464 lines
58 KiB
XML
464 lines
58 KiB
XML
<document id="301F1591582B34FEFC5DF1165F491ADB" ID-DOI="10.1080/00222933.2017.1391959" ID-ISSN="1464-5262" ID-Zenodo-Dep="5174516" ID-ZooBank="9E21390E-3FD4-40EB-9442-31BC92A76B4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628558890842" checkinUser="felipe" docAuthor="Ruane, Sara, Richards, Stephen J., McVay, John D., Tjaturadi, Burhan, Krey, Keliopas & Austin, Christopher C." docDate="2017" docId="03DA87E1D400457504CAE5D087BD9844" docLanguage="en" docName="JNatHist.52.13-16.917-944.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 52 (13 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2017.1391959" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Stegonotus admiraltiensis Ruane & Richards & McVay & Tjaturadi & Krey & Austin 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="939" masterDocId="FFE3FF99D415456D0575E55D851E9975" masterDocTitle="Cryptic and non-cryptic diversity in New Guinea ground snakes of the genus Stegonotus Duméril, Bibron and Duméril, 1854: a description of four new species (Squamata: Colubridae)" masterLastPageNumber="944" masterPageNumber="917" pageNumber="937" updateTime="1699046540698" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="052CF838E5A71070CB042A26C57793FE">Cryptic and non-cryptic diversity in New Guinea ground snakes of the genus Stegonotus Duméril, Bibron and Duméril, 1854: a description of four new species (Squamata: Colubridae)</mods:title>
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<mods:namePart id="8D692DCFE093531C7C24626A96AFD8D0">Ruane, Sara</mods:namePart>
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<mods:namePart id="EED8007BAB22C9F05E6FE027803D95C8">Richards, Stephen J.</mods:namePart>
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<mods:namePart id="79BE1B6F7A85C9D05492337FC4BC8A8A">McVay, John D.</mods:namePart>
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<mods:namePart id="CE7BD2DDC2E262AB074F8D0F1A47796A">Tjaturadi, Burhan</mods:namePart>
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<mods:namePart id="980E8CB81DEAEB0F7EA5B0CB20B19A1B">Krey, Keliopas</mods:namePart>
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<mods:namePart id="53A1B2738672BCB697F701E47CC24D63">Austin, Christopher C.</mods:namePart>
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<mods:title id="A512ABC66913A21AD93927AB186E91CE">Journal of Natural History</mods:title>
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<mods:date id="C9F75C4644A9A143674A94605E5C55F2">2017</mods:date>
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<mods:title id="B881626F09BD750A211ACB700811B95E">J. Nat. Hist.</mods:title>
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<mods:number id="F709645671635B234EBA6B21AEAF6F90">13 - 16</mods:number>
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<heading id="D084819BD400457804CAE5D0867B99D2" box="[447,869,141,167]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="937" reason="5">
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<taxonomicName id="4C734D74D400457804CAE5D087E199D2" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[447,767,141,167]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="937" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
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<emphasis id="B907EAE5D400457804CAE5D087E199D2" bold="true" box="[447,767,141,167]" italics="true" pageId="21" pageNumber="937">Stegonotus admiraltiensis</emphasis>
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<taxonomicNameLabel id="A234579ED4004578067DE5D3867B99D2" box="[776,869,142,167]" pageId="21" pageNumber="937" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C369657CD40045780774E5EC863C99BF" box="[513,802,177,202]" pageId="21" pageNumber="937" type="description">
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<paragraph id="8BCC36F7D40045780774E5EC863C99BF" blockId="21.[447,869,141,202]" box="[513,802,177,202]" pageId="21" pageNumber="937">
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(
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<figureCitation id="13482A72D4004578077CE5EC878B99BF" box="[521,661,177,202]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStartId-0="6.[160,225,1511,1534]" captionStartId-1="9.[160,225,1677,1700]" captionStartId-2="14.[160,225,1613,1636]" captionTargetBox-0="[194,1121,142,1480]" captionTargetBox-1="[217,1128,144,1646]" captionTargetBox-2="[334,981,142,1582]" captionTargetId-0="figure-1@6.[193,1121,142,1480]" captionTargetId-1="graphics-2@9.[256,1128,623,1646]" captionTargetId-2="figure-1@14.[333,981,142,1582]" captionTargetPageId-0="6" captionTargetPageId-1="9" captionTargetPageId-2="14" captionText-0="Figure 2. Photographs of Stegonotus species with dorsal, ventral and lateral head views from left to right: (a) Stegonotus batjanensis USNM 237129, (b) Stegonotus borneensis FMNH 251054, (c) Stegonotus muelleri LSUMZ 41802, (d) Stegonotus florensis WAM 104606, (e) Stegonotus guentheri AM 129712, (f) Stegonotus heterurus BPBM 22556, (g) Stegonotus cucullatus LSUMZ 94371. Photos from SR. (h) Stegonotus modestus LSUMZ 92339, (i) Stegonotus iridis sp. nov. SJR 7514, (j) Stegonotus diehli LSUMZ 92344 (k) Stegonotus melanolabiatus sp. nov. AMS R115361 (l) Stegonotus derooijae sp. nov. SAMA R70467 (m) Stegonotus admiraltiensis sp. nov. LSUMZ 93597, (n) Stegonotus parvus LSUMZ 92333. Photos from Sara Ruane and Stephen J. Richards." captionText-1="Figure 3. Map of New Guinea sampling localities for Stegonotus (a) and mtDNA + nucDNA maximum likelihood gene tree (b). Symbols for species/clades match those on the map; * indicates bootstrap values ≥ 70. For some localities, symbols were adjusted for better visibility and so localities are approximate; see Appendix 1. for exact locality information." captionText-2="Figure 4. Photographs of three Stegonotus species described here in life; (a) Stegonotus iridis sp. nov. SAMA R70466 (b) Stegonotus derooijae sp. nov. MZB.Ophi.3288 (c) Stegonotus admiraltiensis sp. nov. from Manus Island, unregistered. Photos from Stephen J. Richards." figureDoi-0="http://doi.org/10.5281/zenodo.5174520" figureDoi-1="http://doi.org/10.5281/zenodo.5174522" figureDoi-2="http://doi.org/10.5281/zenodo.5174525" httpUri-0="https://zenodo.org/record/5174520/files/figure.png" httpUri-1="https://zenodo.org/record/5174522/files/figure.png" httpUri-2="https://zenodo.org/record/5174525/files/figure.png" pageId="21" pageNumber="937">Figures 2−4</figureCitation>
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,
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<figureCitation id="13482A72D400457807D0E5EC87AD99BF" box="[677,691,177,202]" captionStart="Figure 8" captionStartId="22.[160,225,1382,1405]" captionTargetBox="[163,1153,142,1351]" captionTargetId="figure-61@22.[163,1153,142,1351]" captionTargetPageId="22" captionText="Figure 8. Photographs of holotype Stegonotus admiraltiensis sp. nov. LSUMZ 93598 showing (clockwise from top left) dorsal of body, ventral of body, dorsal of head, lateral of head. Photos from Sara Ruane." figureDoi="http://doi.org/10.5281/zenodo.5174533" httpUri="https://zenodo.org/record/5174533/files/figure.png" pageId="21" pageNumber="937">8</figureCitation>
