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<document id="FA43CA25D0681379678DA58B1B452B0B" ID-DOI="10.11646/zootaxa.4012.1.9" ID-GBIF-Dataset="4c0d8376-71b9-42d1-8dbc-08e0ff16338a" ID-ISSN="1175-5326" ID-Zenodo-Dep="244632" ID-ZooBank="55822610-6FE6-4218-A412-A080F3218F51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461239579310" checkinUser="plazi" docAuthor="Keith, Denis, Sabatinelli, Guido &amp; Uliana, Marco" docDate="2015" docId="03DA87884C3BFFE4FF0FB193FCB06DED" docLanguage="en" docName="zt04012p180.pdf" docOrigin="Zootaxa 4012 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pharaonus (Pharaonus) fasciculatus Burmeister 1844" docType="treatment" docVersion="8" lastPageNumber="171" masterDocId="FFE3FFF04C3FFFE0FF98B26BFF826853" masterDocTitle="Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa" masterLastPageNumber="180" masterPageNumber="167" pageNumber="171" updateTime="1698605019073" updateUser="plazi">
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<mods:title id="4226C2B3FAE3095C6683B84D463392DD">Synopsis of the genus Pharaonus (Coleoptera: Scarabaeidae: Rutelinae), with descriptions of new taxa</mods:title>
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<mods:namePart id="46C31DB324E9E198D648F6C4B475D845">Keith, Denis</mods:namePart>
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<mods:namePart id="9E5107E3EAD8D1005CE8A4CF4477F9DB">Sabatinelli, Guido</mods:namePart>
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<mods:namePart id="20ACD64E12C40670A6599D209411B18F">Uliana, Marco</mods:namePart>
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<mods:title id="C8BF3DB1B79575D8FA25EF344DE9C30B">Zootaxa</mods:title>
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<mods:date id="9462940D9153D7EB6F977EDF15B39A62">2015</mods:date>
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<treatment id="03DA87884C3BFFE4FF0FB193FCB06DED" ID-DOI="http://doi.org/10.5281/zenodo.5694704" ID-GBIF-Taxon="119632619" ID-Zenodo-Dep="5694704" LSID="urn:lsid:plazi:treatment:03DA87884C3BFFE4FF0FB193FCB06DED" httpUri="http://treatment.plazi.org/id/03DA87884C3BFFE4FF0FB193FCB06DED" lastPageNumber="171" pageId="4" pageNumber="171">
<subSubSection id="C36965154C3BFFE4FF0FB193FEA26C60" pageId="4" pageNumber="171" type="nomenclature">
<paragraph id="8BCC369E4C3BFFE4FF0FB193FCC66C41" blockId="4.[151,836,1016,1075]" box="[151,836,1016,1042]" pageId="4" pageNumber="171">
<heading id="D08481F24C3BFFE4FF0FB193FCC66C41" bold="true" box="[151,836,1016,1042]" fontSize="11" level="1" pageId="4" pageNumber="171" reason="1">
<taxonomicName id="4C734D1D4C3BFFE4FF0FB193FCC66C41" authority="Burmeister, 1844" authorityName="Burmeister" authorityYear="1844" box="[151,836,1016,1042]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="fasciculatus" subGenus="Pharaonus">
<emphasis id="B907EA8C4C3BFFE4FF0FB193FCC66C41" bold="true" box="[151,836,1016,1042]" pageId="4" pageNumber="171">
<emphasis id="B907EA8C4C3BFFE4FF0FB193FE9F6C41" bold="true" box="[151,285,1016,1042]" italics="true" pageId="4" pageNumber="171">Pharaonus</emphasis>
(
<emphasis id="B907EA8C4C3BFFE4FEB5B193FE306C41" bold="true" box="[301,434,1016,1042]" italics="true" pageId="4" pageNumber="171">Pharaonus</emphasis>
)
<emphasis id="B907EA8C4C3BFFE4FE5CB193FDD76C41" bold="true" box="[452,597,1016,1042]" italics="true" pageId="4" pageNumber="171">fasciculatus</emphasis>
(
<bibRefCitation id="EFE24B6F4C3BFFE4FDFCB193FCB96C41" author="Burmeister" box="[612,827,1016,1042]" pageId="4" pageNumber="171" refString="Burmeister, H. C. C. (1844) Handbuch der Entomologie. Vierter Band, Erste Abtheilung. Coleoptera Lamellicornia Anthobia et Phyllophaga systellochela. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1844">Burmeister, 1844</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCC369E4C3BFFE4FF0FB671FEA26C60" blockId="4.[151,836,1016,1075]" box="[151,288,1050,1075]" pageId="4" pageNumber="171">
(
<figureCitation id="13482A1B4C3BFFE4FF07B671FF6C6C61" box="[159,238,1050,1075]" captionStart="FIGURES 7 10" captionStartId="5.[151,264,1955,1977]" captionTargetBox="[211,1377,193,1914]" captionTargetId="figure@5.[196,1390,193,1933]" captionTargetPageId="5" captionText="FIGURES 7 10. Habitus of Pharaonus (Pharaonus) species. 7: P. fasciculatus, male (Egypt). 8: P. saharicus, male paratype (Tamrit, Tassili nAjjer, Algeria). 9: P. varicoloreus, male (environs of Palymra, Syria). 10: P. varicoloreus, female (Birecik, Turkey)." httpUri="https://zenodo.org/record/244635/files/figure.png" pageId="4" pageNumber="171">Figs. 7</figureCitation>
,
<figureCitation id="13482A1B4C3BFFE4FF62B670FE956C60" box="[250,279,1051,1075]" captionStart="FIGURES 15 23" captionStartId="8.[151,264,1847,1869]" captionTargetBox="[174,1402,241,1792]" captionTargetId="figure@8.[174,1405,193,1813]" captionTargetPageId="8" captionText="FIGURES 15 23. Genitalia of Pharaonus. 15 19: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Pharaonus): 15: P. adelphus, paratype (a-c) and holotype (d) (Kerman, Iran); 16: P. caucasicus (Gorovan, Armenia); 17: P. fasciculatus (Egypt); 18: P. farsensis, holotype (Darab, Iran); 19: P. saharicus, paratype (Tamrit, Tassili nAjjer, Algeria); 20: P. varicoloreus (Halfeti, Birecik, Adiyaman, Turkey). 21: female genital sclerites of P. adelphus (Kerman, Bakram-djord [= Bahramjerd], Iran). 22 23: aedeagus in dorsal, ventral, and lateral view of Pharaonus (Tamerlanius): 22: P. lederi (Koron 15 km Ekoc. Dzejran, Buckara, Uzbekistan); 23: P. semenowi (Kul'dzhuktau, Kizilkum desert, Uzbekistan)." httpUri="https://zenodo.org/record/244638/files/figure.png" pageId="4" pageNumber="171">17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36965154C3BFFE4FF0FB609FB686CB4" pageId="4" pageNumber="171" type="materials_examined">
<paragraph id="8BCC369E4C3BFFE4FF0FB609FB686CB4" blockId="4.[151,1437,1122,1471]" pageId="4" pageNumber="171">
<emphasis id="B907EA8C4C3BFFE4FF0FB609FE0A6C28" bold="true" box="[151,392,1122,1147]" pageId="4" pageNumber="171">Examined material.</emphasis>
[
<collectingCountry id="F364760E4C3BFFE4FE02B609FE616C28" box="[410,483,1122,1147]" name="Libya" pageId="4" pageNumber="171">Libya</collectingCountry>
], Trip. [Tripolitania], Sirte,
<date id="FFCD105E4C3BFFE4FCB4B609FC236C28" box="[812,929,1122,1147]" pageId="4" pageNumber="171" value="1936-05-21">21.V.1936</date>
, leg. R.e C. Koch:
<specimenCount id="9D75FD174C3BFFE4FB15B609FB3E6C29" box="[1165,1212,1122,1147]" pageId="4" pageNumber="171" type="generic">5 ♂</specimenCount>
(
<collectionCode id="ED62AE5B4C3BFFE4FB57B608FAAB6C28" box="[1231,1321,1123,1147]" pageId="4" pageNumber="171">MSNM</collectionCode>
,
<collectionCode id="ED62AE5B4C3BFFE4FAA1B608FA136C28" box="[1337,1425,1123,1147]" httpUri="http://grbio.org/cool/vphy-ev73" name="Mie University Culture Collection (Culture Collection, Laboratory of Plant Pathology)" pageId="4" pageNumber="171">MUCC</collectionCode>
). [
<collectingCountry id="F364760E4C3BFFE4FF05B6EDFF646CCC" box="[157,230,1158,1183]" name="Libya" pageId="4" pageNumber="171">Libya</collectingCountry>
], Trip. [Tripolitania], Sirte,
<date id="FFCD105E4C3BFFE4FDB0B6EDFD1F6CCC" box="[552,669,1158,1183]" pageId="4" pageNumber="171" value="1936-05-25">25.V.1936</date>
, leg. R.e C. Koch:
<specimenCount id="9D75FD174C3BFFE4FC1AB6ECFC2D6CCD" box="[898,943,1159,1183]" pageId="4" pageNumber="171" type="generic">1 ♂</specimenCount>
(
<collectionCode id="ED62AE5B4C3BFFE4FC59B6ECFB9F6CCC" box="[961,1053,1159,1183]" pageId="4" pageNumber="171">MSNM</collectionCode>
).
<collectingCountry id="F364760E4C3BFFE4FBAAB6ECFBFA6CCC" box="[1074,1144,1159,1183]" name="Egypt" pageId="4" pageNumber="171">Egypt</collectingCountry>
, ante 1892, [coll.?] H.W. Bates:
<specimenCount id="9D75FD174C3BFFE4FF70B6C0FEF86C90" box="[232,378,1194,1219]" pageId="4" pageNumber="171" type="generic">8 specimens</specimenCount>
, (
<collectionCode id="ED62AE5B4C3BFFE4FE09B6C0FE616C90" box="[401,483,1195,1219]" pageId="4" pageNumber="171">DKCC</collectionCode>
,
<collectionCode id="ED62AE5B4C3BFFE4FE6BB6C0FDCC6C90" LSID="urn:lsid:biocol.org:col:34988" box="[499,590,1195,1219]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="171">MNHN</collectionCode>
); Eg. [
<collectingCountry id="F364760E4C3BFFE4FD06B6C0FD6A6C90" box="[670,744,1195,1219]" name="Egypt" pageId="4" pageNumber="171">Egypt</collectingCountry>
], Kirdassah [=Kirdasah],
<date id="FFCD105E4C3BFFE4FB8DB6C1FBC16C90" box="[1045,1091,1194,1219]" pageId="4" pageNumber="171">6.V.</date>
[19]33, leg. W. Wittmer:
<specimenCount id="9D75FD174C3BFFE4FAF6B6C1FA1E6C91" box="[1390,1436,1194,1219]" pageId="4" pageNumber="171" type="generic">5 ♂</specimenCount>
(
<collectionCode id="ED62AE5B4C3BFFE4FF06B6A4FF7A6CB4" box="[158,248,1231,1255]" pageId="4" pageNumber="171">MSNM</collectionCode>
,
<collectionCode id="ED62AE5B4C3BFFE4FE9CB6A4FEDE6CB4" box="[260,348,1231,1255]" httpUri="http://grbio.org/cool/vphy-ev73" name="Mie University Culture Collection (Culture Collection, Laboratory of Plant Pathology)" pageId="4" pageNumber="171">MUCC</collectionCode>
);
<collectingCountry id="F364760E4C3BFFE4FEF6B6A5FE2D6CB4" box="[366,431,1230,1255]" name="Israel" pageId="4" pageNumber="171">Israel</collectingCountry>
, North of Jericho, Ein Duyuk,
<date id="FFCD105E4C3BFFE4FC90B6A4FCFF6CB4" box="[776,893,1231,1255]" pageId="4" pageNumber="171" value="1971-05-28">28.V.1971</date>
, leg. Fammishes:
<specimenCount id="9D75FD174C3BFFE4FBDFB6A4FBF66CB5" box="[1095,1140,1231,1255]" pageId="4" pageNumber="171" type="generic">1 ♂</specimenCount>
. (
<collectionCode id="ED62AE5B4C3BFFE4FB1FB6A4FB5C6CB4" box="[1159,1246,1231,1255]" pageId="4" pageNumber="171">TAUM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36965154C3BFFE4FF5FB699FC806D58" box="[199,770,1266,1291]" pageId="4" pageNumber="171" type="distribution">
<paragraph id="8BCC369E4C3BFFE4FF5FB699FC806D58" blockId="4.[151,1437,1122,1471]" box="[199,770,1266,1291]" pageId="4" pageNumber="171">
<emphasis id="B907EA8C4C3BFFE4FF5FB699FEE26D58" bold="true" box="[199,352,1266,1291]" pageId="4" pageNumber="171">Distribution.</emphasis>
<collectingCountry id="F364760E4C3BFFE4FEFFB699FE3F6D58" box="[359,445,1266,1291]" name="Tunisia" pageId="4" pageNumber="171">Tunisia</collectingCountry>
,
<collectingCountry id="F364760E4C3BFFE4FE50B699FD8E6D58" box="[456,524,1266,1291]" name="Libya" pageId="4" pageNumber="171">Libya</collectingCountry>
,
<collectingCountry id="F364760E4C3BFFE4FD8FB698FDDF6D58" box="[535,605,1267,1291]" name="Egypt" pageId="4" pageNumber="171">Egypt</collectingCountry>
,
<collectingCountry id="F364760E4C3BFFE4FDF0B699FD336D58" box="[616,689,1266,1291]" name="Sudan" pageId="4" pageNumber="171">Sudan</collectingCountry>
,
<collectingCountry id="F364760E4C3BFFE4FD24B699FD7C6D58" box="[700,766,1266,1291]" name="Israel" pageId="4" pageNumber="171">Israel</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36965154C3BFFE4FF5FB77DFCB06DED" pageId="4" pageNumber="171" type="discussion">
<paragraph id="8BCC369E4C3BFFE4FF5FB77DFE6E6D00" blockId="4.[151,1437,1122,1471]" pageId="4" pageNumber="171">
<emphasis id="B907EA8C4C3BFFE4FF5FB77DFEB96D7C" bold="true" box="[199,315,1302,1327]" pageId="4" pageNumber="171">Remarks.</emphasis>
First record for
<collectingCountry id="F364760E4C3BFFE4FE66B77DFDC26D7C" box="[510,576,1302,1327]" name="Israel" pageId="4" pageNumber="171">Israel</collectingCountry>
. The examined specimen was previously identified as
<taxonomicName id="4C734D1D4C3BFFE4FB22B77CFAE76D7C" box="[1210,1381,1302,1327]" class="Insecta" family="Rutelidae" genus="Pharaonus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="171" phylum="Arthropoda" rank="species" species="varicoloreus">
<emphasis id="B907EA8C4C3BFFE4FB22B77CFAE76D7C" box="[1210,1381,1302,1327]" italics="true" pageId="4" pageNumber="171">P. varicoloreus</emphasis>
</taxonomicName>
(see discussion under that species).
</paragraph>
<paragraph id="8BCC369E4C3BFFE4FF5FB734FCB06DED" blockId="4.[151,1437,1122,1471]" pageId="4" pageNumber="171">
An informal record published on the web (
<bibRefCitation id="EFE24B6F4C3BFFE4FD3FB735FCF06D24" author="Ringenbach" box="[679,882,1374,1399]" pageId="4" pageNumber="171" refString="Ringenbach, J. C. (2006) Beetles and Rock Art in Libya. Available from: http: // jcringenbach. free. fr / website / beetles / scarabaeidae / Pharaonus _ fasciculatus. htm (accessed 17 February 2013)" type="journal article" year="2006">Ringenbach 2006</bibRefCitation>
), documented by the photograph of the specimen (a male, black form, from Sabratah,
<date id="FFCD105E4C3BFFE4FDB1B7E9FD1D6DC8" box="[553,671,1410,1435]" pageId="4" pageNumber="171" value="2005-05-06">06.V.2005</date>
, leg. P. Weill) should be added to the first records for
<collectingCountry id="F364760E4C3BFFE4FB69B7E9FAB06DC8" box="[1265,1330,1410,1435]" name="Libya" pageId="4" pageNumber="171">Libya</collectingCountry>
provided by
<bibRefCitation id="EFE24B6F4C3BFFE4FF22B7CDFEBF6DEC" author="Kral" box="[186,317,1446,1471]" pageId="4" pageNumber="171" refString="Kral, D. (2012) Pharaonus saharicus sp. nov. (Coleoptera: Scarabaeidae: Rutelinae) from the Central Sahara (Algeria). Klapalekiana, 48, 127 - 136." type="journal article" year="2012">Král (2012)</bibRefCitation>
and to those mentioned in the present paper.
</paragraph>
</subSubSection>
</treatment>
</document>