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<document id="46CBB33D15688150E7BC4A002B3453CA" ID-CLB-Dataset="25919" ID-DOI="10.11646/zootaxa.4660.1.1" ID-GBIF-Dataset="2334efd2-e346-48ff-85ff-0bc8661538b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3377134" ID-ZooBank="DA70C158-7FB0-4B46-9A1D-EA32D24B19B6" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566808464076" checkinUser="plazi" docAuthor="Moreno-González, Jairo A., O, Ranulfo González &amp; D, Eduardo Flórez" docDate="2019" docId="03B387E632591033FF3EF940DA9D3C87" docLanguage="en" docName="zootaxa.4660.1.1.imd" docOrigin="Zootaxa 4660 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Tityus (Archaeotityus) Lourenco 2006" docType="treatment" docVersion="12" lastPageNumber="8" masterDocId="FF8AFF9E325C1034FFA9FFD5D8493D28" masterDocTitle="Taxonomic revision of the Colombian Tityus (Archaeotityus) (Scorpiones, Buthidae) species: a morphological and morphometric approach, with a description of a new species" masterLastPageNumber="94" masterPageNumber="1" pageNumber="6" updateTime="1698752490567" updateUser="ExternalLinkService">
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<mods:title id="1856F7507F28CBA4B5BBB63887CFD70F">Taxonomic revision of the Colombian Tityus (Archaeotityus) (Scorpiones, Buthidae) species: a morphological and morphometric approach, with a description of a new species</mods:title>
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<mods:namePart id="40552C88644DC3A846C6B9461828F528">Moreno-González, Jairo A.</mods:namePart>
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<mods:namePart id="FBBFA156D08AF35AAA34F6E51002ED32">O, Ranulfo González</mods:namePart>
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<mods:namePart id="6D6E173F0D939619F46C8B72AAA54C66">D, Eduardo Flórez</mods:namePart>
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<treatment id="03B387E632591033FF3EF940DA9D3C87" ID-DOI="http://doi.org/10.5281/zenodo.4648574" ID-GBIF-Taxon="159243652" ID-Zenodo-Dep="4648574" LSID="urn:lsid:plazi:treatment:03B387E632591033FF3EF940DA9D3C87" httpUri="http://treatment.plazi.org/id/03B387E632591033FF3EF940DA9D3C87" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="C300657B32591031FF3EF940DAC13B87" box="[151,648,1685,1712]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BA536F032591031FF3EF940DAC13B87" blockId="5.[151,648,1685,1712]" box="[151,648,1685,1712]" pageId="5" pageNumber="6">
<heading id="D0ED819C32591031FF3EF940DAC13B87" bold="true" box="[151,648,1685,1712]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="B96EEAE232591031FF3EF940DAC13B87" bold="true" box="[151,648,1685,1712]" pageId="5" pageNumber="6">
Subgenus
<taxonomicName id="4C1A4D7332591031FEBEF940DAC13B87" authority="Lourenco, 2006" authorityName="Lourenco" authorityYear="2006" box="[279,648,1685,1712]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE232591031FEBEF940D9F63B98" bold="true" box="[279,447,1685,1712]" italics="true" pageId="5" pageNumber="6">Archaeotityus</emphasis>
<bibRefCitation id="EF8B4B0132591031FE6FF940DAC13B87" author="Lourenco, W. R. &amp; Jesus-Junior, M. M. B. &amp; Limeira-De-Oliveira, F." box="[454,648,1685,1711]" pageId="5" pageNumber="6" pagination="117 - 120" refId="ref50272" refString="Lourenco, W. R., Jesus-Junior, M. M. B. &amp; Limeira-De-Oliveira, F. (2006) A new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae) from the state of Maranh &quot; o in Brazil. Boletin Sociedad Entomologica Aragonesa, 38, 117 - 120." type="journal article" year="2006">Lourenço, 2006</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<paragraph id="8BA536F032591031FF3EF90CDB413BD9" blockId="5.[151,776,1752,1777]" box="[151,776,1752,1777]" pageId="5" pageNumber="6">
<taxonomicName id="4C1A4D7332591031FF3EF90CDA5A3BD9" authority="Lourenco 2006: 56 - 58" authorityName="Lourenco" authorityYear="2006" box="[151,531,1752,1777]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE232591031FF3EF90CD96A3BD9" box="[151,291,1753,1777]" italics="true" pageId="5" pageNumber="6">Archaeotityus</emphasis>
<bibRefCitation id="EF8B4B0132591031FE80F90FDA5A3BD9" author="Lourenco, W. R. &amp; Jesus-Junior, M. M. B. &amp; Limeira-De-Oliveira, F." box="[297,531,1752,1777]" pageId="5" pageNumber="6" pagination="117 - 120" refId="ref50272" refString="Lourenco, W. R., Jesus-Junior, M. M. B. &amp; Limeira-De-Oliveira, F. (2006) A new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae) from the state of Maranh &quot; o in Brazil. Boletin Sociedad Entomologica Aragonesa, 38, 117 - 120." type="journal article" year="2006">Lourenço 2006: 56–58</bibRefCitation>
</taxonomicName>
, 60, figures 1, 2, 16, 17.
</paragraph>
</subSubSection>
<subSubSection id="C300657B32591033FF3EF8C2DA9D3C87" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BA536F032591031FF3EF8C2DBD93A07" blockId="5.[151,912,1814,1839]" box="[151,912,1814,1839]" pageId="5" pageNumber="6">
<emphasis id="B96EEAE232591031FF3EF8C2D96A3A07" bold="true" box="[151,291,1814,1839]" pageId="5" pageNumber="6">
<typeStatus id="54A1885232591031FF3EF8C2D8853A07" box="[151,204,1815,1839]" pageId="5" pageNumber="6">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C1A4D7332591031FE83F8C2DA1B3A07" authority="Koch, 1844" authorityName="C. L. Koch" authorityYear="1844" box="[298,594,1814,1839]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="clathratus">
<emphasis id="B96EEAE232591031FE83F8C2D99D3A06" box="[298,468,1814,1838]" italics="true" pageId="5" pageNumber="6">Tityus clathratus</emphasis>
Koch, 1844
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, designated by
<bibRefCitation id="EF8B4B0132591031FD46F8C2DBC53A07" author="Lourenco, W. R. &amp; Jesus-Junior, M. M. B. &amp; Limeira-De-Oliveira, F." box="[751,908,1814,1839]" pageId="5" pageNumber="6" pagination="117 - 120" refId="ref50272" refString="Lourenco, W. R., Jesus-Junior, M. M. B. &amp; Limeira-De-Oliveira, F. (2006) A new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae) from the state of Maranh &quot; o in Brazil. Boletin Sociedad Entomologica Aragonesa, 38, 117 - 120." type="journal article" year="2006">Lourenço 2006</bibRefCitation>
.
</paragraph>
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<emphasis id="B96EEAE232591031FF3EF889D9DF3A5F" bold="true" box="[151,406,1884,1911]" pageId="5" pageNumber="6">Diagnosis (emended).</emphasis>
The members of
<taxonomicName id="4C1A4D7332591031FDF4F88BDB1D3A5F" authority="" authorityName="Lourenco" authorityYear="2006" box="[605,852,1885,1911]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE232591031FDF4F88BDAE93A5F" box="[605,672,1886,1911]" italics="true" pageId="5" pageNumber="6">Tityus</emphasis>
(
<emphasis id="B96EEAE232591031FD19F888DB033A5F" box="[688,842,1885,1911]" italics="true" pageId="5" pageNumber="6">Archaeotityus</emphasis>
)
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can be easily recognized by the following character state combinations: (1) total length between
<quantity id="4CE29B1532591031FD2FF855DAB63AB3" box="[646,767,1920,1947]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="4.3" metricValueMin="2.5" pageId="5" pageNumber="6" unit="mm" value="34.0" valueMax="43.0" valueMin="25.0">25–43 mm</quantity>
(
<figureCitation id="13212A7532591031FCA5F854DB303AB3" box="[780,889,1921,1947]" captionStart="FIGURE 4" captionStartId="12.[151,250,1941,1966]" captionTargetBox="[319,1262,169,1900]" captionTargetId="figure@12.[307,1280,166,1922]" captionTargetPageId="12" captionText="FIGURE 4. Tityus (Archaeotityus) bastosi Lourenço, 1984, habitus. A, B. Female (ICN-As-740). A. Dorsal aspect. B. Ventral aspect. C, E. Male (San Martín, Meta) (ICN-As-738). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Puerto López, Meta) (ICN-As-741). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377142/files/figure.png" pageId="5" pageNumber="6">Figures 4</figureCitation>
,
<figureCitation id="13212A7532591031FC2DF854DBEB3AB3" box="[900,930,1921,1947]" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="5" pageNumber="6">21</figureCitation>
,
<figureCitation id="13212A7532591031FC07F854DB853AB3" box="[942,972,1921,1947]" captionStart="FIGURE 24" captionStartId="40.[151,250,1942,1967]" captionTargetBox="[175,1426,611,1897]" captionTargetId="figure@40.[151,1436,597,1923]" captionTargetPageId="40" captionText="FIGURE 24. Tityus (Archaeotityus) guane sp. nov., habitus. A, B. Male holotype (ICN-As-1003). A. Dorsal aspect. B. Ventral aspect. C, D. Female paratype (ICN-As-1003). C. Dorsal aspect. B. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377183/files/figure.png" pageId="5" pageNumber="6">24</figureCitation>
,
<figureCitation id="13212A7532591031FC7EF854DBBC3AB3" box="[983,1013,1921,1947]" captionStart="FIGURE 31" captionStartId="54.[151,250,1932,1957]" captionTargetBox="[172,1391,176,1901]" captionTargetId="figure@54.[162,1425,166,1912]" captionTargetPageId="54" captionText="FIGURE 31. Tityus (Archaeotityus) parvulus Kraepelin, 1914, habitus. A, B. Female (ICN-As-1004). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Chinchiná, Caldas) (ICN-As-1004). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Chinchiná, Caldas) (ICN-As-1004). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377199/files/figure.png" pageId="5" pageNumber="6">31</figureCitation>
,
<figureCitation id="13212A7532591031FBA8F854DC563AB3" box="[1025,1055,1921,1947]" captionStart="FIGURE 41" captionStartId="68.[151,250,1937,1962]" captionTargetBox="[344,1241,189,1898]" captionTargetId="figure@68.[338,1249,166,1917]" captionTargetPageId="68" captionText="FIGURE 41. Tityus (Archaeotityus) tayrona Lourenço, 1991, habitus. A, B. Female (ICN-As-181). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Zambrano, Bolívar) (ICN-As-181). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Zambrano, Bolívar). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377219/files/figure.png" pageId="5" pageNumber="6">41</figureCitation>
,
<tableCitation id="C698034B32591031FB83F854DCC03AB3" box="[1066,1161,1921,1947]" captionStart="TABLE 4" captionStartId="45.[151,244,495,522]" captionTargetBox="[160,1951,534,1667]" captionText="TABLE 4. Measurements (mm) of Tityus guane sp. nov." httpUri="http://table.plazi.org/id/DF65667832711019FF3EFE3ADB5A3F22" pageId="5" pageNumber="6" tableUuid="DF65667832711019FF3EFE3ADB5A3F22">Tables 4</tableCitation>
,
<tableCitation id="C698034B32591031FB3CF854DCEC3AB3" box="[1173,1189,1921,1947]" captionStart="TABLE 9" captionStartId="85.[151,239,488,514]" captionTargetBox="[155,2014,561,1666]" captionText="TABLE 9. Measurements (mm) of Tityus (Archaeotityus) bastosi Lourenço, 1991, and Tityus (Archaeotityus) columbianus (Thorell, 1876). Abbreviations: F, females; M, males; SD, standard deviation." httpUri="http://table.plazi.org/id/DF65667832091061FF3EFE3DD9D93F0E" pageId="5" pageNumber="6" tableUuid="DF65667832091061FF3EFE3DD9D93F0E">9</tableCitation>
,
<tableCitation id="C698034B32591031FB19F854DC863AB3" box="[1200,1231,1921,1947]" captionStart="TABLE 10" captionStartId="87.[151,239,507,533]" captionTargetBox="[155,2019,580,1647]" captionText="TABLE 10. Measurements (mm) of Tityus (Archaeotityus) parvulus Kraepelin, 1914, and Tityus (Archaeotityus) tayrona. Abbreviations: F, females; M, males; SD, standard deviation." httpUri="http://table.plazi.org/id/DF656678320B1063FF3EFE2ED8A13F11" pageId="5" pageNumber="6" tableUuid="DF656678320B1063FF3EFE2ED8A13F11">10</tableCitation>
); (2) general body coloration with a dark yellow background slightly to densely covered with variegated brown or reddish-brown spots (
<figureCitation id="13212A7532591031FF36F81CD9473ACB" box="[159,270,1993,2019]" captionStart="FIGURE 4" captionStartId="12.[151,250,1941,1966]" captionTargetBox="[319,1262,169,1900]" captionTargetId="figure@12.[307,1280,166,1922]" captionTargetPageId="12" captionText="FIGURE 4. Tityus (Archaeotityus) bastosi Lourenço, 1984, habitus. A, B. Female (ICN-As-740). A. Dorsal aspect. B. Ventral aspect. C, E. Male (San Martín, Meta) (ICN-As-738). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Puerto López, Meta) (ICN-As-741). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377142/files/figure.png" pageId="5" pageNumber="6">Figures 4</figureCitation>
,
<figureCitation id="13212A7532591031FEB5F81CD9733ACB" box="[284,314,1993,2019]" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="5" pageNumber="6">21</figureCitation>
,
<figureCitation id="13212A7532591031FEE0F81CD92E3ACB" box="[329,359,1993,2019]" captionStart="FIGURE 24" captionStartId="40.[151,250,1942,1967]" captionTargetBox="[175,1426,611,1897]" captionTargetId="figure@40.[151,1436,597,1923]" captionTargetPageId="40" captionText="FIGURE 24. Tityus (Archaeotityus) guane sp. nov., habitus. A, B. Male holotype (ICN-As-1003). A. Dorsal aspect. B. Ventral aspect. C, D. Female paratype (ICN-As-1003). C. Dorsal aspect. B. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377183/files/figure.png" pageId="5" pageNumber="6">24</figureCitation>
,
<figureCitation id="13212A7532591031FEDCF81CD9DA3ACB" box="[373,403,1993,2019]" captionStart="FIGURE 31" captionStartId="54.[151,250,1932,1957]" captionTargetBox="[172,1391,176,1901]" captionTargetId="figure@54.[162,1425,166,1912]" captionTargetPageId="54" captionText="FIGURE 31. Tityus (Archaeotityus) parvulus Kraepelin, 1914, habitus. A, B. Female (ICN-As-1004). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Chinchiná, Caldas) (ICN-As-1004). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Chinchiná, Caldas) (ICN-As-1004). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377199/files/figure.png" pageId="5" pageNumber="6">31</figureCitation>
,
<figureCitation id="13212A7532591031FE08F81CD9893ACB" box="[417,448,1993,2019]" captionStart="FIGURE 41" captionStartId="68.[151,250,1937,1962]" captionTargetBox="[344,1241,189,1898]" captionTargetId="figure@68.[338,1249,166,1917]" captionTargetPageId="68" captionText="FIGURE 41. Tityus (Archaeotityus) tayrona Lourenço, 1991, habitus. A, B. Female (ICN-As-181). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Zambrano, Bolívar) (ICN-As-181). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Zambrano, Bolívar). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377219/files/figure.png" pageId="5" pageNumber="6">41</figureCitation>
); (3) metasomal segments I–IV ventral carinae parallel (
<figureCitation id="13212A7532591031FBE3F81CDC8A3ACB" box="[1098,1219,1993,2019]" captionStart="FIGURE 1" captionStartId="9.[151,250,1939,1964]" captionTargetBox="[243,1344,166,1920]" captionTargetId="figure@9.[243,1344,166,1920]" captionTargetPageId="9" captionText="FIGURE 1. Tityus (Archaeotityus) Lourenço, 2006, metasomal segments I–V showing distribution of macrosetae.A. Segments I–V, ventral aspect. B. Segment V, lateral aspect. Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian." httpUri="https://zenodo.org/record/3377136/files/figure.png" pageId="5" pageNumber="6">Figure 1A</figureCitation>
); (4) pectines with 11–20 teeth (
<tableCitation id="C698034B325A1032FE8FFF4CD9323D9B" box="[294,379,153,179]" captionStart="TABLE 2" captionStartId="8.[151,239,478,502]" captionText="TABLE 2. Pectinal tooth counts of Colombian Tityus (Archaeotityus) species. Note: Both pectines of each examined specimen were counted. Abbreviations: F, female; M, male; Juv, juvenile; SD, standard deviation." pageId="6" pageNumber="7">Table 2</tableCitation>
); (5) females with pectinal basal middle lamellae not dilated; (6) dorsal margins of the movable fingers with 11–15 oblique rows of denticles (
<tableCitation id="C698034B325A1032FD3DFF68DAA03DFF" box="[660,745,189,215]" captionStart="TABLE 3" captionStartId="28.[151,239,730,754]" captionText="TABLE 3. Number of movable finger oblique rows of denticles among Colombian Tityus (Archaeotityus) species. Note: Both movable fingers of each examined specimen were included. Abbreviations: F, female; M, male; Juv, juvenile; SD, standard deviation." pageId="6" pageNumber="7">Table 3</tableCitation>
); and (7)
<taxonomicName id="4C1A4D73325A1032FCFCFF68DBD13DFF" box="[853,920,189,215]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="6" pageNumber="84" phylum="Arthropoda" rank="genus">telson</taxonomicName>
with a medium-sized to large-sized pyramidal subaculear tubercle (
<figureCitation id="13212A75325A1032FED6FF34D9B03DD3" box="[383,505,225,251]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="6" pageNumber="7">Figures 19</figureCitation>
,
<figureCitation id="13212A75325A1032FDACFF34DA6C3DD3" box="[517,549,225,251]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="6" pageNumber="7">20</figureCitation>
).
</paragraph>
<paragraph id="8BA536F0325A1032FF6EFED1DD333CDF" blockId="6.[151,1437,152,503]" pageId="6" pageNumber="7">
<emphasis id="B96EEAE2325A1032FF6EFED1D9243C37" bold="true" box="[199,365,260,287]" pageId="6" pageNumber="7">Comparisons.</emphasis>
<taxonomicName id="4C1A4D73325A1032FEDCFED3DA243C37" authority="" authorityName="Lourenco" authorityYear="2006" box="[373,621,261,287]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE2325A1032FEDCFED3D9F13C37" box="[373,440,262,287]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FE63FED0DA2D3C37" box="[458,612,261,287]" italics="true" pageId="6" pageNumber="7">Archaeotityus</emphasis>
)
</taxonomicName>
is more similar to
<taxonomicName id="4C1A4D73325A1032FCEEFED3DC663C37" authority="" authorityName="Armas &amp; Marcano Fondeur" authorityYear="1992" box="[839,1071,261,287]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Caribetityus">
<emphasis id="B96EEAE2325A1032FCEEFED3DBC33C37" box="[839,906,262,287]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FC35FED0DC6F3C37" box="[924,1062,261,287]" italics="true" pageId="6" pageNumber="7">Caribetityus</emphasis>
)
</taxonomicName>
and
<taxonomicName id="4C1A4D73325A1032FBC1FED3DD1E3C37" authority="" authorityName="Lourenco" authorityYear="2006" box="[1128,1367,261,287]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Brazilotityus">
<emphasis id="B96EEAE2325A1032FBC1FED3DCE23C37" box="[1128,1195,262,287]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FB14FED0DD053C37" box="[1213,1356,261,287]" italics="true" pageId="6" pageNumber="7">Brazilotityus</emphasis>
)
</taxonomicName>
, sharing with them a small body size (
<quantity id="4CE29B15325A1032FDA6FEFCDAC23C6B" box="[527,651,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="4.3" metricValueMin="1.8" pageId="6" pageNumber="7" unit="mm" value="30.5" valueMax="43.0" valueMin="18.0">18–43 mm</quantity>
) and a similar color pattern that is composed of a yellow background covered with brown or reddish-brown spots.
<taxonomicName id="4C1A4D73325A1032FD37FE9BDBD13C4F" authority="" authorityName="Lourenco" authorityYear="2006" box="[670,920,333,359]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE2325A1032FD37FE9BDAA83C4F" box="[670,737,334,359]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FD5DFE98DBC73C4F" box="[756,910,333,359]" italics="true" pageId="6" pageNumber="7">Archaeotityus</emphasis>
)
</taxonomicName>
differs from
<taxonomicName id="4C1A4D73325A1032FB9DFE9BDD543C4F" authority="" authorityName="Armas &amp; Marcano Fondeur" authorityYear="1992" box="[1076,1309,333,359]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Caribetityus">
<emphasis id="B96EEAE2325A1032FB9DFE9BDC3E3C4F" box="[1076,1143,334,359]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FB23FE98DD5D3C4F" box="[1162,1300,333,359]" italics="true" pageId="6" pageNumber="7">Caribetityus</emphasis>
)
</taxonomicName>
and
<taxonomicName id="4C1A4D73325A1032FAF0FE9BD9703CA3" authority="" authorityName="Lourenco" authorityYear="2006" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Brazilotityus">
<emphasis id="B96EEAE2325A1032FAF0FE9BDDD53C4F" box="[1369,1436,334,359]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FF08FEA4D9793CA3" box="[161,304,369,395]" italics="true" pageId="6" pageNumber="7">Brazilotityus</emphasis>
)
</taxonomicName>
by having a medium-sized to large-sized pyramidal subaculear tubercle, while,
<taxonomicName id="4C1A4D73325A1032FB18FEA7DDD53CA3" authority="" authorityName="Lourenco" authorityYear="2006" box="[1201,1436,369,395]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Brazilotityus">
<emphasis id="B96EEAE2325A1032FB18FEA7DCBD3CA3" box="[1201,1268,370,395]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FAADFEA4DDDA3CA3" box="[1284,1427,369,395]" italics="true" pageId="6" pageNumber="7">Brazilotityus</emphasis>
)
</taxonomicName>
has a well-developed acute tubercle, and
<taxonomicName id="4C1A4D73325A1032FDC0FE43DB193C87" authority="" authorityName="Armas &amp; Marcano Fondeur" authorityYear="1992" box="[617,848,405,431]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Caribetityus">
<emphasis id="B96EEAE2325A1032FDC0FE43DAE53C87" box="[617,684,406,431]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FD14FE40DB0E3C87" box="[701,839,405,431]" italics="true" pageId="6" pageNumber="7">Caribetityus</emphasis>
)
</taxonomicName>
exhibits an acute tubercle which can be well-developed, vestigial or absent. Also,
<taxonomicName id="4C1A4D73325A1032FE5FFE6FDAA43CFB" authority="" authorityName="Lourenco" authorityYear="2006" box="[502,749,441,467]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE2325A1032FE5FFE6FDA703CFB" box="[502,569,442,467]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FDE0FE6CDAAA3CFB" box="[585,739,441,467]" italics="true" pageId="6" pageNumber="7">Archaeotityus</emphasis>
)
</taxonomicName>
can be easily distinguished from
<taxonomicName id="4C1A4D73325A1032FBCCFE6FDD193CFB" authority="" authorityName="Lourenco" authorityYear="2006" box="[1125,1360,441,467]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Brazilotityus">
<emphasis id="B96EEAE2325A1032FBCCFE6FDCE13CFB" box="[1125,1192,442,467]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FB11FE6CDD0E3CFB" box="[1208,1351,441,467]" italics="true" pageId="6" pageNumber="7">Brazilotityus</emphasis>
)
</taxonomicName>
by the presence of well-developed pectinal fulcra, while in
<taxonomicName id="4C1A4D73325A1032FD4AFE0BDB873CDF" authority="" authorityName="Lourenco" authorityYear="2006" box="[739,974,477,503]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Brazilotityus">
<emphasis id="B96EEAE2325A1032FD4AFE0BDB6F3CDF" box="[739,806,478,503]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FC9FFE08DB8C3CDF" box="[822,965,477,503]" italics="true" pageId="6" pageNumber="7">Brazilotityus</emphasis>
)
</taxonomicName>
pectinal fulcra are absent or vestigial.
</paragraph>
<caption id="DF656678325A1032FF3EFDF3DB313F17" box="[151,888,550,575]" pageId="6" pageNumber="7">
<paragraph id="8BA536F0325A1032FF3EFDF3DB313F17" blockId="6.[151,888,550,575]" box="[151,888,550,575]" pageId="6" pageNumber="7">
<emphasis id="B96EEAE2325A1032FF3EFDF3D9433F16" bold="true" box="[151,266,550,574]" pageId="6" pageNumber="7">TABLE 1.</emphasis>
Updated checklist of the subgenus
<taxonomicName id="4C1A4D73325A1032FD2CFDF2DB3A3F17" authority="" authorityName="Lourenco" authorityYear="2006" box="[645,883,551,575]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE2325A1032FD2CFDF2DA8C3F16" box="[645,709,551,574]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FD7CFDF2DB213F17" box="[725,872,551,575]" italics="true" pageId="6" pageNumber="7">Archaeotityus</emphasis>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA536F0325A1032FF0FFD9FD9723A2A" pageId="6" pageNumber="7">
<table id="F91AC450325AEFCBFF36FD9FDDCF3A2A" box="[159,1414,586,1794]" gridcols="2" gridrows="24" pageId="6" pageNumber="7">
<tr id="352A34B2325AEFCBFF36FD9FDDCF3F4B" box="[159,1414,586,611]" gridrow="0" pageId="6" pageNumber="7">
<th id="76FB5DCE325AEFCBFF36FD9FDA0C3F4B" box="[159,581,586,611]" gridcol="0" gridrow="0" pageId="6" pageNumber="7">
<emphasis id="B96EEAE2325A1032FF0FFD9FD8BC3F4B" bold="true" box="[166,245,586,611]" pageId="6" pageNumber="7">Species</emphasis>
</th>
<th id="76FB5DCE325AEFCBFDF0FD9FDDCF3F4B" box="[601,1414,586,611]" gridcol="1" gridrow="0" pageId="6" pageNumber="7">
<emphasis id="B96EEAE2325A1032FDC9FD9FDCEF3F4B" bold="true" box="[608,1190,586,611]" pageId="6" pageNumber="7">Distribution (Country: department, state or province)</emphasis>
</th>
</tr>
<tr id="352A34B2325AEFCBFF36FDA5DDCF3F85" box="[159,1414,624,685]" gridrow="1" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FDA5DA0C3F85" box="[159,581,624,685]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFDA7D9943FA1" authority="Pocock, 1897" authorityName="Pocock" authorityYear="1897" box="[166,477,624,649]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="atriventer">
<emphasis id="B96EEAE2325A1032FF0FFDA7D9053FA1" box="[166,332,626,649]" italics="true" pageId="6" pageNumber="7">Tityus atriventer</emphasis>
Pocock, 1897
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FDA5DDCF3F85" box="[601,1414,624,685]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">Grenade: Grenade Island; Saint Vincent and the Grenadines, and the Grenadines: Union Island</td>
</tr>
<tr id="352A34B2325AEFCBFF36FD60DDCF3FC6" box="[159,1414,693,750]" gridrow="2" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FD60DA0C3FC6" box="[159,581,693,750]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFD63D99E3FE6" authority="Lourenco, 1984" authorityName="Lourenco" authorityYear="1984" box="[166,471,693,718]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bastosi">
<emphasis id="B96EEAE2325A1032FF0FFD63D9793FE5" box="[166,304,693,717]" italics="true" pageId="6" pageNumber="7">Tityus bastosi</emphasis>
<bibRefCitation id="EF8B4B01325A1032FE9EFD63D99E3FE6" author="Lourenco, W. R." box="[311,471,694,718]" pageId="6" pageNumber="7" pagination="349 - 360" refId="ref49629" refString="Lourenco, W. R. (1984) Analyse taxonomique des Scorpions du groupe Tityus clathratus Koch, 1845 (Scorpiones, Buthidae). Bulletin Museum National d' Histoire Naturelle, Paris, 4 e series, 6, 349 - 360." type="journal article" year="1984">Lourenço, 1984</bibRefCitation>
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FD60DDCF3FC6" box="[601,1414,693,750]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">Brazil: Amazonas; Colombia: Boyacá, Caquetá, Casanare, Guaviare, Meta Ecuador: Morona Santiago, Napo and Sucumbíos; Peru: Loreto and San Martín</td>
</tr>
<tr id="352A34B2325AEFCBFF36FD23DDCF3E27" box="[159,1414,758,783]" gridrow="3" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FD23DA0C3E27" box="[159,581,758,783]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFD2DD9853E27" authority="Koch, 1836" authorityName="Koch" authorityYear="1836" box="[166,460,758,783]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="clathratus">
<emphasis id="B96EEAE2325A1032FF0FFD2DD9063E27" box="[166,335,759,783]" italics="true" pageId="6" pageNumber="7">Tityus clathratus</emphasis>
<bibRefCitation id="EF8B4B01325A1032FEFFFD23D9853E27" author="Koch, C. L." box="[342,460,758,783]" pageId="6" pageNumber="7" pagination="17 - 104" refId="ref49274" refString="Koch, C. L. (1836) Die Arachniden. N ʾ rnberg: C. H. Zehsche Buchhandlung, 3 (1 - 5), 17 - 104." type="journal article" year="1836">Koch, 1836</bibRefCitation>
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FD23DDCF3E27" box="[601,1414,758,783]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">Venezuela: Falcón, Carabobo, Aragua, Miranda, Distrito Capital; Guyana</td>
</tr>
<tr id="352A34B2325AEFCBFF36FCC9DDCF3E1D" box="[159,1414,796,821]" gridrow="4" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FCC9DA0C3E1D" box="[159,581,796,821]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFCC8DA423E1D" authority="(Thorell, 1876)" authorityName="Thorell" authorityYear="1876" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[166,523,796,821]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="columbianus">
<emphasis id="B96EEAE2325A1032FF0FFCC8D9203E1D" box="[166,361,797,821]" italics="true" pageId="6" pageNumber="7">Tityus columbianus</emphasis>
(
<bibRefCitation id="EF8B4B01325A1032FEDEFCC9DA4D3E1D" author="Thorell, T." box="[375,516,796,821]" pageId="6" pageNumber="7" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
)
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FCC9DDCF3E1D" box="[601,1414,796,821]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">Colombia: Cundinamarca, Boyacá, Bolívar, Nariño, Tolima</td>
</tr>
<tr id="352A34B2325AEFCBFF36FC97DDCF3E73" box="[159,1414,834,859]" gridrow="5" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FC97DA0C3E73" box="[159,581,834,859]" gridcol="0" gridrow="5" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFC96D9AC3E73" authority="Lourenco, 2012" authorityName="Lourenco" authorityYear="2012" box="[166,485,834,859]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="grahami">
<emphasis id="B96EEAE2325A1032FF0FFC96D9773E72" box="[166,318,834,858]" italics="true" pageId="6" pageNumber="7">Tityus grahami</emphasis>
Lourenço, 2012
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FC97DDCF3E73" box="[601,1414,834,859]" gridcol="1" gridrow="5" pageId="6" pageNumber="7">Brazil: Amazonas</td>
</tr>
<tr id="352A34B2325AEFCBFF36FCBDDDCF3EA9" box="[159,1414,872,897]" gridrow="6" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FCBDDA0C3EA9" box="[159,581,872,897]" gridcol="0" gridrow="6" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFCBCD96F3EA8" authority="Moreno-González &amp; O &amp; D, 2019" authorityName="Moreno-González &amp; O &amp; D" authorityYear="2019" box="[166,294,873,896]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guane" status="sp. nov.">
<emphasis id="B96EEAE2325A1032FF0FFCBCD96F3EA8" box="[166,294,873,896]" italics="true" pageId="6" pageNumber="7">Tityus guane</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D5799325A1032FE84FCBDD92F3EA8" box="[301,358,872,896]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B96EEAE2325A1032FE84FCBDD92F3EA8" bold="true" box="[301,358,872,896]" pageId="6" pageNumber="7">sp. n.</emphasis>
</taxonomicNameLabel>
</td>
<td id="76FB5DCE325AEFCBFDF0FCBDDDCF3EA9" box="[601,1414,872,897]" gridcol="1" gridrow="6" pageId="6" pageNumber="7">Colombia: Santander</td>
</tr>
<tr id="352A34B2325AEFCBFF36FC58DDCF3E8E" box="[159,1414,909,934]" gridrow="7" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FC58DA0C3E8E" box="[159,581,909,934]" gridcol="0" gridrow="7" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFC5ADA5B3E8E" authority="Mello-Leitao, 1934" authorityName="Mello-Leitao" authorityYear="1934" box="[166,530,909,934]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="indecisus">
<emphasis id="B96EEAE2325A1032FF0FFC5AD90F3E8E" box="[166,326,910,934]" italics="true" pageId="6" pageNumber="7">Tityus indecisus</emphasis>
Mello-Leitão, 1934
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FC58DDCF3E8E" box="[601,1414,909,934]" gridcol="1" gridrow="7" pageId="6" pageNumber="7">Argentina: Jujuy; Brazil: Matto Grosso</td>
</tr>
<tr id="352A34B2325AEFCBFF36FC66DDCF3EE4" box="[159,1414,947,972]" gridrow="8" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FC66DA0C3EE4" box="[159,581,947,972]" gridcol="0" gridrow="8" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFC60D9A53EE4" authority="Borelli, 1899" authorityName="Borelli" authorityYear="1899" box="[166,492,947,972]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B96EEAE2325A1032FF0FFC60D9293EE4" box="[166,352,948,972]" italics="true" pageId="6" pageNumber="7">Tityus intermedius</emphasis>
Borelli, 1899
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FC66DDCF3EE4" box="[601,1414,947,972]" gridcol="1" gridrow="8" pageId="6" pageNumber="7">Colombia:?; Ecuador: Ibarra</td>
</tr>
<tr id="352A34B2325AEFCBFF36FC0CDDCF3EDA" box="[159,1414,985,1010]" gridrow="9" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FC0CDA0C3EDA" box="[159,581,985,1010]" gridcol="0" gridrow="9" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFC0FD9AA3EDA" authority="Lourenco, 2005" authorityName="Lourenco" authorityYear="2005" box="[166,483,985,1010]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="julianae">
<emphasis id="B96EEAE2325A1032FF0FFC0FD9723EDA" box="[166,315,986,1010]" italics="true" pageId="6" pageNumber="7">Tityus julianae</emphasis>
<bibRefCitation id="EF8B4B01325A1032FEEBFC0FD9AA3EDA" author="Lourenco, W. R." box="[322,483,985,1010]" pageId="6" pageNumber="7" pagination="221 - 227" refId="ref50058" refString="Lourenco, W. R. (2005) A new species of Tityus Koch, 1836 (Scorpiones, Buthidae) from Ecuador. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 14 (171), 221 - 227." type="journal article" year="2005">Lourenço, 2005</bibRefCitation>
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FC0CDDCF3EDA" box="[601,1414,985,1010]" gridcol="1" gridrow="9" pageId="6" pageNumber="7">Ecuador: Esmeraldas</td>
</tr>
<tr id="352A34B2325AEFCBFF36FC2ADDCF3930" box="[159,1414,1023,1048]" gridrow="10" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FC2ADA0C3930" box="[159,581,1023,1048]" gridcol="0" gridrow="10" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFBD5D9933930" authority="Kovarik, 2005" authorityName="Kovarik" authorityYear="2005" box="[166,474,1023,1048]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kaderkai">
<emphasis id="B96EEAE2325A1032FF0FFBD5D908393F" box="[166,321,1023,1047]" italics="true" pageId="6" pageNumber="7">Tityus kaderkai</emphasis>
<bibRefCitation id="EF8B4B01325A1032FEEEFC2AD9933930" author="Kovarik, F." box="[327,474,1023,1048]" pageId="6" pageNumber="7" pagination="58 - 61" refId="ref49308" refString="Kovarik, F. (2005) Novy druh stira Tityus kaderkai sp. n. v terariu. Arachnologie, 58 - 61." type="book chapter" year="2005">Kovarik, 2005</bibRefCitation>
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FC2ADDCF3930" box="[601,1414,1023,1048]" gridcol="1" gridrow="10" pageId="6" pageNumber="7">Bolivia: Yucumo</td>
</tr>
<tr id="352A34B2325AEFCBFF36FBF0DDCF3916" box="[159,1414,1061,1086]" gridrow="11" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FBF0DA0C3916" box="[159,581,1061,1086]" gridcol="0" gridrow="11" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFBF3D9D13915" authority="Giltay, 1928" authorityName="Giltay" authorityYear="1928" box="[166,408,1061,1086]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lutzi">
<emphasis id="B96EEAE2325A1032FF0FFBF3D95D3915" box="[166,276,1061,1085]" italics="true" pageId="6" pageNumber="7">Tityus lutzi</emphasis>
Giltay, 1928
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FBF0DDCF3916" box="[601,1414,1061,1086]" gridcol="1" gridrow="11" pageId="6" pageNumber="7">Brazil: Cuiabá, Mato Grosso</td>
</tr>
<tr id="352A34B2325AEFCBFF36FB9FDDCF394B" box="[159,1414,1098,1123]" gridrow="12" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FB9FDA0C394B" box="[159,581,1098,1123]" gridcol="0" gridrow="12" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFB99D981394B" authority="Lourenco, 2012" authorityName="Lourenco" authorityYear="2012" box="[166,456,1099,1123]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mana">
<emphasis id="B96EEAE2325A1032FF0FFB99D969394B" box="[166,288,1100,1123]" italics="true" pageId="6" pageNumber="7">Tityus mana</emphasis>
Lourenço, 2012
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FB9FDDCF394B" box="[601,1414,1098,1123]" gridcol="1" gridrow="12" pageId="6" pageNumber="7">French Guyana: Pisti Mana- Les Hattes</td>
</tr>
<tr id="352A34B2325AEFCBFF36FBA5DDCF3985" box="[159,1414,1136,1197]" gridrow="13" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FBA5DA0C3985" box="[159,581,1136,1197]" gridcol="0" gridrow="13" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFBA7DA5A39A0" authority="Lourenco, Jesus" authorityName="Lourenco, Jesus" box="[166,531,1137,1161]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="maranhensis">
<emphasis id="B96EEAE2325A1032FF0FFBA7D92039A1" box="[166,361,1137,1161]" italics="true" pageId="6" pageNumber="7">Tityus maranhensis</emphasis>
Lourenço, Jesus
</taxonomicName>
&amp; Limeira-de-Oliveira, 2006
</td>
<td id="76FB5DCE325AEFCBFDF0FBA5DDCF3985" box="[601,1414,1136,1197]" gridcol="1" gridrow="13" pageId="6" pageNumber="7">Brazil: Maranhão</td>
</tr>
<tr id="352A34B2325AEFCBFF36FB60DDCF39E6" box="[159,1414,1205,1230]" gridrow="14" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FB60DA0C39E6" box="[159,581,1205,1230]" gridcol="0" gridrow="14" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFB63DA4439E6" authority="Borelli, 1901" authorityName="Borelli" authorityYear="1901" box="[166,525,1205,1230]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mattogrosensis">
<emphasis id="B96EEAE2325A1032FF0FFB63D9C939E5" box="[166,384,1206,1229]" italics="true" pageId="6" pageNumber="7">Tityus mattogrosensis</emphasis>
Borelli, 1901
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FB60DDCF39E6" box="[601,1414,1205,1230]" gridcol="1" gridrow="14" pageId="6" pageNumber="7">Brazil: Bahia, Goias, Matto Grosso, Minas Gerais</td>
</tr>
<tr id="352A34B2325AEFCBFF36FB0EDDCF39DC" box="[159,1414,1243,1268]" gridrow="15" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FB0EDA0C39DC" box="[159,581,1243,1268]" gridcol="0" gridrow="15" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFB09DA6539DB" authority="Lutz &amp; Mello, 1922" authorityName="Lutz &amp; Mello" authorityYear="1922" box="[166,556,1243,1268]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="microcystis">
<emphasis id="B96EEAE2325A1032FF0FFB09D91339DB" box="[166,346,1244,1267]" italics="true" pageId="6" pageNumber="7">Tityus microcystis</emphasis>
Lutz &amp; Mello, 1922
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FB0EDDCF39DC" box="[601,1414,1243,1268]" gridcol="1" gridrow="15" pageId="6" pageNumber="7">Brazil: Marianna, Minas Gerais</td>
</tr>
<tr id="352A34B2325AEFCBFF36FAD4DDCF3816" box="[159,1414,1281,1342]" gridrow="16" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FAD4DA0C3816" box="[159,581,1281,1342]" gridcol="0" gridrow="16" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFAD7D9C33832" authority="Francke" authorityName="Francke &amp; Stockwell" authorityYear="1987" box="[166,394,1281,1306]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocelote">
<emphasis id="B96EEAE2325A1032FF0FFAD7D9783832" box="[166,305,1282,1306]" italics="true" pageId="6" pageNumber="7">Tityus ocelote</emphasis>
Francke
</taxonomicName>
&amp; Stockwell, 1987
</td>
<td id="76FB5DCE325AEFCBFDF0FAD4DDCF3816" box="[601,1414,1281,1342]" gridcol="1" gridrow="16" pageId="6" pageNumber="7">Costa Rica: Heredia, Limón, Puntarenas; Panamá: Bocas del Toro</td>
</tr>
<tr id="352A34B2325AEFCBFF36FA93DDCF38A8" box="[159,1414,1350,1408]" gridrow="17" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FA93DA0C38A8" box="[159,581,1350,1408]" gridcol="0" gridrow="17" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFA92DA6F3877" authority="Kraepelin, 1895" authorityName="Kraepelin" authorityYear="1895" box="[166,550,1350,1375]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="paraguayensis">
<emphasis id="B96EEAE2325A1032FF0FFA92D9353876" box="[166,380,1351,1374]" italics="true" pageId="6" pageNumber="7">Tityus paraguayensis</emphasis>
Kraepelin, 1895
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FA93DDCF38A8" box="[601,1414,1350,1408]" gridcol="1" gridrow="17" pageId="6" pageNumber="7">Argentina: Chaco, Formosa, Santa Fe, Buenos Aires Brazil: Matto Grosso, Pará; Paraguay: Chaco</td>
</tr>
<tr id="352A34B2325AEFCBFF36FA52DDCF3888" box="[159,1414,1415,1440]" gridrow="18" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FA52DA0C3888" box="[159,581,1415,1440]" gridcol="0" gridrow="18" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFA5CD9B23888" authority="(Kraepelin, 1914)" authorityName="Kraepelin" authorityYear="1914" baseAuthorityName="Kraepelin" baseAuthorityYear="1914" box="[166,507,1415,1440]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B96EEAE2325A1032FF0FFA5CD9083888" box="[166,321,1416,1440]" italics="true" pageId="6" pageNumber="7">Tityus parvulus</emphasis>
(
<bibRefCitation id="EF8B4B01325A1032FEE7FA52D9BD3888" author="Kraepelin, K." box="[334,500,1415,1440]" pageId="6" pageNumber="7" pagination="15 - 28" refId="ref49461" refString="Kraepelin, K. (1914) Beitrag zur Kenntnis der Skorpione und Pedipalpen Columbiens. Memoires de la Societe des Sciences naturelles, Neuchatel, 5, 15 - 28." type="journal article" year="1914">Kraepelin, 1914</bibRefCitation>
)
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FA52DDCF3888" box="[601,1414,1415,1440]" gridcol="1" gridrow="18" pageId="6" pageNumber="7">Colombia: Antioquia, Caldas, Cauca, Quindío, Risaralda, Valle del Cauca</td>
</tr>
<tr id="352A34B2325AEFCBFF36FA78DDCF38EE" box="[159,1414,1453,1478]" gridrow="19" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FA78DA0C38EE" box="[159,581,1453,1478]" gridcol="0" gridrow="19" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFA7BD98138EE" authority="Pocock, 1893" authorityName="Pocock" authorityYear="1893" box="[166,456,1453,1478]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B96EEAE2325A1032FF0FFA7BD97E38EE" box="[166,311,1454,1478]" italics="true" pageId="6" pageNumber="7">Tityus pusillus</emphasis>
Pocock, 1893
</taxonomicName>
</td>
<td id="76FB5DCE325AEFCBFDF0FA78DDCF38EE" box="[601,1414,1453,1478]" gridcol="1" gridrow="19" pageId="6" pageNumber="7">Brazil: Alagoas, Bahia, Paraiba, Pernambuco, Rio Grande do Norte, Sergipe</td>
</tr>
<tr id="352A34B2325AEFCBFF36FA06DDCF3B38" box="[159,1414,1491,1552]" gridrow="20" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36FA06DDCF3B38" box="[159,1414,1491,1552]" colspan="2" colspanRight="1" gridcol="0" gridrow="20" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FFA01D9B738C4" authority="Rojas-Runjaic" authorityName="Rojas-Runjaic &amp; Armas" authorityYear="2007" box="[166,510,1491,1516]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rondonorum">
<emphasis id="B96EEAE2325A1032FF0FFA01D92F38C3" box="[166,358,1491,1515]" italics="true" pageId="6" pageNumber="7">Tityus rondonorum</emphasis>
Rojas-Runjaic
</taxonomicName>
&amp; Venezuela: Barinas Armas, 2007
</td>
</tr>
<tr id="352A34B2325AEFCBFF36F9C2DDCF3B50" box="[159,1414,1559,1656]" gridrow="21" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36F9C2DDCF3B50" box="[159,1414,1559,1656]" colspan="2" colspanRight="1" gridcol="0" gridrow="21" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF0FF9CCD99A3B18" authority="Pocock, 1897" authorityName="Pocock" authorityYear="1897" box="[166,467,1559,1584]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="silvestris">
<emphasis id="B96EEAE2325A1032FF0FF9CCD90B3B18" box="[166,322,1560,1584]" italics="true" pageId="6" pageNumber="7">Tityus silvestris</emphasis>
Pocock, 1897
</taxonomicName>
Brazil: Acre, Amapa, Amazonas, Matto Grosso, Pará French Guyana: Cayenne, Saut Pararé; Ecuador: Oreyana. Suriname. Peru: Loreto, San Martín, Madre de Dios
</td>
</tr>
<tr id="352A34B2325AEFCBFF36F955DDCF3B95" box="[159,1414,1664,1725]" gridrow="22" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36F955DDCF3B95" box="[159,1414,1664,1725]" colspan="2" colspanRight="1" gridcol="0" gridrow="22" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF36F954D99F3BB1" authority="Lourenco, 1991" authorityName="Lourenco" authorityYear="1991" baseAuthorityName="Lourenco" baseAuthorityYear="1999" box="[159,470,1665,1689]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B96EEAE2325A1032FF36F954D9663BB1" box="[159,303,1665,1689]" italics="true" pageId="6" pageNumber="7">Tityus tayrona</emphasis>
<bibRefCitation id="EF8B4B01325A1032FE9FF954D99F3BB1" author="Lourenco, W. R." box="[310,470,1665,1689]" pageId="6" pageNumber="7" pagination="275 - 288" refId="ref49674" refString="Lourenco, W. R. (1991) Les scorpions de Colombie, II. Les faunes des regions de Santa Marta et de la. Cordillere Orientale. Approche biogeographique. Senckenbergiana Biologica, 71 (4), 275 - 288." type="journal article" year="1991">Lourenço, 1991</bibRefCitation>
</taxonomicName>
Colombia: Antioquia, Atlántico, Bolívar, Caldas, Cesar, Córdoba, Magdalena, Sucre; Panamá: Panamá; Venezuela: Trujillo
</td>
</tr>
<tr id="352A34B2325AEFCBFF36F910DDCF3A2A" box="[159,1414,1733,1794]" gridrow="23" pageId="6" pageNumber="7">
<td id="76FB5DCE325AEFCBFF36F910DDCF3A2A" box="[159,1414,1733,1794]" colspan="2" colspanRight="1" gridcol="0" gridrow="23" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D73325A1032FF36F913DA623BF5" authority="Kovarik, Teruel, Lowe" authorityName="Kovarik, Teruel, Lowe" box="[159,555,1733,1758]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="wachtelii">
<emphasis id="B96EEAE2325A1032FF36F913D9773BF5" box="[159,318,1733,1757]" italics="true" pageId="6" pageNumber="7">Tityus wachtelii</emphasis>
Kovarik, Teruel, Lowe
</taxonomicName>
&amp; Peru: Panguana, Huanuco Friedrich, 2015
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA536F0325A1033FF6EF8F8DD7D3C4F" blockId="6.[151,1437,1837,2044]" lastBlockId="7.[151,1436,153,431]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B96EEAE2325A1032FF6EF8F8DAB13A60" bold="true" box="[199,760,1837,1864]" pageId="6" pageNumber="7">
Hemispermatophores of
<taxonomicName id="4C1A4D73325A1032FE59F8F8DAB83A60" authority="" authorityName="Lourenco" authorityYear="2006" box="[496,753,1837,1864]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE2325A1032FE59F8F8DA7E3A6F" bold="true" box="[496,567,1837,1863]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FDE1F8F8DAA13A60" bold="true" box="[584,744,1837,1864]" italics="true" pageId="6" pageNumber="7">Archaeotityus</emphasis>
)
</taxonomicName>
.
</emphasis>
The male hemispermatophore of
<taxonomicName id="4C1A4D73325A1032FBDEF8FBDD263A60" authority="" authorityName="Lourenco" authorityYear="2006" box="[1143,1391,1837,1864]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE2325A1032FBDEF8FBDCF33A6F" box="[1143,1210,1838,1863]" italics="true" pageId="6" pageNumber="7">Tityus</emphasis>
(
<emphasis id="B96EEAE2325A1032FB62F8F8DD2C3A6F" box="[1227,1381,1837,1863]" italics="true" pageId="6" pageNumber="7">Archaeotityus</emphasis>
)
</taxonomicName>
has only been described for three species:
<taxonomicName id="4C1A4D73325A1032FDF0F887DB523A43" authorityName="Kraepelin" authorityYear="1895" box="[601,795,1874,1899]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="paraguayensis">
<emphasis id="B96EEAE2325A1032FDF0F887DB523A43" box="[601,795,1874,1899]" italics="true" pageId="6" pageNumber="7">T. paraguayensis</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8B4B01325A1032FC84F884DC283A44" author="Ojanguren-Affilastro, A. A." box="[813,1121,1873,1900]" pageId="6" pageNumber="7" pagination="75 - 247" refId="ref51025" refString="Ojanguren-Affilastro, A. A. (2005) Estudio monografico de los escorpiones de la Republica Argentina. Revista Iberica de Aracnologia, 11, 75 - 247." type="journal article" year="2005">Ojanguren-Affilastro 2005</bibRefCitation>
: figures 43, 44);
<taxonomicName id="4C1A4D73325A1032FA85F887DDD53A43" authorityName="Francke &amp; Stockwell" authorityYear="1987" box="[1324,1436,1873,1899]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ocelote">
<emphasis id="B96EEAE2325A1032FA85F887DDD53A43" box="[1324,1436,1873,1899]" italics="true" pageId="6" pageNumber="7">T. ocelote</emphasis>
</taxonomicName>
(Francke &amp; Stockwell 1991: 24, figure 104); and
<taxonomicName id="4C1A4D73325A1032FD6FF8A3DB003AA7" box="[710,841,1909,1935]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="wachtelii">
<emphasis id="B96EEAE2325A1032FD6FF8A3DB003AA7" box="[710,841,1909,1935]" italics="true" pageId="6" pageNumber="7">T. wachtelii</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8B4B01325A1032FCFEF8A0DC2B3AB8" author="Kovarik, F. &amp; Teruel, R. &amp; Lowe, G. &amp; Friedrich, S." box="[855,1122,1909,1936]" pageId="6" pageNumber="7" pagination="1 - 40" refId="ref49398" refString="Kovarik, F., Teruel, R., Lowe, G. &amp; Friedrich, S. (2015) Four new scorpion species (Scorpiones: Buthidae) from Amazonian Peru. Euscorpius, 210, 1 - 40. https: // doi. org / 10.18590 / euscorpius. 2015. vol 2015. iss 210. 1" type="journal article" year="2015">
Kovarik
<emphasis id="B96EEAE2325A1032FC14F8A3DBBC3AA7" box="[957,1013,1909,1935]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2015: 34
</bibRefCitation>
, figures 142–145). Here we describe the hemispermatophore for
<taxonomicName id="4C1A4D73325A1032FD9EF84FDAEB3A9B" authorityName="Lourenco" authorityYear="1984" box="[567,674,1945,1971]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bastosi">
<emphasis id="B96EEAE2325A1032FD9EF84FDAEB3A9B" box="[567,674,1945,1971]" italics="true" pageId="6" pageNumber="7">T. bastosi</emphasis>
</taxonomicName>
(
<figureCitation id="13212A75325A1032FD1BF84CDB6D3A9C" box="[690,804,1945,1972]" captionStart="FIGURE 13" captionStartId="23.[151,250,1901,1926]" captionTargetBox="[169,1435,193,1881]" captionTargetId="figure@23.[151,1435,166,1881]" captionTargetPageId="23" captionText="FIGURE 13. Tityus (Archaeotityus) bastosi Lourenço, 1984, dextral hemispermatophore, showing capsule details, male (ICN- As-741). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (San Martín, Meta) (ICN-As-738). E. Male (Villavicencio, Meta) (MZUSP 71740). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377161/files/figure.png" pageId="6" pageNumber="7">Figure 13</figureCitation>
),
<taxonomicName id="4C1A4D73325A1032FC93F84FDBAF3A9B" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[826,998,1945,1971]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="columbianus">
<emphasis id="B96EEAE2325A1032FC93F84FDBAF3A9B" box="[826,998,1945,1971]" italics="true" pageId="6" pageNumber="7">T. columbianus</emphasis>
</taxonomicName>
(
<figureCitation id="13212A75325A1032FC5FF84CDC213A9C" box="[1014,1128,1945,1972]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="6" pageNumber="7">Figure 23</figureCitation>
),
<taxonomicName id="4C1A4D73325A1032FBD4F84FDC973A9B" authority="Moreno-González &amp; O &amp; D, 2019" authorityName="Moreno-González &amp; O &amp; D" authorityYear="2019" box="[1149,1246,1946,1971]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="guane" status="sp. nov.">
<emphasis id="B96EEAE2325A1032FBD4F84FDC973A9B" box="[1149,1246,1946,1971]" italics="true" pageId="6" pageNumber="7">T. guane</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D5799325A1032FB4EF84FDD0B3A9C" box="[1255,1346,1946,1972]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B96EEAE2325A1032FB4EF84FDD0B3A9C" bold="true" box="[1255,1346,1946,1972]" pageId="6" pageNumber="7">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13212A75325A1032FAF8F84CD8FF3AF0" captionStart="FIGURE 27" captionStartId="43.[151,250,1071,1096]" captionTargetBox="[172,1419,194,1045]" captionTargetId="figure@43.[151,1436,181,1052]" captionTargetPageId="43" captionText="FIGURE 27. Tityus (Archaeotityus) guane sp. nov., dextral hemispermatophore, showing capsule details, male (Los Santos, Santander) (MZSP 71736). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Los Santos, Santander) (MZSP 71736). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377189/files/figure.png" pageId="6" pageNumber="7">Figure 27</figureCitation>
),
<taxonomicName id="4C1A4D73325A1032FF62F86BD9013AFF" baseAuthorityName="Kraepelin" baseAuthorityYear="1914" box="[203,328,1981,2007]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="B96EEAE2325A1032FF62F86BD9013AFF" box="[203,328,1981,2007]" italics="true" pageId="6" pageNumber="7">T. parvulus</emphasis>
</taxonomicName>
(
<figureCitation id="13212A75325A1032FEFEF868D9813AF0" box="[343,456,1981,2008]" captionStart="FIGURE 36" captionStartId="59.[151,250,1335,1360]" captionTargetBox="[151,1436,181,1310]" captionTargetId="figure@59.[151,1436,181,1310]" captionTargetPageId="59" captionText="FIGURE 36. Tityus (Archaeotityus) parvulus Kraepelin, 1914, male pedipalp chela. A–C. Male (Chinchiná, Caldas) (ICN-As- 1004). A. External aspect. B. Dorsal aspect. C. Ventral aspect. Scale bar= 2.0 mm." httpUri="https://zenodo.org/record/3377209/files/figure.png" pageId="6" pageNumber="7">Figure 36</figureCitation>
), and
<taxonomicName id="4C1A4D73325A1032FDA5F86BDA373AFF" authorityName="Lourenco" authorityYear="1991" baseAuthorityName="Lourenco" baseAuthorityYear="1999" box="[524,638,1982,2007]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B96EEAE2325A1032FDA5F86BDA373AFF" box="[524,638,1982,2007]" italics="true" pageId="6" pageNumber="7">T. tayrona</emphasis>
</taxonomicName>
(
<figureCitation id="13212A75325A1032FD25F868DAB73AF0" box="[652,766,1981,2008]" captionStart="FIGURE 43" captionStartId="74.[151,250,1941,1966]" captionTargetBox="[274,1313,166,1921]" captionTargetId="figure@74.[274,1313,166,1921]" captionTargetPageId="74" captionText="FIGURE 43. Tityus (Archaeotityus) tayrona Lourenço, 1991, male pedipalp chela. A–C. Male (Zambrano, Bolívar) (ICN-As- 181). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Male (Zambrano, Bolívar) (ICN-As-181). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377223/files/figure.png" pageId="6" pageNumber="7">Figure 43</figureCitation>
). All mentioned species share the following character states related to hemispermatophore morphology: (1) hemispermatophore thin and sclerotized; (2) stalk flagelliform, folded once over itself (
<figureCitation id="13212A75325B1033FED8FF4CD9A23D9B" box="[369,491,153,179]" captionStart="FIGURE 13" captionStartId="23.[151,250,1901,1926]" captionTargetBox="[169,1435,193,1881]" captionTargetId="figure@23.[151,1435,166,1881]" captionTargetPageId="23" captionText="FIGURE 13. Tityus (Archaeotityus) bastosi Lourenço, 1984, dextral hemispermatophore, showing capsule details, male (ICN- As-741). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (San Martín, Meta) (ICN-As-738). E. Male (Villavicencio, Meta) (MZUSP 71740). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377161/files/figure.png" pageId="7" pageNumber="8">Figures 13</figureCitation>
,
<figureCitation id="13212A75325B1033FE5FFF4CDA5D3D9B" box="[502,532,153,179]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="7" pageNumber="8">23</figureCitation>
,
<figureCitation id="13212A75325B1033FD89FF4CDA773D9B" box="[544,574,153,179]" captionStart="FIGURE 27" captionStartId="43.[151,250,1071,1096]" captionTargetBox="[172,1419,194,1045]" captionTargetId="figure@43.[151,1436,181,1052]" captionTargetPageId="43" captionText="FIGURE 27. Tityus (Archaeotityus) guane sp. nov., dextral hemispermatophore, showing capsule details, male (Los Santos, Santander) (MZSP 71736). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Los Santos, Santander) (MZSP 71736). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377189/files/figure.png" pageId="7" pageNumber="8">27</figureCitation>
,
<figureCitation id="13212A75325B1033FDE0FF4CDA213D9B" box="[585,616,153,179]" captionStart="FIGURE 37" captionStartId="61.[151,250,1012,1037]" captionTargetBox="[211,1376,292,984]" captionTargetId="figure@61.[202,1386,270,985]" captionTargetPageId="61" captionText="FIGURE 37. Tityus (Archaeotityus) parvulus Kraepelin, 1914, dextral hemispermatophore, showing capsule details, male (Villamaría, Caldas) (MZSP 71738). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Chinchiná, Caldas) (ICN-As-1004). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377211/files/figure.png" pageId="7" pageNumber="8">37</figureCitation>
); (3) pedicel narrow and flat (
<figureCitation id="13212A75325B1033FC1BFF4CDC653D9B" box="[946,1068,153,179]" captionStart="FIGURE 13" captionStartId="23.[151,250,1901,1926]" captionTargetBox="[169,1435,193,1881]" captionTargetId="figure@23.[151,1435,166,1881]" captionTargetPageId="23" captionText="FIGURE 13. Tityus (Archaeotityus) bastosi Lourenço, 1984, dextral hemispermatophore, showing capsule details, male (ICN- As-741). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (San Martín, Meta) (ICN-As-738). E. Male (Villavicencio, Meta) (MZUSP 71740). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377161/files/figure.png" pageId="7" pageNumber="8">Figures 13</figureCitation>
,
<figureCitation id="13212A75325B1033FB91FF4CDC1F3D9B" box="[1080,1110,153,179]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="7" pageNumber="8">23</figureCitation>
,
<figureCitation id="13212A75325B1033FBC8FF4CDC363D9B" box="[1121,1151,153,179]" captionStart="FIGURE 27" captionStartId="43.[151,250,1071,1096]" captionTargetBox="[172,1419,194,1045]" captionTargetId="figure@43.[151,1436,181,1052]" captionTargetPageId="43" captionText="FIGURE 27. Tityus (Archaeotityus) guane sp. nov., dextral hemispermatophore, showing capsule details, male (Los Santos, Santander) (MZSP 71736). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Los Santos, Santander) (MZSP 71736). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377189/files/figure.png" pageId="7" pageNumber="8">27</figureCitation>
,
<figureCitation id="13212A75325B1033FB23FF4CDCE33D9C" box="[1162,1194,153,180]" captionStart="FIGURE 36" captionStartId="59.[151,250,1335,1360]" captionTargetBox="[151,1436,181,1310]" captionTargetId="figure@59.[151,1436,181,1310]" captionTargetPageId="59" captionText="FIGURE 36. Tityus (Archaeotityus) parvulus Kraepelin, 1914, male pedipalp chela. A–C. Male (Chinchiná, Caldas) (ICN-As- 1004). A. External aspect. B. Dorsal aspect. C. Ventral aspect. Scale bar= 2.0 mm." httpUri="https://zenodo.org/record/3377209/files/figure.png" pageId="7" pageNumber="8">36</figureCitation>
); (4) pedal flexure inconspicuous (
<figureCitation id="13212A75325B1033FE86FF68D9E03DFF" box="[303,425,189,215]" captionStart="FIGURE 13" captionStartId="23.[151,250,1901,1926]" captionTargetBox="[169,1435,193,1881]" captionTargetId="figure@23.[151,1435,166,1881]" captionTargetPageId="23" captionText="FIGURE 13. Tityus (Archaeotityus) bastosi Lourenço, 1984, dextral hemispermatophore, showing capsule details, male (ICN- As-741). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (San Martín, Meta) (ICN-As-738). E. Male (Villavicencio, Meta) (MZUSP 71740). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377161/files/figure.png" pageId="7" pageNumber="8">Figures 13</figureCitation>
,
<figureCitation id="13212A75325B1033FE1DFF68D99B3DFF" box="[436,466,189,215]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="7" pageNumber="8">23</figureCitation>
,
<figureCitation id="13212A75325B1033FE74FF68D9B23DFF" box="[477,507,189,215]" captionStart="FIGURE 27" captionStartId="43.[151,250,1071,1096]" captionTargetBox="[172,1419,194,1045]" captionTargetId="figure@43.[151,1436,181,1052]" captionTargetPageId="43" captionText="FIGURE 27. Tityus (Archaeotityus) guane sp. nov., dextral hemispermatophore, showing capsule details, male (Los Santos, Santander) (MZSP 71736). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Los Santos, Santander) (MZSP 71736). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377189/files/figure.png" pageId="7" pageNumber="8">27</figureCitation>
,
<figureCitation id="13212A75325B1033FDAFFF68DA6D3DFF" box="[518,548,189,215]" captionStart="FIGURE 37" captionStartId="61.[151,250,1012,1037]" captionTargetBox="[211,1376,292,984]" captionTargetId="figure@61.[202,1386,270,985]" captionTargetPageId="61" captionText="FIGURE 37. Tityus (Archaeotityus) parvulus Kraepelin, 1914, dextral hemispermatophore, showing capsule details, male (Villamaría, Caldas) (MZSP 71738). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Chinchiná, Caldas) (ICN-As-1004). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377211/files/figure.png" pageId="7" pageNumber="8">37</figureCitation>
,
<figureCitation id="13212A75325B1033FD86FF68DA063DFF" box="[559,591,189,215]" captionStart="FIGURE 44" captionStartId="75.[151,250,1554,1579]" captionTargetBox="[164,1423,230,1497]" captionTargetId="figure@75.[151,1436,181,1497]" captionTargetPageId="75" captionText="FIGURE 44. Tityus (Archaeotityus) tayrona Lourenço, 1991, dextral hemispermatophore, showing capsule details (Zambrano, Bolívar) (ICN-As-181). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (La Dorada, Caldas) (ICN-As-1006). E. Male (Valledupar, César) (MZUSP 71735). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377225/files/figure.png" pageId="7" pageNumber="8">44</figureCitation>
); (5) stem occupying 2/3 of the hemispermatophore total length (
<figureCitation id="13212A75325B1033FAB4FF68DDDE3DFF" box="[1309,1431,189,215]" captionStart="FIGURE 13" captionStartId="23.[151,250,1901,1926]" captionTargetBox="[169,1435,193,1881]" captionTargetId="figure@23.[151,1435,166,1881]" captionTargetPageId="23" captionText="FIGURE 13. Tityus (Archaeotityus) bastosi Lourenço, 1984, dextral hemispermatophore, showing capsule details, male (ICN- As-741). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (San Martín, Meta) (ICN-As-738). E. Male (Villavicencio, Meta) (MZUSP 71740). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377161/files/figure.png" pageId="7" pageNumber="8">Figures 13</figureCitation>
,
<figureCitation id="13212A75325B1033FF3EFF34D8FC3DD3" box="[151,181,225,251]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="7" pageNumber="8">23</figureCitation>
,
<figureCitation id="13212A75325B1033FF68FF34D8963DD3" box="[193,223,225,251]" captionStart="FIGURE 27" captionStartId="43.[151,250,1071,1096]" captionTargetBox="[172,1419,194,1045]" captionTargetId="figure@43.[151,1436,181,1052]" captionTargetPageId="43" captionText="FIGURE 27. Tityus (Archaeotityus) guane sp. nov., dextral hemispermatophore, showing capsule details, male (Los Santos, Santander) (MZSP 71736). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Los Santos, Santander) (MZSP 71736). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377189/files/figure.png" pageId="7" pageNumber="8">27</figureCitation>
,
<figureCitation id="13212A75325B1033FF42FF34D9403DD3" box="[235,265,225,251]" captionStart="FIGURE 37" captionStartId="61.[151,250,1012,1037]" captionTargetBox="[211,1376,292,984]" captionTargetId="figure@61.[202,1386,270,985]" captionTargetPageId="61" captionText="FIGURE 37. Tityus (Archaeotityus) parvulus Kraepelin, 1914, dextral hemispermatophore, showing capsule details, male (Villamaría, Caldas) (MZSP 71738). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Chinchiná, Caldas) (ICN-As-1004). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377211/files/figure.png" pageId="7" pageNumber="8">37</figureCitation>
,
<figureCitation id="13212A75325B1033FEBCFF34D97C3DD3" box="[277,309,225,251]" captionStart="FIGURE 44" captionStartId="75.[151,250,1554,1579]" captionTargetBox="[164,1423,230,1497]" captionTargetId="figure@75.[151,1436,181,1497]" captionTargetPageId="75" captionText="FIGURE 44. Tityus (Archaeotityus) tayrona Lourenço, 1991, dextral hemispermatophore, showing capsule details (Zambrano, Bolívar) (ICN-As-181). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (La Dorada, Caldas) (ICN-As-1006). E. Male (Valledupar, César) (MZUSP 71735). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377225/files/figure.png" pageId="7" pageNumber="8">44</figureCitation>
); and (6) basal hook curved towards the posterior region (
<figureCitation id="13212A75325B1033FC15FF34DC033DD3" box="[956,1098,225,251]" captionStart="FIGURE 13" captionStartId="23.[151,250,1901,1926]" captionTargetBox="[169,1435,193,1881]" captionTargetId="figure@23.[151,1435,166,1881]" captionTargetPageId="23" captionText="FIGURE 13. Tityus (Archaeotityus) bastosi Lourenço, 1984, dextral hemispermatophore, showing capsule details, male (ICN- As-741). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (San Martín, Meta) (ICN-As-738). E. Male (Villavicencio, Meta) (MZUSP 71740). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377161/files/figure.png" pageId="7" pageNumber="8">Figures 13B</figureCitation>
,
<figureCitation id="13212A75325B1033FBFCFF34DCCF3DD3" box="[1109,1158,225,251]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="7" pageNumber="8">23B</figureCitation>
,
<figureCitation id="13212A75325B1033FB3BFF34DC8A3DD3" box="[1170,1219,225,251]" captionStart="FIGURE 27" captionStartId="43.[151,250,1071,1096]" captionTargetBox="[172,1419,194,1045]" captionTargetId="figure@43.[151,1436,181,1052]" captionTargetPageId="43" captionText="FIGURE 27. Tityus (Archaeotityus) guane sp. nov., dextral hemispermatophore, showing capsule details, male (Los Santos, Santander) (MZSP 71736). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Los Santos, Santander) (MZSP 71736). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377189/files/figure.png" pageId="7" pageNumber="8">27B</figureCitation>
,
<figureCitation id="13212A75325B1033FB66FF34DD493DD3" box="[1231,1280,225,251]" captionStart="FIGURE 37" captionStartId="61.[151,250,1012,1037]" captionTargetBox="[211,1376,292,984]" captionTargetId="figure@61.[202,1386,270,985]" captionTargetPageId="61" captionText="FIGURE 37. Tityus (Archaeotityus) parvulus Kraepelin, 1914, dextral hemispermatophore, showing capsule details, male (Villamaría, Caldas) (MZSP 71738). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Chinchiná, Caldas) (ICN-As-1004). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377211/files/figure.png" pageId="7" pageNumber="8">37B</figureCitation>
,
<figureCitation id="13212A75325B1033FAA2FF34DD773DD3" box="[1291,1342,225,251]" captionStart="FIGURE 44" captionStartId="75.[151,250,1554,1579]" captionTargetBox="[164,1423,230,1497]" captionTargetId="figure@75.[151,1436,181,1497]" captionTargetPageId="75" captionText="FIGURE 44. Tityus (Archaeotityus) tayrona Lourenço, 1991, dextral hemispermatophore, showing capsule details (Zambrano, Bolívar) (ICN-As-181). A. Posterior view. B. Lateral view. C. Anterior view. D, E. Capsule variations, anterior view. D. Male (La Dorada, Caldas) (ICN-As-1006). E. Male (Valledupar, César) (MZUSP 71735). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377225/files/figure.png" pageId="7" pageNumber="8">44B</figureCitation>
). While, these character states are homogeneous, interspecific variations in the capsule region and intraspecific variations in the shape of the basal hook can occur (see variability). Determination of the diagnostic value of hemispermatophore morphology at the specific level within
<taxonomicName id="4C1A4D73325B1033FDFFFE9BDB053C4F" authority="" authorityName="Lourenco" authorityYear="2006" box="[598,844,333,359]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="B96EEAE2325B1033FDFFFE9BDAD03C4F" box="[598,665,334,359]" italics="true" pageId="7" pageNumber="8">Tityus</emphasis>
(
<emphasis id="B96EEAE2325B1033FD00FE98DB0A3C4F" box="[681,835,333,359]" italics="true" pageId="7" pageNumber="8">Archaeotityus</emphasis>
)
</taxonomicName>
requires examination of additional species.
</paragraph>
<paragraph id="8BA536F0325B1033FF6EFEA4DAC93CA3" blockId="7.[151,1436,153,431]" box="[199,640,368,395]" pageId="7" pageNumber="8">
<emphasis id="B96EEAE2325B1033FF6EFEA4D9DB3CA3" bold="true" box="[199,402,368,395]" pageId="7" pageNumber="8">Included species.</emphasis>
23 species (
<tableCitation id="C698034B325B1033FDB2FEA4DA273CA3" box="[539,622,369,395]" captionStart="TABLE 1" captionStartId="6.[151,239,550,574]" captionText="TABLE 1. Updated checklist of the subgenus Tityus (Archaeotityus)." pageId="7" pageNumber="8">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BA536F0325B1033FF6EFE41DA9D3C87" blockId="7.[151,1436,153,431]" box="[199,724,404,431]" pageId="7" pageNumber="8">
<emphasis id="B96EEAE2325B1033FF6EFE41D9283C87" bold="true" box="[199,353,404,431]" pageId="7" pageNumber="8">Distribution.</emphasis>
<collectingCountry id="F30D7660325B1033FECEFE40D9A83C87" box="[359,481,405,431]" name="Costa Rica" pageId="7" pageNumber="8">Costa Rica</collectingCountry>
to Central
<collectingCountry id="F30D7660325B1033FDF4FE40DA993C87" box="[605,720,405,431]" name="Argentina" pageId="7" pageNumber="8">Argentina</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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