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,
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<tableCitation id="C6F1034CD400457807B7E5EC860799BF" box="[706,793,177,202]" captionStart="Table 1" captionStartId="10.[133,189,620,643]" captionText="Table 1. Morphological characters of Stegonotus examined." pageId="21" pageNumber="937">Table 1</tableCitation>
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)
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</paragraph>
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<subSubSection id="C369657CD400457805D5E5A9875E9879" box="[160,576,244,268]" pageId="21" pageNumber="937" type="reference_group">
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<taxonomicName id="4C734D74D400457805D5E5A9875E9879" authority="(Schlegel 1837)" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[160,576,244,268]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="937" phylum="Chordata" rank="species" species="modestus">
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<emphasis id="B907EAE5D400457805D5E5A984949879" box="[160,394,244,268]" italics="true" pageId="21" pageNumber="937">Stegonotus modestus</emphasis>
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(
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<bibRefCitation id="EFE24B06D400457804EDE5A987269879" author="Schlegel H" box="[408,568,244,268]" pageId="21" pageNumber="937" refId="ref12789" refString="Schlegel H. 1837. Essai sur la physionomie des serpents [Essay on the physiognomy of snakes]. Amsterdam: MH Schonekat." type="book" year="1837">Schlegel 1837</bibRefCitation>
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)
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</taxonomicName>
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<typeStatus id="54C88855D400457805D5E46484159826" box="[160,267,313,339]" pageId="21" pageNumber="937" type="holotype">Holotype</typeStatus>
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<materialsCitation id="3B1B3CAAD400457805D5E400867798CF" collectingDate="2001-09-01" collectionCode="LSUMZ, CCA" collectorName="Christopher C. Austin" country="Papua New Guinea" elevation="58" latitude="-2.3283334" location="Penchal Village on Rambutyo Island" longLatPrecision="1" longitude="147.76666" pageId="21" pageNumber="937" specimenCode="LSUMZ 93598, CCA 2087" specimenCount="1" stateProvince="Manus" typeStatus="holotype">
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<specimenCode id="DBD59E8CD400457805D5E40084239800" box="[160,317,349,373]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="21" pageNumber="937">LSUMZ 93598</specimenCode>
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(field number
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<specimenCode id="DBD59E8CD4004578049FE40087409800" box="[490,606,349,373]" collectionCode="CCA" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="21" pageNumber="937">CCA 2087</specimenCode>
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), adult male. Collected by
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on
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<collectingDate id="EF89E9DFD400457805B0E4DD848B98ED" box="[197,405,384,408]" pageId="21" pageNumber="937" value="2001-09-01">1 September 2001</collectingDate>
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from
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,
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<collectingRegion id="49B7F815D40045780608E4DD812C98ED" box="[893,1074,384,408]" country="Papua New Guinea" name="Manus" pageId="21" pageNumber="937">Manus Province</collectingRegion>
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,
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<collectingCountry id="F3647667D40045780148E4DD843398CF" name="Papua New Guinea" pageId="21" pageNumber="937">Papua New Guinea</collectingCountry>
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,
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elevation
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asl
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, (
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,
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<geoCoordinate id="EE475030D400457807C2E4FF867E98CF" box="[695,864,418,442]" degrees="147.7666667" direction="east" orientation="longitude" pageId="21" pageNumber="937" precision="1" value="147.76666">147.7666667°E</geoCoordinate>
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).
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<typeStatus id="54C88855D400457805D5E4BA840B9B74" box="[160,277,487,513]" pageId="21" pageNumber="937" type="paratype">Paratypes</typeStatus>
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<paragraph id="8BCC36F7D400457805D5E756813F9BA5" blockId="21.[160,1156,487,720]" pageId="21" pageNumber="937">
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<specimenCode id="DBD59E8CD400457805D5E75684249B56" box="[160,314,523,547]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="21" pageNumber="937">LSUMZ 93599</specimenCode>
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(field number
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<specimenCode id="DBD59E8CD40045780495E75687519B56" box="[480,591,523,547]" collectionCode="CCA" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="21" pageNumber="937">CCA 2092</specimenCode>
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) and
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<specimenCode id="DBD59E8CD400457807F8E75686399B56" box="[653,807,523,547]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="21" pageNumber="937">LSUMZ 93600</specimenCode>
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(field number
|
||
<specimenCode id="DBD59E8CD400457806B8E75681209B56" box="[973,1086,523,547]" collectionCode="CCA" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="21" pageNumber="937">CCA 2097</specimenCode>
|
||
), both collected by
|
||
<collectorName id="26865321D4004578044FE77384C29B33" box="[314,476,558,582]" pageId="21" pageNumber="937">Christopher C</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1B3CAAD400457804A9E773819E9BFE" collectingDate="2001-09-03" collectionCode="LSUMZ, CCA" country="Papua New Guinea" elevation="100" latitude="-2.3405" location="Austin" longLatPrecision="1" longitude="147.7945" municipality="Penchal Village on Rambutyo Island" pageId="21" pageNumber="937" specimenCode="LSUMZ 93597, CCA 2042" specimenCount="1" stateProvince="Manus" typeStatus="paratype">
|
||
.
|
||
<location id="8EAC602CD40045780499E773872D9B33" LSID="urn:lsid:plazi:treatment:03DA87E1D400457504CAE5D087BD9844:8EAC602CD40045780499E773872D9B33" box="[492,563,558,582]" country="Papua New Guinea" latitude="-2.3405" longLatPrecision="1" longitude="147.7945" municipality="Penchal Village on Rambutyo Island" name="Austin" pageId="21" pageNumber="937" stateProvince="Manus">Austin</location>
|
||
on
|
||
<date id="FFCD1037D4004578071CE773865C9B33" box="[617,834,558,582]" pageId="21" pageNumber="937" value="2001-09-03">
|
||
<collectingDate id="EF89E9DFD4004578071CE773865C9B33" box="[617,834,558,582]" pageId="21" pageNumber="937" value="2001-09-03">3 September 2001</collectingDate>
|
||
</date>
|
||
from northeast of
|
||
<collectingMunicipality id="6BA8AC8DD4004578015EE77384FE9B1D" pageId="21" pageNumber="937">Penchal Village on Rambutyo Island</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49B7F815D40045780498E70D87BB9B1D" box="[493,677,592,616]" country="Papua New Guinea" name="Manus" pageId="21" pageNumber="937">Manus Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3647667D400457807C6E70D868C9B1D" box="[691,914,592,616]" name="Papua New Guinea" pageId="21" pageNumber="937">Papua New Guinea</collectingCountry>
|
||
,
|
||
<elevation id="005ED1C4D400457806EAE70D819D9B1D" box="[927,1155,592,616]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="937" unit="m" value="100.0">
|
||
elevation
|
||
<quantity id="4C8B9B12D40045780164E70D81459B1D" box="[1041,1115,592,616]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="937" unit="m" value="100.0">100 m</quantity>
|
||
asl
|
||
</elevation>
|
||
(
|
||
<geoCoordinate id="EE475030D400457805D3E72E842D9BFE" box="[166,307,627,651]" degrees="2.3405000" direction="south" orientation="latitude" pageId="21" pageNumber="937" precision="1" value="-2.3405">2.3405000°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE475030D40045780434E72E84F79BFE" box="[321,489,627,651]" degrees="147.7945000" direction="east" orientation="longitude" pageId="21" pageNumber="937" precision="1" value="147.7945">147.7945000°E</geoCoordinate>
|
||
);
|
||
<specimenCode id="DBD59E8CD40045780488E72E87859BFE" box="[509,667,627,651]" collectionCode="LSUMZ" country="USA" httpUri="http://grbio.org/cool/ew6z-dwdm" name="Louisiana State University, Musuem of Zoology" pageId="21" pageNumber="937">LSUMZ 93597</specimenCode>
|
||
(field number
|
||
<specimenCode id="DBD59E8CD4004578063BE72E86DD9BFE" box="[846,963,627,651]" collectionCode="CCA" country="Germany" httpUri="http://grbio.org/cool/mqei-3i36" name="Culture Collection of Algae at the University of Marburg" pageId="21" pageNumber="937">CCA 2042</specimenCode>
|
||
), collected by C
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1B3CAAD400457801F5E72E81039BA5" collectingDate="2001-08-29" country="Papua New Guinea" elevation="10" latitude="-2.0326667" location="Austin" longLatPrecision="1" longitude="147.43417" municipality="Peyon Village on Los Negros Island" pageId="21" pageNumber="937" specimenCount="1" stateProvince="Manus" typeStatus="paratype">
|
||
.
|
||
<location id="8EAC602CD400457805D5E7CB85F99BDB" LSID="urn:lsid:plazi:treatment:03DA87E1D400457504CAE5D087BD9844:8EAC602CD400457805D5E7CB85F99BDB" box="[160,231,662,686]" country="Papua New Guinea" latitude="-2.0326667" longLatPrecision="1" longitude="147.43417" municipality="Peyon Village on Los Negros Island" name="Austin" pageId="21" pageNumber="937" stateProvince="Manus">Austin</location>
|
||
on
|
||
<date id="FFCD1037D40045780457E7CB84FC9BDB" box="[290,482,662,686]" pageId="21" pageNumber="937" value="2001-08-29">
|
||
<collectingDate id="EF89E9DFD40045780457E7CB84FC9BDB" box="[290,482,662,686]" pageId="21" pageNumber="937" value="2001-08-29">29 August 2001</collectingDate>
|
||
</date>
|
||
from near
|
||
<collectingMunicipality id="6BA8AC8DD40045780700E7CB81369BDB" box="[629,1064,662,686]" pageId="21" pageNumber="937">Peyon Village on Los Negros Island</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49B7F815D4004578014CE7CB841D9BA5" country="Papua New Guinea" name="Manus" pageId="21" pageNumber="937">Manus Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3647667D4004578047AE7E584F59BA5" box="[271,491,696,720]" name="Papua New Guinea" pageId="21" pageNumber="937">Papua New Guinea</collectingCountry>
|
||
,
|
||
<elevation id="005ED1C4D40045780483E7E587D69BA5" box="[502,712,696,720]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="937" unit="m" value="10.0">
|
||
elevation
|
||
<quantity id="4C8B9B12D40045780713E7E587BF9BA5" box="[614,673,696,720]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="937" unit="m" value="10.0">10 m</quantity>
|
||
asl
|
||
</elevation>
|
||
(
|
||
<geoCoordinate id="EE475030D400457807A3E7E5867D9BA5" box="[726,867,696,720]" degrees="2.0326667" direction="south" orientation="latitude" pageId="21" pageNumber="937" precision="1" value="-2.0326667">2.0326667°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE475030D4004578061AE7E581069BA5" box="[879,1048,696,720]" degrees="147.4341667" direction="east" orientation="longitude" pageId="21" pageNumber="937" precision="1" value="147.43417">147.4341667°E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369657CD400457805D5E7A081369D7C" pageId="21" pageNumber="937" type="diagnosis">
|
||
<paragraph id="8BCC36F7D400457805D5E7A0840A9A62" blockId="21.[160,1155,765,1033]" box="[160,276,765,791]" pageId="21" pageNumber="937">
|
||
<emphasis id="B907EAE5D400457805D5E7A0840A9A62" bold="true" box="[160,276,765,791]" italics="true" pageId="21" pageNumber="937">Diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D400457805D5E67C81369D7C" blockId="21.[160,1155,765,1033]" pageId="21" pageNumber="937">
|
||
<taxonomicName id="4C734D74D400457805D5E67C84A79A4C" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[160,441,801,825]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="21" pageNumber="937" phylum="Chordata" rank="species" species="admiraltiensis">
|
||
<emphasis id="B907EAE5D400457805D5E67C84A79A4C" box="[160,441,801,825]" italics="true" pageId="21" pageNumber="937">Stegonotus admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from its congeners by the following combination of characters: a reddish-tan unpatterned dorsum with an immaculate white or cream venter that may be barely darker approaching and continuing posteriorly onto the subcaudal scales, 17 or 19 dorsal scale rows at midbody, the presence of two apical pits on most or all dorsal scales, 202−214 ventral scales and 93−98 divided subcaudal scales, two preocular scales, eight supralabial scales, nine or ten infralabial scales, with the supralabial and infralabial scales cream with little or no dark pigmentation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369657CD400457A05D5E168813E9B81" lastPageId="23" lastPageNumber="939" pageId="21" pageNumber="937" type="description">
|
||
<paragraph id="8BCC36F7D400457805D5E16884A69D3A" blockId="21.[160,1156,1077,1727]" box="[160,440,1077,1103]" pageId="21" pageNumber="937">
|
||
<emphasis id="B907EAE5D400457805D5E16884A69D3A" bold="true" box="[160,440,1077,1103]" italics="true" pageId="21" pageNumber="937">
|
||
Description of
|
||
<typeStatus id="54C88855D40045780425E16884A69D3A" box="[336,440,1077,1103]" pageId="21" pageNumber="937" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D400457B05D5E107846B9F7A" blockId="21.[160,1156,1077,1727]" lastBlockId="22.[160,1155,1492,1551]" lastPageId="22" lastPageNumber="938" pageId="21" pageNumber="937">
|
||
Adult male in excellent state of preservation, tail complete, short (~
|
||
<quantity id="4C8B9B12D400457806C5E10786E29D07" box="[944,1020,1114,1138]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="937" unit="cm" value="10.0">10 cm</quantity>
|
||
) ventral slit midbody for DNA tissue sample, hemipenes partially everted. Snout–vent length
|
||
<quantity id="4C8B9B12D400457805D5E1C2841B9DC2" box="[160,261,1183,1207]" metricMagnitude="-1" metricUnit="m" metricValue="6.58" pageId="21" pageNumber="937" unit="mm" value="658.0">658 mm</quantity>
|
||
with 208 ventral scales, tail length
|
||
<quantity id="4C8B9B12D400457807B9E1C2862F9DC2" box="[716,817,1183,1207]" metricMagnitude="-1" metricUnit="m" metricValue="1.85" pageId="21" pageNumber="937" unit="mm" value="185.0">185 mm</quantity>
|
||
with 98 divided subcaudal scales. Anal scale single. Head length
|
||
<quantity id="4C8B9B12D4004578071BE19F87D89DAF" box="[622,710,1218,1242]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="21" pageNumber="937" unit="mm" value="27.0">27 mm</quantity>
|
||
, as measured from the snout to the rear of the quadrate, width
|
||
<quantity id="4C8B9B12D40045780493E1B887249D88" box="[486,570,1253,1277]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="937" unit="mm" value="15.0">15 mm</quantity>
|
||
, as measured across the widest point of the head anterior to the quadrate. Eyes
|
||
<quantity id="4C8B9B12D40045780765E05A87749C6A" box="[528,618,1287,1311]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="21" pageNumber="937" unit="mm" value="2.9">2.9 mm</quantity>
|
||
horizontal diameter, pupil shape subcircular. Supralabials eight with the fourth and fifth in contact with the eye. Infralabials 10 with first pair in contact behind mental, infralabials one to six in contact with inframaxillaries. Rostral broader than high,
|
||
<quantity id="4C8B9B12D400457807DBE03287E89CF2" box="[686,758,1391,1415]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="937" unit="mm" value="5.0">5 mm</quantity>
|
||
wide/
|
||
<quantity id="4C8B9B12D40045780636E03286BF9CF2" box="[835,929,1391,1415]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="937" unit="mm" value="2.5">2.5 mm</quantity>
|
||
high, visible from above. Nasal in contact with first supralabial. Single loreal, in contact with nasal, preoculars, prefrontal and supralabials two and three. Circumoculars five: one supraocular, two preoculars, zero suboculars and two postoculars. Anterior temporals two. Dorsal surface of head includes pair of internasals (width
|
||
<quantity id="4C8B9B12D40045780628E0A786A69F67" box="[861,952,1530,1554]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="21" pageNumber="937" unit="mm" value="2.9">2.9 mm</quantity>
|
||
/length of suture
|
||
<quantity id="4C8B9B12D400457805D5E34085E59F40" box="[160,251,1565,1589]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="937" unit="mm" value="1.5">1.5 mm</quantity>
|
||
), pair of prefrontals (width 4.0 mm/length of suture 4.0 mm), pair of supraoculars (width
|
||
<quantity id="4C8B9B12D4004578043CE36284B89F22" box="[329,422,1599,1623]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="21" pageNumber="937" unit="mm" value="2.5">2.5 mm</quantity>
|
||
/length
|
||
<quantity id="4C8B9B12D4004578077DE36287769F22" box="[520,616,1599,1623]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="21" pageNumber="937" unit="mm" value="4.8">4.8 mm</quantity>
|
||
), frontal longer than wide (length
|
||
<quantity id="4C8B9B12D40045780168E36281649F22" box="[1053,1146,1599,1623]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="21" pageNumber="937" unit="mm" value="5.9">5.9 mm</quantity>
|
||
/ anterior width
|
||
<quantity id="4C8B9B12D40045780422E33F84A89F0F" box="[343,438,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="21" pageNumber="937" unit="mm" value="5.4">5.4 mm</quantity>
|
||
), pair of parietals (length of suture
|
||
<quantity id="4C8B9B12D40045780606E33F86CD9F0F" box="[883,979,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="21" pageNumber="937" unit="mm" value="6.5">6.5 mm</quantity>
|
||
). Two pairs of inframaxillaries, (anterior inframaxillary suture length
|
||
<quantity id="4C8B9B12D40045780664E3D886759FE8" box="[785,875,1669,1693]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="21" pageNumber="937" unit="mm" value="2.8">2.8 mm</quantity>
|
||
, posterior inframaxillary suture length
|
||
<quantity id="4C8B9B12D4004578043FE3FA84B69FCA" box="[330,424,1703,1727]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="21" pageNumber="937" unit="mm" value="1.7">1.7 mm</quantity>
|
||
), posterior inframaxillaries in contact with each other. Dorsal scale rows 17−19−15 at 15th ventral from anterior, midbody, and 15th ventral anterior to cloaca.
|
||
</paragraph>
|
||
<caption id="DF0C667FD403457B05D5E03B81769CEF" ID-DOI="http://doi.org/10.5281/zenodo.5174533" ID-Zenodo-Dep="5174533" httpUri="https://zenodo.org/record/5174533/files/figure.png" pageId="22" pageNumber="938" startId="22.[160,225,1382,1405]" targetBox="[163,1153,142,1351]" targetPageId="22">
|
||
<paragraph id="8BCC36F7D403457B05D5E03B81769CEF" blockId="22.[160,1155,1381,1434]" pageId="22" pageNumber="938">
|
||
<emphasis id="B907EAE5D403457B05D5E03B85E39C08" bold="true" box="[160,253,1382,1405]" pageId="22" pageNumber="938">Figure 8.</emphasis>
|
||
Photographs of holotype
|
||
<taxonomicName id="4C734D74D403457B048EE03887F89C09" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[507,742,1381,1404]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="938" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D403457B048EE03887F89C09" box="[507,742,1381,1404]" italics="true" pageId="22" pageNumber="938">Stegonotus admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED403457B079BE03B86279C08" box="[750,825,1382,1405]" pageId="22" pageNumber="938" rank="species">sp. nov.</taxonomicNameLabel>
|
||
LSUMZ 93598 showing (clockwise from top left) dorsal of body, ventral of body, dorsal of head, lateral of head. Photos from Sara Ruane.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCC36F7D403457B05D5E36084109F22" blockId="22.[160,1155,1597,1727]" box="[160,270,1597,1623]" pageId="22" pageNumber="938">
|
||
<emphasis id="B907EAE5D403457B05D5E36084109F22" bold="true" box="[160,270,1597,1623]" italics="true" pageId="22" pageNumber="938">Variation</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D403457A05D5E33F84EC9999" blockId="22.[160,1155,1597,1727]" lastBlockId="23.[160,1155,142,236]" lastPageId="23" lastPageNumber="939" pageId="22" pageNumber="938">
|
||
<typeStatus id="54C88855D403457B05D5E33F84139F0F" box="[160,269,1634,1658]" pageId="22" pageNumber="938" type="paratype">Paratypes</typeStatus>
|
||
similar to the
|
||
<typeStatus id="54C88855D403457B04B5E33F873A9F0F" box="[448,548,1634,1658]" pageId="22" pageNumber="938" type="holotype">holotype</typeStatus>
|
||
excepting the following: LSUMZ 93597 and LSUMZ 93599 with 17 rather than 19 dorsal scale rows midbody; LSUMZ 93599 with nine rather than ten infralabial scales; LSUMZ 93597 with slightly darker/grey wash on the subcaudal scales compared to the
|
||
<typeStatus id="54C88855D402457A04DEE5D3870C99D3" box="[427,530,142,166]" pageId="23" pageNumber="939" type="holotype">holotype</typeStatus>
|
||
; LSUMZ 93597 with 214 ventrals, 94 subcaudals; LSUMZ 93599 with 206 ventrals, 93 subcaudals, LSUMZ 93600 with 202 ventrals, 43 subcaudals (missing large portion of tail).
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D402457A05D5E44A84749844" blockId="23.[160,1155,279,444]" box="[160,362,279,305]" pageId="23" pageNumber="939">
|
||
<emphasis id="B907EAE5D402457A05D5E44A84749844" bold="true" box="[160,362,279,305]" italics="true" pageId="23" pageNumber="939">Coloration in life</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D402457A05D5E461860298C9" blockId="23.[160,1155,279,444]" pageId="23" pageNumber="939">Dorsal ground colour in life dark brown along the vertebral scale row and fading to yellowish lighter tan/light brown ventrally but with no distinct pattern down entire dorsal side of body and tail. Dorsum of head dark brown, ventrals and subcaudals cream with no patterns. Colour of iris in life dark brown-black.</paragraph>
|
||
<paragraph id="8BCC36F7D402457A05D5E4BA84C99B74" blockId="23.[160,1156,487,756]" box="[160,471,487,513]" pageId="23" pageNumber="939">
|
||
<emphasis id="B907EAE5D402457A05D5E4BA84C99B74" bold="true" box="[160,471,487,513]" italics="true" pageId="23" pageNumber="939">Coloration in preservative</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D402457A05D5E751813E9B81" blockId="23.[160,1156,487,756]" pageId="23" pageNumber="939">Overall coloration reddish brown in preservation and lighter in colour approaching the ventrals with no pattern down the dorsal side of body and tail. Colour of iris opaque grey in preservation. Dorsum of head, including rostral, internasals, prefrontals, frontal, supraoculars and parietals reddish brown. Supralabials one to five cream, with six to eight cream with a light wash of reddish brown infringing on the anterior of the scale where the supralabials come into contact with the temporals. Ventral scales cream with no pattern, subcaudals cream with no pattern. Ventral scales show iridescence under lights.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369657CD402457505D5E67D87BD9844" lastPageId="24" lastPageNumber="940" pageId="23" pageNumber="939" type="etymology">
|
||
<paragraph id="8BCC36F7D402457A05D5E67D84039A4F" blockId="23.[160,1156,800,929]" box="[160,285,800,826]" pageId="23" pageNumber="939">
|
||
<emphasis id="B907EAE5D402457A05D5E67D84039A4F" bold="true" box="[160,285,800,826]" italics="true" pageId="23" pageNumber="939">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D402457A05D5E619849E9AD4" blockId="23.[160,1156,800,929]" pageId="23" pageNumber="939">
|
||
The species epithet refers to the Admiralty Islands where this species is found, an archipelago of 18 islands to the north of New
|
||
<collectingCountry id="F3647667D402457A07AFE63A86329A0A" box="[730,812,871,895]" name="Guinea-Bissau" pageId="23" pageNumber="939">Guinea</collectingCountry>
|
||
, comprising
|
||
<collectingRegion id="49B7F815D402457A06B3E63A819E9A0A" box="[966,1152,871,895]" country="Papua New Guinea" name="Manus" pageId="23" pageNumber="939">Manus Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3647667D402457A05D5E6D484629AD4" box="[160,380,905,929]" name="Papua New Guinea" pageId="23" pageNumber="939">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D402457A05D5E69087039A92" blockId="23.[160,1156,973,1346]" box="[160,541,973,999]" pageId="23" pageNumber="939">
|
||
<emphasis id="B907EAE5D402457A05D5E69087039A92" bold="true" box="[160,541,973,999]" italics="true" pageId="23" pageNumber="939">Natural history and distribution</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D402457A05D5E6AF819A9C37" blockId="23.[160,1156,973,1346]" pageId="23" pageNumber="939">
|
||
As with all New
|
||
<collectingCountry id="F3647667D402457A043BE6AF84829D7F" box="[334,412,1010,1034]" name="Guinea-Bissau" pageId="23" pageNumber="939">Guinea</collectingCountry>
|
||
<emphasis id="B907EAE5D402457A04D7E6AF87D19D7F" box="[418,719,1010,1034]" italics="true" pageId="23" pageNumber="939">
|
||
<taxonomicName id="4C734D74D402457A04D7E6AF87049D7F" authorityName="Dumeril, Bibron and Dumeril" authorityYear="1854" box="[418,538,1010,1034]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="genus">Stegonotus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C734D74D402457A0751E6AF87D19D7F" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[548,719,1010,1034]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">S. admiraltiensis</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A234579ED402457A07A0E6AF86349D7F" box="[725,810,1010,1034]" pageId="23" pageNumber="939" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is nocturnal. All
|
||
<specimenCount id="9D75FD7ED402457A06A9E6AF819D9D7F" box="[988,1155,1010,1034]" pageId="23" pageNumber="939" type="generic">four specimens</specimenCount>
|
||
were collected at night as they were moving on the forest floor on the leaf litter. Typical for
|
||
<taxonomicName id="4C734D74D402457A05D5E16A84099D3A" authorityName="Dumeril, Bibron and Dumeril" authorityYear="1854" box="[160,279,1079,1103]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="genus">
|
||
<emphasis id="B907EAE5D402457A05D5E16A84099D3A" box="[160,279,1079,1103]" italics="true" pageId="23" pageNumber="939">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
, once picked up
|
||
<taxonomicName id="4C734D74D402457A04B8E16A87699D3A" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[461,631,1079,1103]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D402457A04B8E16A87699D3A" box="[461,631,1079,1103]" italics="true" pageId="23" pageNumber="939">S. admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED402457A0708E16A87CF9D3A" box="[637,721,1079,1103]" pageId="23" pageNumber="939" rank="species">sp. nov.</taxonomicNameLabel>
|
||
rapidly attempted to bite the collector’ s hand and writhed violently in a side-to-side motion. Nothing more is known about the natural history or ecology of
|
||
<taxonomicName id="4C734D74D402457A0493E121878D9DE1" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[486,659,1148,1172]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D402457A0493E121878D9DE1" box="[486,659,1148,1172]" italics="true" pageId="23" pageNumber="939">S. admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED402457A07EEE12187EC9DE1" box="[667,754,1148,1172]" pageId="23" pageNumber="939" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<taxonomicName id="4C734D74D402457A078FE12181129DE1" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[762,1036,1148,1172]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D402457A078FE12181129DE1" box="[762,1036,1148,1172]" italics="true" pageId="23" pageNumber="939">Stegonotus admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED402457A0161E12181729DE1" box="[1044,1132,1148,1172]" pageId="23" pageNumber="939" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is found on the islands of Rambutyo and Los Negros in
|
||
<collectingRegion id="49B7F815D402457A07AAE1C286939DC2" box="[735,909,1183,1207]" country="Papua New Guinea" name="Manus" pageId="23" pageNumber="939">Manus Province</collectingRegion>
|
||
of
|
||
<collectingCountry id="F3647667D402457A06DBE1C2819E9DC2" box="[942,1152,1183,1207]" name="Papua New Guinea" pageId="23" pageNumber="939">Papua New Guinea</collectingCountry>
|
||
. Based on descriptions of similar specimens from
|
||
<collectingRegion id="49B7F815D402457A0791E19F86339DAF" box="[740,813,1218,1242]" country="Papua New Guinea" name="Manus" pageId="23" pageNumber="939">Manus</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFE24B06D402457A0635E19F86EC9DAF" author="McDowell SB" box="[832,1010,1218,1242]" pageId="23" pageNumber="939" pagination="6 - 26" refId="ref12286" refString="McDowell SB. 1972. The species of Stegonotus (Serpentes, Colubridae) in Papua New Guinea. Zool Meded. 47: 6 - 26." type="journal article" year="1972">McDowell 1972</bibRefCitation>
|
||
) and recent collections there by SJR that will be registered in the near future (
|
||
<bibRefCitation id="EFE24B06D402457A06DFE1B985C89C6A" author="Richards SJ & Aplin K" pageId="23" pageNumber="939" pagination="31 - 37" refId="ref12689" refString="Richards SJ, Aplin K. 2015. Herpetofauna of Manus and Mussau Islands. In: Whitmore N, editor. A rapid biodiversity survey of Papua New Guinea' s Manus and Mussau islands. Goroka: Wildlife Conservation Society Papua New Guinea Program; p. 31 - 37." type="book chapter" year="2015">Richards and Aplin 2015</bibRefCitation>
|
||
), we conclude that
|
||
<taxonomicName id="4C734D74D402457A04C8E05A87759C6A" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[445,619,1287,1311]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D402457A04C8E05A84D79C6A" box="[445,457,1287,1311]" italics="true" pageId="23" pageNumber="939">S</emphasis>
|
||
.
|
||
<emphasis id="B907EAE5D402457A04ADE05A87759C6A" box="[472,619,1287,1311]" italics="true" pageId="23" pageNumber="939">admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED402457A0703E05A87D19C6A" box="[630,719,1287,1311]" pageId="23" pageNumber="939" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is also found on
|
||
<collectingRegion id="49B7F815D402457A06EEE05A86FA9C6A" box="[923,996,1287,1311]" country="Papua New Guinea" name="Manus" pageId="23" pageNumber="939">Manus</collectingRegion>
|
||
Island proper and that all specimens of
|
||
<taxonomicName id="4C734D74D402457A04C3E07787339C37" authorityName="Dumeril, Bibron and Dumeril" authorityYear="1854" box="[438,557,1322,1346]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="genus">
|
||
<emphasis id="B907EAE5D402457A04C3E07787339C37" box="[438,557,1322,1346]" italics="true" pageId="23" pageNumber="939">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
from the Admiralty Islands are
|
||
<taxonomicName id="4C734D74D402457A06F5E07781349C37" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[896,1066,1322,1346]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D402457A06F5E07786929C37" box="[896,908,1322,1346]" italics="true" pageId="23" pageNumber="939">S</emphasis>
|
||
.
|
||
<emphasis id="B907EAE5D402457A06E2E07781349C37" box="[919,1066,1322,1346]" italics="true" pageId="23" pageNumber="939">admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED402457A0145E077819A9C37" box="[1072,1156,1322,1346]" pageId="23" pageNumber="939" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D402457A05D5E03084249CF2" blockId="23.[160,1156,1389,1727]" box="[160,314,1389,1415]" pageId="23" pageNumber="939">
|
||
<emphasis id="B907EAE5D402457A05D5E03084249CF2" bold="true" box="[160,314,1389,1415]" italics="true" pageId="23" pageNumber="939">Comparisons</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCC36F7D402457505D5E0CF87BD9844" blockId="23.[160,1156,1389,1727]" lastBlockId="24.[160,1157,142,305]" lastPageId="24" lastPageNumber="940" pageId="23" pageNumber="939">
|
||
The Manus population of
|
||
<taxonomicName id="4C734D74D402457A04BAE0CF87569CDF" authorityName="Dumeril, Bibron and Dumeril" authorityYear="1854" box="[463,584,1426,1450]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="genus">
|
||
<emphasis id="B907EAE5D402457A04BAE0CF87569CDF" box="[463,584,1426,1450]" italics="true" pageId="23" pageNumber="939">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
was thought to be conspecific with
|
||
<taxonomicName id="4C734D74D402457A068CE0CF81609CDF" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[1017,1150,1426,1450]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="B907EAE5D402457A068CE0CF81609CDF" box="[1017,1150,1426,1450]" italics="true" pageId="23" pageNumber="939">S. modestus</emphasis>
|
||
</taxonomicName>
|
||
; however, in our phylogeny it is not sister to
|
||
<taxonomicName id="4C734D74D402457A07DCE0E886339CB8" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[681,813,1461,1485]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="B907EAE5D402457A07DCE0E886339CB8" box="[681,813,1461,1485]" italics="true" pageId="23" pageNumber="939">S. modestus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13482A72D402457A064BE0E886839CB8" box="[830,925,1461,1485]" captionStart="Figure 3" captionStartId="9.[160,225,1677,1700]" captionTargetBox="[217,1128,144,1646]" captionTargetId="graphics-2@9.[256,1128,623,1646]" captionTargetPageId="9" captionText="Figure 3. Map of New Guinea sampling localities for Stegonotus (a) and mtDNA + nucDNA maximum likelihood gene tree (b). Symbols for species/clades match those on the map; * indicates bootstrap values ≥ 70. For some localities, symbols were adjusted for better visibility and so localities are approximate; see Appendix 1. for exact locality information." figureDoi="http://doi.org/10.5281/zenodo.5174522" httpUri="https://zenodo.org/record/5174522/files/figure.png" pageId="23" pageNumber="939">Figure 3</figureCitation>
|
||
). Due to the poorly supported node, it is unclear what the relationship of
|
||
<taxonomicName id="4C734D74D402457A0675E08A86AE9C9A" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[768,944,1495,1519]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis">
|
||
<emphasis id="B907EAE5D402457A0675E08A86129C9A" box="[768,780,1495,1519]" italics="true" pageId="23" pageNumber="939">S</emphasis>
|
||
.
|
||
<emphasis id="B907EAE5D402457A066CE08A86AE9C9A" box="[793,944,1495,1519]" italics="true" pageId="23" pageNumber="939">admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
is to other species in the genus.
|
||
<taxonomicName id="4C734D74D402457A044EE0A7874D9F67" authority="Ruane & Richards & McVay & Tjaturadi & Krey & Austin, 2017" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[315,595,1530,1554]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis" status="sp. nov.">
|
||
<emphasis id="B907EAE5D402457A044EE0A7874D9F67" box="[315,595,1530,1554]" italics="true" pageId="23" pageNumber="939">Stegonotus admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A234579ED402457A072EE0A787AC9F67" box="[603,690,1530,1554]" pageId="23" pageNumber="939" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs morphologically from
|
||
<taxonomicName id="4C734D74D402457A0177E0A7819D9F67" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[1026,1155,1530,1554]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="B907EAE5D402457A0177E0A7819D9F67" box="[1026,1155,1530,1554]" italics="true" pageId="23" pageNumber="939">S. modestus</emphasis>
|
||
</taxonomicName>
|
||
by having no or little dark pigmentation on the subcaudal scales and in lacking extensive dark/grey pigmentation on the supralabial and infralabial scales, which is seen in some
|
||
<emphasis id="B907EAE5D402457A0431E33F87EC9F0F" box="[324,754,1634,1658]" italics="true" pageId="23" pageNumber="939">
|
||
<taxonomicName id="4C734D74D402457A0431E33F84D49F0F" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[324,458,1634,1658]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="modestus">S. modestus</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C734D74D402457A04ADE33F87EC9F0F" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[472,754,1634,1658]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis">Stegonotus admiraltiensis</taxonomicName>
|
||
</emphasis>
|
||
also has two apicals pits on all or most dorsal scales, but these are absent in
|
||
<taxonomicName id="4C734D74D402457A07D7E3D886399FE8" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[674,807,1669,1693]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="B907EAE5D402457A07D7E3D886399FE8" box="[674,807,1669,1693]" italics="true" pageId="23" pageNumber="939">S. modestus</emphasis>
|
||
</taxonomicName>
|
||
. Superficially,
|
||
<taxonomicName id="4C734D74D402457A06BAE3D8819D9FE8" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[975,1155,1669,1693]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="admiraltiensis">
|
||
<emphasis id="B907EAE5D402457A06BAE3D8819D9FE8" box="[975,1155,1669,1693]" italics="true" pageId="23" pageNumber="939">S. admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles
|
||
<taxonomicName id="4C734D74D402457A04DAE3FA870F9FCA" authorityName="LSUMZ" baseAuthorityName="Meyer" baseAuthorityYear="1874" box="[431,529,1703,1727]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="939" phylum="Chordata" rank="species" species="parvus">
|
||
<emphasis id="B907EAE5D402457A04DAE3FA870F9FCA" box="[431,529,1703,1727]" italics="true" pageId="23" pageNumber="939">S. parvus</emphasis>
|
||
</taxonomicName>
|
||
. Both species are more tan/brown than the grey/black typical of most New
|
||
<collectingCountry id="F3647667D40D457504E5E5D384C199D3" box="[400,479,142,166]" name="Guinea-Bissau" pageId="24" pageNumber="940">Guinea</collectingCountry>
|
||
<taxonomicName id="4C734D74D40D4575049DE5D3877F99D3" authorityName="Dumeril, Bibron and Dumeril" authorityYear="1854" box="[488,609,142,166]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="940" phylum="Chordata" rank="genus">
|
||
<emphasis id="B907EAE5D40D4575049DE5D3877F99D3" box="[488,609,142,166]" italics="true" pageId="24" pageNumber="940">Stegonotus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13482A72D40D45750707E5D387CE99D3" box="[626,720,142,166]" captionStart="Figure 2" captionStartId="6.[160,225,1511,1534]" captionTargetBox="[194,1121,142,1480]" captionTargetId="figure-1@6.[193,1121,142,1480]" captionTargetPageId="6" captionText="Figure 2. Photographs of Stegonotus species with dorsal, ventral and lateral head views from left to right: (a) Stegonotus batjanensis USNM 237129, (b) Stegonotus borneensis FMNH 251054, (c) Stegonotus muelleri LSUMZ 41802, (d) Stegonotus florensis WAM 104606, (e) Stegonotus guentheri AM 129712, (f) Stegonotus heterurus BPBM 22556, (g) Stegonotus cucullatus LSUMZ 94371. Photos from SR. (h) Stegonotus modestus LSUMZ 92339, (i) Stegonotus iridis sp. nov. SJR 7514, (j) Stegonotus diehli LSUMZ 92344 (k) Stegonotus melanolabiatus sp. nov. AMS R115361 (l) Stegonotus derooijae sp. nov. SAMA R70467 (m) Stegonotus admiraltiensis sp. nov. LSUMZ 93597, (n) Stegonotus parvus LSUMZ 92333. Photos from Sara Ruane and Stephen J. Richards." figureDoi="http://doi.org/10.5281/zenodo.5174520" httpUri="https://zenodo.org/record/5174520/files/figure.png" pageId="24" pageNumber="940">Figure 2</figureCitation>
|
||
), and they both have an unpatterned cream venter and no grey mottling on the labial scales. Apart from genetic differences,
|
||
<taxonomicName id="4C734D74D40D457505D5E589844C9999" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[160,338,212,236]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="940" phylum="Chordata" rank="species" species="admiraltiensis">
|
||
<emphasis id="B907EAE5D40D457505D5E589844C9999" box="[160,338,212,236]" italics="true" pageId="24" pageNumber="940">S. admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C734D74D40D4575049DE58987529999" authorityName="LSUMZ" baseAuthorityName="Meyer" baseAuthorityYear="1874" box="[488,588,212,236]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="940" phylum="Chordata" rank="species" species="parvus">
|
||
<emphasis id="B907EAE5D40D4575049DE58987529999" box="[488,588,212,236]" italics="true" pageId="24" pageNumber="940">S. parvus</emphasis>
|
||
</taxonomicName>
|
||
in having apical pits on all or most dorsal scales, (absent in
|
||
<taxonomicName id="4C734D74D40D4575045CE5AB848A987B" authorityName="LSUMZ" baseAuthorityName="Meyer" baseAuthorityYear="1874" box="[297,404,246,270]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="940" phylum="Chordata" rank="species" species="parvus">
|
||
<emphasis id="B907EAE5D40D4575045CE5AB848A987B" box="[297,404,246,270]" italics="true" pageId="24" pageNumber="940">S. parvus</emphasis>
|
||
</taxonomicName>
|
||
), as well as a higher number of ventral scales (
|
||
<quantity id="4C8B9B12D40D4575068FE5AB819D987B" box="[1018,1155,246,270]" metricMagnitude="0" metricUnit="m" metricValue="5.283200000000001" metricValueMax="5.435600000000001" metricValueMin="5.130800000000001" pageId="24" pageNumber="940" unit="in" value="208.0" valueMax="214.0" valueMin="202.0">202−214 in</quantity>
|
||
<taxonomicName id="4C734D74D40D457505D5E444844C9844" authorityName="Ruane & Richards & McVay & Tjaturadi & Krey & Austin" authorityYear="2017" box="[160,338,281,305]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="940" phylum="Chordata" rank="species" species="admiraltiensis">
|
||
<emphasis id="B907EAE5D40D457505D5E444844C9844" box="[160,338,281,305]" italics="true" pageId="24" pageNumber="940">S. admiraltiensis</emphasis>
|
||
</taxonomicName>
|
||
versus
|
||
<quantity id="4C8B9B12D40D457504DFE44487349844" box="[426,554,281,305]" metricMagnitude="0" metricUnit="m" metricValue="4.5847" metricValueMax="4.9784" metricValueMin="4.191" pageId="24" pageNumber="940" unit="in" value="180.5" valueMax="196.0" valueMin="165.0">165−196 in</quantity>
|
||
<taxonomicName id="4C734D74D40D45750746E44487889844" authorityName="LSUMZ" baseAuthorityName="Meyer" baseAuthorityYear="1874" box="[563,662,281,305]" class="Reptilia" family="Colubridae" genus="Stegonotus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="940" phylum="Chordata" rank="species" species="parvus">
|
||
<emphasis id="B907EAE5D40D45750746E44487889844" box="[563,662,281,305]" italics="true" pageId="24" pageNumber="940">S. parvus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |