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<document id="777B4EFABA1AA08B2B80FF6F443C0A52" ID-CLB-Dataset="8861" ID-DOI="10.11646/zootaxa.4869.1.1" ID-GBIF-Dataset="c285848a-ccaf-4f48-8e05-7f9cf6acf225" ID-ISSN="1175-5326" ID-Zenodo-Dep="4418246" ID-ZooBank="C34E9D0C-336A-4F4B-A670-2F342470839D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609812691373" checkinUser="plazi" docAuthor="Dawah, Hassan A., Abdullah, Mohammed A. &amp; Deeming, John C." docDate="2020" docId="03EF183F2C03FFB1FF3A624AFAD60560" docLanguage="en" docName="zootaxa.4869.1.1.pdf" docOrigin="Zootaxa 4869 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atherigona (Atherigona) reversura Villeneuve 1936" docType="treatment" docVersion="7" lastPageNumber="27" masterDocId="FFD660472C19FFABFFAD6108FF960050" masterDocTitle="The Muscidae (Diptera) of Saudi Arabia, descriptions of two new species, new records and updated list of species" masterLastPageNumber="54" masterPageNumber="1" pageNumber="27" updateTime="1698852512324" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="387890E62D11E79AF45154E9958BD9D3">The Muscidae (Diptera) of Saudi Arabia, descriptions of two new species, new records and updated list of species</mods:title>
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<mods:namePart id="E103E20E6441894ABE618030F025D3BB">Abdullah, Mohammed A.</mods:namePart>
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<mods:affiliation id="A77B393E6E752BC6C801F407A0816E23">Department of Biology, College of Science, King Khalid University, PO Box 9004, Abha- 61413, Kingdom of Saudi Arabia. mohd _ robiya @ hotmail. com; https: // orcid. org / 0000 - 0002 - 3323 - 3623</mods:affiliation>
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<mods:affiliation id="6BA001E7B55458E0C8DD74D01CBD3A97">National Museum of Wales, Department of Natural Sciences, Entomology Section, Cardiff, CF 10 3 NP, UK. john. deeming @ museumwales. ac. uk; https: // orcid. org / 0000 - 0002 - 6703 - 8180</mods:affiliation>
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<taxonomicName id="4C46D2AA2C03FFB1FF3A624AFD60030C" ID-CoL="8B52J" authority="Villeneuve, 1936" authorityName="Villeneuve" authorityYear="1936" box="[151,758,834,861]" class="Insecta" family="Muscidae" genus="Atherigona" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="reversura" subGenus="Atherigona">
<emphasis id="B932753B2C03FFB1FF3A624AFD60030C" bold="true" box="[151,758,834,861]" pageId="26" pageNumber="27">
<emphasis id="B932753B2C03FFB1FF3A624AFE8C030D" bold="true" box="[151,282,834,861]" italics="true" pageId="26" pageNumber="27">Atherigona</emphasis>
(
<emphasis id="B932753B2C03FFB1FE87624AFE3B030D" bold="true" box="[298,429,834,861]" italics="true" pageId="26" pageNumber="27">Atherigona</emphasis>
)
<emphasis id="B932753B2C03FFB1FE10624BFDBD030D" bold="true" box="[445,555,835,861]" italics="true" pageId="26" pageNumber="27">reversura</emphasis>
<bibRefCitation id="EFD7D4D82C03FFB1FD9E624AFD60030C" author="Villeneuve, J." box="[563,758,834,860]" pageId="26" pageNumber="27" pagination="1 - 13" refId="ref37380" refString="Villeneuve, J. (1936) 52. Dipteren. 16. Muscidae. In: Schwedisch-chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping-chang. Insekten gesammelt vom schwedischen Art der Expedition Dr. David Hummerl 1927 - 1930. Arkiv for Zoologi, Stockholm, 27 A (34), pp. 1 - 13." type="journal article" year="1936">Villeneuve, 1936</bibRefCitation>
