167 lines
17 KiB
XML
167 lines
17 KiB
XML
<document id="0CC50E03B60253D7B0A9F42CCD6859C8" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFB71F27FF50FDCBFA558DF7" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax rigida Beccari 1907" docType="treatment" docVersion="5" lastPageNumber="73" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="73" updateTime="1696265644834" updateUser="diego">
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<mods:mods id="DF5CF66B5BDB1D2E728B68AF9BDCD2AD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="22E27C6FFD9A7B7961CAD9337CDE2A20">
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<mods:title id="1FF324D45A5A6D0164D6BF824DA268B9">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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</mods:titleInfo>
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<mods:name id="2B3866A8C17CB65522F9E8A68F4B7A2E" type="personal">
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<mods:role id="B20FC9EB6B34F2DD9B5B2F3B303B6A4E">
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<mods:roleTerm id="EA729062158397A055AF66E09B4D1D85">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F437059872518892823914FB7A278982">Henderson, Andrew</mods:namePart>
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<mods:typeOfResource id="22F963651E90CB219C2627483753C0C0">text</mods:typeOfResource>
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<mods:titleInfo id="171A5F44CEE8DF0D28D5FAA393271A2A">
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<mods:title id="FE3219B638BEE39B07823041985A30B7">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="197C1AD1CB187F452D87875BCD2DBDF6">
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<mods:date id="E45169F654A5969B8737A253D4949997">2023</mods:date>
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<mods:detail id="68EAD0577C3E4300770114DB5857C72B" type="pubDate">
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<mods:number id="8DEC271D95DAE851846E0C65392B8FDE">2023-09-19</mods:number>
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</mods:detail>
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<mods:detail id="D476AD9C94D06DD1824212E307D3C62C" type="volume">
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<mods:number id="3949CF3260AAC9392E01FA99F50D2E7B">614</mods:number>
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</mods:detail>
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<mods:detail id="E4E5609F8CD34515984965E2955EA5E9" type="issue">
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<mods:number id="37ADC4EC56E72DC25AC00A22ACFAC6CC">1</mods:number>
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</mods:detail>
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<mods:extent id="887B9A24DA82DAC2BD86C158B75D78AB" unit="page">
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<mods:start id="9D2D86869CAE5C33737EBA0105570FA2">1</mods:start>
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<mods:end id="7E147EF7EC746EA081C699391B9C5D8F">115</mods:end>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="C65467C3191FB9763C352F5273225D6A">
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<mods:url id="FC9341032A2BC6DEFDC77353DBC9EFA1">http://dx.doi.org/10.11646/phytotaxa.614.1.1</mods:url>
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</mods:location>
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<mods:classification id="D0992843980A95BD6939D60F892E7477">journal article</mods:classification>
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<mods:identifier id="53A55920801C21138B52A1BE409EEF96" type="DOI">10.11646/phytotaxa.614.1.1</mods:identifier>
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<mods:identifier id="62AFFF16F194BA152D8407603682C673" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="82D71AA83C2A9C9C04F4AF44E8C3192E" type="Zenodo-Dep">8389307</mods:identifier>
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</mods:mods>
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<treatment id="03B387DAFFB71F27FF50FDCBFA558DF7" LSID="urn:lsid:plazi:treatment:03B387DAFFB71F27FF50FDCBFA558DF7" httpUri="http://treatment.plazi.org/id/03B387DAFFB71F27FF50FDCBFA558DF7" lastPageNumber="73" pageId="72" pageNumber="73">
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<subSubSection id="C3006547FFB71F27FF50FDCBFC2E8A97" box="[136,983,626,653]" pageId="72" pageNumber="73" type="nomenclature">
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<paragraph id="8BA536CCFFB71F27FF50FDCBFC2E8A97" blockId="72.[136,1452,626,761]" box="[136,983,626,653]" pageId="72" pageNumber="73">
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<emphasis id="B96EEADEFFB71F27FF50FDCBFC2E8A97" bold="true" box="[136,983,626,653]" pageId="72" pageNumber="73">
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1.31.
