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<document id="A6D0E08E13F6F65B686451D3DEE50B42" ID-DOI="10.37520/aemnp.2024.017" ID-ISSN="1804-6487" ID-Zenodo-Dep="14655612" ID-ZooBank="E3DA805D-15F4-46DF-93C2-85C512C1ECBA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736957711734" checkinUser="felipe" docAuthor="Yamane, Seiki, Hosoishi, Shingo &amp; Ito, Fuminori" docDate="2024" docId="0380D165191AFFC7FC74FB4F531C3464" docLanguage="en" docName="ActaEntMusNatPra.64.2.249-267.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 64 (2)" docSource="https://doi.org/10.37520/aemnp.2024.017" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Anochetus Mayr 1861" docType="treatment" docVersion="2" lastPageNumber="253" masterDocId="FFB9A91D1919FFC3FF9AFFCE56033C6B" masterDocTitle="Taxonomic study on the queens of the Japanese ponerine genera, with a redescription of Ectomomyrmex horni restituted as a valid species (Hymenoptera: Formicidae)" masterLastPageNumber="267" masterPageNumber="249" pageNumber="252" updateTime="1736958290155" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="6285CA6EAE9162A1B1FF95FDA4320ADA">Taxonomic study on the queens of the Japanese ponerine genera, with a redescription of Ectomomyrmex horni restituted as a valid species (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="33C7BAE804B186A006E03CF7EF639332">Yamane, Seiki</mods:namePart>
<mods:affiliation id="2EE68280B6C5BD3E243EECB660437384">Haruyama 1054 -</mods:affiliation>
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<mods:namePart id="34C81622444A58AE10D3E2683EE7ED60">Hosoishi, Shingo</mods:namePart>
<mods:affiliation id="9463B41EE6224D9B41BFCFFA7B0C2802">Institute of Tropical Agriculture, Kyushu University, Motooka 744, Nishi-ku, Fukuoka 819 - 0395, Japan</mods:affiliation>
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<mods:namePart id="6AB02DF5F101CCF96D9DC3FE1C6FC28F">Ito, Fuminori</mods:namePart>
<mods:affiliation id="7827D3769433AF9A7F419D92CFF7C144">Faculty of Agriculture, Kagawa University, Ikenobe, Miki, Kagawa Pref. 761 - 0795, Japan</mods:affiliation>
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<mods:title id="CB2A77A497BAA9A7F42A09BEB60232E9">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="2176B4F6034A59FFFEF4727D3120BCB9">2024</mods:date>
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<mods:number id="30C2BDCDC50994A0B426AD279AF5E32D">2024-10-01</mods:number>
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<mods:number id="1F0F0B62D83098283F401CBC554E9773">64</mods:number>
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<treatment id="0380D165191AFFC7FC74FB4F531C3464" LSID="urn:lsid:plazi:treatment:0380D165191AFFC7FC74FB4F531C3464" httpUri="http://treatment.plazi.org/id/0380D165191AFFC7FC74FB4F531C3464" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252">
<subSubSection id="C33333F8191AFFC0FC74FB4F52EA38F3" box="[1006,1257,1153,1176]" pageId="3" pageNumber="252" type="nomenclature">
<paragraph id="8B966073191AFFC0FC74FB4F52EA38F3" blockId="3.[1006,1257,1153,1203]" box="[1006,1257,1153,1176]" pageId="3" pageNumber="252">
<heading id="D0DED71F191AFFC0FC74FB4F52EA38F3" box="[1006,1257,1153,1176]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="252" reason="3">
<taxonomicName id="4C291BF0191AFFC0FC74FB4F52EA38F3" ID-CoL="WNN" authority="Mayr, 1861" authorityName="Mayr" authorityYear="1861" box="[1006,1257,1153,1176]" class="Insecta" family="Formicidae" genus="Anochetus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="B95DBC61191AFFC0FC74FB4F52EA38F3" bold="true" box="[1006,1257,1153,1176]" pageId="3" pageNumber="252">
<emphasis id="B95DBC61191AFFC0FC74FB4F526038F3" bold="true" box="[1006,1123,1153,1176]" italics="true" pageId="3" pageNumber="252">Anochetus</emphasis>
