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<document id="A73BE3D9C66E39F14ECCC033A83A1626" ID-CLB-Dataset="29024" ID-DOI="10.11646/zootaxa.4459.2.2" ID-GBIF-Dataset="634e23da-84c4-46a4-8bb8-94fcf1637160" ID-ISSN="1175-5326" ID-Zenodo-Dep="1458548" ID-ZooBank="73510C3B-6482-4760-8173-B6E701839940" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539288809625" checkinUser="plazi" docAuthor="Sheth, Sayali D., Ghate, Hemant V. &amp; Hájek, Jiří" docDate="2018" docId="03BC3747FF8FFF85FF5E1289FB27FAD9" docLanguage="en" docName="zootaxa.4459.2.2.pdf" docOrigin="Zootaxa 4459 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus sociennus J. Balfour-Browne in Guignot 1952" docType="treatment" docVersion="7" lastPageNumber="257" masterDocId="FF854F3FFF9BFF93FFC91143FFF8FFA8" masterDocTitle="Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="260" masterPageNumber="235" pageNumber="255" updateTime="1698578320975" updateUser="ExternalLinkService">
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<mods:title id="DEE7669B45FBB6032C3D08F27FF2AD4A">Copelatus Erichson, 1832 from Maharashtra, India, with description of three new species and notes on other taxa of the genus (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="CA3E0229E372EE4909F1898B15445AA4">Sheth, Sayali D.</mods:namePart>
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<mods:namePart id="F541A40E98297785AFD8537F5A1867A5">Ghate, Hemant V.</mods:namePart>
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<mods:namePart id="90D2B922F73EFBBD691406C116E0AAF0">Hájek, Jiří</mods:namePart>
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<mods:date id="946F5F5176EC9A544B992B8E32C4A68F">2018</mods:date>
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<treatment id="03BC3747FF8FFF85FF5E1289FB27FAD9" ID-DOI="http://doi.org/10.5281/zenodo.5966551" ID-GBIF-Taxon="148692682" ID-Zenodo-Dep="5966551" LSID="urn:lsid:plazi:treatment:03BC3747FF8FFF85FF5E1289FB27FAD9" httpUri="http://treatment.plazi.org/id/03BC3747FF8FFF85FF5E1289FB27FAD9" lastPageId="22" lastPageNumber="257" pageId="20" pageNumber="255">
<subSubSection id="C30FD5DAFF8FFF87FF5E1289FEDFFBAE" pageId="20" pageNumber="255" type="nomenclature">
<paragraph id="8BAA8651FF8FFF87FF5E1289FD3FFC4C" blockId="20.[151,711,970,1030]" box="[151,711,970,996]" pageId="20" pageNumber="255">
<heading id="D0E2313DFF8FFF87FF5E1289FD3FFC4C" bold="true" box="[151,711,970,996]" fontSize="11" level="1" pageId="20" pageNumber="255" reason="1">
<taxonomicName id="4C15FDD2FF8FFF87FF5E1289FD3FFC4C" ID-CoL="Y3FW" authority="J. Balfour-Browne, 1952" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[151,711,970,996]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FF5E1289FD3FFC4C" bold="true" box="[151,711,970,996]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF5E1289FE77FC4C" bold="true" box="[151,399,970,996]" italics="true" pageId="20" pageNumber="255">Copelatus sociennus</emphasis>
J. Balfour-Browne, 1952
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF5E12AEFEDFFBAE" blockId="20.[151,711,970,1030]" box="[151,295,1005,1030]" pageId="20" pageNumber="255">
(
<figureCitation id="132E9AD4FF8FFF87FF5612AEFEE7FBAE" box="[159,287,1005,1030]" captionStart="FIGURES 49–51" captionStartId="23.[151,263,1951,1974]" captionTargetBox="[192,1412,971,1918]" captionTargetId="figure@23.[151,1436,913,1931]" captionTargetPageId="23" captionText="FIGURES 49–51. Copelatus sociennus. 49. male habitus (Nepal); 50. median lobe in lateral view (redrawn from original description of C. bangalorensis); 51. same (paratype of C. sociennus). Scale bar 0.5 mm (Figs 50–51)." httpUri="https://zenodo.org/record/1458572/files/figure.png" pageId="20" pageNumber="255">Figs 49–51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8FFF87FF5E1573FCE8FBCC" pageId="20" pageNumber="255" type="reference_group">
<paragraph id="8BAA8651FF8FFF87FF5E1573FCF2FBEE" blockId="20.[151,784,1072,1124]" box="[151,778,1072,1094]" pageId="20" pageNumber="255">
<treatmentCitationGroup id="AB05A17FFF8FFF87FF5E1573FCF2FBEE" box="[151,778,1072,1094]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF5E1573FCFEFBEE" ID-CoL="Y3FW" authority="J. Balfour-Browne in Guignot, 1952: 26" authorityName="J. Balfour-Browne in Guignot" authorityPageNumber="26" authorityYear="1952" box="[151,774,1072,1094]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FF5E1573FE90FBEE" box="[151,360,1072,1094]" italics="true" pageId="20" pageNumber="255">Copelatus sociennus</emphasis>