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<taxonomicName id="4C1A4D4FFFB71F27FF1FFDCBFC288A97" ID-CoL="5ZFW6" authority="(Grisebach & Wendland) Beccari (1907: 299)" authorityName="Beccari" authorityPageNumber="299" authorityYear="1907" baseAuthorityName="Grisebach & Wendland" box="[199,977,626,653]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B96EEADEFFB71F27FF1FFDCBFE568A96" bold="true" box="[199,431,626,653]" italics="true" pageId="72" pageNumber="73">Coccothrinax rigida</emphasis>
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(Grisebach & Wendland)
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<bibRefCitation id="EF8B4B3DFFB71F27FD32FDCBFC288A97" author="Beccari, O." box="[746,977,626,652]" pageId="72" pageNumber="73" pagination="1 - 343" refId="ref60874" refString="Beccari, O. (1907) Le palme americane della tribu delle Corypheae. Webbia 2: 1 - 343. https: // doi. org / 10.5962 / bhl. title. 15531" type="journal article" year="1907">Beccari (1907: 299)</bibRefCitation>
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</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFB71F27FC06FDCBFE728AAB" pageId="72" pageNumber="73" type="reference_group">
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<paragraph id="8BA536CCFFB71F27FC06FDCBFE728AAB" blockId="72.[136,1452,626,761]" pageId="72" pageNumber="73">
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<taxonomicName id="4C1A4D4FFFB71F27FC06FDCBFA768A97" authority="Grisebach & Wendland" authorityName="Grisebach & Wendland" box="[990,1423,626,652]" class="Liliopsida" family="Arecaceae" genus="Thrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B96EEADEFFB71F27FC06FDCBFB7B8A97" box="[990,1154,626,652]" italics="true" pageId="72" pageNumber="73">Thrinax rigida</emphasis>
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Grisebach & Wendland
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</taxonomicName>
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in
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<bibRefCitation id="EF8B4B3DFFB71F27FF50FD2FFE718AAB" author="Grisebach, A." box="[136,392,662,688]" pageId="72" pageNumber="73" refId="ref61997" refString="Grisebach, A. (1866) Catalogus plantarum cubensium. Apud Guilielmum Engelmann, Lipsiae, 301 pp." type="book" year="1866">Grisebach (1866: 221)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFB71F27FE4AFD2EFD168AE3" pageId="72" pageNumber="73" type="materials_examined">
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<paragraph id="8BA536CCFFB71F27FE4AFD2EFD168AE3" blockId="72.[136,1452,626,761]" pageId="72" pageNumber="73">
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<typeStatus id="54A1886EFFB71F27FE4AFD2EFDFB8AAA" box="[402,514,663,689]" pageId="72" pageNumber="73">Lectotype</typeStatus>
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(designated by
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<bibRefCitation id="EF8B4B3DFFB71F27FD6CFD2EFCCE8AAB" author="Moya, C." box="[692,823,662,689]" pageId="72" pageNumber="73" pagination="1 - 63" refId="ref63249" refString="Moya, C. (2020) An annotated checklist of Cuban palms 2. Coccothrinax, Pt. 1: 1816 - 1939. Nomenclature, typification, and distribution. PalmArbor 4: 1 - 63." type="journal article" year="2020">Moya 2020</bibRefCitation>
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):—
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<collectingCountry id="F30D765CFFB71F27FCB9FD2FFC4A8AAB" box="[865,947,662,688]" name="Cuba" pageId="72" pageNumber="73">CUBA</collectingCountry>
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. La Catalina near Sagua de Tánamo, no date,
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<emphasis id="B96EEADEFFB71F27FF50FD02FECB8ACF" box="[136,306,698,725]" italics="true" pageId="72" pageNumber="73">C. Wright 3220</emphasis>
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(
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<typeStatus id="54A1886EFFB71F27FEE7FD03FE518ACF" box="[319,424,698,724]" pageId="72" pageNumber="73">lectotype</typeStatus>
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G
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<emphasis id="B96EEADEFFB71F27FE1FFD02FE178ACF" box="[455,494,699,724]" italics="true" pageId="72" pageNumber="73">n.v.</emphasis>
