334D23C03BFFB2A6AB0507432DF9C9" box="[397,814,1590,1615]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C03BFFB2A6AB050741F8F9C8" box="[397,507,1591,1614]" italics="true" pageId="4" pageNumber="467">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03BFFB2A52A050642B3F9C9" box="[524,688,1590,1615]" italics="true" pageId="4" pageNumber="467">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03BFFB2A5E60506432DF9C9" box="[704,814,1590,1615]" italics="true" pageId="4" pageNumber="467">shutjensis</emphasis>
|
||
</taxonomicName>
|
||
in dorsal view (
|
||
<figureCitation id="13082A25C03BFFB2A4C30506442CF9C9" box="[997,1071,1590,1615]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="4" pageNumber="467">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="13082A25C03BFFB2A31D05074454F9C9" box="[1083,1111,1591,1615]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">13</figureCitation>
|
||
) is short and subtrapezoidal, with sides strongly rounded and convergent anteriorly, the pronotum is broadest near base. The pronotal base bears one pair of small but distinct lateral antebasal pits (
|
||
<figureCitation id="13082A25C03BFFB2A5E9054E4321F911" box="[719,802,1662,1687]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 13</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03BFFB2A40B054E4352F911" box="[813,849,1662,1687]" italics="true" pageId="4" pageNumber="467">lab</emphasis>
|
||
). The general shape of pronotum is similar to that of
|
||
<taxonomicName id="4C334D23C03BFFB2A7B105924104F93D" box="[151,263,1698,1723]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A7B105924104F93D" box="[151,263,1698,1723]" italics="true" pageId="4" pageNumber="467">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, except for the strongly rounded sides.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C03BFFB2A7E105F641E6F815" blockId="4.[151,1436,618,1939]" pageId="4" pageNumber="467">
|
||
The prosternum (
|
||
<figureCitation id="13082A25C03BFFB2A6AD05F641DDF959" box="[395,478,1734,1759]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 16</figureCitation>
|
||
) is short; the prosternal process (
|
||
<figureCitation id="13082A25C03BFFB2A47C05F643ACF959" box="[858,943,1734,1759]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 16</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03BFFB2A49A05F743E0F958" box="[956,995,1735,1758]" italics="true" pageId="4" pageNumber="467">psp</emphasis>
|
||
) rudimentary, carinate; the notosternal sutures (
|
||
<figureCitation id="13082A25C03BFFB2A7DF05DA4153F885" box="[249,336,1770,1795]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 16</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03BFFB2A67905DC4180F885" box="[351,387,1772,1795]" italics="true" pageId="4" pageNumber="467">nss</emphasis>
|
||
) seem complete (but they are only partly visible in the studied specimen); the hypomeral ridges (
|
||
<figureCitation id="13082A25C03BFFB2A7CD043E413CF8A1" box="[235,319,1806,1831]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="4" pageNumber="467">Fig. 16</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C03BFFB2A66D043E4173F8A1" box="[331,368,1806,1831]" italics="true" pageId="4" pageNumber="467">hyr</emphasis>
|
||
) are obliterated anteriorly and posteriorly (i.e., not connected to notosternal sutures or anterior prothoracic margin anteriorly and not connected to the posterior hypomeral margin posteriorly); the sides of prothorax are covered with thick bristles. These are characters similar to those known in
|
||
<taxonomicName id="4C334D23C03BFFB2A38104664514F8E9" box="[1191,1303,1878,1903]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C03BFFB2A38104664514F8E9" box="[1191,1303,1878,1903]" italics="true" pageId="4" pageNumber="467">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, except for incomplete hypomeral ridges.
|
||
</paragraph>
|
||
<caption id="DF4C6628C03AFFB3A7B104324189F834" httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="5" pageNumber="468" targetBox="[151,1436,193,1772]" targetPageId="5">
|
||
<paragraph id="8B8C36A0C03AFFB3A7B104324189F834" blockId="5.[151,1436,1794,1970]" pageId="5" pageNumber="468">
|
||
<emphasis id="B947EAB2C03AFFB3A7B104324150F89E" bold="true" box="[151,339,1794,1817]" pageId="5" pageNumber="468">FIGURES 13–16.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C334D23C03AFFB3A55E04344435F89F" authority="Franz." authorityName="Franz." box="[632,1078,1795,1817]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C03AFFB3A55E043442DFF89F" box="[632,732,1796,1817]" italics="true" pageId="5" pageNumber="468">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C03AFFB3A5CC0433437DF89F" box="[746,894,1795,1817]" italics="true" pageId="5" pageNumber="468">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C03AFFB3A4AA043343F3F89F" box="[908,1008,1795,1817]" italics="true" pageId="5" pageNumber="468">shutjensis</emphasis>
|
||
Franz.