</emphasis>
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(
<figureCitation id="137DB5AC2C03FFB1FCA8624AFCC3030C" box="[773,853,834,860]" captionStart="FIGURE" captionStartId="28.[151,251,1934,1958]" captionTargetBox="[154,1421,1022,1910]" captionTargetId="figure-304@28.[153,1434,1011,1911]" captionTargetPageId="28" captionText="FIGURE. 3741. Muscidae species habitus: 37) Atherigona (Atherigona) reversura Villeneuve, 1936, Male; 38) Atherigona (Atherigona) soccata Rondani, 1871, Male; 39) Atherigona (Atherigona) theodori Hennig, Female; 40) Atherigona (Atherigona) varia (Meigen, 1826), Male; 41) Atherigona (Acritochaeta) afrotropicalis Deeming sp.n., Male." figureDoi="http://doi.org/10.5281/zenodo.4418274" httpUri="https://zenodo.org/record/4418274/files/figure.png" pageId="26" pageNumber="27">Fig. 37</figureCitation>
)xxx
</paragraph>
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<treatmentCitation id="0AE78F382C03FFB1FF3A626EFDED03D0" author="Villeneuve, J." box="[151,635,870,896]" page="13" pageId="26" pageNumber="27" year="1936">
<taxonomicName id="4C46D2AA2C03FFB1FF3A626EFDED03D0" ID-CoL="8B52J" authority="Villeneuve, 1936: 13" authorityName="Villeneuve" authorityPageNumber="13" authorityYear="1936" box="[151,635,870,896]" class="Insecta" family="Muscidae" genus="Atherigona" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="reversura">
<emphasis id="B932753B2C03FFB1FF3A626EFE1E03D0" box="[151,392,870,896]" italics="true" pageId="26" pageNumber="27">Atherigona reversura</emphasis>
<bibRefCitation id="EFD7D4D82C03FFB1FE23626EFDED03D0" author="Villeneuve, J." box="[398,635,870,896]" pageId="26" pageNumber="27" pagination="1 - 13" refId="ref37380" refString="Villeneuve, J. (1936) 52. Dipteren. 16. Muscidae. In: Schwedisch-chinesische wissenschaftliche Expedition nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. Sven Hedin und Prof. Su Ping-chang. Insekten gesammelt vom schwedischen Art der Expedition Dr. David Hummerl 1927 - 1930. Arkiv for Zoologi, Stockholm, 27 A (34), pp. 1 - 13." type="journal article" year="1936">Villeneuve, 1936: 13</bibRefCitation>
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.
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</paragraph>
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<emphasis id="B932753B2C03FFB1FF6A6282FE5403F4" bold="true" box="[199,450,906,932]" pageId="26" pageNumber="27">Specimen examined.</emphasis>
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,
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,
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,
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fruit farm,
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,
<collectingMethod id="5207D13E2C03FFB1FBC76282FB6803F5" box="[1130,1278,906,933]" pageId="26" pageNumber="27">Malaise trap</collectingMethod>
,
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.
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.Dawah. (CERS).
</paragraph>
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<emphasis id="B932753B2C03FFB1FF6A62DAFEF703BC" bold="true" box="[199,353,978,1004]" pageId="26" pageNumber="27">Distribution.</emphasis>
This species was previously recorded from
<collectingCountry id="F351E9B92C03FFB1FCF562DAFC7D03BC" box="[856,1003,978,1004]" name="Saudi Arabia" pageId="26" pageNumber="27">Saudi Arabia</collectingCountry>
by
<bibRefCitation id="EFD7D4D82C03FFB1FBB562DAFADD03BC" author="Dawah, H. A. &amp; Abdullah, M. A." box="[1048,1355,978,1004]" pageId="26" pageNumber="27" pagination="373 - 396" refId="ref31297" refString="Dawah, H. A. &amp; Abdullah, M. A. (2009) The Muscidae (Diptera: Brachycera: Muscomorpha) of south-western Saudi Arabia. Fauna of Arabia, 24, 373 - 396." type="journal article" year="2009">Dawah &amp; Abdullah (2009)</bibRefCitation>
. It was described from
<collectingCountry id="F351E9B92C03FFB1FEE562FEFE190440" box="[328,399,1014,1040]" name="China" pageId="26" pageNumber="27">China</collectingCountry>
. In the Middle East it is known from
<collectingCountry id="F351E9B92C03FFB1FC9A62FEFCED0440" box="[823,891,1014,1040]" name="Oman" pageId="26" pageNumber="27">Oman</collectingCountry>
and
<collectingCountry id="F351E9B92C03FFB1FC1E62FEFBD30440" box="[947,1093,1014,1040]" name="Saudi Arabia" pageId="26" pageNumber="27">Saudi Arabia</collectingCountry>
and the
<collectingCountry id="F351E9B92C03FFB1FB0A62FEFA0B0440" box="[1191,1437,1014,1040]" name="United Arab Emirates" pageId="26" pageNumber="27">United Arab Emirates</collectingCountry>
(
<bibRefCitation id="EFD7D4D82C03FFB1FF326512FEDF0464" author="Deeming, J. C." box="[159,329,1050,1076]" pageId="26" pageNumber="27" refId="ref31673" refString="Deeming, J. C. (2008) Order Diptera, family Muscidae. In: van Harten, A. (Ed.), Arthropod Fauna of the United Arab Emirates. Vol. 1. Dar Al-Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, pp. 714 - 723." type="book" year="2008">Deeming 2008</bibRefCitation>
;
<bibRefCitation id="EFD7D4D82C03FFB1FEF86512FDF60464" author="El-Hawagry, M. S. &amp; Abdel-Dayem, M. S. &amp; Elgharbawy, A. A. &amp; Al Dhafer, H. M." box="[341,608,1050,1077]" pageId="26" pageNumber="27" pagination="107 - 139" refId="ref32055" refString="El-Hawagry, M. S., Abdel-Dayem, M. S., Elgharbawy, A. A. &amp; Al Dhafer, H. M. (2016) A preliminary account of the fly fauna in Jabal Shada al-A'la Nature Reserve, Saudi Arabia, with new records and biogeographical remarks (Diptera, Insecta). Zookeys, 636, 107 - 139. https: // doi. org / 10.3897 / zookeys. 636.9905" type="journal article" year="2016">
El-Hawagry
<emphasis id="B932753B2C03FFB1FE486513FD800464" box="[485,534,1050,1076]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. 2016
</bibRefCitation>
;
<bibRefCitation id="EFD7D4D82C03FFB1FDC66512FD3E0464" author="El-Hawagry, M. S. &amp; Abdel-Dayem, M. S. &amp; Al Dhafer, H. M." box="[619,680,1050,1076]" pageId="26" pageNumber="27" pagination="1499 - 1530" refId="ref32135" refString="El-Hawagry, M. S., Abdel-Dayem, M. S., El-Sonbati &amp; Al Dhafer, H. M. (2017) A preliminary account of the fly fauna in Gard Raydah Nature Reserve, Kingdom of Saudi Arabia, with new records and biogeographical remarks (Diptera: Insecta). Journal of Natural History, 51 (25 - 26), 1499 - 1530. https: // doi. org / 10.1080 / 00222933.2017.1347299" type="journal article" year="2017">2017</bibRefCitation>
).
</paragraph>
<paragraph id="8BF9A9292C03FFB1FF6A6536FAD60560" blockId="26.[151,1437,834,1329]" pageId="26" pageNumber="27">
<emphasis id="B932753B2C03FFB1FF6A6536FE250408" bold="true" box="[199,435,1086,1112]" pageId="26" pageNumber="27">Biological remarks.</emphasis>
<taxonomicName id="4C46D2AA2C03FFB1FE106537FDDD0408" authorityName="Villeneuve" authorityYear="1936" box="[445,587,1087,1113]" class="Insecta" family="Muscidae" genus="Atherigona" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="reversura">
<emphasis id="B932753B2C03FFB1FE106537FDDD0408" box="[445,587,1087,1113]" italics="true" pageId="26" pageNumber="27">A. reversura</emphasis>
</taxonomicName>
is a serious pest which can cause severe damage to plants. Although the larvae of
<taxonomicName id="4C46D2AA2C03FFB1FEAD656BFE1C042C" authorityName="Villeneuve" authorityYear="1936" box="[256,394,1123,1149]" class="Insecta" family="Muscidae" genus="Atherigona" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="reversura">
<emphasis id="B932753B2C03FFB1FEAD656BFE1C042C" box="[256,394,1123,1149]" italics="true" pageId="26" pageNumber="27">A. reversura</emphasis>
</taxonomicName>
known as Bermudagrass breeds mainly in Bermudagrass (
<taxonomicName id="4C46D2AA2C03FFB1FBBB656AFB74042C" authorityName="C.H.Persoon" authorityYear="1805" box="[1046,1250,1122,1148]" class="Actinopterygii" family="Cynodontidae" genus="Cynodon" kingdom="Animalia" order="Characiformes" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="dactylon">
<emphasis id="B932753B2C03FFB1FBBB656AFB74042C" box="[1046,1250,1122,1148]" italics="true" pageId="26" pageNumber="27">Cynodon dactylon</emphasis>
</taxonomicName>
(L.) which is regarded as invasive pest species in
<collectingCountry id="F351E9B92C03FFB1FDBC658EFDD104F0" box="[529,583,1158,1184]" name="United States of America" pageId="26" pageNumber="27">USA</collectingCountry>
(
<bibRefCitation id="EFD7D4D82C03FFB1FDFB658EFCDE04F0" author="Grzywacz, A. &amp; Pape, T. &amp; Hudson, W. G. &amp; Gomez, S." box="[598,840,1158,1185]" pageId="26" pageNumber="27" pagination="1055 - 1067" refId="ref32662" refString="Grzywacz, A., Pape, T., Hudson, W. G. &amp; Gomez, S. (2013) Morphology and immature stages of Atherigona reversura (Diptera: Muscidae) with notes on the recent invasion of North America. Journal of Natural History, 47 (15 - 16), 1055 - 1067. https: // doi. org / 10.1080 / 00222933.2012.742244" type="journal article" year="2013">
Grzywacz
<emphasis id="B932753B2C03FFB1FD62658FFC9604F0" box="[719,768,1158,1184]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. 2013
</bibRefCitation>
; Patitucci
<emphasis id="B932753B2C03FFB1FC16658FFC7A04F0" box="[955,1004,1158,1184]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. 2016), other
<taxonomicName id="4C46D2AA2C03FFB1FB2A658FFB7504F1" box="[1159,1251,1159,1185]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
host plants have been listed by
<bibRefCitation id="EFD7D4D82C03FFB1FE9A65A3FDD10495" author="Pont, A. C. &amp; Magpayo, F. R." box="[311,583,1194,1221]" pageId="26" pageNumber="27" pagination="1 - 123" refId="ref35055" refString="Pont, A. C. &amp; Magpayo, F. R. (1995) Muscid shoot-flies of the Philippine Islands (Diptera: Muscidae, genus Atherigona Rondani). Bulletin of Entomological Research, Supplement Series, Supplement No. 3, 1 - 123. https: // doi. org / 10.1017 / S 1367426900000321" type="book chapter" year="1995">Pont &amp; Magpayo (1995)</bibRefCitation>
such as
<taxonomicName id="4C46D2AA2C03FFB1FD0965A2FC150494" authorityName="Link" authorityYear="1833" baseAuthorityName="Linnaeus" box="[676,899,1194,1220]" class="Liliopsida" family="Poaceae" genus="Echinochloa" kingdom="Plantae" order="Poales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="colona">
<emphasis id="B932753B2C03FFB1FD0965A2FC150494" box="[676,899,1194,1220]" italics="true" pageId="26" pageNumber="27">Echinochloa colona</emphasis>
</taxonomicName>
(L.) Link,
<taxonomicName id="4C46D2AA2C03FFB1FC5065A2FA890494" authority="Gaertn" authorityName="Gaertn" box="[1021,1311,1194,1220]" class="Liliopsida" family="Poaceae" genus="Eleusine" kingdom="Plantae" order="Poales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="coracana">
<emphasis id="B932753B2C03FFB1FC5065A2FB5B0494" box="[1021,1229,1194,1220]" italics="true" pageId="26" pageNumber="27">Eleusine coracana</emphasis>
Gaertn
</taxonomicName>
,
<taxonomicName id="4C46D2AA2C03FFB1FA8165A2FED704B8" authority="(Retz.)" baseAuthorityName="Retz." class="Liliopsida" family="Poaceae" genus="Eriochloa" kingdom="Plantae" order="Poales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="procera">
<emphasis id="B932753B2C03FFB1FA8165A2FF7904B8" italics="true" pageId="26" pageNumber="27">Eriochloa procera</emphasis>
(Retz.)
</taxonomicName>
,
<taxonomicName id="4C46D2AA2C03FFB1FEE265C6FDED04B9" authority="(Rottler)" baseAuthorityName="Rottler" box="[335,635,1230,1257]" class="Liliopsida" family="Poaceae" genus="Sehima" kingdom="Plantae" order="Poales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="nervosum">
<emphasis id="B932753B2C03FFB1FEE265C6FD8204B8" box="[335,532,1230,1256]" italics="true" pageId="26" pageNumber="27">Sehima nervosum</emphasis>
(Rottler)
</taxonomicName>
,
<emphasis id="B932753B2C03FFB1FD2A65C6FCD404B8" box="[647,834,1230,1256]" italics="true" pageId="26" pageNumber="27">
Sorghum
<taxonomicName id="4C46D2AA2C03FFB1FD5C65C6FCD404B8" box="[753,834,1230,1256]" class="Insecta" family="Muscidae" genus="Spilogaster" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bicolor">bicolor</taxonomicName>
</emphasis>
and
<taxonomicName id="4C46D2AA2C03FFB1FCDB65C7FA9504B8" authority="L. Grzywacz et al. (2013)" authorityName="L. Grzywacz" authorityYear="2013" box="[886,1283,1230,1257]" class="Liliopsida" family="Poaceae" genus="Zea" kingdom="Plantae" order="Poales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="mays">
<emphasis id="B932753B2C03FFB1FCDB65C7FC7604B8" box="[886,992,1231,1256]" italics="true" pageId="26" pageNumber="27">Zea mays</emphasis>
L.
<bibRefCitation id="EFD7D4D82C03FFB1FBA965C6FA9504B8" author="Grzywacz, A. &amp; Pape, T. &amp; Hudson, W. G. &amp; Gomez, S." box="[1028,1283,1230,1256]" pageId="26" pageNumber="27" pagination="1055 - 1067" refId="ref32662" refString="Grzywacz, A., Pape, T., Hudson, W. G. &amp; Gomez, S. (2013) Morphology and immature stages of Atherigona reversura (Diptera: Muscidae) with notes on the recent invasion of North America. Journal of Natural History, 47 (15 - 16), 1055 - 1067. https: // doi. org / 10.1080 / 00222933.2012.742244" type="journal article" year="2013">
Grzywacz
<emphasis id="B932753B2C03FFB1FBD665C7FB2504B8" box="[1147,1203,1230,1256]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
(2013)
</bibRefCitation>
</taxonomicName>
described the immature stages of
<taxonomicName id="4C46D2AA2C03FFB1FEDD65FBFE6D055C" authorityName="Villeneuve" authorityYear="1936" box="[368,507,1267,1293]" class="Insecta" family="Muscidae" genus="Atherigona" kingdom="Animalia" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="reversura">
<emphasis id="B932753B2C03FFB1FEDD65FBFE6D055C" box="[368,507,1267,1293]" italics="true" pageId="26" pageNumber="27">A. reversura</emphasis>
</taxonomicName>
but the life cycle and the biology has yet to be researched. Since its initial introduction to the
<collectingCountry id="F351E9B92C03FFB1FEBD641EFECB0560" box="[272,349,1302,1328]" name="United States of America" pageId="26" pageNumber="27">U.S.A.</collectingCountry>
it has in a very short time increased its distribution from
<collectingCountry id="F351E9B92C03FFB1FC72641EFBA50560" box="[991,1075,1302,1328]" name="Canada" pageId="26" pageNumber="27">Canada</collectingCountry>
to
<collectingCountry id="F351E9B92C03FFB1FBFA641EFB020560" box="[1111,1172,1302,1328]" name="Chile" pageId="26" pageNumber="27">Chile</collectingCountry>
and
<collectingCountry id="F351E9B92C03FFB1FB65641EFAAD0560" box="[1224,1339,1302,1328]" name="Argentina" pageId="26" pageNumber="27">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>