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, G image!, isolectotypes BH!, FI!, GH!, GOET
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<emphasis id="B96EEADEFFB71F27FBDAFD02FBD68ACF" box="[1026,1071,699,724]" italics="true" pageId="72" pageNumber="73">
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<taxonomicNameLabel id="A25D57A5FFB71F27FBDAFD02FBD38ACF" box="[1026,1066,699,724]" pageId="72" pageNumber="73" rank="variety">n.v.</taxonomicNameLabel>
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,
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</emphasis>
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GOET image!, HAC
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<emphasis id="B96EEADEFFB71F27FAF9FD02FABE8ACF" box="[1313,1351,699,724]" italics="true" pageId="72" pageNumber="73">n.v.</emphasis>
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, K!, MO
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<emphasis id="B96EEADEFFB71F27FF50FD66FF568AE3" box="[136,175,735,760]" italics="true" pageId="72" pageNumber="73">n.v.</emphasis>
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, MO image!, P
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<emphasis id="B96EEADEFFB71F27FEBAFD66FE718AE3" box="[354,392,735,760]" italics="true" pageId="72" pageNumber="73">n.v.</emphasis>
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, P image!, YU
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<emphasis id="B96EEADEFFB71F27FDEEFD66FDA58AE3" box="[566,604,735,760]" italics="true" pageId="72" pageNumber="73">n.v.</emphasis>
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, YU image!)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFB71F27FF50FC9FFD1F8D0F" pageId="72" pageNumber="73" type="description">
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<paragraph id="8BA536CCFFB71F27FF50FC9FFD1F8D0F" blockId="72.[136,1452,806,1517]" pageId="72" pageNumber="73">
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<emphasis id="B96EEADEFFB71F27FF50FC9FFF298B5B" bold="true" box="[136,208,806,832]" pageId="72" pageNumber="73">Stems</emphasis>
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length and diameter not recorded, branching not recorded.
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<emphasis id="B96EEADEFFB71F27FC5DFC9FFC2E8B5B" bold="true" box="[901,983,806,832]" pageId="72" pageNumber="73">Leaves</emphasis>
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not recorded; leaf sheath fibers
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<quantity id="4CE29B29FFB71F27FA8BFC9FFA558B5A" box="[1363,1452,806,833]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="72" pageNumber="73" unit="mm" value="0.7">0.7 mm</quantity>
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diameter, stout, woody, loosely woven, ± joined or briefly free at the apices; petioles 4.0 mm diameter just below the apex; palmans
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<quantity id="4CE29B29FFB71F27FE87FCD7FE548B92" box="[351,429,878,905]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="72" pageNumber="73" unit="cm" value="2.5">2.5 cm</quantity>
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long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades not wedge-shaped; segments number not recorded, the middle ones
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<quantity id="4CE29B29FFB71F27FA97FC2BFA528BB6" box="[1359,1451,914,941]" metricMagnitude="-1" metricUnit="m" metricValue="2.45" pageId="72" pageNumber="73" unit="cm" value="24.5">24.5 cm</quantity>
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long and
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<quantity id="4CE29B29FFB71F27FF2AFC0FFEC68BCA" box="[242,319,950,977]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="72" pageNumber="73" unit="cm" value="1.7">1.7 cm</quantity>
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wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively short and broad, abruptly narrowed (shoulder) toward the apex, otherwise parallel-sided, often strongly folded, stiff and leathery, the apices briefly splitting; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially, without indumentum, scales, or wax abaxially, without transverse veinlets.