J. Balfour-Browne in
<treatmentCitation id="0AB4A040FF8FFF87FD831573FCFEFBEE" author="" box="[586,774,1072,1094]" page="26" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FD831573FD18FBEE" box="[586,736,1072,1094]" pageId="20" pageNumber="255" refId="ref15555" refString="Guignot, F. (1952) Description de dytiscides inedits de la collection Regimbart. Revue Francaise d'Entomologie, 19, 17 - 31." type="journal article">Guignot, 1952</bibRefCitation>
: 26
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF5E150DFCE8FBCC" blockId="20.[151,784,1072,1124]" box="[151,784,1102,1124]" pageId="20" pageNumber="255">
<treatmentCitationGroup id="AB05A17FFF8FFF87FF5E150DFCE8FBCC" box="[151,784,1102,1124]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF5E150DFD9AFBCC" ID-CoL="8L472" authority="Vazirani, 1970: 311" authorityName="Vazirani" authorityPageNumber="311" authorityYear="1970" box="[151,610,1102,1124]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="bangalorensis">
<emphasis id="B9615A43FF8FFF87FF5E150DFE6AFBCC" box="[151,402,1102,1124]" italics="true" pageId="20" pageNumber="255">Copelatus bangalorensis</emphasis>
<treatmentCitation id="0AB4A040FF8FFF87FE50150DFD9AFBCC" author="" box="[409,610,1102,1124]" page="311" pageId="20" pageNumber="255">
<bibRefCitation id="EF84FBA0FF8FFF87FE50150DFDC8FBCC" box="[409,560,1102,1124]" pageId="20" pageNumber="255" refId="ref16506" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4, 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article">Vazirani, 1970</bibRefCitation>
: 311
</treatmentCitation>
</taxonomicName>
;
<emphasis id="B9615A43FF8FFF87FDA5150CFCF2FBCC" bold="true" box="[620,778,1103,1124]" pageId="20" pageNumber="255">new synonymy</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8FFF87FF5E15CDFCE3F976" pageId="20" pageNumber="255" type="materials_examined">
<paragraph id="8BAA8651FF8FFF87FF5E15CDFB13FB0F" blockId="20.[151,1437,1166,2025]" box="[151,1259,1166,1191]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF5E15CDFEB0FB0F" bold="true" box="[151,328,1166,1191]" pageId="20" pageNumber="255">
<typeStatus id="54AE38F3FF8FFF87FF5E15CDFF2AFB0F" box="[151,210,1166,1191]" pageId="20" pageNumber="255">Type</typeStatus>
localities.
</emphasis>
<taxonomicName id="4C15FDD2FF8FFF87FE8715CDFD9DFB0F" authorityName="Vazirani" authorityYear="1970" box="[334,613,1166,1191]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="bangalorensis">
<emphasis id="B9615A43FF8FFF87FE8715CDFD9DFB0F" box="[334,613,1166,1191]" italics="true" pageId="20" pageNumber="255">Copelatus bangalorensis</emphasis>
</taxonomicName>
: &quot;
<collectingCountry id="F302C6C1FF8FFF87FDB415CDFD45FB0F" box="[637,701,1166,1191]" name="India" pageId="20" pageNumber="255">India</collectingCountry>
: Bangalore&quot;.
<taxonomicName id="4C15FDD2FF8FFF87FC9D15CCFC1AFB0E" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[852,994,1167,1190]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FC9D15CCFC1AFB0E" box="[852,994,1167,1190]" italics="true" pageId="20" pageNumber="255">C. sociennus</emphasis>
</taxonomicName>
: &quot;
<collectingCountry id="F302C6C1FF8FFF87FC3215CDFBBDFB0F" box="[1019,1093,1166,1191]" name="China" pageId="20" pageNumber="255">China</collectingCountry>
: (
<collectingCountry id="F302C6C1FF8FFF87FB9E15CDFB2EFB0F" box="[1111,1238,1166,1191]" name="Hong Kong" pageId="20" pageNumber="255">Hongkong</collectingCountry>
)&quot;.
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF0E15ECFD17FB42" blockId="20.[151,1437,1166,2025]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF0E15ECFE81FB60" bold="true" box="[199,377,1199,1224]" pageId="20" pageNumber="255">Type material.</emphasis>
<taxonomicName id="4C15FDD2FF8FFF87FE4D15ECFD67FB60" authorityName="Vazirani" authorityYear="1970" box="[388,671,1199,1224]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="bangalorensis">
<emphasis id="B9615A43FF8FFF87FE4D15ECFD67FB60" box="[388,671,1199,1224]" italics="true" pageId="20" pageNumber="255">Copelatus bangalorensis</emphasis>
</taxonomicName>
: Holotype ♂, deposited in Muséum national d'Histoire naturelle, Paris, France (not found; A. Mantilleri, pers. comm.).