Mayr, 1861
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33333F8191AFFC7FBBAFB6E57E83B84" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252" type="description">
<paragraph id="8B966073191AFFC0FBBAFB6E52BB38D8" blockId="3.[1006,1257,1153,1203]" box="[1056,1208,1184,1203]" pageId="3" pageNumber="252">
(
<figureCitation id="13127CF6191AFFC0FBBCFB6E526F38D8" box="[1062,1132,1184,1203]" captionStart="Fig" captionStartId="4.[151,182,1459,1478]" captionTargetBox="[151,1439,189,1445]" captionTargetId="figure-248@4.[148,1444,186,1448]" captionTargetPageId="4" captionText="Fig. 2. Habitus (queen) in profile view.A – Hypoponera beppin Terayama, 1999 (Yakusugiland, Yaku-shima, Ôsumi Is., Kagoshima-ken, 22.viii.2006, Y. Kuboki leg., Litter No. 3); B – Anochetus shohki Terayama, 1996 (Oganzaki, Sakieda, Ishigaki-jima, Yaeyama Is., Okinawa-ken, 10.iv.2022, H. Matsumura leg., MH97, emeraged in vii.2022 from captive colony); C – Brachyponera luteipes (Mayr, 1862) (Shirahama-rindo, Iriomote-jima, Okinawa-ken, 11.xi.2018, Sk.Yamane leg., JP18-SKY-154); D – Euponera sakishimensis (Terayama, 1999) (Funaura, Iriomote-jima,Yaeyana Is., Okinawa-ken, 9.v.1979, K. Ogata leg.); E – Leptogenys confucii Forel, 1912 (Mikyo, Tokuno-shima, Amami Is., Kagoshima-ken, 6.iii.2010, Sk. Yamane leg., M10-SKY-12); F – Odontomachus cf. monticola Emery, 1892 (Miyanoura, Yaku-shima, Ôsumi Is., Kagoshima-ken, 16.ix.2008, collector unknown)." figureDoi="http://doi.org/10.5281/zenodo.14655616" httpUri="https://zenodo.org/record/14655616/files/figure.png" pageId="3" pageNumber="252">Figs 2B</figureCitation>
,
<figureCitation id="13127CF6191AFFC0FBEFFB6E528C38D8" box="[1141,1167,1184,1203]" captionStart="Fig" captionStartId="7.[151,182,1326,1345]" captionTargetBox="[157,1431,189,1296]" captionTargetId="figure-347@7.[151,1438,183,1302]" captionTargetPageId="7" captionText="Fig. 4. Habitus (queen) in dorsal view. A – Hypoponera beppin Terayama, 1999; B – Anochetus shohki Terayama, 1996; C – Brachyponera luteipes (Mayr, 1862); D – Euponera sakishimensis (Terayama, 1999); E – Leptogenys confucii Forel, 1912; F – Odontomachus cf. monticola Emery, 1892. (Collection data same as in Fig. 2)." figureDoi="http://doi.org/10.5281/zenodo.14655624" httpUri="https://zenodo.org/record/14655624/files/figure.png" pageId="3" pageNumber="252">4B</figureCitation>
,
<figureCitation id="13127CF6191AFFC0FB02FB6E52B138D8" box="[1176,1202,1184,1203]" captionStart="Fig" captionStartId="10.[152,183,1890,1909]" captionTargetBox="[153,1429,190,1868]" captionTargetId="figure-13@10.[151,1436,189,1875]" captionTargetPageId="10" captionText="Fig. 6. Head (queen) in full-face view. A – Hypoponera beppin Terayama, 1999; B – Anochetus shohki Terayama, 1996; C – Brachyponera luteipes (Mayr, 1862); D – Euponera sakishimensis (Terayama, 1999); E – Leptogenys confucii Forel, 1912; F – Odontomachus cf. monticola Emery, 1892. (Collection data same as in Fig. 2)." figureDoi="http://doi.org/10.5281/zenodo.14655630" httpUri="https://zenodo.org/record/14655630/files/figure.png" pageId="3" pageNumber="252">6B</figureCitation>
)
</paragraph>
<paragraph id="8B966073191AFFC0FCA6FB0555AA3909" blockId="3.[828,1437,1227,2082]" pageId="3" pageNumber="252">
<emphasis id="B95DBC61191AFFC0FCA6FB0555C93889" bold="true" box="[828,970,1227,1250]" pageId="3" pageNumber="252">Recognition.</emphasis>
Relatively small ants with total length measuring
<quantity id="4CD1CD96191AFFC0FCE7FB2552013969" box="[893,1026,1259,1282]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="3" pageNumber="252" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6–3.8 mm</quantity>
; posterior margin of head in full-face view deeply and broadly emarginate; mandible linear and very long, apically strongly curved inwardly; petiole scale-like.
</paragraph>
<paragraph id="8B966073191AFFC7FCA6FAA557DF3AA4" blockId="3.[828,1437,1227,2082]" lastBlockId="4.[151,760,1656,2063]" lastPageId="4" lastPageNumber="253" pageId="3" pageNumber="252">
<emphasis id="B95DBC61191AFFC0FCA6FAA555C439E9" bold="true" box="[828,967,1387,1410]" pageId="3" pageNumber="252">Description.</emphasis>
Relatively small ants with total length measuring ca.
<quantity id="4CD1CD96191AFFC0FC06FA45521939C9" box="[924,1050,1419,1442]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="3" pageNumber="252" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6–3.8 mm</quantity>
and head width 0.94–1.00 mm in the single Japanese species
<taxonomicName id="4C291BF0191AFFC0FBA4FA65535A39A9" authority="Terayama, 1996" authorityName="Terayama" authorityYear="1996" box="[1086,1369,1451,1474]" class="Insecta" family="Formicidae" genus="Anochetus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="252" phylum="Arthropoda" rank="species" species="shohki">
<emphasis id="B95DBC61191AFFC0FBA4FA6552A139A9" box="[1086,1186,1451,1474]" italics="true" pageId="3" pageNumber="252">A. shohki</emphasis>
Terayama, 1996
</taxonomicName>
. Head as long as broad, broadest at level of eye, with posterior margin deeply and broadly emarginate; lateral margin behind eye shallowly concave; nuchal carina (occipital carina) running behind ‘vertex’ continuously curved, without anterior V-shaped infolding; median furrow (line) absent on vertex. Eye with rather long sparse hairs. Ocelli situated near posterior margin of head; distance between posterior ocellus and posterior margin of head about as long as distance between posterior ocelli. Mandible linear and long, inserted medially on front of cranium, apically strongly curved inwardly; in the single Japanese species apical portion with two major teeth that are arranged vertically; the lower tooth with additional smaller tooth near base. Metanotum flanked by deep anterior and posterior furrows. Opening of propodeal spiracle oval. Petiole subsquamiform, in profile view tapering apicad. Constriction between gastral segments I and II moderate; pretergite and presternite of segment II transversely striate; cinctus distinctly defined, cross-ribbed (forming series of elongate punctures). Midtibia without apical spur; hindtibia with pectinate spur only. For the structure of mesosoma, see also TERAYAMA (1996: fig. 25).
</paragraph>
<caption id="DF5630FB191DFFC7FF0DFA7D530A3A20" ID-DOI="http://doi.org/10.5281/zenodo.14655616" ID-Zenodo-Dep="14655616" httpUri="https://zenodo.org/record/14655616/files/figure.png" pageId="4" pageNumber="253" startId="4.[151,182,1459,1478]" targetBox="[151,1439,189,1445]" targetPageId="4" targetType="figure">
<paragraph id="8B966073191DFFC7FF0DFA7D530A3A20" blockId="4.[151,1437,1459,1611]" pageId="4" pageNumber="253">
Fig. 2. Habitus (queen) in profile view.A –
<taxonomicName id="4C291BF0191DFFC7FD91FA7D554739AD" authority="Terayama, 1999" authorityName="Terayama" authorityYear="1999" box="[523,836,1459,1478]" class="Insecta" family="Formicidae" genus="Hypoponera" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="beppin">
<emphasis id="B95DBC61191DFFC7FD91FA7D54B539AD" box="[523,694,1459,1478]" italics="true" pageId="4" pageNumber="253">Hypoponera beppin</emphasis>
Terayama, 1999
</taxonomicName>
(Yakusugiland, Yaku-shima, Ôsumi Is., Kagoshima-ken, 22.viii.2006, Y. Kuboki leg., Litter No. 3); B –
<taxonomicName id="4C291BF0191DFFC7FE29FA0054D5398A" authority="Terayama, 1996" authorityName="Terayama" authorityYear="1996" box="[435,726,1486,1505]" class="Insecta" family="Formicidae" genus="Anochetus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="shohki">
<emphasis id="B95DBC61191DFFC7FE29FA00544A398A" box="[435,585,1486,1505]" italics="true" pageId="4" pageNumber="253">Anochetus shohki</emphasis>
Terayama, 1996
</taxonomicName>
(Oganzaki, Sakieda, Ishigaki-jima, Yaeyama Is., Okinawa-ken, 10.iv.2022, H. Matsumura leg., MH97, emeraged in vii.2022 from captive colony); C –
<taxonomicName id="4C291BF0191DFFC7FD4BFA2652003990" authority="(Mayr, 1862)" baseAuthorityName="Mayr" baseAuthorityYear="1862" box="[721,1027,1512,1531]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="B95DBC61191DFFC7FD4BFA2655933990" box="[721,912,1512,1531]" italics="true" pageId="4" pageNumber="253">Brachyponera luteipes</emphasis>
(Mayr, 1862)
</taxonomicName>
(Shirahama-rindo, Iriomote-jima, Okinawa-ken, 11.xi.2018, Sk.Yamane leg., JP18-SKY-154); D –
<taxonomicName id="4C291BF0191DFFC7FDADF9CD559B3A7D" authority="(Terayama, 1999)" baseAuthorityName="Terayama" baseAuthorityYear="1999" box="[567,920,1539,1558]" class="Insecta" family="Formicidae" genus="Euponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="sakishimensis">
<emphasis id="B95DBC61191DFFC7FDADF9CD55033A7D" box="[567,768,1539,1558]" italics="true" pageId="4" pageNumber="253">Euponera sakishimensis</emphasis>
(Terayama, 1999)
</taxonomicName>
(Funaura, Iriomote-jima,Yaeyana Is., Okinawa-ken, 9.v.1979, K. Ogata leg.); E –
<taxonomicName id="4C291BF0191DFFC7FED8F9D054543A5A" authority="Forel, 1912" authorityName="Forel" authorityYear="1912" box="[322,599,1566,1585]" class="Insecta" family="Formicidae" genus="Leptogenys" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="confucii">
<emphasis id="B95DBC61191DFFC7FED8F9D057ED3A5A" box="[322,494,1566,1585]" italics="true" pageId="4" pageNumber="253">Leptogenys confucii</emphasis>
Forel, 1912
</taxonomicName>
(Mikyo, Tokuno-shima, Amami Is., Kagoshima-ken, 6.iii.2010, Sk. Yamane leg., M10-SKY-12); F –
<taxonomicName id="4C291BF0191DFFC7FF22F9F654213A20" authority="Emery, 1892" authorityName="Emery" authorityYear="1892" box="[184,546,1592,1611]" class="Insecta" family="Formicidae" genus="Odontomachus" isUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B95DBC61191DFFC7FF22F9F6573A3A20" box="[184,313,1592,1611]" italics="true" pageId="4" pageNumber="253">Odontomachus</emphasis>
cf.
<emphasis id="B95DBC61191DFFC7FEC0F9F657AC3A20" box="[346,431,1592,1611]" italics="true" pageId="4" pageNumber="253">monticola</emphasis>
Emery, 1892
</taxonomicName>
(Miyanoura, Yaku-shima, Ôsumi Is., Kagoshima-ken, 16.ix.2008, collector unknown).
</paragraph>
</caption>
<paragraph id="8B966073191DFFC7FF0DF91657E83B84" blockId="4.[151,760,1656,2063]" pageId="4" pageNumber="253">
<emphasis id="B95DBC61191DFFC7FF0DF916575D3A84" bold="true" box="[151,350,1752,1776]" pageId="4" pageNumber="253">Caste differences.</emphasis>
In
<taxonomicName id="4C291BF0191DFFC7FEE5F91657E13A84" authorityName="Terayama" authorityYear="1996" box="[383,482,1752,1775]" class="Insecta" family="Formicidae" genus="Anochetus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="shohki">
<emphasis id="B95DBC61191DFFC7FEE5F91657E13A84" box="[383,482,1752,1775]" italics="true" pageId="4" pageNumber="253">A. shohki</emphasis>
</taxonomicName>
the queen differs from the worker in having the following characters: body larger (HW 0. 94–1.00 mm in the queen
<emphasis id="B95DBC61191DFFC7FD97F8D654203B44" box="[525,547,1816,1839]" italics="true" pageId="4" pageNumber="253">vs</emphasis>
ca.
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in the worker); eye larger (maximum diameter
<quantity id="4CD1CD96191DFFC7FDDEF8F654A23B24" box="[580,673,1848,1871]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="253" unit="mm" value="0.2">0.20 mm</quantity>
<emphasis id="B95DBC61191DFFC7FD3DF8F654BE3B24" box="[679,701,1848,1871]" italics="true" pageId="4" pageNumber="253">vs</emphasis>
.
<quantity id="4CD1CD96191DFFC7FD53F8F656C53B04" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="253" unit="mm" value="0.13">0.13 mm</quantity>
); three ocelli present; meso- and metathorax with developed wings and associated structures; mesoscutellum differentiated from mesoscutum; groove dividing mesopleuron into upper and lower sections much more conspicuous than in the worker.
