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<document id="8899D07CF0B2F6B22CFAD5D6834E0DD3" ID-CLB-Dataset="28528" ID-DOI="10.11646/zootaxa.4556.1.1" ID-GBIF-Dataset="9ffb3429-ba6c-4d23-a88e-781dde361e56" ID-ISSN="1175-5326" ID-Zenodo-Dep="2585204" ID-ZooBank="7C55D5CB-5A0F-4DCE-A5A7-755339CF45F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1551120081848" checkinUser="plazi" docAuthor="Qiu, Jian-Yue, Xu, Hao &amp; Chen, Li" docDate="2019" docId="03CA3425FFB2A40AC2E9FEAA161D1571" docLanguage="en" docName="zootaxa.4556.1.1.pdf" docOrigin="Zootaxa 4556 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Macronotops sexmaculatus" docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="FFF34C5DFF90A429C27EFF3E140B123D" masterDocTitle="A revision of the rare flower beetle genus Macronotops Krikken (Coleoptera: Scarabaeidae: Cetoniinae) from Asia with biological notes" masterLastPageNumber="65" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1550769681392" originalUpdateUser="plazi" pageNumber="35" updateTime="1698700146227" updateUser="ExternalLinkService">
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<mods:title id="48B97BFB0F02AF10ED1F17B5544D08A4">A revision of the rare flower beetle genus Macronotops Krikken (Coleoptera: Scarabaeidae: Cetoniinae) from Asia with biological notes</mods:title>
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<treatment id="03CA3425FFB2A40AC2E9FEAA161D1571" ID-DOI="http://doi.org/10.5281/zenodo.5934017" ID-GBIF-Taxon="152633818" ID-Zenodo-Dep="5934017" LSID="urn:lsid:plazi:treatment:03CA3425FFB2A40AC2E9FEAA161D1571" httpUri="http://treatment.plazi.org/id/03CA3425FFB2A40AC2E9FEAA161D1571" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
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<taxonomicName id="4C63FEB0FFB2A40BC2E9FEAA16951393" ID-CoL="3X78X" authority="(Kraatz, 1894)" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[151,670,404,430]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB2A40BC2E9FEAA16951393" bold="true" box="[151,670,404,430]" pageId="34" pageNumber="35">
<emphasis id="B9175921FFB2A40BC2E9FEAA15E91393" bold="true" box="[151,482,404,430]" italics="true" pageId="34" pageNumber="35">Macronotops sexmaculatus</emphasis>
(
<bibRefCitation id="EFF2F8C2FFB2A40BC38FFEAA169D1393" author="Kraatz, G." box="[497,662,404,430]" pageId="34" pageNumber="35" pagination="141" refId="ref34691" refString="Kraatz, G. (1894) Pleuronota 6 - maculata n. sp. Deutsche Entomologische Zeitschrift, 1894 (1), 141." type="journal article" year="1894">Kraatz, 1894</bibRefCitation>
)
</emphasis>
</taxonomicName>
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<subSubSection id="C379D6B8FFB2A40BC2E9FE8810C213F2" box="[151,1225,438,463]" pageId="34" pageNumber="35" type="description">
<paragraph id="8BDC8533FFB2A40BC2E9FE8810C213F2" blockId="34.[151,1225,404,463]" box="[151,1225,438,463]" pageId="34" pageNumber="35">
(
<figureCitation id="135899B6FFB2A40BC2E1FE88152D13F2" box="[159,294,438,463]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="34" pageNumber="35">Figs. 1415</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC34CFE89157113F2" box="[306,378,439,463]" captionStart="FIGURES 2447" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 2447. Protibia of Macronotops (male and female). 24, M. biserratus new species; 2526, M. dianensis new species; 2728, M. fulvoguttatus; 2930, M. fulvopilosus; 3132, M. miksici new species; 3334, M. nigropubescens; 3536, M. olivaceofuscus; 3738, M. sexmaculatus; 3940, M. vuilleti; 41, Macronotops sp.; 4243, M. curvimarginatus; 4445, M. medogensis new species; 4647, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="34" pageNumber="35">3738</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC3F8FE8815C413F2" box="[390,463,438,463]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="34" pageNumber="35">6162</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC3A4FE89162813F2" box="[474,547,439,463]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="34" pageNumber="35">9293</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC051FE89165113F2" box="[559,602,439,463]" captionStart="FIGURES 111122" captionStartId="16.[151,264,1744,1767]" captionTargetBox="[156,1403,544,1706]" captionTargetId="figure@16.[151,1436,525,1723]" captionTargetPageId="16" captionText="FIGURES 111122. Male genitalia of Macronotops (apical view). 111, M. biserratus new species; 112, M. dianensis new species; 113, M. fulvoguttatus; 114, M. fulvopilosus; 115, M. miksici new species; 116, M. nigropubescens; 117, M. olivaceofuscus; 118, M. sexmaculatus; 119, M. vuilleti. 120, M. curvimarginatus; 121, M. medogensis new species; 122, M. ovaliceps. Arrows indicate the membrane between the parameres." httpUri="https://zenodo.org/record/2585220/files/figure.png" pageId="34" pageNumber="35">118</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC01BFE89169913F2" box="[613,658,439,463]" captionStart="FIGURES 122134" captionStartId="17.[151,264,1757,1780]" captionTargetBox="[168,1424,207,1720]" captionTargetId="figure@17.[151,1436,193,1737]" captionTargetPageId="17" captionText="FIGURES 122134. Male genitalia of Macronotops (lateral view). 123, M. biserratus new species; 124, M. dianensis new species; 125, M. fulvoguttatus; 126, M. fulvopilosus; 127, M. miksici new species; 128, M. nigropubescens; 129, M. olivaceofuscus; 130, M. sexmaculatus; 131, M. vuilleti; 132, M. curvimarginatus; 133, M. medogensis new species; 134, M. ovaliceps." httpUri="https://zenodo.org/record/2585222/files/figure.png" pageId="34" pageNumber="35">130</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC0E3FE88170A13F2" box="[669,769,438,463]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="34" pageNumber="35">165168</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC173FE89177913F2" box="[781,882,439,463]" captionStart="FIGURES 215226" captionStartId="51.[151,264,1940,1963]" captionTargetBox="[165,1421,187,1916]" captionTargetId="figure@51.[151,1436,181,1920]" captionTargetPageId="51" captionText="FIGURES 215226. Habitus of Macronotops (male in left, female in right; oblique view). 215216, M. olivaceofuscus; 217 218, M. sexmaculatus; 219220, M. vuilleti; 221222, M. curvimarginatus; 223224, M. medogensis new species; 225226, M. ovaliceps." httpUri="https://zenodo.org/record/2585256/files/figure.png" pageId="34" pageNumber="35">217218</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC103FE8917E913F2" box="[893,994,439,463]" captionStart="FIGURES 227234" captionStartId="52.[151,264,1245,1268]" captionTargetBox="[153,1432,197,1212]" captionTargetId="figure@52.[151,1436,193,1225]" captionTargetPageId="52" captionText="FIGURES 227234. Type specimens and labels of Macronotops. 227229, M. fulvopilosus, male holotype (provided by Antoine Mantilleri); 230234, M. sexmaculatus, female holotype (provided by Antoine Mantilleri)." httpUri="https://zenodo.org/record/2585258/files/figure.png" pageId="34" pageNumber="35">230234</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC193FE88105913F2" box="[1005,1106,438,463]" captionStart="FIGURES 258260" captionStartId="55.[151,264,1245,1268]" captionTargetBox="[151,1436,193,1224]" captionTargetId="figure@55.[151,1436,193,1224]" captionTargetPageId="55" captionText="FIGURES 258260. Known distribution of Macronotops. 258, Distribution for the whole genus; 259, map of SE Xizang, China; 260, map of W Sichuan, China. Syncretic markers indicate identical or very close localities; type localities of the known species are underlined, that of M. olivaceofuscus is missing because the precise type locality is unknown." httpUri="https://zenodo.org/record/2585264/files/figure.png" pageId="34" pageNumber="35">258259</figureCitation>
,
<figureCitation id="135899B6FFB2A40BC623FE8810CA13F2" box="[1117,1217,438,463]" captionStart="FIGURES 276281" captionStartId="58.[151,264,1812,1835]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@58.[151,1436,193,1792]" captionTargetPageId="58" captionText="FIGURES 276281. Habitat of Macronotops. 276277, habitat of M. sexmaculatus and M. medogensis new species in Valley near Bononggong of Mêdog (Xizang, China); 278, an adult of M. sexmaculatus feeding on sap of bamboo (photograph by Yang He); 279281, habitat of M. vuilleti in Mount Huanglianshan (Yunnan, China), and larva in fallen rotten wood." httpUri="https://zenodo.org/record/2585270/files/figure.png" pageId="34" pageNumber="35">276278</figureCitation>
)
</paragraph>
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<taxonomicName id="4C63FEB0FFB2A40BC2E9FEC016421029" ID-CoL="77RR4" authority="Kraatz, 1894: 141" authorityName="Kraatz" authorityPageNumber="141" authorityYear="1894" box="[151,585,510,532]" class="Insecta" family="Scarabaeidae" genus="Pleuronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis id="B9175921FFB2A40BC2E9FEC015871029" box="[151,396,510,532]" italics="true" pageId="34" pageNumber="35">Pleuronota sexmaculata</emphasis>
<treatmentCitation id="0AC2A322FFB2A40BC3EDFEC116421029" author="Kraatz, G." box="[403,585,510,532]" page="141" pageId="34" pageNumber="35" year="1894">
<bibRefCitation id="EFF2F8C2FFB2A40BC3EDFEC116111029" author="Kraatz, G." box="[403,538,510,532]" pageId="34" pageNumber="35" pagination="141" refId="ref34691" refString="Kraatz, G. (1894) Pleuronota 6 - maculata n. sp. Deutsche Entomologische Zeitschrift, 1894 (1), 141." type="journal article" year="1894">Kraatz, 1894</bibRefCitation>
: 141
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</taxonomicName>
(
<typeStatus id="54D83B91FFB2A40BC028FEC016881029" box="[598,643,510,532]" pageId="34" pageNumber="35">type</typeStatus>
locality: Darjeeling, N.
<collectingCountry id="F374C5A3FFB2A40BC107FEC017B81029" box="[889,947,510,532]" name="India" pageId="34" pageNumber="35">India</collectingCountry>
)*;
<treatmentCitation id="0AC2A322FFB2A40BC1AEFEC010761029" author="Schoch, G." box="[976,1149,510,532]" page="37" pageId="34" pageNumber="35" year="1896">
<bibRefCitation id="EFF2F8C2FFB2A40BC1AEFEC0105C1029" author="Schoch, G." box="[976,1111,510,532]" pageId="34" pageNumber="35" refId="ref35852" refString="Schoch, G. (1896) Catalogus systematicus Cetonidarum et Trichiidarum ad huc cognitarum. published not identified, Zurich, 95 pp." type="book" year="1896">Schoch 1896</bibRefCitation>
: 37
</treatmentCitation>
;
<treatmentCitation id="0AC2A322FFB2A40BC6F9FEC011621029" author="Schenkling, S." box="[1159,1385,510,532]" page="144" pageId="34" pageNumber="35" year="1921">
<bibRefCitation id="EFF2F8C2FFB2A40BC6F9FEC0113D1029" author="Schenkling, S." box="[1159,1334,510,532]" pageId="34" pageNumber="35" pagination="1 - 431" refId="ref35805" refString="Schenkling, S. (1921) Scarabaeidae: Cetoniinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Fol. XXI. Scarabaeidae III Pars. 72. W. Junk, Berlin, pp. 1 - 431." type="book chapter" year="1921">Schenkling 1921</bibRefCitation>
: 144
</treatmentCitation>
.
<taxonomicName id="4C63FEB0FFB2A40BC2E9FD2015E2100E" authority="(Kraatz)" baseAuthorityName="Kraatz" box="[151,489,541,563]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis id="B9175921FFB2A40BC2E9FD201581100E" box="[151,394,541,563]" italics="true" pageId="34" pageNumber="35">Macronota sexmaculata</emphasis>
(Kraatz)
</taxonomicName>
:
<treatmentCitation id="0AC2A322FFB2A40BC38DFD23169F100E" author="Arrow, G. J." box="[499,660,541,563]" page="47" pageId="34" pageNumber="35" year="1910">
<bibRefCitation id="EFF2F8C2FFB2A40BC38DFD23167A100E" author="Arrow, G. J." box="[499,625,541,563]" pageId="34" pageNumber="35" refId="ref34094" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma). Lamellicornia (Cetoniinae and Dynastinae). Taylor &amp; Francis, London, 322 pp." type="book" year="1910">Arrow 1910</bibRefCitation>
: 47
</treatmentCitation>
(partim, specimens from
<collectingCountry id="F374C5A3FFB2A40BC1E7FD2317EC100E" box="[921,999,541,563]" name="Bhutan" pageId="34" pageNumber="35">Bhutan</collectingCountry>
);
<treatmentCitation id="0AC2A322FFB2A40BC186FD2310CF100E" author="Bourgoin, A." box="[1016,1220,541,563]" page="137" pageId="34" pageNumber="35" year="1916">
<bibRefCitation id="EFF2F8C2FFB2A40BC186FD23109F100E" author="Bourgoin, A." box="[1016,1172,541,563]" pageId="34" pageNumber="35" pagination="133 - 137" refId="ref34313" refString="Bourgoin, A. (1916) Description de trois Macronota nouveaux (Col. Scarabaeidae). Bulletin de la Societe Entomologique de France Paris, 1916 (7), 133 - 137." type="journal article" year="1916">Bourgoin 1916</bibRefCitation>
: 137
</treatmentCitation>
(remarks).
<taxonomicName id="4C63FEB0FFB2A40BC2E9FD03160B106F" authority="(Kraatz)" baseAuthorityName="Kraatz" box="[151,512,572,594]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis id="B9175921FFB2A40BC2E9FD0315AA106F" box="[151,417,572,594]" italics="true" pageId="34" pageNumber="35">Macronotops sexmaculata</emphasis>
(Kraatz)
</taxonomicName>
:
<treatmentCitation id="0AC2A322FFB2A40BC074FD0216C2106F" author="Krikken, J." box="[522,713,572,594]" page="208" pageId="34" pageNumber="35" year="1977">
<bibRefCitation id="EFF2F8C2FFB2A40BC074FD021692106F" author="Krikken, J." box="[522,665,572,594]" pageId="34" pageNumber="35" pagination="199 - 209" refId="ref34797" refString="Krikken, J. (1977) The Asian genus Pleuronota Kraatz and allied forms: a clarification (Coleoptera: Cetoniidae). Zoologische Mededelingen, 51 (13), 199 - 209." type="journal article" year="1977">Krikken 1977</bibRefCitation>
: 208
</treatmentCitation>
(partim);
<treatmentCitation id="0AC2A322FFB2A40BC151FD0217D6106F" author="Krajcik, M." box="[815,989,572,594]" page="90" pageId="34" pageNumber="35" year="1998">
<bibRefCitation id="EFF2F8C2FFB2A40BC151FD0217B3106F" author="Krajcik, M." box="[815,952,572,594]" pageId="34" pageNumber="35" refId="ref34717" refString="Krajcik, M. (1998) Cetoniidae of the World Catalogue-Part I. Typos Studio, Most, 96 pp." type="book" year="1998">Krajčík 1998</bibRefCitation>
: 90
</treatmentCitation>
;
<treatmentCitation id="0AC2A322FFB2A40BC196FD0210A4106F" author="Smetana, A." box="[1000,1199,572,594]" page="307" pageId="34" pageNumber="35" year="2006">
<bibRefCitation id="EFF2F8C2FFB2A40BC196FD021077106F" author="Smetana, A." box="[1000,1148,572,594]" pageId="34" pageNumber="35" pagination="283 - 313" refId="ref35877" refString="Smetana, A. (2006) Subfamily Cetoniinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera Folume 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Apollo Books, Stenstrup, pp. 283 - 313." type="book chapter" year="2006">Smetana 2006</bibRefCitation>
: 307
</treatmentCitation>
;
<treatmentCitation id="0AC2A322FFB2A40BC6C7FD02117F106F" author="Krajcik, M." box="[1209,1396,572,594]" page="152" pageId="34" pageNumber="35" year="2012">
<bibRefCitation id="EFF2F8C2FFB2A40BC6C7FD021149106F" author="Krajcik, M." box="[1209,1346,572,594]" pageId="34" pageNumber="35" pagination="1 - 278" refId="ref34772" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement, 5, 1 - 278." type="journal article" year="2012">Krajčík 2012</bibRefCitation>
: 152
</treatmentCitation>
.
<taxonomicName id="4C63FEB0FFB2A40BC2E9FD651601104D" ID-CoL="3X78X" authority="(Kraatz)" authorityName="Kraatz" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[151,522,602,624]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB2A40BC2E9FD6515A0104D" box="[151,427,602,624]" italics="true" pageId="34" pageNumber="35">Macronotops sexmaculatus</emphasis>
(Kraatz)
</taxonomicName>
:
<treatmentCitation id="0AC2A322FFB2A40BC06DFD6416C4104D" author="Bezdek, A." box="[531,719,602,624]" page="403" pageId="34" pageNumber="35" year="2016">
<bibRefCitation id="EFF2F8C2FFB2A40BC06DFD641696104D" author="Bezdek, A." box="[531,669,602,624]" pageId="34" pageNumber="35" pagination="367 - 412" refId="ref34250" refString="Bezdek, A. (2016) Subfamily Cetoniinae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera, Folume 3, Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, pp. 367 - 412." type="book chapter" year="2016">Bezděk 2016</bibRefCitation>
: 403
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB2A40BC2E9FDA315021797" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BDC8533FFB2A40BC2E9FDA31526111F" blockId="34.[151,1437,669,2026]" pageId="34" pageNumber="35">
<emphasis id="B9175921FFB2A40BC2E9FDA315CE108B" bold="true" box="[151,453,669,694]" pageId="34" pageNumber="35">
Type material.
<materialsCitation id="3B0B8F6EFFB2A40BC326FDA315CE108B" ID-GBIF-Occurrence="2004970356" box="[344,453,669,694]" collectionCode="MNHN" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="54D83B91FFB2A40BC326FDA315CE108B" box="[344,453,669,694]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
labeled: (
<specimenCount id="9D654EBAFFB2A40BC03BFDA01653108B" box="[581,600,670,694]" pageId="34" pageNumber="35" type="female"></specimenCount>
, MNHN,
<figureCitation id="135899B6FFB2A40BC0A8FDA3178A108B" box="[726,897,669,694]" captionStart="FIGURES 227234" captionStartId="52.[151,264,1245,1268]" captionTargetBox="[153,1432,197,1212]" captionTargetId="figure@52.[151,1436,193,1225]" captionTargetPageId="52" captionText="FIGURES 227234. Type specimens and labels of Macronotops. 227229, M. fulvopilosus, male holotype (provided by Antoine Mantilleri); 230234, M. sexmaculatus, female holotype (provided by Antoine Mantilleri)." httpUri="https://zenodo.org/record/2585258/files/figure.png" pageId="34" pageNumber="35">Figs. 230234</figureCitation>
), Darjeeling (Watkins.) [pale yellow label] // unknown to me [pale yellow label] // 6-maculata Krtz. [pale yellow label] // Kraatz visit 1893. [pale yellow label] / /
<typeStatus id="54D83B91FFB2A40BC2D9FDD8153010C3" box="[167,315,742,766]" pageId="34" pageNumber="35" type="holotype">HOLOTYPE</typeStatus>
[red label] //
<materialsCitation id="3B0B8F6EFFB2A40BC3A9FDD81522111F" ID-GBIF-Occurrence="2004970377" collectionCode="MNHN" pageId="34" pageNumber="35" specimenCode="EC4377" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D83B91FFB2A40BC3A9FDD8166510C3" box="[471,622,742,766]" pageId="34" pageNumber="35" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C63FEB0FFB2A40BC007FDDB102F10C3" authority="Kraatz, 1894" authorityName="Kraatz" authorityYear="1894" box="[633,1060,741,766]" class="Insecta" family="Scarabaeidae" genus="Pleuronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis id="B9175921FFB2A40BC007FDDB178110C3" box="[633,906,741,766]" italics="true" pageId="34" pageNumber="35">Pleuronota sexmaculata</emphasis>
<bibRefCitation id="EFF2F8C2FFB2A40BC1ECFDD8102F10C3" author="Kraatz, G." box="[914,1060,742,766]" pageId="34" pageNumber="35" pagination="141" refId="ref34691" refString="Kraatz, G. (1894) Pleuronota 6 - maculata n. sp. Deutsche Entomologische Zeitschrift, 1894 (1), 141." type="journal article" year="1894">Kraatz, 1894</bibRefCitation>
</taxonomicName>
[white label] //
<collectionCode id="ED721DF6FFB2A40BC6A5FDD8113B10C3" box="[1243,1328,742,766]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="35">MNHN</collectionCode>
/
<specimenCode id="DBC52D48FFB2A40BC73EFDD8119710C3" box="[1344,1436,742,766]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="35">EC4377</specimenCode>
[white label]
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC8533FFB2A40BC2B9FC1317DF118F" blockId="34.[151,1437,669,2026]" pageId="34" pageNumber="35">
The female
<typeStatus id="54D83B91FFB2A40BC332FC1315A5117B" box="[332,430,813,838]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
<materialsCitation id="3B0B8F6EFFB2A40BC3CBFC1317DB118F" ID-GBIF-Occurrence="2004970388" collectionCode="MNHN, BMNH" country="India" county="He" location="Oberthur Collection" municipality="This" pageId="34" pageNumber="35" specimenCount="1" typeStatus="holotype">
originally belonged to
<location id="8EBCD3E8FFB2A40BC0CDFC13179D117B" LSID="urn:lsid:plazi:treatment:03CA3425FFB2A40AC2E9FEAA161D1571:8EBCD3E8FFB2A40BC0CDFC13179D117B" box="[691,918,813,838]" country="India" county="He" municipality="This" name="Oberthur Collection" pageId="34" pageNumber="35">Oberthür Collection</location>
, and now is deposited in
<collectionCode id="ED721DF6FFB2A40BC6D1FC1011031178" box="[1199,1288,814,837]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="35">MNHN</collectionCode>
.
<collectingMunicipality id="6BB81F49FFB2A40BC76DFC13114E117B" box="[1299,1349,813,838]" pageId="34" pageNumber="35">This</collectingMunicipality>
species was originally described based on a female, and
<bibRefCitation id="EFF2F8C2FFB2A40BC09DFC6C178F1157" author="Arrow, G. J." box="[739,900,850,874]" pageId="34" pageNumber="35" refId="ref34094" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma). Lamellicornia (Cetoniinae and Dynastinae). Taylor &amp; Francis, London, 322 pp." type="book" year="1910">Arrow (1910)</bibRefCitation>
given the first description of male based on specimens from
<collectingCountry id="F374C5A3FFB2A40BC32EFC4B158111B3" box="[336,394,885,910]" name="India" pageId="34" pageNumber="35">India</collectingCountry>
and
<collectingCountry id="F374C5A3FFB2A40BC3BEFC4B161E11B3" box="[448,533,885,910]" name="Bhutan" pageId="34" pageNumber="35">Bhutan</collectingCountry>
.
<collectingCounty id="62BDFDBFFFB2A40BC061FC48164B11B0" box="[543,576,886,909]" pageId="34" pageNumber="35">He</collectingCounty>
claimed that no female was examined, but the label date of a male specimen in
<collectionCode id="ED721DF6FFB2A40BC2CAFCA41503118C" box="[180,264,922,945]" pageId="34" pageNumber="35">BMNH</collectionCode>
indicated the fact that he compared it with the female
<typeStatus id="54D83B91FFB2A40BC114FCA717DB118F" box="[874,976,921,946]" pageId="34" pageNumber="35" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC8533FFB2A40BC2B9FC8315021797" blockId="34.[151,1437,669,2026]" pageId="34" pageNumber="35">
<emphasis id="B9175921FFB2A40BC2B9FC83162411EB" bold="true" box="[199,559,957,982]" pageId="34" pageNumber="35">Additional material examined</emphasis>
(
<specimenCount id="9D654EBAFFB2A40BC03EFC80168211EB" box="[576,649,958,982]" pageId="34" pageNumber="35" type="male">18♂♂</specimenCount>
,
<specimenCount id="9D654EBAFFB2A40BC0E8FC8316C411EB" box="[662,719,957,982]" pageId="34" pageNumber="35" type="female">6♀♀</specimenCount>
).
<emphasis id="B9175921FFB2A40BC098FC8317B811EB" bold="true" box="[742,947,957,982]" pageId="34" pageNumber="35">
<collectingCountry id="F374C5A3FFB2A40BC098FC83174C11EB" box="[742,839,957,982]" name="China" pageId="34" pageNumber="35">CHINA</collectingCountry>
:
<collectingRegion id="49A74BD1FFB2A40BC12BFC8317A611EB" box="[853,941,957,982]" country="China" name="Xizang" pageId="34" pageNumber="35">Xizang</collectingRegion>
:
</emphasis>
<specimenCount id="9D654EBAFFB2A40BC1C5FC8017D511EB" box="[955,990,958,982]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(QCCC),
<date id="FFDDA3F3FFB2A40BC62BFC8010EB11EB" box="[1109,1248,958,982]" pageId="34" pageNumber="35" value="2014-07-30">30.VII.2014</date>
, Mêdog County, Guang-Lin XIE;
<specimenCount id="9D654EBAFFB2A40BC329FCDC157111C7" box="[343,378,994,1018]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(QCCC),
<date id="FFDDA3F3FFB2A40BC38EFCDF167C11C7" box="[496,631,993,1018]" pageId="34" pageNumber="35" value="2016-08-07">7.VIII.2016</date>
, Yarlung Tsangpo Grand Canyon Nature Reserve, Renqinbeng Temple, Mêdog County, Zhi-Lin CHEN;
<specimenCount id="9D654EBAFFB2A40BC06CFB38163E1623" box="[530,565,1030,1054]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(CCCC),
<date id="FFDDA3F3FFB2A40BC0D3FB3B17481623" box="[685,835,1029,1054]" pageId="34" pageNumber="35" value="2016-08-20">20.VIII.2016</date>
, Mêdog County,
<quantity id="4C9B28D6FFB2A40BC672FB3B10661620" box="[1036,1133,1029,1054]" metricMagnitude="0" metricUnit="m" metricValue="1.526" pageId="34" pageNumber="35" unit="m" value="1.526">1,526 m</quantity>
, Xiao-Dong YANG;
<specimenCount id="9D654EBAFFB2A40BC71AFB3811971623" box="[1380,1436,1030,1054]" pageId="34" pageNumber="35" type="male">2♂♂</specimenCount>
(QCCC),
<date id="FFDDA3F3FFB2A40BC37AFB141591167F" box="[260,410,1066,1090]" pageId="34" pageNumber="35" value="2017-08-18">18.VIII.2017</date>
, Yarlung Tsangpo Grand Canyon Nature Reserve, Pêlung, Nyingchi County,
<quantity id="4C9B28D6FFB2A40BC77CFB17116A167C" box="[1282,1377,1065,1090]" metricMagnitude="0" metricUnit="m" metricValue="2.05" pageId="34" pageNumber="35" unit="m" value="2.05">2,050 m</quantity>
, Shi- Wei GUO;
<specimenCount id="9D654EBAFFB2A40BC364FB70155E165B" box="[282,341,1102,1126]" pageId="34" pageNumber="35" type="male">2♂♂</specimenCount>
,
<specimenCount id="9D654EBAFFB2A40BC31CFB70158E165B" box="[354,389,1102,1126]" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
(QCCC),
<date id="FFDDA3F3FFB2A40BC382FB70169A165B" box="[508,657,1102,1126]" pageId="34" pageNumber="35" value="2017-08-12">12.VIII.2017</date>
, Yarlung Tsangpo Grand Canyon Nature Reserve, Pêlung, Nyingchi County,
<quantity id="4C9B28D6FFB2A40BC289FB4F155D16B4" box="[247,342,1137,1162]" metricMagnitude="0" metricUnit="m" metricValue="2.05" pageId="34" pageNumber="35" unit="m" value="2.05">2,050 m</quantity>
, Jian-Yue QIU &amp; Hao XU.
<emphasis id="B9175921FFB2A40BC0EBFB4F16EC16B7" bold="true" box="[661,743,1137,1162]" pageId="34" pageNumber="35">
<collectingCountry id="F374C5A3FFB2A40BC0EBFB4F16EC16B7" box="[661,743,1137,1162]" name="India" pageId="34" pageNumber="35">INDIA</collectingCountry>
</emphasis>
:
<specimenCount id="9D654EBAFFB2A40BC086FB4C173916B7" box="[760,818,1138,1162]" pageId="34" pageNumber="35" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D654EBAFFB2A40BC141FB4C176916B7" box="[831,866,1138,1162]" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
(MNHN), British Bootan Padong, L. Durel 1913;
<specimenCount id="9D654EBAFFB2A40BC2E9FBA814B11693" box="[151,186,1174,1198]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(MNHN), British Bootang L. Durel 1899;
<specimenCount id="9D654EBAFFB2A40BC0DAFBA816CD1693" box="[676,710,1174,1198]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(MNHN), British India
<collectingRegion id="49A74BD1FFB2A40BC1A2FBAB10261693" box="[988,1069,1173,1198]" country="India" name="Sikkim" pageId="34" pageNumber="35">Sikkim</collectingRegion>
Lachen-Lachung
<date id="FFDDA3F3FFB2A40BC684FBA811661693" box="[1274,1389,1174,1198]" pageId="34" pageNumber="35" value="1933-08">VIII.1933</date>
;
<specimenCount id="9D654EBAFFB2A40BC707FBA811971693" box="[1401,1436,1174,1198]" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
(MNHN),
<collectingRegion id="49A74BD1FFB2A40BC371FB87156B16EF" box="[271,352,1209,1234]" country="India" name="Sikkim" pageId="34" pageNumber="35">Sikkim</collectingRegion>
Lachen 1932 Chasseurs indigenes;
<specimenCount id="9D654EBAFFB2A40BC080FB84172A16EF" box="[766,801,1210,1234]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(BMNH), British Bootang Maria Basti 1899, Oberthur Coll. 1909-159 //
<emphasis id="B9175921FFB2A40BC322FBE0165516CB" box="[348,606,1245,1270]" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="4C63FEB0FFB2A40BC322FBE015DC16C8" authorityName="Hoffmannsegg" authorityYear="1817" box="[348,471,1246,1269]" class="Insecta" family="Scarabaeidae" genus="Macronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Macronota</taxonomicName>
6-maculata
</emphasis>
Kr., Compared with
<typeStatus id="54D83B91FFB2A40BC137FBE0177616CB" box="[841,893,1246,1270]" pageId="34" pageNumber="35">type</typeStatus>
. G. J. A.;
<specimenCount id="9D654EBAFFB2A40BC199FBE0100116CB" box="[999,1034,1246,1270]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(BMNH), British Bootang L. Durel 1899 // Bourgoin Coll. B. M. 1938-252;
<specimenCount id="9D654EBAFFB2A40BC017FA3C16871727" box="[617,652,1282,1306]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(BMNH), British Bootang L. Durel 1899 //
<taxonomicName id="4C63FEB0FFB2A40BC6F4FA3F14B11700" authority="Kr." authorityName="Kraatz" authorityYear="1894" class="Insecta" family="Scarabaeidae" genus="Pleuronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis id="B9175921FFB2A40BC6F4FA3F11971727" box="[1162,1436,1281,1306]" italics="true" pageId="34" pageNumber="35">Pleuronota sexmaculata</emphasis>
Kr.
</taxonomicName>
// Bourgoin Coll. B. M. 1938-252;
<specimenCount id="9D654EBAFFB2A40BC036FA18166B1703" box="[584,608,1318,1342]" pageId="34" pageNumber="35" type="female">1♀</specimenCount>
(BMNH), Environs de Kurseong, R. P. Bretaudeau, Oberthur Coll. 1909- 159;
<emphasis id="B9175921FFB2A40BC2A6FA771545175F" bold="true" box="[216,334,1353,1378]" pageId="34" pageNumber="35">
<collectingCountry id="F374C5A3FFB2A40BC2A6FA771545175F" box="[216,334,1353,1378]" name="Bhutan" pageId="34" pageNumber="35">BHUTAN</collectingCountry>
</emphasis>
:
<specimenCount id="9D654EBAFFB2A40BC31BFA741583175F" box="[357,392,1354,1378]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(MNHN), Bhutan //
<taxonomicName id="4C63FEB0FFB2A40BC0E9FA771053175C" authority="Kr. G. Ruter" authorityName="Kr. G. Ruter" box="[663,1112,1353,1378]" class="Insecta" family="Scarabaeidae" genus="Pleuronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis id="B9175921FFB2A40BC0E9FA7717BB175F" box="[663,944,1353,1378]" italics="true" pageId="34" pageNumber="35">Pleuronota sexmaculata</emphasis>
Kr. G. Ruter
</taxonomicName>
det. 1977 //
<taxonomicName id="4C63FEB0FFB2A40BC776FA74158C17BB" authority="(Kraatz)" baseAuthorityName="Kraatz" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sexmaculata">
<emphasis id="B9175921FFB2A40BC776FA74152817BB" italics="true" pageId="34" pageNumber="35">Macronotops sexmaculata</emphasis>
(Kraatz)
</taxonomicName>
J. Ph. Legrand det. 2010;
<specimenCount id="9D654EBAFFB2A40BC0C9FA5016D017BB" box="[695,731,1390,1414]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
,
<specimenCount id="9D654EBAFFB2A40BC099FA50171417BB" box="[743,799,1390,1414]" pageId="34" pageNumber="35" type="female">2♀♀</specimenCount>
(MNHN), Bhutan;
<specimenCount id="9D654EBAFFB2A40BC184FA50101617BB" box="[1018,1053,1390,1414]" pageId="34" pageNumber="35" type="male">1♂</specimenCount>
(MNHN), Brit. Buthan Pankasary Hill 1932.
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB2A40AC2B9FA8B11121105" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="description">
<paragraph id="8BDC8533FFB2A40AC2B9FA8B150610B9" blockId="34.[151,1437,669,2026]" lastBlockId="35.[151,1437,151,1869]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<emphasis id="B9175921FFB2A40BC2B9FA8B163117F3" bold="true" box="[199,570,1461,1486]" pageId="34" pageNumber="35">Redescription (male). General:</emphasis>
Body length 17.018.5 mm; width 7.08.0 mm, widest at humeral umbone, gradually narrowed backward. Body dark brown to black. Surface with fulvous, reddish-brown, brown and light yellow setae, and yellow tomentous maculae.
<emphasis id="B9175921FFB2A40BC0E4FAC316E9142B" bold="true" box="[666,738,1533,1558]" pageId="34" pageNumber="35">Head:</emphasis>
Brown. Dorsal surface with dense, setiferous punctures; setae long, fulvous, and longer on frons. Anterior margin of clypeus nearly straight; anterior margin and clypeolateral ridge slightly raised. Frons with a slightly raised longitudinal ridge (
<figureCitation id="135899B6FFB2A40BC1DBF97B17F31463" box="[933,1016,1605,1630]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="34" pageNumber="35">Fig. 14</figureCitation>
). Maxillary palpus and labial palpus brown. Antenna yellowish-brown; antennal club very long, about twice length of antennomeres 27 combined; inner side of antennomere 8 with short, dense, fulvous setae (
<figureCitation id="135899B6FFB2A40BC13AF9B3179F149B" box="[836,916,1677,1702]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="34" pageNumber="35">Fig. 14</figureCitation>
). Ventral surface clad with long, fulvous setae (
<figureCitation id="135899B6FFB2A40BC2E1F98F14F514F7" box="[159,254,1713,1738]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="34" pageNumber="35">Fig. 166</figureCitation>
).
<emphasis id="B9175921FFB2A40BC369F98F159214F7" bold="true" box="[279,409,1713,1738]" pageId="34" pageNumber="35">Pronotum:</emphasis>
Dark brown. Widest at base, basomedian area depressed. Lateral margin curved; posterior margin distinctly protruded. Surface densely clad with long, fulvous and dark brown setae (
<figureCitation id="135899B6FFB2A40BC6DDF9EB110914D3" box="[1187,1282,1749,1774]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="34" pageNumber="35">Fig. 165</figureCitation>
).
<emphasis id="B9175921FFB2A40BC765F9EB119714D3" bold="true" box="[1307,1436,1749,1774]" pageId="34" pageNumber="35">Scutellum:</emphasis>
Dark brown to black, with sparse, light yellow setae.
<emphasis id="B9175921FFB2A40BC099F9C71740152F" bold="true" box="[743,843,1785,1810]" pageId="34" pageNumber="35">Elytron:</emphasis>
Black; with posthumeral macula, lateral macula, and median macula, but without distal macula. Surface densely clad with long setae; setae longer between sutural and discolateral costae; setae light yellow on maculae; setae brown on lateral declivity; setae fulvous on the rest of elytron; humeral umbone glabrous (
<figureCitation id="135899B6FFB2A40BC055F85B16811543" box="[555,650,1893,1918]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="34" pageNumber="35">Fig. 165</figureCitation>
).
<emphasis id="B9175921FFB2A40BC0DCF85B17451543" bold="true" box="[674,846,1893,1918]" pageId="34" pageNumber="35">Mesepimeron:</emphasis>
black, with light yellow setae.
<emphasis id="B9175921FFB2A40BC6CEF85B1535159F" bold="true" pageId="34" pageNumber="35">Metepisternum and metepimeron:</emphasis>
dark brown, with brown setae.
<emphasis id="B9175921FFB2A40BC0AFF8B7174A159F" bold="true" box="[721,833,1929,1954]" pageId="34" pageNumber="35">Sternum:</emphasis>
Black, preprosternum with sparse, setae on side. Mesosternum with sparse, light yellow setae. Mesometasternal process short, glabrous, black, apex rounded; mesometasternal suture depressed, with long, dense, fulvous setae. Metasternum with sparse, fulvous and brown setae; middle portion glabrous (
<figureCitation id="135899B6FFB3A40AC07DFFA91668128D" box="[515,611,151,176]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Fig. 166</figureCitation>
).
<emphasis id="B9175921FFB3A40AC002FFA916F2128D" bold="true" box="[636,761,151,176]" pageId="35" pageNumber="36">Pygidium:</emphasis>
Dark brown. Surface with dense, fulvous and light yellow setae; a large, round, yellow macula in middle (
<figureCitation id="135899B6FFB3A40AC0A6FF82173B12E9" box="[728,816,188,213]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="35" pageNumber="36">Fig. 92</figureCitation>
).
<emphasis id="B9175921FFB3A40AC132FF8217C112E8" bold="true" box="[844,970,188,213]" pageId="35" pageNumber="36">Abdomen:</emphasis>
Reddish-brown to dark brown. With a longitudinal groove in median. Six abdominal sternites visible; ventral surface of sternites sparsely clad with yellow setae; setae on sternites IIV short, yellow, but longer on sides; setae near posterior margin of sternite VI long, dense, fulvous; sternite VII with sparse short setae on sides. Sternites IIV with a small yellow macula close to posterior margin on each side; surface densely clad with yellow and black setae in dorsal and lateral portion (
<figureCitation id="135899B6FFB3A40AC710FE7214CA13B5" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Fig. 166</figureCitation>
).
<emphasis id="B9175921FFB3A40AC2A5FE51151713B5" bold="true" box="[219,284,367,392]" pageId="35" pageNumber="36">Legs:</emphasis>
Slender, simple. Metacoxa black, with long, brown setae. Femora and tibia dark brown, with long, fulvous setae. Protibia with 3 sharp, large teeth, without tiny tooth (
<figureCitation id="135899B6FFB3A40AC1EAFEAA17EE1391" box="[916,997,404,429]" captionStart="FIGURES 2447" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 2447. Protibia of Macronotops (male and female). 24, M. biserratus new species; 2526, M. dianensis new species; 2728, M. fulvoguttatus; 2930, M. fulvopilosus; 3132, M. miksici new species; 3334, M. nigropubescens; 3536, M. olivaceofuscus; 3738, M. sexmaculatus; 3940, M. vuilleti; 41, Macronotops sp.; 4243, M. curvimarginatus; 4445, M. medogensis new species; 4647, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="35" pageNumber="36">Fig. 37</figureCitation>
); mesotibia and metatibia with a spine near the middle of outer margin; metatibia with a row of brush-like, fulvous setae along inner margin (
<figureCitation id="135899B6FFB3A40AC750FE8914C913C8" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Fig. 165 166</figureCitation>
); inner margin of metatibia convex in distal portion (
<figureCitation id="135899B6FFB3A40AC169FEE2177213C8" box="[791,889,476,501]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Fig. 166</figureCitation>
); dorsal tooth of metatibia longer than basitarsus of metatarsus, outer tooth absent (
<figureCitation id="135899B6FFB3A40AC069FEC116711025" box="[535,634,511,536]" captionStart="FIGURES 159164" captionStartId="33.[151,264,1632,1655]" captionTargetBox="[159,1408,206,1578]" captionTargetId="figure@33.[151,1436,193,1611]" captionTargetPageId="33" captionText="FIGURES 159164. Habitus of Macronotops olivaceofuscus (dorsal and ventral view). 159160, male (Mount Simianshan, W Chongqing, China); 161, male (Wudaoxia, W Hubei, China); 162163, female (Mount Simianshan, W Chongqing, China); 164, female (Mount Daweishan, S Yunnan, China)." httpUri="https://zenodo.org/record/2585236/files/figure.png" pageId="35" pageNumber="36">Fig. 162</figureCitation>
). Tarsi reddish-brown; basitarsus of metatarsus with a cluster of dense, long setae (
<figureCitation id="135899B6FFB3A40AC360FD1A15881000" box="[286,387,548,573]" captionStart="FIGURES 159164" captionStartId="33.[151,264,1632,1655]" captionTargetBox="[159,1408,206,1578]" captionTargetId="figure@33.[151,1436,193,1611]" captionTargetPageId="33" captionText="FIGURES 159164. Habitus of Macronotops olivaceofuscus (dorsal and ventral view). 159160, male (Mount Simianshan, W Chongqing, China); 161, male (Wudaoxia, W Hubei, China); 162163, female (Mount Simianshan, W Chongqing, China); 164, female (Mount Daweishan, S Yunnan, China)." httpUri="https://zenodo.org/record/2585236/files/figure.png" pageId="35" pageNumber="36">Fig. 162</figureCitation>
).
<emphasis id="B9175921FFB3A40AC3E4FD1A161A1000" bold="true" box="[410,529,548,573]" pageId="35" pageNumber="36">Genitalia:</emphasis>
Parameres broad, short; apex rounded, slightly expanded. Interparameral split wide; membrane almost absent. Median lobe wide at base, filiform apically; nearly equal to the length of paramere (
<figureCitation id="135899B6FFB3A40AC2E1FD5214F710B9" box="[159,252,620,645]" captionStart="FIGURES 111122" captionStartId="16.[151,264,1744,1767]" captionTargetBox="[156,1403,544,1706]" captionTargetId="figure@16.[151,1436,525,1723]" captionTargetPageId="16" captionText="FIGURES 111122. Male genitalia of Macronotops (apical view). 111, M. biserratus new species; 112, M. dianensis new species; 113, M. fulvoguttatus; 114, M. fulvopilosus; 115, M. miksici new species; 116, M. nigropubescens; 117, M. olivaceofuscus; 118, M. sexmaculatus; 119, M. vuilleti. 120, M. curvimarginatus; 121, M. medogensis new species; 122, M. ovaliceps. Arrows indicate the membrane between the parameres." httpUri="https://zenodo.org/record/2585220/files/figure.png" pageId="35" pageNumber="36">Fig. 118</figureCitation>
).
</paragraph>
<paragraph id="8BDC8533FFB3A40AC2B0FDB116E110D2" blockId="35.[151,1437,151,1869]" pageId="35" pageNumber="36">
<emphasis id="B9175921FFB3A40AC2B0FDB115261095" bold="true" box="[206,301,655,680]" pageId="35" pageNumber="36">Female.</emphasis>
Body length 17.520.0 mm; width 7.59.0 mm. Except for sexual dimorphism characteristics of the genus, dorsal teeth of metatibia slightly larger (
<figureCitation id="135899B6FFB3A40AC0D2FD8A16F610F0" box="[684,765,692,717]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="35" pageNumber="36">Fig. 62</figureCitation>
), setae on mesotibia and metatibia shorter (
<figureCitation id="135899B6FFB3A40AC694FD8A118610F0" box="[1258,1421,692,717]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Figs. 167168</figureCitation>
), and basitarsus of metatarsus with less setae (
<figureCitation id="135899B6FFB3A40AC0F4FDE916D110CD" box="[650,730,727,752]" captionStart="FIGURES 4865" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[187,1411,183,1839]" captionTargetId="figure@11.[187,1411,183,1839]" captionTargetPageId="11" captionText="FIGURES 4865. Metatibia and basitarsus of metatarsus of the Macronotops sexmaculatus species group (male in left, female in right). 48, M. biserratus new species; 4950, M. dianensis new species; 5152, M. fulvoguttatus; 5354, M. fulvopilosus; 5556, M. miksici new species; 5758, M. nigropubescens;5960, M. olivaceofuscus; 6162, M. sexmaculatus; 6364, M. vuilleti; 65, Macronotops sp. Red arrows indicate the difference in the dorsal and outer tooth of the metatibia in females; yellow arrows indicate the difference in the dorsal and outer tooth of the metatibia in males; blue arrows indicate the density of setae on the basitarsus of the metatarsus." httpUri="https://zenodo.org/record/2585210/files/figure.png" pageId="35" pageNumber="36">Fig. 62</figureCitation>
).
</paragraph>
<paragraph id="8BDC8533FFB3A40AC2B9FDC211121105" blockId="35.[151,1437,151,1869]" pageId="35" pageNumber="36">
<emphasis id="B9175921FFB3A40AC2B9FDC215461128" bold="true" box="[199,333,764,789]" pageId="35" pageNumber="36">Variability.</emphasis>
The yellow macula in the middle of elytra and on sternite III to VI are absent in some individuals (
<figureCitation id="135899B6FFB3A40AC2E1FC2114F51105" box="[159,254,799,824]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Fig. 166</figureCitation>
). Specimens from Brahmaputra Valley usually have more fulvous setae on the dorsal surface.
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB3A40AC2B9FC7A15A81641" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="8BDC8533FFB3A40AC2B9FC7A15341609" blockId="35.[151,1437,151,1869]" pageId="35" pageNumber="36">
<emphasis id="B9175921FFB3A40AC2B9FC7A15DA1160" bold="true" box="[199,465,836,861]" pageId="35" pageNumber="36">Differential diagnosis.</emphasis>
Length of clypeus almost equal to width. Antenna yellowish-brown (
<figureCitation id="135899B6FFB3A40AC77CFC7A11871160" box="[1282,1420,836,861]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="35" pageNumber="36">Figs. 1415</figureCitation>
); male antennal club very long (
<figureCitation id="135899B6FFB3A40AC395FC59163011BD" box="[491,571,871,896]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="35" pageNumber="36">Fig. 14</figureCitation>
). Elytra dark brown, with posthumeral maculae, median maculae, and lateral maculae (
<figureCitation id="135899B6FFB3A40AC37BFCB215A11198" box="[261,426,908,933]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Figs. 165, 167</figureCitation>
). Abdominal sternites IIV usually with a small, yellow macula on each side (
<figureCitation id="135899B6FFB3A40AC755FCB211801198" box="[1323,1419,908,933]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Fig. 168</figureCitation>
); male abdomen with a longitudinal groove. Pygidium with a large, yellow macula (
<figureCitation id="135899B6FFB3A40AC63BFC9110C611F5" box="[1093,1229,943,968]" captionStart="FIGURES 7298" captionStartId="14.[151,264,1921,1944]" captionTargetBox="[151,1436,195,1900]" captionTargetId="figure@14.[151,1436,195,1900]" captionTargetPageId="14" captionText="FIGURES 7298. Pygidium of the Macronotops sexmaculatus species group (size of macula and color of setae). 7273, M. biserratus new species; 7475, M. dianensis new species; 7679, M. fulvoguttatus; 8083, M. fulvopilosus; 8485, M. miksici new species; 8689, M. nigropubescens; 9091, M. olivaceofuscus; 9293, M. sexmaculatus; 9497, M. vuilleti; 98, Macronotops sp." httpUri="https://zenodo.org/record/2585216/files/figure.png" pageId="35" pageNumber="36">Figs. 9293</figureCitation>
). Protibia without tiny tooth (
<figureCitation id="135899B6FFB3A40AC36AFCEA159611D1" box="[276,413,980,1005]" captionStart="FIGURES 2447" captionStartId="10.[151,264,1767,1790]" captionTargetBox="[151,1436,193,1746]" captionTargetId="figure@10.[151,1436,193,1746]" captionTargetPageId="10" captionText="FIGURES 2447. Protibia of Macronotops (male and female). 24, M. biserratus new species; 2526, M. dianensis new species; 2728, M. fulvoguttatus; 2930, M. fulvopilosus; 3132, M. miksici new species; 3334, M. nigropubescens; 3536, M. olivaceofuscus; 3738, M. sexmaculatus; 3940, M. vuilleti; 41, Macronotops sp.; 4243, M. curvimarginatus; 4445, M. medogensis new species; 4647, M. ovaliceps. Arrows indicate the angle between large teeth and the outer margin of protibia." httpUri="https://zenodo.org/record/2585208/files/figure.png" pageId="35" pageNumber="36">Figs. 3738</figureCitation>
); inner margin of metatibia convex in distal portion (
<figureCitation id="135899B6FFB3A40AC185FCEA109411D0" box="[1019,1183,980,1005]" captionStart="FIGURES 165168" captionStartId="36.[151,264,1978,2001]" captionTargetBox="[167,1403,193,1938]" captionTargetId="figure@36.[167,1403,193,1938]" captionTargetPageId="36" captionText="FIGURES 165168. Habitus of Macronotops sexmaculatus (dorsal and ventral view). 165166, male (Mêdog, SE Xizang, China); 167168, female (Mêdog, SE Xizang, China). Arrow indicates inner margin of metatibia in distal portion." httpUri="https://zenodo.org/record/2585238/files/figure.png" pageId="35" pageNumber="36">Figs. 166, 168</figureCitation>
); dorsal tooth of male metatibia longer than basitarsus of metatarsus (
<figureCitation id="135899B6FFB3A40AC0D9FCC916FC162D" box="[679,759,1015,1040]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="35" pageNumber="36">Fig. 6 1</figureCitation>
); basitarsus of male metatarsus with a cluster of dense, long setae (
<figureCitation id="135899B6FFB3A40AC2A0FB2215251609" box="[222,302,1052,1077]" captionStart="FIGURES 123" captionStartId="9.[151,264,1838,1861]" captionTargetBox="[151,1436,195,1817]" captionTargetId="figure@9.[151,1436,195,1817]" captionTargetPageId="9" captionText="FIGURES 123. Head of Macronotops (male in left, female in right). 1, M. biserratus new species; 23, M. dianensis new species; 45, M. fulvoguttatus; 67, M. fulvopilosus; 89, M. miksici new species; 1011, M. nigropubescens; 1213, M. olivaceofuscus; 1415, M. sexmaculatus; 1617, M. vuilleti; 1819, M. curvimarginatus; 2021, M. medogensis new species; 2223, M. ovaliceps. Arrows emphasize the length of antenna club." httpUri="https://zenodo.org/record/2585206/files/figure.png" pageId="35" pageNumber="36">Fig. 6 1</figureCitation>
).
</paragraph>
<paragraph id="8BDC8533FFB3A40AC2B9FB0115A81641" blockId="35.[151,1437,151,1869]" pageId="35" pageNumber="36">
This species highly resembles
<taxonomicName id="4C63FEB0FFB3A40AC060FB7F169F1665" authority="Qiu &amp; Xu &amp; Chen, 2019" authorityName="Qiu &amp; Xu &amp; Chen" authorityYear="2019" box="[542,660,1087,1112]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="miksici" status="sp. nov.">
<emphasis id="B9175921FFB3A40AC060FB7F169F1665" box="[542,660,1087,1112]" italics="true" pageId="35" pageNumber="36">M. miksici</emphasis>
</taxonomicName>
<emphasis id="B9175921FFB3A40AC0E4FB01172E1665" bold="true" box="[666,805,1087,1112]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A224E45AFFB3A40AC0E4FB01172E1665" box="[666,805,1087,1112]" pageId="35" pageNumber="36" rank="species">new species</taxonomicNameLabel>
</emphasis>
, the differences are provided in differential diagnosis of
<taxonomicName id="4C63FEB0FFB3A40AC2E9FB5B15001640" authority="Qiu &amp; Xu &amp; Chen, 2019" authorityName="Qiu &amp; Xu &amp; Chen" authorityYear="2019" box="[151,267,1124,1149]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="miksici" status="sp. nov.">
<emphasis id="B9175921FFB3A40AC2E9FB5B15001640" box="[151,267,1124,1149]" italics="true" pageId="35" pageNumber="36">M. miksici</emphasis>
</taxonomicName>
<emphasis id="B9175921FFB3A40AC36DFB5A15971640" bold="true" box="[275,412,1124,1149]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A224E45AFFB3A40AC36DFB5A15971640" box="[275,412,1124,1149]" pageId="35" pageNumber="36" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB3A40AC2B9FBB910861731" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BDC8533FFB3A40AC2B9FBB917B6169D" blockId="35.[151,1437,151,1869]" box="[199,957,1159,1184]" pageId="35" pageNumber="36">
<emphasis id="B9175921FFB3A40AC2B9FBB9156B169D" bold="true" box="[199,352,1159,1184]" pageId="35" pageNumber="36">Distribution.</emphasis>
<collectingCountry id="F374C5A3FFB3A40AC319FBB915A1169D" box="[359,426,1159,1184]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
(
<emphasis id="B9175921FFB3A40AC3C4FBB916AF169D" bold="true" box="[442,676,1159,1184]" pageId="35" pageNumber="36">new country record</emphasis>
):
<collectingRegion id="49A74BD1FFB3A40AC0C2FBB9171B169D" box="[700,784,1159,1184]" country="China" name="Xizang" pageId="35" pageNumber="36">Xizang</collectingRegion>
;
<collectingCountry id="F374C5A3FFB3A40AC162FBB91752169D" box="[796,857,1159,1184]" name="India" pageId="35" pageNumber="36">India</collectingCountry>
;
<collectingCountry id="F374C5A3FFB3A40AC11BFBB917B1169D" box="[869,954,1159,1184]" name="Bhutan" pageId="35" pageNumber="36">Bhutan</collectingCountry>
.
</paragraph>
<paragraph id="8BDC8533FFB3A40AC2B9FB9210861731" blockId="35.[151,1437,151,1869]" pageId="35" pageNumber="36">
<emphasis id="B9175921FFB3A40AC2B9FB92158D16F8" bold="true" box="[199,390,1196,1221]" pageId="35" pageNumber="36">Natural history.</emphasis>
In the Yarlung Tsangpo Grand Canyon, adults of
<taxonomicName id="4C63FEB0FFB3A40AC1C5FB93107316F8" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[955,1144,1196,1221]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB3A40AC1C5FB93107316F8" box="[955,1144,1196,1221]" italics="true" pageId="35" pageNumber="36">M. sexmaculatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63FEB0FFB3A40AC6F8FB93116F16F9" baseAuthorityName="Ma" baseAuthorityYear="1993" box="[1158,1380,1197,1220]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="curvimarginatus">
<emphasis id="B9175921FFB3A40AC6F8FB93116F16F9" box="[1158,1380,1197,1220]" italics="true" pageId="35" pageNumber="36">M. curvimarginatus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C63FEB0FFB3A40AC2E9FBEF153416D5" authority="Qiu &amp; Xu &amp; Chen, 2019" authorityName="Qiu &amp; Xu &amp; Chen" authorityYear="2019" box="[151,319,1231,1256]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="medogensis" status="sp. nov.">
<emphasis id="B9175921FFB3A40AC2E9FBEF153416D5" box="[151,319,1231,1256]" italics="true" pageId="35" pageNumber="36">M. medogensis</emphasis>
</taxonomicName>
<emphasis id="B9175921FFB3A40AC33BFBF115C416D5" bold="true" box="[325,463,1231,1256]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A224E45AFFB3A40AC33BFBF115C416D5" box="[325,463,1231,1256]" pageId="35" pageNumber="36" rank="species">new species</taxonomicNameLabel>
</emphasis>
were observed flying around bushes in the sunshine after the rain and looked like large bees, and an adult of
<taxonomicName id="4C63FEB0FFB3A40AC3FBFBCB164A1730" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[389,577,1268,1293]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB3A40AC3FBFBCB164A1730" box="[389,577,1268,1293]" italics="true" pageId="35" pageNumber="36">M. sexmaculatus</emphasis>
</taxonomicName>
was found feeding on the sap of bamboo (
<figureCitation id="135899B6FFB3A40AC660FBCA108B1731" box="[1054,1152,1268,1293]" captionStart="FIGURES 276281" captionStartId="58.[151,264,1812,1835]" captionTargetBox="[151,1436,193,1792]" captionTargetId="figure@58.[151,1436,193,1792]" captionTargetPageId="58" captionText="FIGURES 276281. Habitat of Macronotops. 276277, habitat of M. sexmaculatus and M. medogensis new species in Valley near Bononggong of Mêdog (Xizang, China); 278, an adult of M. sexmaculatus feeding on sap of bamboo (photograph by Yang He); 279281, habitat of M. vuilleti in Mount Huanglianshan (Yunnan, China), and larva in fallen rotten wood." httpUri="https://zenodo.org/record/2585270/files/figure.png" pageId="35" pageNumber="36">Fig. 278</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C379D6B8FFB3A40AC2B9FA29161D1571" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BDC8533FFB3A40AC2B9FA29161D1571" blockId="35.[151,1437,151,1869]" pageId="35" pageNumber="36">
<emphasis id="B9175921FFB3A40AC2B9FA291530170D" bold="true" box="[199,315,1303,1328]" pageId="35" pageNumber="36">Remarks.</emphasis>
<taxonomicName id="4C63FEB0FFB3A40AC33DFA27167C170D" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[323,631,1303,1328]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB3A40AC33DFA27167C170D" box="[323,631,1303,1328]" italics="true" pageId="35" pageNumber="36">Macronotops sexmaculatus</emphasis>
</taxonomicName>
was originally described based on a female specimen from Darjeeling (
<bibRefCitation id="EFF2F8C2FFB3A40AC2E0FA0215391769" author="Kraatz, G." box="[158,306,1340,1364]" pageId="35" pageNumber="36" pagination="141" refId="ref34691" refString="Kraatz, G. (1894) Pleuronota 6 - maculata n. sp. Deutsche Entomologische Zeitschrift, 1894 (1), 141." type="journal article" year="1894">Kraatz 1894</bibRefCitation>
), where located in the south side of the Himalayas. After checking the
<typeStatus id="54D83B91FFB3A40AC6F6FA0210E61768" box="[1160,1261,1340,1365]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
,
<bibRefCitation id="EFF2F8C2FFB3A40AC682FA0211971769" author="Arrow, G. J." box="[1276,1436,1340,1364]" pageId="35" pageNumber="36" refId="ref34094" refString="Arrow, G. J. (1910) The Fauna of British India (Including Ceylon and Burma). Lamellicornia (Cetoniinae and Dynastinae). Taylor &amp; Francis, London, 322 pp." type="book" year="1910">Arrow (1910)</bibRefCitation>
provided a short description of male, and listed this species from
<collectingCountry id="F374C5A3FFB3A40AC101FA6117D81745" box="[895,979,1375,1400]" name="Bhutan" pageId="35" pageNumber="36">Bhutan</collectingCountry>
.
<bibRefCitation id="EFF2F8C2FFB3A40AC1A1FA6110741745" author="Miksic, R." box="[991,1151,1375,1400]" pageId="35" pageNumber="36" pagination="208 - 210" refId="ref35268" refString="Miksic, R. (1971) Notizien uber ostasiatische Cetoniiden. (72. Beitrag zur Kenntnis der Scarabaeiden). Bulletin Scientifique, Conseil des Academies de la RPF Yougoslavie. Section A. Sciences Naturelles, Techniques et Medicales, 16 (7 - 8), 208 - 210." type="journal article" year="1971">Mikšić (1971)</bibRefCitation>
briefly commented most members of the genus
<taxonomicName id="4C63FEB0FFB3A40AC3EAFABA16A017A0" authority="sensu Miksic" authorityName="Miksic" box="[404,683,1412,1437]" class="Insecta" family="Scarabaeidae" genus="Pleuronota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B9175921FFB3A40AC3EAFABA161A17A0" box="[404,529,1412,1437]" italics="true" pageId="35" pageNumber="36">Pleuronota</emphasis>
sensu Mikšić
</taxonomicName>
(
<emphasis id="B9175921FFB3A40AC0C2FABB177217A1" box="[700,889,1413,1436]" italics="true" pageId="35" pageNumber="36">
i.e.
<taxonomicName id="4C63FEB0FFB3A40AC09BFABB177217A1" box="[741,889,1413,1436]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Macronotops</taxonomicName>
</emphasis>
), and figured parameres of
<taxonomicName id="4C63FEB0FFB3A40AC6CEFABB116717A0" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[1200,1388,1412,1437]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB3A40AC6CEFABB116717A0" box="[1200,1388,1412,1437]" italics="true" pageId="35" pageNumber="36">M. sexmaculatus</emphasis>
</taxonomicName>
and his new species for comparison. This is the first time structures of the male genitalia of this genus to be figured (
<figureCitation id="135899B6FFB3A40AC2E1FAF2150917D8" box="[159,258,1484,1509]" captionStart="FIGURES 243257" captionStartId="54.[151,264,1899,1922]" captionTargetBox="[161,1405,197,1872]" captionTargetId="figure@54.[161,1405,193,1876]" captionTargetPageId="54" captionText="FIGURES 243257. Type specimens and labels of Macronotops. 243246, M. ovaliceps, male holotype; 247, M. curvimarginatus (modified from Ma 1993); 248, Pleuronota subsexmaculata (modified from Ma 1992a); 249, P. hefengensis (modified from Ma 1992a); 250, M. miksici (modified from Mikšić 1971); 251257, M. miksici, male paratype (Assam, NE India)." httpUri="https://zenodo.org/record/2585262/files/figure.png" pageId="35" pageNumber="36">Fig. 250</figureCitation>
), and the figure of
<taxonomicName id="4C63FEB0FFB3A40AC3A8FAF3169917D8" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[470,658,1484,1509]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB3A40AC3A8FAF3169917D8" box="[470,658,1484,1509]" italics="true" pageId="35" pageNumber="36">M. sexmaculatus</emphasis>
</taxonomicName>
was cited by
<bibRefCitation id="EFF2F8C2FFB3A40AC152FAF3179717D9" author="Ma, W. Z." box="[812,924,1484,1508]" pageId="35" pageNumber="36" pagination="179 - 183" refId="ref35037" refString="Ma, W. Z. (1993) Three new species of Pleuronota from China (Coleoptera: Cetoniidae). Sinozoologia, 10, 179 - 183. [in Chinese]" type="journal article" year="1993">Ma (1993</bibRefCitation>
,
<bibRefCitation id="EFF2F8C2FFB3A40AC1D6FAF217EA17D8" author="Ma, W. Z." box="[936,993,1484,1509]" pageId="35" pageNumber="36" refId="ref35071" refString="Ma, W. Z. (1995) Economic Insect Fauna of China, Folume 46 Coleoptera: Cetoniidae, Trichiidae, and Falgidae. Science and Technology Press, Beijing, 210 pp. [in Chinese]" type="book" year="1995">1995</bibRefCitation>
) who erroneously recorded this species widely distributed in
<collectingCountry id="F374C5A3FFB3A40AC3E6FAD115D01435" box="[408,475,1519,1544]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
(see remarks section of
<taxonomicName id="4C63FEB0FFB3A40AC175FACF17D01435" authorityName=", Miksic" authorityYear="1976" box="[779,987,1519,1544]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="olivaceofuscus">
<emphasis id="B9175921FFB3A40AC175FACF17D01435" box="[779,987,1519,1544]" italics="true" pageId="35" pageNumber="36">M. olivaceofuscus</emphasis>
</taxonomicName>
). After examining the parameres and external features, some individuals collected from southeastern
<collectingRegion id="49A74BD1FFB3A40AC1E6F92A17E21410" box="[920,1001,1556,1581]" country="China" name="Xizang" pageId="35" pageNumber="36">Xizang</collectingRegion>
are confirmed conspecific with the specimens of
<taxonomicName id="4C63FEB0FFB3A40AC344F90715F0146D" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[314,507,1591,1616]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB3A40AC344F90715F0146D" box="[314,507,1591,1616]" italics="true" pageId="35" pageNumber="36">M. sexmaculatus</emphasis>
</taxonomicName>
from Sikkim and
<collectingCountry id="F374C5A3FFB3A40AC0A7F9091726146D" box="[729,813,1591,1616]" name="Bhutan" pageId="35" pageNumber="36">Bhutan</collectingCountry>
. These specimens represented the first record of this species in
<collectingCountry id="F374C5A3FFB3A40AC36DF96215521448" box="[275,345,1628,1653]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
; the parameres, however, without membrane, are not identical to the figure in
<bibRefCitation id="EFF2F8C2FFB3A40AC68CF96211921449" author="Miksic, R." box="[1266,1433,1628,1653]" pageId="35" pageNumber="36" pagination="208 - 210" refId="ref35268" refString="Miksic, R. (1971) Notizien uber ostasiatische Cetoniiden. (72. Beitrag zur Kenntnis der Scarabaeiden). Bulletin Scientifique, Conseil des Academies de la RPF Yougoslavie. Section A. Sciences Naturelles, Techniques et Medicales, 16 (7 - 8), 208 - 210." type="journal article" year="1971">Mikšić (1971)</bibRefCitation>
. Mikšićs figure was actually drawn in accordance with a single male from
<collectingRegion id="49A74BD1FFB3A40AC670F9BE105614A5" box="[1038,1117,1664,1688]" country="India" name="Assam" pageId="35" pageNumber="36">Assam</collectingRegion>
,
<collectingCountry id="F374C5A3FFB3A40AC612F94110A314A5" box="[1132,1192,1663,1688]" name="India" pageId="35" pageNumber="36">India</collectingCountry>
, and it unexpectedly matchs the parameres of several specimens we collected from Xima (Sima) in western Yunnan (
<figureCitation id="135899B6FFB3A40AC756F99A119B1480" box="[1320,1424,1700,1725]" captionStart="FIGURES 111122" captionStartId="16.[151,264,1744,1767]" captionTargetBox="[156,1403,544,1706]" captionTargetId="figure@16.[151,1436,525,1723]" captionTargetPageId="16" captionText="FIGURES 111122. Male genitalia of Macronotops (apical view). 111, M. biserratus new species; 112, M. dianensis new species; 113, M. fulvoguttatus; 114, M. fulvopilosus; 115, M. miksici new species; 116, M. nigropubescens; 117, M. olivaceofuscus; 118, M. sexmaculatus; 119, M. vuilleti. 120, M. curvimarginatus; 121, M. medogensis new species; 122, M. ovaliceps. Arrows indicate the membrane between the parameres." httpUri="https://zenodo.org/record/2585220/files/figure.png" pageId="35" pageNumber="36">Fig. 115</figureCitation>
). Moreover, an undissected male from
<collectingRegion id="49A74BD1FFB3A40AC02CF9F6169514DD" box="[594,670,1736,1760]" country="India" name="Assam" pageId="35" pageNumber="36">Assam</collectingRegion>
was examined in MFNB (
<figureCitation id="135899B6FFB3A40AC19EF9F9108C14DD" box="[992,1159,1735,1760]" captionStart="FIGURES 243257" captionStartId="54.[151,264,1899,1922]" captionTargetBox="[161,1405,197,1872]" captionTargetId="figure@54.[161,1405,193,1876]" captionTargetPageId="54" captionText="FIGURES 243257. Type specimens and labels of Macronotops. 243246, M. ovaliceps, male holotype; 247, M. curvimarginatus (modified from Ma 1993); 248, Pleuronota subsexmaculata (modified from Ma 1992a); 249, P. hefengensis (modified from Ma 1992a); 250, M. miksici (modified from Mikšić 1971); 251257, M. miksici, male paratype (Assam, NE India)." httpUri="https://zenodo.org/record/2585262/files/figure.png" pageId="35" pageNumber="36">Figs. 251257</figureCitation>
) which bears the same external morphological characters. This fact revealed that a similar species is distributed in northeastern
<collectingCountry id="F374C5A3FFB3A40AC74CF9D211671538" box="[1330,1388,1772,1797]" name="India" pageId="35" pageNumber="36">India</collectingCountry>
and southwestern
<collectingCountry id="F374C5A3FFB3A40AC34DF83115721515" box="[307,377,1807,1832]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
, and here we named it as
<taxonomicName id="4C63FEB0FFB3A40AC0DBF82F17101515" authority="Qiu &amp; Xu &amp; Chen, 2019" authorityName="Qiu &amp; Xu &amp; Chen" authorityYear="2019" box="[677,795,1807,1832]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="miksici" status="sp. nov.">
<emphasis id="B9175921FFB3A40AC0DBF82F17101515" box="[677,795,1807,1832]" italics="true" pageId="35" pageNumber="36">M. miksici</emphasis>
</taxonomicName>
<emphasis id="B9175921FFB3A40AC15AF83117BB1515" bold="true" box="[804,944,1807,1832]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A224E45AFFB3A40AC15AF83117BB1515" box="[804,944,1807,1832]" pageId="35" pageNumber="36" rank="species">new species</taxonomicNameLabel>
</emphasis>
; while
<taxonomicName id="4C63FEB0FFB3A40AC679F82F10CE1515" baseAuthorityName="Kraatz" baseAuthorityYear="1894" box="[1031,1221,1807,1832]" class="Insecta" family="Scarabaeidae" genus="Macronotops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B9175921FFB3A40AC679F82F10CE1515" box="[1031,1221,1807,1832]" italics="true" pageId="35" pageNumber="36">M. sexmaculatus</emphasis>
</taxonomicName>
is only distributed along the south Himalayas region.
</paragraph>
</subSubSection>
</treatment>
</document>