|
||
</taxonomicName>
|
||
Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4C6628C039FFB0A7B107FC451CFAA6" httpUri="https://zenodo.org/record/263217/files/figure.png" pageId="6" pageNumber="469" targetBox="[151,1436,193,1206]" targetPageId="6">
|
||
<paragraph id="8B8C36A0C039FFB0A7B107FC451CFAA6" blockId="6.[151,1436,1228,1312]" pageId="6" pageNumber="469">
|
||
<emphasis id="B947EAB2C039FFB0A7B107FC415AFB64" bold="true" box="[151,345,1228,1251]" pageId="6" pageNumber="469">FIGURES 17–19.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C334D23C039FFB0A5B207FE4467FB64" authority="Franz." authorityName="Franz." box="[660,1124,1228,1251]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C039FFB0A5B207FE42FBFB65" box="[660,760,1230,1251]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C039FFB0A42A07FC43A3FB64" box="[780,928,1228,1250]" italics="true" pageId="6" pageNumber="469">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C039FFB0A49207FC441BFB64" box="[948,1048,1228,1250]" italics="true" pageId="6" pageNumber="469">shutjensis</emphasis>
|
||
Franz.
|
||
</taxonomicName>
|
||
Pterothorax and abdomen in ventral view (17); right (18) and left (19) mesocoxal cavity in ventral view. Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; st3–8, abdominal sternites 3–8; v3, metaventrite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C36A0C039FFB0A7E1067C43D0FA73" blockId="6.[151,1437,1356,2029]" pageId="6" pageNumber="469">
|
||
The mesoventral structures of
|
||
<taxonomicName id="4C334D23C039FFB0A53A067E43BEFAE3" box="[540,957,1356,1381]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C039FFB0A53A067E4289FAE3" box="[540,650,1358,1381]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C039FFB0A5BD067C433DFAE3" box="[667,830,1356,1381]" italics="true" pageId="6" pageNumber="469">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C039FFB0A469067C43BEFAE3" box="[847,957,1356,1381]" italics="true" pageId="6" pageNumber="469">shutjensis</emphasis>
|
||
</taxonomicName>
|
||
differ from those of any hitherto reviewed subgenus of
|
||
<taxonomicName id="4C334D23C039FFB0A60306424190FA0F" box="[293,403,1394,1417]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A60306424190FA0F" box="[293,403,1394,1417]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
in the mesoventral intercoxal process (
|
||
<figureCitation id="13082A25C039FFB0A468064043A3FA0F" box="[846,928,1392,1417]" captionStart="FIGURES 17 – 19" captionStartId="6.[151,264,1228,1251]" captionTargetBox="[151,1436,193,1206]" captionTargetId="figure@6.[151,1436,193,1207]" captionTargetPageId="6" captionText="FIGURES 17 – 19. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Pterothorax and abdomen in ventral view (17); right (18) and left (19) mesocoxal cavity in ventral view. Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; st 3 – 8, abdominal sternites 3 – 8; v 3, metaventrite." httpUri="https://zenodo.org/record/263217/files/figure.png" pageId="6" pageNumber="469">Fig. 17</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C039FFB0A48B064243E4FA0F" box="[941,999,1394,1417]" italics="true" pageId="6" pageNumber="469">msvp</emphasis>
|
||
), which in a cross-section is T-shaped, i.e., its ventral surface is expanded laterally, much broader than in any
|
||
<taxonomicName id="4C334D23C039FFB0A4C006A64457FA2B" box="[998,1108,1430,1453]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A4C006A64457FA2B" box="[998,1108,1430,1453]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
, the mesoventral process is broadened posteriorly and coarsely sculptured. The anterior portion of mesoventrite, where lateral impressions functioning as procoxal rests (
|
||
<figureCitation id="13082A25C039FFB0A6CF06EC423FFA73" box="[489,572,1500,1525]" captionStart="FIGURES 17 – 19" captionStartId="6.[151,264,1228,1251]" captionTargetBox="[151,1436,193,1206]" captionTargetId="figure@6.[151,1436,193,1207]" captionTargetPageId="6" captionText="FIGURES 17 – 19. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Pterothorax and abdomen in ventral view (17); right (18) and left (19) mesocoxal cavity in ventral view. Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; st 3 – 8, abdominal sternites 3 – 8; v 3, metaventrite." httpUri="https://zenodo.org/record/263217/files/figure.png" pageId="6" pageNumber="469">Fig. 17</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C039FFB0A56E06EE426EFA73" box="[584,621,1502,1525]" italics="true" pageId="6" pageNumber="469">pcr</emphasis>
|
||
) are located, is unusually short.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C039FFB0A7E10530417FF9E7" blockId="6.[151,1437,1356,2029]" pageId="6" pageNumber="469">
|
||
The mesoventral cavities (
|
||
<figureCitation id="13082A25C039FFB0A6CB0530426EF99F" box="[493,621,1536,1561]" captionStart="FIGURES 17 – 19" captionStartId="6.[151,264,1228,1251]" captionTargetBox="[151,1436,193,1206]" captionTargetId="figure@6.[151,1436,193,1207]" captionTargetPageId="6" captionText="FIGURES 17 – 19. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Pterothorax and abdomen in ventral view (17); right (18) and left (19) mesocoxal cavity in ventral view. Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; st 3 – 8, abdominal sternites 3 – 8; v 3, metaventrite." httpUri="https://zenodo.org/record/263217/files/figure.png" pageId="6" pageNumber="469">Figs 18–19</figureCitation>
|
||
) of
|
||
<taxonomicName id="4C334D23C039FFB0A5BC05324439F99F" box="[666,1082,1536,1561]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C039FFB0A5BC0532430BF99F" box="[666,776,1538,1561]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C039FFB0A43F053043BEF99F" box="[793,957,1536,1561]" italics="true" pageId="6" pageNumber="469">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C039FFB0A4EA05304439F99F" box="[972,1082,1536,1561]" italics="true" pageId="6" pageNumber="469">shutjensis</emphasis>
|
||
</taxonomicName>
|
||
are unusual in having posterior 'curtains' composed of modified setae; similar fringes are located on mesocoxae; such structures are not known in other scydmaenines.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C039FFB0A7E1055C436EF977" blockId="6.[151,1437,1356,2029]" pageId="6" pageNumber="469">
|
||
The metaventral intercoxal process of
|
||
<taxonomicName id="4C334D23C039FFB0A552055E4417F903" box="[628,1044,1644,1669]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C039FFB0A552055E42E1F903" box="[628,738,1646,1669]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C039FFB0A5D4055C4396F903" box="[754,917,1644,1669]" italics="true" pageId="6" pageNumber="469">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C039FFB0A480055C4417F903" box="[934,1044,1644,1669]" italics="true" pageId="6" pageNumber="469">shutjensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13082A25C039FFB0A305055C4475F903" box="[1059,1142,1644,1669]" captionStart="FIGURES 17 – 19" captionStartId="6.[151,264,1228,1251]" captionTargetBox="[151,1436,193,1206]" captionTargetId="figure@6.[151,1436,193,1207]" captionTargetPageId="6" captionText="FIGURES 17 – 19. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Pterothorax and abdomen in ventral view (17); right (18) and left (19) mesocoxal cavity in ventral view. Abbreviations: msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest; st 3 – 8, abdominal sternites 3 – 8; v 3, metaventrite." httpUri="https://zenodo.org/record/263217/files/figure.png" pageId="6" pageNumber="469">Fig. 17</figureCitation>
|
||
;
|
||
<emphasis id="B947EAB2C039FFB0A3A4055E44BBF903" box="[1154,1208,1646,1669]" italics="true" pageId="6" pageNumber="469">mtvp</emphasis>
|
||
), is clearly different from structures found previously in
|
||
<taxonomicName id="4C334D23C039FFB0A50C05A2429BF92F" box="[554,664,1682,1705]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A50C05A2429BF92F" box="[554,664,1682,1705]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
. The process is narrow and forming a pair of long spines touching at middle and projecting posteriorly, so that the metacoxae are narrowly separated. In
|
||
<taxonomicName id="4C334D23C039FFB0A370058644C7F94B" box="[1110,1220,1718,1741]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A370058644C7F94B" box="[1110,1220,1718,1741]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
the metacoxae are distinctly separated and the metaventral process lacks the spines.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C039FFB0A7E105CC44E9F8BE" blockId="6.[151,1437,1356,2029]" pageId="6" pageNumber="469">
|
||
The elytra (
|
||
<figureCitation id="13082A25C039FFB0A66D05CC4195F893" box="[331,406,1788,1813]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="6" pageNumber="469">Figs 4</figureCitation>
|
||
,
|
||
<figureCitation id="13082A25C039FFB0A68205CD41C2F893" box="[420,449,1789,1813]" captionStart="FIGURES 13 – 16" captionStartId="5.[151,264,1794,1817]" captionTargetBox="[151,1436,193,1772]" captionTargetId="figure@5.[151,1436,193,1773]" captionTargetPageId="5" captionText="FIGURES 13 – 16. Morphological structures of Euconnus (Himaloconnus) shutjensis Franz. Head, prothorax and elytral base in dorsal view (13); head in ventral view (14); impression of submentum (15); prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; fr, frons; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hyr, hypomeral ridge; lab, lateral antebasal pit; mn, mentum; mxp, maxillary palp; nss, notosternal suture; occ, occipital constriction; psp, prosternal process; ptp, posterior tentorial pit; smn, submentum; vt, vertex;?, unknown structure of mouthparts." httpUri="https://zenodo.org/record/263216/files/figure.png" pageId="6" pageNumber="469">13</figureCitation>
|
||
) are oval, elongate, each with two distinct basal foveae, each with sparse setae inside. Foveae known in all previously studied subgenera of
|
||
<taxonomicName id="4C334D23C039FFB0A5CB04124358F8BF" box="[749,859,1826,1849]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A5CB04124358F8BF" box="[749,859,1826,1849]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
, when well-developed, are asetose.
|
||
</paragraph>
|
||
<paragraph id="8B8C36A0C039FFB0A7E10474434BF822" blockId="6.[151,1437,1356,2029]" pageId="6" pageNumber="469">
|
||
The aedeagus (
|
||
<figureCitation id="13082A25C039FFB0A65A047441E4F8DB" box="[380,487,1860,1885]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1554,1577]" captionTargetBox="[158,1426,180,1516]" captionTargetId="figure@2.[151,1436,170,1533]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Euconnus (Napoconnus) kubienai Franz (1 – 3); Euconnus (Himaloconnus) shutjensis Franz (4 – 6); and Euconnus (Nepaloconnus) khumbuensis Franz (7 – 9). Dorsal habitus of holotype male (1, 4, 7) and aedeagus in ventral (2, 5, 8) and lateral (3, 6, 9) views." httpUri="https://zenodo.org/record/263214/files/figure.png" pageId="6" pageNumber="469">Figs 5–6</figureCitation>
|
||
) of
|
||
<taxonomicName id="4C334D23C039FFB0A507047643CCF8DB" box="[545,975,1860,1885]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="shutjensis" subGenus="Himaloconnus">
|
||
<emphasis id="B947EAB2C039FFB0A5070476428CF8DB" box="[545,655,1862,1885]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
(
|
||
<emphasis id="B947EAB2C039FFB0A58104744349F8DB" box="[679,842,1860,1885]" italics="true" pageId="6" pageNumber="469">Himaloconnus</emphasis>
|
||
)
|
||
<emphasis id="B947EAB2C039FFB0A447047443CCF8DB" box="[865,975,1860,1885]" italics="true" pageId="6" pageNumber="469">shutjensis</emphasis>
|
||
</taxonomicName>
|
||
is symmetrical, with free and slender parameres, similar to copulatory organs in many genera of Glandulariini, including
|
||
<taxonomicName id="4C334D23C039FFB0A364045A44B3F807" box="[1090,1200,1898,1921]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A364045A44B3F807" box="[1090,1200,1898,1921]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
; it does not bear any unique characters that could be used to define
|
||
<taxonomicName id="4C334D23C039FFB0A5B804BC4342F823" box="[670,833,1932,1957]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A5B804BC4342F823" box="[670,833,1932,1957]" italics="true" pageId="6" pageNumber="469">Himaloconnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C329652BC039FFB1A7E1048042CCFE9B" lastPageId="7" lastPageNumber="470" pageId="6" pageNumber="469" type="discussion">
|
||
<paragraph id="8B8C36A0C039FFB1A7E1048042CCFE9B" blockId="6.[151,1437,1356,2029]" lastBlockId="7.[151,1437,151,680]" lastPageId="7" lastPageNumber="470" pageId="6" pageNumber="469">
|
||
<emphasis id="B947EAB2C039FFB0A7E10480415CF84F" bold="true" box="[199,351,1968,1993]" pageId="6" pageNumber="469">Conclusions.</emphasis>
|
||
Morphological structures of
|
||
<taxonomicName id="4C334D23C039FFB0A59F0480435FF84F" box="[697,860,1968,1993]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A59F0480435FF84F" box="[697,860,1968,1993]" italics="true" pageId="6" pageNumber="469">Himaloconnus</emphasis>
|
||
</taxonomicName>
|
||
are similar to those of
|
||
<taxonomicName id="4C334D23C039FFB0A35A048244E9F84F" box="[1148,1258,1970,1993]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A35A048244E9F84F" box="[1148,1258,1970,1993]" italics="true" pageId="6" pageNumber="469">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
, especially the subgenus
|
||
<taxonomicName id="4C334D23C039FFB0A62A04E4417FF86B" box="[268,380,2004,2029]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C039FFB0A62A04E4417FF86B" box="[268,380,2004,2029]" italics="true" pageId="6" pageNumber="469">Napochus</emphasis>
|
||
</taxonomicName>
|
||
, but the narrow metaventral intercoxal process with a pair of posteriorly directed spines excludes this taxon from
|
||
<taxonomicName id="4C334D23C038FFB1A69203A94221FF36" box="[436,546,153,176]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C038FFB1A69203A94221FF36" box="[436,546,153,176]" italics="true" pageId="7" pageNumber="470">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
. The setose basal elytral foveae are not known in any
|
||
<taxonomicName id="4C334D23C038FFB1A3AE03A944F5FF36" box="[1160,1270,153,176]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C038FFB1A3AE03A944F5FF36" box="[1160,1270,153,176]" italics="true" pageId="7" pageNumber="470">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
revised so far, the mouthparts with enigmatic lateral lobes and mesocoxal cavities with setal 'curtains' are unique for
|
||
<taxonomicName id="4C334D23C038FFB1A7B103EF4139FF7E" box="[151,314,223,248]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C038FFB1A7B103EF4139FF7E" box="[151,314,223,248]" italics="true" pageId="7" pageNumber="470">Himaloconnus</emphasis>
|
||
</taxonomicName>
|
||
, and the broad mesoventral intercoxal process is also not known in
|
||
<taxonomicName id="4C334D23C038FFB1A35C03D144EBFF7E" box="[1146,1256,225,248]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C038FFB1A35C03D144EBFF7E" box="[1146,1256,225,248]" italics="true" pageId="7" pageNumber="470">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
. Consequently,
|
||
<taxonomicName id="4C334D23C038FFB1A7B102344139FE9B" box="[151,314,260,285]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C038FFB1A7B102344139FE9B" box="[151,314,260,285]" italics="true" pageId="7" pageNumber="470">Himaloconnus</emphasis>
|
||
</taxonomicName>
|
||
is here elevated to the generic rank.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C329652BC038FFB1A7E1021743C4FD21" pageId="7" pageNumber="470" type="diagnosis">
|
||
<paragraph id="8B8C36A0C038FFB1A7E1021743C4FD21" blockId="7.[151,1437,151,680]" pageId="7" pageNumber="470">
|
||
<emphasis id="B947EAB2C038FFB1A7E1021743ABFEC6" bold="true" box="[199,936,295,320]" pageId="7" pageNumber="470">
|
||
Emended diagnosis of the genus
|
||
<taxonomicName id="4C334D23C038FFB1A561021742F1FEC6" box="[583,754,295,320]" class="Insecta" family="Staphylinidae" genus="Himaloconnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C038FFB1A561021742F1FEC6" bold="true" box="[583,754,295,320]" italics="true" pageId="7" pageNumber="470">Himaloconnus</emphasis>
|
||
</taxonomicName>
|
||
, status revised.
|
||
</emphasis>
|
||
Body approximately
|
||
<taxonomicName id="4C334D23C038FFB1A3BC02174509FEC6" box="[1178,1290,295,320]" class="Insecta" family="Staphylinidae" genus="Napochus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B947EAB2C038FFB1A3BC02174509FEC6" box="[1178,1290,295,320]" italics="true" pageId="7" pageNumber="470">Napochus</emphasis>
|
||
</taxonomicName>
|
||
-like, i.e., the head with long tempora, pronotum subtrapezoidal, broadest near base and strongly narrowing anteriorly, and oval elytra, but the sides of pronotum strongly rounded; head and pronotum with thick bristles. Antenna with distinct club composed of antennomeres IX–XI; hypostomal ridges extending to the level of posterior tentorial pits, but their ends located laterally in relation to pits; mouthparts with rounded lateral lobes flanking prementum, which are presumably associated with hypopharynx; pronotum with one pair of small lateral pits, lacking sublateral carinae; hypomeral ridges incomplete, obliterated anteriorly and posteriorly; prosternum with rudimentary prosternal carina; mesoventral intercoxal process carinate, T-shaped in cross-section, broad and strongly elevated, broadening posteriorly, anteriorly not connected to anterior ridge of mesoventrite; metaventral intercoxal process narrowly separating metacoxae, with a pair of posteriorly directed, pointed spines adjacent at middle; each elytron with two distinct but small and setose basal foveae; aedeagus with free parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |