treatments-xml/data/03/DA/87/03DA87C4FF8D1762FF4C6861AB37F8C2.xml
2024-06-21 12:22:17 +02:00

219 lines
24 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="A6654A01FEF993627E4C7BE7DBADA0DD" ID-DOI="10.11646/zootaxa.3919.1.6" ID-GBIF-Dataset="4c40552e-e778-4d8e-a6b7-9662436cadc2" ID-ISSN="1175-5326" ID-Zenodo-Dep="244587" ID-ZooBank="F711CA99-1B2C-4E18-9F4B-7521D38D2303" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460989857372" checkinUser="plazi" docAuthor="Smith, Ian M., Cook, David R. &amp; Gerecke, Reinhard" docDate="2015" docId="03DA87C4FF8D1762FF4C6861AB37F8C2" docLanguage="en" docName="zt03919p156.pdf" docOrigin="Zootaxa 3919 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Forelia Haller 1882" docType="treatment" docVersion="9" lastPageNumber="116" masterDocId="FFE3FFBCFF881767FFDB6A61AA1CFF8D" masterDocTitle="Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)" masterLastPageNumber="156" masterPageNumber="111" pageNumber="116" updateTime="1698561557693" updateUser="plazi">
<mods:mods id="506354E16F32E1DFA9168EDB244F1006" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EDF17A1127C724096AE5B4ABFA82120E">
<mods:title id="335667D33749DEDEC7A416EC367B67B8">Revision of the status of some genus-level water mite taxa in the families Pionidae Thor, 1900, Aturidae Thor, 1900, and Nudomideopsidae Smith, 1990 (Acari: Hydrachnidiae)</mods:title>
</mods:titleInfo>
<mods:name id="D030435CA34AD9C67A3BCEF5D56E17D0" type="personal">
<mods:role id="5496DAEAA4CE1158A0FBFF43D3C33DA1">
<mods:roleTerm id="4EC1E1C44DC033FF931E10CC9C2368FF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DE2361A7B764E1EFC0DF17EFF0FDAE75">Smith, Ian M.</mods:namePart>
</mods:name>
<mods:name id="EC169F9C1EFAE61366FE5B0D8F4E0656" type="personal">
<mods:role id="5593CC490316E1F76DB3730D0AD4E1E1">
<mods:roleTerm id="9494FA20696372CE025984458CFCEDAD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E311FE1AF1C9AA0F5A639C95312DD107">Cook, David R.</mods:namePart>
</mods:name>
<mods:name id="64B811CF412FD062955D59AEB24659FE" type="personal">
<mods:role id="9FDA481B9B57CDBBE30298C8FE65EE45">
<mods:roleTerm id="712427AB388E3DEAE9BF174E0F5D4929">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B123091401282D04E9169A3D1764873C">Gerecke, Reinhard</mods:namePart>
</mods:name>
<mods:typeOfResource id="F5A6285CDDFEB3623AC4683AAAE4407F">text</mods:typeOfResource>
<mods:relatedItem id="347DB40BBA722D1BA49FA09E117EFA1E" type="host">
<mods:titleInfo id="90308637A9AB953764834001519D8912">
<mods:title id="42A39E1B5E02A104FED170FDDB30173C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6CC7E32846E594489F511E35EF6D5CC6">
<mods:date id="8DC734B1369A6B3B4B7B98D504AA47DF">2015</mods:date>
<mods:detail id="87634B8466ABE37A3C6E8D1D6EB2BD28" type="volume">
<mods:number id="604A7D2F0EE9216B762D6CAA852F535C">3919</mods:number>
</mods:detail>
