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<document id="A56782DEB5558155EF81FDEE616107DA" ID-ISSN="1175­5334" ID-ZooBank="505937B0-9F57-4068-82E6-8553826DD5AA" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629822157238" checkinUser="felipe" docAuthor="Judson, Sarah W. &amp; Nelson, C. Riley" docDate="2012" docId="03B387E715468117FF5AF996FD19524A" docLanguage="en" docName="zt03541p118.pdf" docOrigin="Zootaxa 3541" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Nemoura cinerea" docType="treatment" docVersion="2" lastPageNumber="34" masterDocId="FF8AFF9F15668136FFCDFFA7FFAF5015" masterDocTitle="3541" masterLastPageNumber="118" masterPageNumber="1" pageNumber="33" updateTime="1699117090469" updateUser="plazi" zenodo-license-document="CLOSED">
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<treatment id="03B387E715468117FF5AF996FD19524A" LSID="urn:lsid:plazi:treatment:03B387E715468117FF5AF996FD19524A" httpUri="http://treatment.plazi.org/id/03B387E715468117FF5AF996FD19524A" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<subSubSection id="C300657A15468116FF5AF996FE935685" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BA536F115468116FF5AF996FD85565F" blockId="32.[151,554,1584,1611]" box="[151,554,1584,1611]" pageId="32" pageNumber="33">
<heading id="D0ED819D15468116FF5AF996FD85565F" bold="true" box="[151,554,1584,1611]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
<taxonomicName id="4C1A4D7215468116FF5AF996FD85565F" ID-CoL="469PR" authority="(Retzius, 1783)" baseAuthorityName="Retzius" baseAuthorityYear="1783" box="[151,554,1584,1611]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315468116FF5AF996FD85565F" bold="true" box="[151,554,1584,1611]" pageId="32" pageNumber="33">
<emphasis id="B96EEAE315468116FF5AF996FEC6565E" bold="true" box="[151,361,1585,1611]" italics="true" pageId="32" pageNumber="33">Nemoura cinerea</emphasis>
(Retzius, 1783)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA536F115468116FF5AF9D0FE935685" blockId="32.[151,433,1655,1744]" box="[151,316,1655,1680]" pageId="32" pageNumber="33">
<heading id="D0ED819D15468116FF5AF9D0FE935685" bold="true" box="[151,316,1655,1680]" fontSize="10" level="2" pageId="32" pageNumber="33" reason="2">
<emphasis id="B96EEAE315468116FF5AF9D0FE935685" bold="true" box="[151,316,1655,1680]" pageId="32" pageNumber="33">SYNONYMY</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C300657A15468116FF5AF93BFE1E56C5" pageId="32" pageNumber="33" type="description">
<paragraph id="8BA536F115468116FF5AF93BFE1E56A4" blockId="32.[151,433,1655,1744]" box="[151,433,1691,1714]" pageId="32" pageNumber="33">
<taxonomicName id="4C1A4D7215468116FF5AF93BFE1E56A4" authority="Retzius, 1783" authorityName="Retzius" authorityYear="1783" box="[151,433,1691,1714]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315468116FF5AF93BFE8F56A4" box="[151,288,1692,1714]" italics="true" pageId="32" pageNumber="33">Perla cinerea</emphasis>
Retzius, 1783
</taxonomicName>
</paragraph>
<paragraph id="8BA536F115468116FF5AF91CFE1E56C5" blockId="32.[151,433,1655,1744]" box="[151,433,1722,1744]" pageId="32" pageNumber="33">
<taxonomicName id="4C1A4D7215468116FF5AF91CFE1E56C5" authority="Illies, 1955" authorityName="Illies" authorityYear="1955" box="[151,433,1722,1744]" class="Insecta" family="Nemouridae" genus="Nemura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315468116FF5AF91CFE9556C5" box="[151,314,1723,1744]" italics="true" pageId="32" pageNumber="33">Nemura cinerea</emphasis>
Illies, 1955
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C300657A15468116FF5AF95AFE435700" box="[151,492,1789,1814]" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BA536F115468116FF5AF95AFE435700" blockId="32.[151,1436,1789,2029]" box="[151,492,1789,1814]" pageId="32" pageNumber="33">
<materialsCitation id="3B723CAC15468116FF5AF95AFE475700" box="[151,488,1789,1814]" country="Sweden" location="Sweden" pageId="32" pageNumber="33" specimenCount="1" typeStatus="holotype">
<emphasis id="B96EEAE315468116FF5AF95AFE2B5703" bold="true" box="[151,388,1789,1814]" pageId="32" pageNumber="33">
<typeStatus id="54A1885315468116FF5AF95AFF4D5700" box="[151,226,1789,1813]" pageId="32" pageNumber="33">TYPE</typeStatus>
LOCALITY:
</emphasis>
<collectingCountry id="F30D766115468116FE41F95AFE475700" box="[396,488,1789,1813]" name="Sweden" pageId="32" pageNumber="33">Sweden</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C300657A15468117FF0AF886FDA75192" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="8BA536F115468117FF0AF886FDA75192" blockId="32.[151,1436,1789,2029]" lastBlockId="33.[151,1436,151,607]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="B96EEAE315468116FF0AF886FEC2572F" bold="true" box="[199,365,1825,1850]" pageId="32" pageNumber="33">DIAGNOSIS:</emphasis>
This species is characterized by the epiproct having apical prongs of the ventral sclerite that are triangular, and come to a point which has a few spines at the apex (
<figureCitation id="13212A7415468116FC69F8E2FBEC5748" box="[932,1091,1861,1885]" captionStart="FIGURES 176180" captionStartId="74.[151,263,983,1006]" captionTargetBox="[151,1434,193,962]" captionTargetId="figure-75@74.[151,1436,193,962]" captionTargetPageId="74" captionText="FIGURES 176180. Nemoura cinerea.176: Habitus (preserved); 177: Male terminalia, dorsal; 178: Male epiproct ventral sclerites, detail; 179: Male paraprocts, ventral; 180: Predicted ENM map with documented occurrences and range estimate." pageId="32" pageNumber="33">Fig. 177, 178</figureCitation>
). In contrast,
<taxonomicName id="4C1A4D7215468116FB2FF8E1FAC15748" authorityName="Morton" authorityYear="1896" box="[1250,1390,1861,1885]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="sahlbergi">
<emphasis id="B96EEAE315468116FB2FF8E1FAC15748" box="[1250,1390,1861,1885]" italics="true" pageId="32" pageNumber="33">N. sahlbergi</emphasis>
</taxonomicName>
has prongs which are more rounded and have more spines at the apex(
<figureCitation id="13212A7415468116FC44F8CEFC455794" box="[905,1002,1897,1921]" captionStart="FIGURES 185189" captionStartId="75.[151,264,983,1006]" captionTargetBox="[151,1436,193,962]" captionTargetId="figure-76@75.[151,1436,193,962]" captionTargetPageId="75" captionText="FIGURES 185189. Nemoura sahlbergi. 185: Habitus (preserved); 186: Male terminalia, dorsal; 187: Male epiproct ventral sclerites, detail; 188: Male paraprocts, ventral; 189: Predicted ENM map with documented occurrences and range estimate." pageId="32" pageNumber="33">Fig. 187</figureCitation>
). Like,
<taxonomicName id="4C1A4D7215468116FB8DF8CDFB1F5794" authorityName="Esben-Petersen" authorityYear="1910" box="[1088,1200,1898,1921]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="arctica">
<emphasis id="B96EEAE315468116FB8DF8CDFB1F5794" box="[1088,1200,1898,1921]" italics="true" pageId="32" pageNumber="33">N. arctica</emphasis>
</taxonomicName>
(
<figureCitation id="13212A7415468116FB0CF8CEFA8C5794" box="[1217,1315,1897,1921]" captionStart="FIGURES 171175" captionStartId="73.[151,263,1978,2001]" captionTargetBox="[177,1421,1221,1965]" captionTargetId="figure-81@73.[177,1421,1221,1965]" captionTargetPageId="73" captionText="FIGURES 171175. Nemoura arctica.171: Habitus (preserved); 172: Male terminalia, dorsal; 173: Male epiproct ventral sclerites, detail; 174: Male paraprocts, ventral; 175: Predicted ENM map with documented occurrences and range estimate." pageId="32" pageNumber="33">Fig. 174</figureCitation>
), the outer (major) lobe of the paraproct of
<taxonomicName id="4C1A4D7215468116FDCFF829FDD857B0" authorityName="Illies" authorityYear="1955" box="[514,631,1934,1957]" class="Insecta" family="Nemouridae" genus="Nemura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315468116FDCFF829FDD857B0" box="[514,631,1934,1957]" italics="true" pageId="32" pageNumber="33">N. cinerea</emphasis>
</taxonomicName>
is convex (
<figureCitation id="13212A7415468116FD34F82AFCF557B0" box="[761,858,1933,1957]" captionStart="FIGURES 176180" captionStartId="74.[151,263,983,1006]" captionTargetBox="[151,1434,193,962]" captionTargetId="figure-75@74.[151,1436,193,962]" captionTargetPageId="74" captionText="FIGURES 176180. Nemoura cinerea.176: Habitus (preserved); 177: Male terminalia, dorsal; 178: Male epiproct ventral sclerites, detail; 179: Male paraprocts, ventral; 180: Predicted ENM map with documented occurrences and range estimate." pageId="32" pageNumber="33">Fig. 179</figureCitation>
), but in
<taxonomicName id="4C1A4D7215468116FC75F829FB8357B0" authorityName="Illies" authorityYear="1955" box="[952,1068,1934,1957]" class="Insecta" family="Nemouridae" genus="Nemura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315468116FC75F829FB8357B0" box="[952,1068,1934,1957]" italics="true" pageId="32" pageNumber="33">N. cinerea</emphasis>
</taxonomicName>
the narrow dividing notch at the apex of the segment creates an asymmetrical structure with an outline in the shape of a mitten (
<figureCitation id="13212A7415468116FB07F816FA8357DC" box="[1226,1324,1969,1993]" captionStart="FIGURES 176180" captionStartId="74.[151,263,983,1006]" captionTargetBox="[151,1434,193,962]" captionTargetId="figure-75@74.[151,1436,193,962]" captionTargetPageId="74" captionText="FIGURES 176180. Nemoura cinerea.176: Habitus (preserved); 177: Male terminalia, dorsal; 178: Male epiproct ventral sclerites, detail; 179: Male paraprocts, ventral; 180: Predicted ENM map with documented occurrences and range estimate." pageId="32" pageNumber="33">Fig. 179</figureCitation>
). Cerci of this species have only 2 apical spines perpendicular to the cercus. Unlike other
<taxonomicName id="4C1A4D7215468116FBECF871FB2657F8" box="[1057,1161,2006,2029]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAE315468116FBECF871FB2657F8" box="[1057,1161,2006,2029]" italics="true" pageId="32" pageNumber="33">Nemoura</emphasis>
</taxonomicName>
, the cerci are twisted in such a way that the black dot in the center of the membranous portion is readily visible ventrally (
<figureCitation id="13212A7415478117FB29FF30FAE950BA" box="[1252,1350,151,175]" captionStart="FIGURES 176180" captionStartId="74.[151,263,983,1006]" captionTargetBox="[151,1434,193,962]" captionTargetId="figure-75@74.[151,1436,193,962]" captionTargetPageId="74" captionText="FIGURES 176180. Nemoura cinerea.176: Habitus (preserved); 177: Male terminalia, dorsal; 178: Male epiproct ventral sclerites, detail; 179: Male paraprocts, ventral; 180: Predicted ENM map with documented occurrences and range estimate." pageId="33" pageNumber="34">Fig. 179</figureCitation>
), rather than dorsally (
<figureCitation id="13212A7415478117FE89FF1BFE0250C1" box="[324,429,188,212]" captionStart="FIGURES 185189" captionStartId="75.[151,264,983,1006]" captionTargetBox="[151,1436,193,962]" captionTargetId="figure-76@75.[151,1436,193,962]" captionTargetPageId="75" captionText="FIGURES 185189. Nemoura sahlbergi. 185: Habitus (preserved); 186: Male terminalia, dorsal; 187: Male epiproct ventral sclerites, detail; 188: Male paraprocts, ventral; 189: Predicted ENM map with documented occurrences and range estimate." pageId="33" pageNumber="34">Fig. 186</figureCitation>
). The female subgenital plate has a broad central bump, but this is not particularly diagnostic because of variability within the species.
<taxonomicName id="4C1A4D7215478117FD3AFF46FC1350E2" baseAuthorityName="Retzius" baseAuthorityYear="1783" box="[759,956,224,248]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315478117FD3AFF46FC1350E2" box="[759,956,224,248]" italics="true" pageId="33" pageNumber="34">Nemoura cinerea</emphasis>
</taxonomicName>
is noted to have rather homogenous hair across the femora with some longer hairs dorsally near the distal margin. However, this character is variably distinguishable from
<taxonomicName id="4C1A4D7215478117FE49FE8EFE5D512A" authorityName="Esben-Petersen" authorityYear="1910" box="[388,498,296,320]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="arctica">
<emphasis id="B96EEAE315478117FE49FE8EFE5D512A" box="[388,498,296,320]" italics="true" pageId="33" pageNumber="34">N. arctica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A4D7215478117FDE5FE8EFD1D5155" authorityName="Morton" authorityYear="1896" box="[552,690,296,320]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sahlbergi">
<emphasis id="B96EEAE315478117FDE5FE8EFD1D5155" box="[552,690,296,320]" italics="true" pageId="33" pageNumber="34">N. sahlbergi</emphasis>
</taxonomicName>
, such that Lillehammer (1988) included
<taxonomicName id="4C1A4D7215478117FBB4FE8EFB42512A" baseAuthorityName="Retzius" baseAuthorityYear="1783" box="[1145,1261,296,320]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315478117FBB4FE8EFB42512A" box="[1145,1261,296,320]" italics="true" pageId="33" pageNumber="34">N. cinerea</emphasis>
</taxonomicName>
in two portions of his key both with and without this
<typeStatus id="54A1885315478117FD8DFEEBFDDF5171" box="[576,624,332,356]" pageId="33" pageNumber="34">type</typeStatus>
of setation.
<taxonomicName id="4C1A4D7215478117FD30FEEAFC6F5171" baseAuthorityName="Retzius" baseAuthorityYear="1783" box="[765,960,333,356]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315478117FD30FEEAFC6F5171" box="[765,960,333,356]" italics="true" pageId="33" pageNumber="34">Nemoura cinerea</emphasis>
</taxonomicName>
nymphs lack hairs on the galea, which
<taxonomicName id="4C1A4D7215478117FA4FFEEAFF4A5192" authorityName="Esben-Petersen" authorityYear="1910" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="arctica">
<emphasis id="B96EEAE315478117FA4FFEEAFF4A5192" italics="true" pageId="33" pageNumber="34">N. arctica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1A4D7215478117FED1FED6FE0A519D" authorityName="Morton" authorityYear="1896" box="[284,421,368,392]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sahlbergi">
<emphasis id="B96EEAE315478117FED1FED6FE0A519D" box="[284,421,368,392]" italics="true" pageId="33" pageNumber="34">N. sahlbergi</emphasis>
</taxonomicName>
possess.
</paragraph>
</subSubSection>
<subSubSection id="C300657A15478117FF0AFE34FEE251DA" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BA536F115478117FF0AFE34FEE251DA" blockId="33.[151,1436,151,607]" pageId="33" pageNumber="34">
<emphasis id="B96EEAE315478117FF0AFE34FDA351B9" bold="true" box="[199,524,403,428]" pageId="33" pageNumber="34">DISTRIBUTION—Global:</emphasis>
Trans-Palearctic—
<emphasis id="B96EEAE315478117FD24FE34FCF351B9" bold="true" box="[745,860,403,428]" pageId="33" pageNumber="34">Regional:</emphasis>
AOB, IDB*—
<emphasis id="B96EEAE315478117FBC0FE34FBCA51B9" bold="true" box="[1037,1125,403,428]" pageId="33" pageNumber="34">Aimag:</emphasis>
AR^, BU^, KhD*, KhG^, SE*, TO^, ZA^.
</paragraph>
</subSubSection>
<subSubSection id="C300657A15478117FF0AFE7CFD19524A" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BA536F115478117FF0AFE7CFD19524A" blockId="33.[151,1436,151,607]" pageId="33" pageNumber="34">
<emphasis id="B96EEAE315478117FF0AFE7CFED451E1" bold="true" box="[199,379,475,500]" pageId="33" pageNumber="34">DISCUSSION:</emphasis>
<taxonomicName id="4C1A4D7215478117FE4EFE7AFDE951E1" baseAuthorityName="Retzius" baseAuthorityYear="1783" box="[387,582,477,500]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315478117FE4EFE7AFDE951E1" box="[387,582,477,500]" italics="true" pageId="33" pageNumber="34">Nemoura cinerea</emphasis>
</taxonomicName>
appears to be restricted to the
<collectingRegion id="49DEF81315478117FC6EFE7BFC5451E1" box="[931,1019,476,500]" country="Mongolia" name="Selenge" pageId="33" pageNumber="34">Selenge</collectingRegion>
River Basin (
<figureCitation id="13212A7415478117FB57FE7BFB5451E1" box="[1178,1275,476,500]" captionStart="FIGURES 176180" captionStartId="74.[151,263,983,1006]" captionTargetBox="[151,1434,193,962]" captionTargetId="figure-75@74.[151,1436,193,962]" captionTargetPageId="74" captionText="FIGURES 176180. Nemoura cinerea.176: Habitus (preserved); 177: Male terminalia, dorsal; 178: Male epiproct ventral sclerites, detail; 179: Male paraprocts, ventral; 180: Predicted ENM map with documented occurrences and range estimate." pageId="33" pageNumber="34">Fig. 180</figureCitation>
) and was also recently documented in this region by Teslenko and Bazova (2009). The synonymic history of
<taxonomicName id="4C1A4D7215478117FB1CFDA6FAE85202" baseAuthorityName="Retzius" baseAuthorityYear="1783" box="[1233,1351,512,536]" class="Insecta" family="Nemouridae" genus="Nemoura" kingdom="Animalia" order="Plecoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B96EEAE315478117FB1CFDA6FAE85202" box="[1233,1351,512,536]" italics="true" pageId="33" pageNumber="34">N. cinerea</emphasis>
</taxonomicName>
is very long and primarily deals with European species, much of which is not relevant to
<collectingCountry id="F30D766115478117FBF3FD83FB015229" box="[1086,1198,548,572]" name="Mongolia" pageId="33" pageNumber="34">Mongolia</collectingCountry>
. For a full history of this species, please refer to DeWalt et al. (2010).
</paragraph>
</subSubSection>
</treatment>
</document>