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF0E15B1FB4FF99F" blockId="20.[151,1437,1166,2025]" pageId="20" pageNumber="255">
<taxonomicName id="4C15FDD2FF8FFF87FF0E15B1FE56FAA3" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[199,430,1266,1291]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FF0E15B1FE56FAA3" box="[199,430,1266,1291]" italics="true" pageId="20" pageNumber="255">Copelatus sociennus</emphasis>
</taxonomicName>
: Holotype ♂ (BMNH), labelled: &quot;833 [hw, round label] // Hong Kong / [on reverse:] 48 / 60 [hw] //
<taxonomicName id="4C15FDD2FF8FFF87FEDA1457FE7BFA85" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[275,387,1300,1325]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/ discoideus Shp [hw] //
<taxonomicName id="4C15FDD2FF8FFF87FD6E1457FCEFFA85" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[679,791,1300,1325]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/ japonicus / ♂ Sharp. / det.J.Balfour-Browne [hw, blue ink] //
<taxonomicName id="4C15FDD2FF8FFF87FF251476FEA4FAE6" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[236,348,1333,1358]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
♂ / chinensis Rég. [hw] / J.Balfour-Browne det. [p] / Det. from Descr. [p]
<date id="FFABA091FF8FFF87FB391476FAC7FAE6" box="[1264,1343,1333,1358]" pageId="20" pageNumber="255" value="1946-02">ii.1946</date>
[hw] // HOLOTYPE ♂ /
<taxonomicName id="4C15FDD2FF8FFF87FEAB1414FE05FAC6" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[354,509,1367,1390]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="B9615A43FF8FFF87FEAB1414FE05FAC6" box="[354,509,1367,1390]" italics="true" pageId="20" pageNumber="255">COPELATUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C15FDD2FF8FFF87FDDF1414FD7CFAC6" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[534,644,1367,1390]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FDDF1414FD7CFAC6" box="[534,644,1367,1390]" italics="true" pageId="20" pageNumber="255">sociennus</emphasis>
</taxonomicName>
/ J.Balfour-Browne, 1952 / labelled by J. Hájek 2016 [p, red label]&quot;. Paratypes: 1♂ (BMNH), labelled: &quot;Bowring / China [hw] // 457 /
<date id="FFABA091FF8FFF87FCBD143BFC31FA38" box="[884,969,1400,1424]" pageId="20" pageNumber="255" value="1949-08-17">17/8/49</date>
[hw, round label] //
<taxonomicName id="4C15FDD2FF8FFF87FB651434FAE4FA38" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[1196,1308,1399,1424]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/ japonicus / ♂ Sharp. / det.J.Balfour-Browne [hw, blue ink] //
<taxonomicName id="4C15FDD2FF8FFF87FD1D14DAFCBCFA1A" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[724,836,1433,1458]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
♂ / chinensis Rég. [hw] / J.Balfour-Browne det. [p] / Det.from Descr. [p]
<date id="FFABA091FF8FFF87FEB114F9FE3FFA7B" box="[376,455,1466,1491]" pageId="20" pageNumber="255" value="1946-02">ii.1946</date>
[hw]&quot;; 1♂ (BMNH), labelled: &quot;Bowring / China [hw] // 457 /
<date id="FFABA091FF8FFF87FB4914F9FB30FA7B" box="[1152,1224,1466,1491]" pageId="20" pageNumber="255" value="1953-08-07">7/8/53</date>
[hw, round label] / /
<taxonomicName id="4C15FDD2FF8FFF87FF6F149FFEEEFA5D" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[166,278,1500,1525]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="260" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/? chinensis Rég. [hw] / J.Balfour-Browne det. [p]&quot;; 1♀ (BMNH), labelled: &quot;Bowring / China [hw] // 457 /
<date id="FFABA091FF8FFF87FF1214BEFEDAF9BE" box="[219,290,1533,1558]" pageId="20" pageNumber="255" value="1952-06-09">9/6/52</date>
[hw, round label]&quot;. All paratypes provided with additional label &quot;PARATYPE ♂ [♀, respectively] /
<taxonomicName id="4C15FDD2FF8FFF87FF5E175CFECAF99E" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[151,306,1567,1590]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="B9615A43FF8FFF87FF5E175CFECAF99E" box="[151,306,1567,1590]" italics="true" pageId="20" pageNumber="255">COPELATUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C15FDD2FF8FFF87FE8E175CFE4DF99E" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[327,437,1567,1590]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FE8E175CFE4DF99E" box="[327,437,1567,1590]" italics="true" pageId="20" pageNumber="255">sociennus</emphasis>
</taxonomicName>
/ J.Balfour-Browne, 1952 / labelled by J. Hájek 2016 [p, red label]&quot;.
</paragraph>
<paragraph id="8BAA8651FF8FFF87FF0E177CFCE3F976" blockId="20.[151,1437,1166,2025]" pageId="20" pageNumber="255">
<materialsCitation id="3B7D8C0CFF8FFF87FF0E177CFE80F9D2" ID-GBIF-Occurrence="1917355436" collectingDate="1997-10-23" collectionCode="JSCL" collectorName="J. Stastny" country="India" location="Rhishikesh" municipality="Uttarakhand" pageId="20" pageNumber="255" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Uttar Pradesh">
<emphasis id="B9615A43FF8FFF87FF0E177CFD90F9F0" bold="true" box="[199,616,1599,1624]" pageId="20" pageNumber="255">
Additional material studied.
<collectingCountry id="F302C6C1FF8FFF87FDD6177CFD9BF9F0" box="[543,611,1599,1624]" name="India" pageId="20" pageNumber="255">India</collectingCountry>
:
</emphasis>
<specimenCount id="9D134DD8FF8FFF87FDA61703FD6AF9F0" box="[623,658,1600,1624]" pageId="20" pageNumber="255" type="male">1♂</specimenCount>
<specimenCount id="9D134DD8FF8FFF87FD501703FD44F9F0" box="[665,700,1600,1624]" pageId="20" pageNumber="255" type="female">1♀</specimenCount>
,
<collectingRegion id="49D148B3FF8FFF87FD031703FC9CF9F0" box="[714,868,1599,1624]" country="India" name="Uttar Pradesh" pageId="20" pageNumber="255">Uttar Pradesh</collectingRegion>
[currently
<collectingMunicipality id="6BCE1C2BFF8FFF87FC29177CFB89F9F0" box="[992,1137,1599,1624]" pageId="20" pageNumber="255">
<collectingRegion id="49D148B3FF8FFF87FC29177CFB89F9F0" box="[992,1137,1599,1624]" country="India" name="Uttarakhand" pageId="20" pageNumber="255">Uttarakhand</collectingRegion>
</collectingMunicipality>
],
<location id="8ECAD08AFF8FFF87FB4B177CFAF9F9F0" LSID="urn:lsid:plazi:treatment:03BC3747FF8FFF85FF5E1289FB27FAD9:8ECAD08AFF8FFF87FB4B177CFAF9F9F0" box="[1154,1281,1599,1624]" country="India" municipality="Uttarakhand" name="Rhishikesh" pageId="20" pageNumber="255" stateProvince="Uttar Pradesh">Rhishikesh</location>
,
<collectingDate id="EFEF5979FF8FFF87FAC51703FA86F9F0" box="[1292,1406,1600,1624]" pageId="20" pageNumber="255" value="1997-10-23">
<date id="FFABA091FF8FFF87FAC51703FA86F9F0" box="[1292,1406,1600,1624]" pageId="20" pageNumber="255" value="1997-10-23">23.x.1997</date>
</collectingDate>
,
<collectorName id="26E0E387FF8FFF87FA431703FF16F9D2" pageId="20" pageNumber="255">J. Šťastný</collectorName>
leg. (
<collectionCode id="ED041E94FF8FFF87FEE51721FE88F9D2" box="[300,368,1634,1658]" pageId="20" pageNumber="255">JSCL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8FFF87FE4A1721FD47F933" ID-GBIF-Occurrence="1917355456" collectingDate="2003-08-04" collectionCode="NMPC" collectorName="Z. Kejval &amp; M. Tryzna" country="India" county="Uttaranchal" elevation="790" location="Yamuna river" municipality="Dakpathar" pageId="20" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Uttarakhand">
<specimenCount id="9D134DD8FF8FFF87FE4A1721FE5FF9D2" box="[387,423,1634,1658]" pageId="20" pageNumber="255" type="male">1♂</specimenCount>
,
<collectingCounty id="62CBFEDDFF8FFF87FE7A1722FDCEF9D2" box="[435,566,1633,1658]" pageId="20" pageNumber="255">Uttaranchal</collectingCounty>
[
<collectingRegion id="49D148B3FF8FFF87FD8A1722FD2EF9D2" box="[579,726,1633,1658]" country="India" name="Uttarakhand" pageId="20" pageNumber="255">Uttarakhand</collectingRegion>
],
<locationDeviation id="9A189F73FF8FFF87FD2E1721FC13F9D2" box="[743,1003,1633,1658]" pageId="20" pageNumber="255">
<quantity id="4CED2BB4FF8FFF87FD2E1721FCD5F9D2" box="[743,813,1633,1658]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="255" unit="km" value="30.0">30 km</quantity>
W of Mussoorie
</locationDeviation>
,
<collectingMunicipality id="6BCE1C2BFF8FFF87FC3F1722FB93F9D2" box="[1014,1131,1633,1658]" pageId="20" pageNumber="255">Dakpathar</collectingMunicipality>
vill.,
<location id="8ECAD08AFF8FFF87FB621721FABCF9D2" LSID="urn:lsid:plazi:treatment:03BC3747FF8FFF85FF5E1289FB27FAD9:8ECAD08AFF8FFF87FB621721FABCF9D2" box="[1195,1348,1633,1658]" country="India" county="Uttaranchal" municipality="Dakpathar" name="Yamuna river" pageId="20" pageNumber="255" stateProvince="Uttarakhand">Yamuna river</location>
,
<elevation id="00386162FF8FFF87FA861721FA6FF9D1" box="[1359,1431,1634,1658]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="20" pageNumber="255" unit="m" value="790.0">
<quantity id="4CED2BB4FF8FFF87FA861721FA6FF9D1" box="[1359,1431,1634,1658]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="20" pageNumber="255" unit="m" value="790.0">790 m</quantity>
</elevation>
,
<collectingDate id="EFEF5979FF8FFF87FF5E17C1FEEBF933" box="[151,275,1666,1691]" pageId="20" pageNumber="255" value="2003-08-04">
<date id="FFABA091FF8FFF87FF5E17C1FEEBF933" box="[151,275,1666,1691]" pageId="20" pageNumber="255" value="2003-08-04">4.viii.2003</date>
</collectingDate>
,
<collectorName id="26E0E387FF8FFF87FED717C0FE71F933" box="[286,393,1666,1691]" pageId="20" pageNumber="255">Z. Kejval</collectorName>
&amp;
<collectorName id="26E0E387FF8FFF87FE6417C0FDDAF933" box="[429,546,1667,1691]" pageId="20" pageNumber="255">M. Trýzna</collectorName>
leg. (
<collectionCode id="ED041E94FF8FFF87FDAB17C0FD40F933" box="[610,696,1667,1691]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="255">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8FFF87FD0217C0FED5F976" ID-GBIF-Occurrence="1917355435" collectingDate="2006-05-31" collectionCode="NMPC" collectorName="V. Ryjacek &amp; D. Kral" country="Nepal" elevation="1387" latitude="27.45" location="Nagarjun forest" longLatPrecision="742" longitude="85.17" municipality="Chikmagalur" pageId="20" pageNumber="255" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Bagmati">
<specimenCount id="9D134DD8FF8FFF87FD0217C0FD16F933" box="[715,750,1667,1691]" pageId="20" pageNumber="255" type="female">1♀</specimenCount>
, Karnataka, Western Ghats Mts.,
<collectingMunicipality id="6BCE1C2BFF8FFF87FBA017C1FB05F933" box="[1129,1277,1666,1691]" pageId="20" pageNumber="255">Chikmagalur</collectingMunicipality>
,
<collectingDate id="EFEF5979FF8FFF87FAC117C1FA81F933" box="[1288,1401,1666,1691]" pageId="20" pageNumber="255" value="2006-05-31">
<date id="FFABA091FF8FFF87FAC117C1FA81F933" box="[1288,1401,1666,1691]" pageId="20" pageNumber="255" value="2006-05-31">31.v.2006</date>
</collectingDate>
,
<collectorName id="26E0E387FF8FFF87FA4D17C0FF0AF915" pageId="20" pageNumber="255">V. Ryjáček</collectorName>
lgt. (
<collectionCode id="ED041E94FF8FFF87FEE617E7FE7DF914" box="[303,389,1700,1724]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="255">NMPC</collectionCode>
).
<emphasis id="B9615A43FF8FFF87FE5117E7FE10F915" bold="true" box="[408,488,1700,1725]" pageId="20" pageNumber="255">
<collectingCountry id="F302C6C1FF8FFF87FE5117E7FE1AF915" box="[408,482,1700,1725]" name="Nepal" pageId="20" pageNumber="255">Nepal</collectingCountry>
:
</emphasis>
<specimenCount id="9D134DD8FF8FFF87FE3917E7FDEBF914" box="[496,531,1700,1724]" pageId="20" pageNumber="255" type="male">2♂</specimenCount>
<specimenCount id="9D134DD8FF8FFF87FDD217E7FDB8F914" box="[539,576,1700,1724]" pageId="20" pageNumber="255" type="female">2♀</specimenCount>
,
<collectingRegion id="49D148B3FF8FFF87FD8717E7FD55F915" box="[590,685,1700,1725]" country="Nepal" name="Bagmati" pageId="20" pageNumber="255">Bagmati</collectingRegion>
Prov.,
<location id="8ECAD08AFF8FFF87FD3617E7FC4AF915" LSID="urn:lsid:plazi:treatment:03BC3747FF8FFF85FF5E1289FB27FAD9:8ECAD08AFF8FFF87FD3617E7FC4AF915" box="[767,946,1700,1725]" country="Nepal" latitude="27.45" longLatPrecision="742" longitude="85.17" municipality="Chikmagalur" name="Nagarjun forest" pageId="20" pageNumber="255" stateProvince="Bagmati">Nagarjun forest</location>
,
<geoCoordinate id="EE21E096FF8FFF87FC7617E7FBEDF915" box="[959,1045,1700,1725]" direction="north" orientation="latitude" pageId="20" pageNumber="255" precision="555" value="27.45">27.45N</geoCoordinate>
,
<geoCoordinate id="EE21E096FF8FFF87FBE817E7FB8CF915" box="[1057,1140,1700,1725]" direction="east" orientation="longitude" pageId="20" pageNumber="255" precision="555" value="85.17">85.17E</geoCoordinate>
,
<elevation id="00386162FF8FFF87FB4817E7FB21F914" box="[1153,1241,1700,1724]" metricMagnitude="3" metricUnit="m" metricValue="1.387" pageId="20" pageNumber="255" unit="m" value="1387.0">
<quantity id="4CED2BB4FF8FFF87FB4817E7FB21F914" box="[1153,1241,1700,1724]" metricMagnitude="3" metricUnit="m" metricValue="1.387" pageId="20" pageNumber="255" unit="m" value="1387.0">1387 m</quantity>
</elevation>
, at light,
<collectorName id="26E0E387FF8FFF87FA8F17E6FA64F915" box="[1350,1436,1700,1725]" pageId="20" pageNumber="255">D. Král</collectorName>
leg. (
<collectionCode id="ED041E94FF8FFF87FF061785FEDDF976" box="[207,293,1734,1758]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="20" pageNumber="255">NMPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7D8C0CFF8FFF87FEF11785FCE3F976" ID-GBIF-Occurrence="1917355448" box="[312,795,1733,1758]" collectingDate="2006-05-31" collectionCode="NHMB" collectorName="J. Farkac" country="Nepal" elevation="1387" latitude="27.45" location="Nagarjun forest" longLatPrecision="742" longitude="85.17" municipality="Chikmagalur" pageId="20" pageNumber="255" specimenCount="1" specimenCount-male="1" stateProvince="Bagmati">
<specimenCount id="9D134DD8FF8FFF87FEF11785FEA4F976" box="[312,348,1734,1758]" pageId="20" pageNumber="255" type="male">1♂</specimenCount>
, same data, but
<collectorName id="26E0E387FF8FFF87FDD81785FD8EF976" box="[529,630,1733,1758]" pageId="20" pageNumber="255">J. Farkač</collectorName>
leg. (
<collectionCode id="ED041E94FF8FFF87FD7C1785FCF7F976" box="[693,783,1734,1758]" country="Switzerland" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="20" pageNumber="255">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30FD5DAFF8FFF87FF0F17A5FAC7F841" pageId="20" pageNumber="255" type="discussion">
<paragraph id="8BAA8651FF8FFF87FF0F17A5FAC7F841" blockId="20.[151,1437,1166,2025]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FF0F17A5FDEAF957" bold="true" box="[198,530,1766,1791]" pageId="20" pageNumber="255">Comments on classification.</emphasis>
<bibRefCitation id="EF84FBA0FF8FFF87FDD417A5FD32F957" box="[541,714,1766,1791]" pageId="20" pageNumber="255" refId="ref16506" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4, 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article">Vazirani (1970)</bibRefCitation>
diagnosed
<taxonomicName id="4C15FDD2FF8FFF87FC8717A5FBA6F957" authorityName="Vazirani" authorityYear="1970" box="[846,1118,1766,1791]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="bangalorensis">
<emphasis id="B9615A43FF8FFF87FC8717A5FBA6F957" box="[846,1118,1766,1791]" italics="true" pageId="20" pageNumber="255">Copelatus bangalorensis</emphasis>
</taxonomicName>
based on the unique dorsal (ventral in the current concept) projection of the median lobe. Indeed, the shape of the median lobe of the species is very characteristic within all
<collectingRegion id="49D148B3FF8FFF87FE57166AFE0EF8EA" box="[414,502,1833,1858]" country="Democratic Republic of the Congo" name="Orientale" pageId="20" pageNumber="255">Oriental</collectingRegion>
<taxonomicName id="4C15FDD2FF8FFF87FE36166AFD95F8EA" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[511,621,1833,1858]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="B9615A43FF8FFF87FE36166AFD95F8EA" box="[511,621,1833,1858]" italics="true" pageId="20" pageNumber="255">Copelatus</emphasis>
</taxonomicName>
. However, the author was apparently not aware of
<taxonomicName id="4C15FDD2FF8FFF87FB681669FAD5F8E9" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[1185,1325,1834,1857]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FB681669FAD5F8E9" box="[1185,1325,1834,1857]" italics="true" pageId="20" pageNumber="255">C. sociennus</emphasis>
</taxonomicName>
described from
<collectingCountry id="F302C6C1FF8FFF87FF1B1608FEACF8CB" box="[210,340,1867,1891]" name="Hong Kong" pageId="20" pageNumber="255">Hong Kong</collectingCountry>
, the median lobe of which has exactly the same dorsal protuberance and ventral projection. Although we were not able to study the
<typeStatus id="54AE38F3FF8FFF87FE25162FFDB3F82D" box="[492,587,1900,1925]" pageId="20" pageNumber="255" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C15FDD2FF8FFF87FDBA162EFCD5F82D" authorityName="Vazirani" authorityYear="1970" box="[627,813,1900,1925]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="bangalorensis">
<emphasis id="B9615A43FF8FFF87FDBA162EFCD5F82D" box="[627,813,1900,1925]" italics="true" pageId="20" pageNumber="255">C. bangalorensis</emphasis>
</taxonomicName>
, the comparison of its description, especially the elytral coloration, and drawing of the median lobe (
<figureCitation id="132E9AD4FF8FFF87FD4D16CEFD2EF80E" box="[644,726,1933,1958]" captionStart="FIGURES 49–51" captionStartId="23.[151,263,1951,1974]" captionTargetBox="[192,1412,971,1918]" captionTargetId="figure@23.[151,1436,913,1931]" captionTargetPageId="23" captionText="FIGURES 49–51. Copelatus sociennus. 49. male habitus (Nepal); 50. median lobe in lateral view (redrawn from original description of C. bangalorensis); 51. same (paratype of C. sociennus). Scale bar 0.5 mm (Figs 50–51)." httpUri="https://zenodo.org/record/1458572/files/figure.png" pageId="20" pageNumber="255">Fig. 50</figureCitation>
), with the
<typeStatus id="54AE38F3FF8FFF87FC8416CDFC83F80E" box="[845,891,1934,1958]" pageId="20" pageNumber="255">type</typeStatus>
material of
<taxonomicName id="4C15FDD2FF8FFF87FBCD16CDFB68F80D" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[1028,1168,1934,1957]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FBCD16CDFB68F80D" box="[1028,1168,1934,1957]" italics="true" pageId="20" pageNumber="255">C. sociennus</emphasis>
</taxonomicName>
(
<figureCitation id="132E9AD4FF8FFF87FB6916CEFB08F80E" box="[1184,1264,1933,1958]" captionStart="FIGURES 49–51" captionStartId="23.[151,263,1951,1974]" captionTargetBox="[192,1412,971,1918]" captionTargetId="figure@23.[151,1436,913,1931]" captionTargetPageId="23" captionText="FIGURES 49–51. Copelatus sociennus. 49. male habitus (Nepal); 50. median lobe in lateral view (redrawn from original description of C. bangalorensis); 51. same (paratype of C. sociennus). Scale bar 0.5 mm (Figs 50–51)." httpUri="https://zenodo.org/record/1458572/files/figure.png" pageId="20" pageNumber="255">Fig. 51</figureCitation>
) and additional material from the region leaves no doubt that the two taxa are conspecific. Therefore, we establish the following synonymy:
<taxonomicName id="4C15FDD2FF8FFF87FEDF1693FCFFF841" authority="J. Balfour-Browne, 1952" authorityName="J. Balfour-Browne in Guignot" authorityYear="1952" box="[278,775,2000,2025]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="sociennus">
<emphasis id="B9615A43FF8FFF87FEDF1693FE0DF840" box="[278,501,2000,2025]" italics="true" pageId="20" pageNumber="255">Copelatus sociennus</emphasis>
J. Balfour-Browne, 1952
</taxonomicName>
=
<taxonomicName id="4C15FDD2FF8FFF87FCED1693FB2CF840" authority="Vazirani, 1970" authorityName="Vazirani" authorityYear="1970" box="[804,1236,2000,2025]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="255" phylum="Arthropoda" rank="species" species="bangalorensis">
<emphasis id="B9615A43FF8FFF87FCED1693FBC9F841" box="[804,1073,2000,2025]" italics="true" pageId="20" pageNumber="255">Copelatus bangalorensis</emphasis>
<bibRefCitation id="EF84FBA0FF8FFF87FBFE1693FB2CF840" box="[1079,1236,2000,2025]" pageId="20" pageNumber="255" refId="ref16506" refString="Vazirani, T. G. (1970) Contributions to the study of aquatic beetles (Coleoptera). VII. A revision of Indian Colymbetinae (Dytiscidae). Oriental Insects, 4, 303 - 362. https: // doi. org / 10.1080 / 00305316.1970.10433967" type="journal article">Vazirani, 1970</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A252E738FF8FFF87FB121693FAC7F841" box="[1243,1343,2000,2025]" pageId="20" pageNumber="255">
<emphasis id="B9615A43FF8FFF87FB121693FAC7F841" bold="true" box="[1243,1343,2000,2025]" pageId="20" pageNumber="255">syn. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<caption id="DF6AD6D9FF8EFF86FF5E1638FC9AF839" box="[151,866,1915,1937]" httpUri="https://zenodo.org/record/1458568/files/figure.png" pageId="21" pageNumber="256" startId="21.[151,249,1915,1937]" targetBox="[190,1401,196,1869]" targetPageId="21">
<paragraph id="8BAA8651FF8EFF86FF5E1638FC9AF839" blockId="21.[151,866,1915,1937]" box="[151,866,1915,1937]" pageId="21" pageNumber="256">
<emphasis id="B9615A43FF8EFF86FF5E1638FEE7F838" bold="true" box="[151,287,1915,1937]" pageId="21" pageNumber="256">FIGURE 45.</emphasis>
Distributional map of
<taxonomicName id="4C15FDD2FF8EFF86FDCC1638FD94F839" authorityName="Sheth &amp; Ghate &amp; Hájek" authorityYear="2018" box="[517,620,1915,1937]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="B9615A43FF8EFF86FDCC1638FD94F839" box="[517,620,1915,1937]" italics="true" pageId="21" pageNumber="256">Copelatus</emphasis>
</taxonomicName>
species in Maharashtra.
</paragraph>
</caption>
<caption id="DF6AD6D9FF8DFF85FF5E1591FCC0FAAF" httpUri="https://zenodo.org/record/1458570/files/figure.png" pageId="22" pageNumber="257" startId="22.[151,263,1234,1257]" targetBox="[180,1407,215,1191]" targetPageId="22">
<paragraph id="8BAA8651FF8DFF85FF5E1591FCC0FAAF" blockId="22.[151,1435,1234,1287]" pageId="22" pageNumber="257">
<emphasis id="B9615A43FF8DFF85FF5E1591FEABFB40" bold="true" box="[151,339,1234,1257]" pageId="22" pageNumber="257">FIGURES 46–48.</emphasis>
<taxonomicName id="4C15FDD2FF8DFF85FE931590FDF6FB41" authorityName="Guignot" authorityYear="1955" box="[346,526,1235,1257]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="freudei">
<emphasis id="B9615A43FF8DFF85FE931590FDF6FB41" box="[346,526,1235,1257]" italics="true" pageId="22" pageNumber="257">Copelatus freudei</emphasis>
</taxonomicName>
. 46. male habitus (Rajasthan); 47. median lobe in lateral view (holotype of
<taxonomicName id="4C15FDD2FF8DFF85FAC61590FA75FB41" authorityName="Vazirani" authorityYear="1974" box="[1295,1421,1235,1257]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="spangleri">
<emphasis id="B9615A43FF8DFF85FAC61590FA75FB41" box="[1295,1421,1235,1257]" italics="true" pageId="22" pageNumber="257">C. spangleri</emphasis>
</taxonomicName>
); 48. same (holotype of
<taxonomicName id="4C15FDD2FF8DFF85FEB315B1FE1AFAAF" authorityName="Vazirani" authorityYear="1975" box="[378,482,1265,1287]" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="257" phylum="Arthropoda" rank="species" species="gibsoni">
<emphasis id="B9615A43FF8DFF85FEB315B1FE1AFAAF" box="[378,482,1265,1287]" italics="true" pageId="22" pageNumber="257">C. gibsoni</emphasis>
</taxonomicName>
). Scale bar 0.5 mm (Figs 47–48).
</paragraph>
</caption>
<subSubSection id="C30FD5DAFF8DFF85FF0E1477FB27FAD9" pageId="22" pageNumber="257" type="distribution">
<paragraph id="8BAA8651FF8DFF85FF0E1477FB27FAD9" blockId="22.[151,1436,1332,1393]" pageId="22" pageNumber="257">
<emphasis id="B9615A43FF8DFF85FF0E1477FE98FAE5" bold="true" box="[199,352,1332,1357]" pageId="22" pageNumber="257">Distribution.</emphasis>
A species occurring in the Indian subcontinent,
<collectingCountry id="F302C6C1FF8DFF85FC5D1477FBFFFAE5" box="[916,1031,1332,1357]" name="Sri Lanka" pageId="22" pageNumber="257">
<collectingRegion id="49D148B3FF8DFF85FC5D1477FC4DFAE5" box="[916,949,1332,1357]" country="United Kingdom" name="Surrey" pageId="22" pageNumber="257">Sri</collectingRegion>
Lanka
</collectingCountry>
, southern
<collectingCountry id="F302C6C1FF8DFF85FBB61477FB3DFAE5" box="[1151,1221,1332,1357]" name="China" pageId="22" pageNumber="257">China</collectingCountry>
, and also reaching continental Southeast Asia. Here, we present the first record from the Indian state of
<collectingRegion id="49D148B3FF8DFF85FB84141BFB23FAD9" box="[1101,1243,1368,1393]" country="India" name="Uttarakhand" pageId="22" pageNumber="257">Uttarakhand</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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