<mods:detail id="68C502B788472BC418DBEC895AF1ED0A" type="issue">
<mods:number id="00ECFEF1220C3B1CF80A3350DA4DDB73">1</mods:number>
</mods:detail>
<mods:extent id="034022E16502E7C9BB69D9C7C3CEC5DE" unit="page">
<mods:start id="C16E78B47DB3946C2B4FEED59DE4E8EF">111</mods:start>
<mods:end id="9478904CEBE50FEF52FCC5C51D6761DD">156</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3CAB026462C94DCED1917410F004218C">journal article</mods:classification>
<mods:identifier id="B10E01CEFA9B3FFE0A038A4A1AF5D3EA" type="DOI">10.11646/zootaxa.3919.1.6</mods:identifier>
<mods:identifier id="89257C4E71BA51C4E1ACC1D2519A801B" type="GBIF-Dataset">4c40552e-e778-4d8e-a6b7-9662436cadc2</mods:identifier>
<mods:identifier id="DF4D1F0D4057BF41641F0F62C0C9E63F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="65006CB4AF4F09A5F26AC8BACCF9A7F5" type="Zenodo-Dep">244587</mods:identifier>
<mods:identifier id="65BDF5A1F7DA142B9B0251DE2A9A18DA" type="ZooBank">F711CA99-1B2C-4E18-9F4B-7521D38D2303</mods:identifier>
</mods:mods>
<treatment id="03DA87C4FF8D1762FF4C6861AB37F8C2" ID-DOI="http://doi.org/10.5281/zenodo.6109822" ID-GBIF-Taxon="119596549" ID-Zenodo-Dep="6109822" LSID="urn:lsid:plazi:treatment:03DA87C4FF8D1762FF4C6861AB37F8C2" httpUri="http://treatment.plazi.org/id/03DA87C4FF8D1762FF4C6861AB37F8C2" lastPageNumber="116" pageId="5" pageNumber="116">
<subSubSection id="C3696559FF8D1762FF4C6861A982FDF6" pageId="5" pageNumber="116" type="nomenclature">
<paragraph id="8BCC36D2FF8D1762FF4C6861ABFAFD97" blockId="5.[151,486,512,538]" box="[151,486,512,538]" pageId="5" pageNumber="116">
<heading id="D08481BEFF8D1762FF4C6861ABFAFD97" bold="true" box="[151,486,512,538]" fontSize="11" level="1" pageId="5" pageNumber="116" reason="1">
<emphasis id="B907EAC0FF8D1762FF4C6861ABFAFD97" bold="true" box="[151,486,512,538]" pageId="5" pageNumber="116">
Genus
<taxonomicName id="4C734D51FF8D1762FF356861ABFAFD97" ID-CoL="4KRJ" authority="Haller, 1882" authorityName="Haller" authorityYear="1882" box="[238,486,512,538]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FF356861AB54FD97" bold="true" box="[238,328,512,538]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
Haller, 1882
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCC36D2FF8D1762FF4C6827A8D1FDD1" blockId="5.[151,1279,582,758]" box="[151,717,582,604]" pageId="5" pageNumber="116">
<taxonomicName id="4C734D51FF8D1762FF4C6827AAFEFDD1" box="[151,226,582,604]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FF4C6827AAFEFDD1" box="[151,226,582,604]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8D1762FE9C6827ABD0FDD1" author="Cook" box="[327,460,582,604]" pageId="5" pageNumber="116" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook, 1974a</bibRefCitation>
, p. 287, figs.
<date id="FFCD1012FF8D1762FD8F6827A8D5FDD1" box="[596,713,582,604]" pageId="5" pageNumber="116" value="1208" valueMax="1213">12081213</date>
.
</paragraph>
<paragraph id="8BCC36D2FF8D1762FF4C6804A982FDF6" blockId="5.[151,1279,582,758]" box="[151,926,613,635]" pageId="5" pageNumber="116">
<taxonomicName id="4C734D51FF8D1762FF4C6804AAFEFDF6" box="[151,226,613,635]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FF4C6804AAFEFDF6" box="[151,226,613,635]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8D1762FE9C6804ABDAFDF6" author="Smith" box="[327,454,613,635]" pageId="5" pageNumber="116" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith, 1976</bibRefCitation>
, pp. 2730, 7980, 9092, figs. 1520, 2331.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8D1762FF4C68E5AE21FD7B" pageId="5" pageNumber="116" type="reference_group">
<paragraph id="8BCC36D2FF8D1762FF4C68E5A8A1FD17" blockId="5.[151,1279,582,758]" box="[151,701,644,666]" pageId="5" pageNumber="116">
<taxonomicName id="4C734D51FF8D1762FF4C68E5AB89FD17" ID-CoL="4KRJ" authority="Wainstein, 1980" authorityName="Wainstein" authorityYear="1980" box="[151,405,644,666]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FF4C68E5AAFEFD17" box="[151,226,644,666]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
:
<bibRefCitation id="EFE24B23FF8D1762FF3468E5AB89FD17" author="Wainstein" box="[239,405,644,666]" pageId="5" pageNumber="116" refString="Wainstein, B. A. (1980) Opredelitel licinok vodjanych klescej. Institut Biologii Vnutrennikh Vod., Nauka, 1 - 238." type="book chapter" year="1980">Wainstein, 1980</bibRefCitation>
</taxonomicName>
, pp. 201204, figs. 694707.
</paragraph>
<paragraph id="8BCC36D2FF8D1762FF4C68C3AE21FD7B" blockId="5.[151,1279,582,758]" pageId="5" pageNumber="116">
<taxonomicName id="4C734D51FF8D1762FF4C68C3AAFEFD35" ID-CoL="4KRJ" box="[151,226,674,696]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FF4C68C3AAFEFD35" box="[151,226,674,696]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8D1762FE9C68C3A801FD35" author="Smith" box="[327,541,674,696]" pageId="5" pageNumber="116" refString="Smith, I. M. &amp; Cook, D. R. (1991) Water mites. Chapter 16. In: Thorp, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. Academic Press, San Diego, CA, pp. 523 - 592. [total page number 911 pp.]" type="book chapter" year="1991">Smith &amp; Cook, 1991</bibRefCitation>
, pp. 554, 576, 578, figs. 16.16816.170, 16.174, 16.177, 16.260, 16.274.
<taxonomicName id="4C734D51FF8D1762FF4C68A0AAFEFD5A" ID-CoL="4KRJ" box="[151,226,705,727]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FF4C68A0AAFEFD5A" box="[151,226,705,727]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8D1762FE9C68A0ABE3FD5A" author="Smith" box="[327,511,705,727]" pageId="5" pageNumber="116" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2001) Water mites and other arachnids. Chapter 16. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 2 nd Edition. Academic Press, Waltham, Massachusetts, pp. 551 - 659. [1056 pp.] http: // dx. doi. org / 10.1016 / B 978 - 012690647 - 9 / 50017 - X" type="book chapter" year="2001">
Smith
<emphasis id="B907EAC0FF8D1762FE5068A3ABABFD5A" box="[395,439,705,727]" italics="true" pageId="5" pageNumber="116">et al</emphasis>
., 2001
</bibRefCitation>
, pp. 581, 612, 615, figs. 132136, 338340.
<taxonomicName id="4C734D51FF8D1762FF4C6881AAFEFD7B" ID-CoL="4KRJ" box="[151,226,736,758]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FF4C6881AAFEFD7B" box="[151,226,736,758]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
(in part):
<bibRefCitation id="EFE24B23FF8D1762FE9C6881ABE3FD7B" author="Smith" box="[327,511,736,758]" pageId="5" pageNumber="116" refString="Smith, I. M., Cook, D. R. &amp; Smith, B. P. (2010) Water mites and other arachnids. Chapter 15. In: Thorpe, J. &amp; Covich, A. (Eds.), Ecology and classification of North American freshwater invertebrates. 3 rd Edition. Academic Press, Waltham, Massachusetts, pp. 485 - 586. [total page number: 1021 pp.] http: // dx. doi. org / 10.1016 / B 978 - 0 - 12 - 374855 - 3.00015 - 7" type="book chapter" year="2010">
Smith
<emphasis id="B907EAC0FF8D1762FE506880ABABFD7B" box="[395,439,736,758]" italics="true" pageId="5" pageNumber="116">et al</emphasis>
., 2010
</bibRefCitation>
, pp. 524, 553, 556, figs. 15.13215.136, 15.33915.341.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8D1762FF4C6943AB7AFA12" pageId="5" pageNumber="116" type="diagnosis">
<paragraph id="8BCC36D2FF8D1762FF4C6943AB7FFB9E" blockId="5.[151,1437,802,1871]" pageId="5" pageNumber="116">
<emphasis id="B907EAC0FF8D1762FF4C6943AB7FFCB7" bold="true" box="[151,355,802,827]" pageId="5" pageNumber="116">
Diagnosis.
<emphasis id="B907EAC0FF8D1762FEC46942AB7FFCB7" bold="true" box="[287,355,803,826]" italics="true" pageId="5" pageNumber="116">Larva</emphasis>
</emphasis>
(modified from
<bibRefCitation id="EFE24B23FF8D1762FDEF6943A8A2FCB6" author="Smith" box="[564,702,802,827]" pageId="5" pageNumber="116" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith 1976</bibRefCitation>
): Character states of the subfamily Foreliinae. Lateral coxal apodeme either nearly parallel to anterior edge of second coxal plate or nearly transverse. Third coxal plate with medial coxal apodeme present and transverse muscle attachment scar present or absent. Excretory pore plate much larger than excretory pore and roughly circular or triangular in shape with setae
<emphasis id="B907EAC0FF8D1762FBD569EEAE29FC2A" box="[1038,1077,911,935]" italics="true" pageId="5" pageNumber="116">ps1</emphasis>
at least slightly anterior to setae
<emphasis id="B907EAC0FF8D1762FF4C69D2AAA2FC46" box="[151,190,947,971]" italics="true" pageId="5" pageNumber="116">ps2</emphasis>
; excretory pore sessile. Numbers of setae on leg segments as follows: IITi 9, 11 (setae Ti12 and Ti13 present), or 12 (setae Ti12, Ti13, and Ti14 present); IIITr 1; IIITi 9, 11 (setae Ti12 and Ti13 present), or 12 (setae Ti12, Ti13, and Ti14 present).
</paragraph>
<paragraph id="8BCC36D2FF8D1762FF1C6E7FAB7AFA12" blockId="5.[151,1437,802,1871]" pageId="5" pageNumber="116">
<emphasis id="B907EAC0FF8D1762FF1C6E7FAB0EFBBA" bold="true" box="[199,274,1054,1079]" italics="true" pageId="5" pageNumber="116">Adults</emphasis>
(modified from
<bibRefCitation id="EFE24B23FF8D1762FE126E7FA857FBBA" author="Smith" box="[457,587,1054,1079]" pageId="5" pageNumber="116" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith 1976</bibRefCitation>
): Character states of subfamily Foreliinae. Idiosoma with dorsal and ventral integument mostly soft and bearing only tiny platelets associated with glandularia and idiosomal setae, or bearing larger plates of various shapes that may be coalesced into shields that occupy entire dorsal and ventral surfaces. Fourth coxal plate with medial edge reduced to angle. Genital field with five or more relatively small acetabula per side borne on tongue-shaped acetabular plates that surround gonopore in males and flank gonopore in females, and that may or may not extend laterally beyond posterolateral angles of fourth coxal plates. Pedipalp segments relatively short and stocky, with tibia bearing a sessile, peg-like seta distomedially. Male third leg with tarsus bearing reduced claw sockets that are terminal in position and small but only slightly modified claws; fourth leg with genu unmodified, lacking both thick setae and a proximoventral projection, tibia relatively short, and tarsus curved dorsally, bearing thick, peg-like setae throughout curved portion, and bearing small but relatively unmodified claws.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8D1762FF1C6FCBAB8AF9A2" pageId="5" pageNumber="116" type="materials_examined">
<paragraph id="8BCC36D2FF8D1762FF1C6FCBA9EDFA4E" blockId="5.[151,1437,802,1871]" box="[199,1009,1450,1475]" pageId="5" pageNumber="116">
<emphasis id="B907EAC0FF8D1762FF1C6FCBAB7EFA4E" bold="true" box="[199,354,1450,1475]" pageId="5" pageNumber="116">
<typeStatus id="54C88870FF8D1762FF1C6FCBAB1EFA4E" box="[199,258,1450,1475]" pageId="5" pageNumber="116">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C734D51FF8D1762FEB26FCBA8B9FA4E" authority="Haller" authorityName="Haller" box="[361,677,1450,1475]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="species" species="cassidiformis">
<emphasis id="B907EAC0FF8D1762FEB26FCBA844FA4E" box="[361,600,1450,1475]" italics="true" pageId="5" pageNumber="116">Forelia cassidiformis</emphasis>
Haller
</taxonomicName>
=
<taxonomicName id="4C734D51FF8D1762FD186FCBA9F1FA4E" authority="Koch" authorityName="Koch" box="[707,1005,1450,1475]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="species" species="variegator">
<emphasis id="B907EAC0FF8D1762FD186FCBA98FFA4E" box="[707,915,1450,1475]" italics="true" pageId="5" pageNumber="116">Forelia variegator</emphasis>
(Koch)
</taxonomicName>
.
</paragraph>
<paragraph id="8BCC36D2FF8D1762FF1C6FAFAB8AF9A2" blockId="5.[151,1437,802,1871]" pageId="5" pageNumber="116">
<emphasis id="B907EAC0FF8D1762FF1C6FAFAB84FA6A" bold="true" box="[199,408,1486,1511]" pageId="5" pageNumber="116">Species included.</emphasis>
Numerous species listed by
<bibRefCitation id="EFE24B23FF8D1762FD2B6FAEA9DFFA6A" author="Viets" box="[752,963,1486,1511]" pageId="5" pageNumber="116" refString="Viets, K. O. (1987) Die Milben des Sußwassers (Hydrachnellae und Halacaridae [part.], Acari). 2. Katalog. Sonderbande des Naturwissenschaftlichen Vereins in Hamburg, 8, 1 - 1012." type="journal article" year="1987">K.O. Viets (1987)</bibRefCitation>
but excluding members of
<taxonomicName id="4C734D51FF8D1762FAD66FAFAF88FA6A" box="[1293,1428,1486,1511]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FAD66FAFAF88FA6A" box="[1293,1428,1486,1511]" italics="true" pageId="5" pageNumber="116">Madawaska</emphasis>
</taxonomicName>
. There are several undescribed species of
<taxonomicName id="4C734D51FF8D1762FDA66F93A8CDF986" box="[637,721,1522,1547]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FDA66F93A8CDF986" box="[637,721,1522,1547]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
from North
<collectingCountry id="F3647642FF8D1762FCB16F93A9D6F986" box="[874,970,1522,1547]" name="United States of America" pageId="5" pageNumber="116">America</collectingCountry>
in the Canadian National Collection of Insects and Arachnids.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8D1762FF1C6C5BABCBF9DE" box="[199,471,1594,1619]" pageId="5" pageNumber="116" type="distribution">
<paragraph id="8BCC36D2FF8D1762FF1C6C5BABCBF9DE" blockId="5.[151,1437,802,1871]" box="[199,471,1594,1619]" pageId="5" pageNumber="116">
<emphasis id="B907EAC0FF8D1762FF1C6C5BAB7CF9DE" bold="true" box="[199,352,1594,1619]" pageId="5" pageNumber="116">Distribution.</emphasis>
Holarctic.
</paragraph>
</subSubSection>
<subSubSection id="C3696559FF8D1762FF1C6C3FAB37F8C2" pageId="5" pageNumber="116" type="discussion">
<paragraph id="8BCC36D2FF8D1762FF1C6C3FAB37F8C2" blockId="5.[151,1437,802,1871]" pageId="5" pageNumber="116">
<emphasis id="B907EAC0FF8D1762FF1C6C3FAB50F9FA" bold="true" box="[199,332,1630,1655]" pageId="5" pageNumber="116">Discussion.</emphasis>
<bibRefCitation id="EFE24B23FF8D1762FE896C3FABF0F9FA" author="Cook" box="[338,492,1630,1655]" pageId="5" pageNumber="116" refString="Cook, D. R. (1974 a) Water mite genera and subgenera. Memoirs of the American Entomological Institute, 21, 1 - 860." type="journal article" year="1974" yearSuffix="a">Cook (1974a)</bibRefCitation>
and
<bibRefCitation id="EFE24B23FF8D1762FDF96C3FA8AAF9FA" author="Smith" box="[546,694,1630,1655]" pageId="5" pageNumber="116" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith (1976)</bibRefCitation>
both included
<taxonomicName id="4C734D51FF8D1762FC876C3FA9FFF9FA" box="[860,995,1630,1655]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FC876C3FA9FFF9FA" box="[860,995,1630,1655]" italics="true" pageId="5" pageNumber="116">Madawaska</emphasis>
</taxonomicName>
as a subgenus of
<emphasis id="B907EAC0FF8D1762FB7C6C3FAF1EF9FA" box="[1191,1282,1630,1655]" italics="true" pageId="5" pageNumber="116">
<taxonomicName id="4C734D51FF8D1762FB7C6C3FAEE1F9FA" box="[1191,1277,1630,1655]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">Forelia</taxonomicName>
.
</emphasis>
<bibRefCitation id="EFE24B23FF8D1762FAD36C3FAF80F9FA" author="Smith" box="[1288,1436,1630,1655]" pageId="5" pageNumber="116" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith (1976)</bibRefCitation>
pointed out that species of
<taxonomicName id="4C734D51FF8D1762FE1E6CE3A804F916" box="[453,536,1666,1691]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FE1E6CE3A804F916" box="[453,536,1666,1691]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
belong to a number of distinct groups that differ in a number of larval and adult character states. Some of these groups may be more closely related to other genera of Foreliinae than to each other. Here we consider
<taxonomicName id="4C734D51FF8D1762FEBA6CABABA8F96E" box="[353,436,1738,1763]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FEBA6CABABA8F96E" box="[353,436,1738,1763]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C734D51FF8D1762FE326CABA86CF96E" box="[489,624,1738,1763]" class="Arachnida" family="Pionidae" genus="Madawaska" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FE326CABA86CF96E" box="[489,624,1738,1763]" italics="true" pageId="5" pageNumber="116">Madawaska</emphasis>
</taxonomicName>
as separate genera. As pointed out by
<bibRefCitation id="EFE24B23FF8D1762FBC56CABAEA8F96E" author="Smith" box="[1054,1204,1738,1763]" pageId="5" pageNumber="116" refString="Smith, I. M. (1976) A study of the systematics of the water mite family Pionidae (Prostigmata: Parasitengona). Memoirs of the Entomological Society of Canada, 98, 1 - 249." type="journal article" year="1976">Smith (1976)</bibRefCitation>
,
<taxonomicName id="4C734D51FF8D1762FB646CABAF0EF96E" box="[1215,1298,1738,1763]" class="Arachnida" family="Pionidae" genus="Forelia" kingdom="Animalia" order="Trombidiformes" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B907EAC0FF8D1762FB646CABAF0EF96E" box="[1215,1298,1738,1763]" italics="true" pageId="5" pageNumber="116">Forelia</emphasis>
</taxonomicName>
as currently constituted appears to be polyphyletic and will be subject to further revision when the species level taxonomy of the North American fauna is sufficiently well known to permit the proposal of species groups that are demonstrably holophyletic.
</paragraph>
</subSubSection>
</treatment>
</document>