</paragraph>
</subSubSection>
<subSubSection id="C33333F8191DFFC7FF0DF836531C3464" pageId="4" pageNumber="253" type="discussion">
<paragraph id="8B966073191DFFC7FF0DF836531C3464" blockId="4.[151,760,1656,2063]" lastBlockId="4.[828,1437,1656,2063]" pageId="4" pageNumber="253">
<emphasis id="B95DBC61191DFFC7FF0DF83657053464" bold="true" box="[151,262,2040,2063]" pageId="4" pageNumber="253">Remarks.</emphasis>
The single Japanese species
<taxonomicName id="4C291BF0191DFFC7FDDFF83655533AE4" authorityName="Terayama" authorityYear="1996" box="[581,848,1656,2063]" class="Insecta" family="Formicidae" genus="Anochetus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="shohki">
<emphasis id="B95DBC61191DFFC7FDDFF83655533AE4" box="[581,848,1656,2063]" italics="true" pageId="4" pageNumber="253">Anochetus shohki</emphasis>
</taxonomicName>
is confined to the Central Ryukyus (Okinawa-jima) and Southern Ryukyus, and found in semi-open sites (TERAYAMA 1996;
<bibRefCitation id="EFB81D82191DFFC7FC6DF97652DA3AA4" author="TERAYAMA M. &amp; KUBOTA S. &amp; EGUCHI K." box="[1015,1241,1720,1743]" pageId="4" pageNumber="253" refId="ref15386" refString="TERAYAMA M., KUBOTA S. &amp; EGUCHI K. 2014: Encyclopedia of Japanese Ants. Asakura-shoten, Tokyo, 48 pls., viii + 278 pp (in Japanese)." type="book" year="2014">TERAYAMA et al. 2014</bibRefCitation>
; HISASUE &amp; TOKU- SHIGE 2021). The colonies are small with a single foundress. Many Asian species of
<taxonomicName id="4C291BF0191DFFC7FB5DF936533A3B64" authorityName="Mayr" authorityYear="1861" box="[1223,1337,1784,1807]" class="Insecta" family="Formicidae" genus="Anochetus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B95DBC61191DFFC7FB5DF936533A3B64" box="[1223,1337,1784,1807]" italics="true" pageId="4" pageNumber="253">Anochetus</emphasis>
</taxonomicName>
have an elongate head and an apically pointed petiole as in
<taxonomicName id="4C291BF0191DFFC7FAC2F8D655A13B24" authorityName="Latreille" authorityYear="1804" class="Insecta" family="Formicidae" genus="Odontomachus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="genus">
<emphasis id="B95DBC61191DFFC7FAC2F8D655A13B24" italics="true" pageId="4" pageNumber="253">Odontomachus</emphasis>
</taxonomicName>
. In the Japanese species the hindwing lacks a jugal lobe, while in other species groups from Asia the jugal lobe is often present. Although in
<taxonomicName id="4C291BF0191DFFC7FB77F8B653513BE4" authorityName="Terayama" authorityYear="1996" box="[1261,1362,1912,1935]" class="Insecta" family="Formicidae" genus="Anochetus" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="shohki">
<emphasis id="B95DBC61191DFFC7FB77F8B653513BE4" box="[1261,1362,1912,1935]" italics="true" pageId="4" pageNumber="253">A. shohki</emphasis>
</taxonomicName>
apicoventral spurs on mid- and hindtibiae are reduced to 0 and 1, respectively, the number of spurs is highly variable in this genus (
<bibRefCitation id="EFB81D82191DFFC7FC20F816524C3B84" author="BROWN W. L." box="[954,1103,2008,2031]" pageId="4" pageNumber="253" pagination="549 - 638" refId="ref14447" refString="BROWN W. L. 1978: Contribution toward a reclassification of the Formicidae. Part VI. Ponerini, Subtribe Odontomachiti, Section B. Genus Anochetus and bibliography. Studia Entomologica (New Series) 20: 549 - 638, 12 pls." type="journal article" year="1978">BROWN 1978</bibRefCitation>
). Thelytokous parthenogenesis is known in dealate queens (
<bibRefCitation id="EFB81D82191DFFC7FBF3F83653173464" author="MIYAZAKI S." box="[1129,1300,2040,2063]" pageId="4" pageNumber="253" pagination="1 - 6" refId="ref14866" refString="MIYAZAKI S. 2023: Thelytokous parthenogenesis in dealate queens of the ponerine ant Anochetus shohki. Asian Myrmecology 16 (e 016006): 1 - 6." type="journal article" year="2023">
<collectingRegion id="49EDAE91191DFFC7FBF3F83652D03464" box="[1129,1235,2040,2063]" country="Japan" name="Miyazaki" pageId="4" pageNumber="253">MIYAZAKI</collectingRegion>
2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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