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<emphasis id="B96EEADEFFB71F27FEB0FBFFFDE88C7B" bold="true" box="[360,529,1094,1120]" pageId="72" pageNumber="73">Inflorescences</emphasis>
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curving, arching, or pendulous amongst the leaves, with few to numerous partial inflorescences; rachis bracts somewhat flattened, loosely sheathing, usually tomentose with a dense tuft of erect hairs at the apex; partial inflorescences not recorded; proximalmost rachillae straight,
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<quantity id="4CE29B29FFB71F27FBC9FB37FBA78CB2" box="[1041,1118,1166,1193]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="72" pageNumber="73" unit="cm" value="5.1">5.1 cm</quantity>
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long and
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<quantity id="4CE29B29FFB71F27FB17FB37FADC8CB2" box="[1231,1317,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="72" pageNumber="73" unit="mm" value="1.7">1.7 mm</quantity>
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diameter in fruit; rachillae glabrous at or near anthesis; stamens 7; fruit pedicels
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<quantity id="4CE29B29FFB71F27FC43FB0BFC0A8CD6" box="[923,1011,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="72" pageNumber="73" unit="mm" value="1.9">1.9 mm</quantity>
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long;
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<emphasis id="B96EEADEFFB71F27FBE6FB0BFB868CD7" bold="true" box="[1086,1151,1202,1228]" pageId="72" pageNumber="73">fruits</emphasis>
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6.0 mm long and
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<quantity id="4CE29B29FFB71F27FA8CFB0BFA558CD6" box="[1364,1452,1202,1229]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="72" pageNumber="73" unit="mm" value="6.8">6.8 mm</quantity>
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diameter, color not recorded; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFB71F27FF65FAA7FE2F8D47" pageId="72" pageNumber="73" type="distribution">
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<paragraph id="8BA536CCFFB71F27FF65FAA7FE2F8D47" blockId="72.[136,1452,806,1517]" pageId="72" pageNumber="73">
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<emphasis id="B96EEADEFFB71F27FF65FAA7FDFA8D23" bold="true" box="[189,515,1310,1336]" pageId="72" pageNumber="73">Distribution and habitat:—</emphasis>
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<collectingCountry id="F30D765CFFB71F27FDDBFAA7FDC78D23" box="[515,574,1310,1336]" name="Cuba" pageId="72" pageNumber="73">Cuba</collectingCountry>
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(
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<collectingRegion id="49DEF82EFFB71F27FD94FAA7FD528D23" box="[588,683,1310,1336]" country="Cuba" name="Holguin" pageId="72" pageNumber="73">Holguín</collectingRegion>
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) (
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<figureCitation id="13212A49FFB71F27FD19FAA7FCEB8D23" box="[705,786,1310,1336]" captionStart="FIGURE 16" captionStartId="71.[136,229,1506,1528]" captionTargetBox="[155,1429,198,1477]" captionTargetId="figure-15@71.[150,1434,193,1482]" captionTargetPageId="71" captionText="FIGURE 16. Distribution of Coccothrinax pseudorigida, C. pumila, C. rigida, and C. salvatoris." figureDoi="http://doi.org/10.5281/zenodo.8389405" httpUri="https://zenodo.org/record/8389405/files/figure.png" pageId="72" pageNumber="73">Fig. 16</figureCitation>
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), habitat and elevation not recorded.
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<bibRefCitation id="EF8B4B3DFFB71F27FB74FAA7FACF8D23" author="Craft, P." box="[1196,1334,1310,1336]" pageId="72" pageNumber="73" refId="ref61560" refString="Craft, P. (2017) The Palms of Cuba. Palm Nut Pages, Florida, 232 pp." type="book" year="2017">Craft (2017)</bibRefCitation>
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wrote that it occurred on limestone hills.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006547FFB71F27FF65FADFFA558DF7" pageId="72" pageNumber="73" type="discussion">
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<paragraph id="8BA536CCFFB71F27FF65FADFFA558DF7" blockId="72.[136,1452,806,1517]" pageId="72" pageNumber="73">
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<emphasis id="B96EEADEFFB71F27FF65FADFFE498D9B" bold="true" box="[189,432,1382,1408]" pageId="72" pageNumber="73">Taxonomic notes:—</emphasis>
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As a preliminary species,
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<taxonomicName id="4C1A4D4FFFB71F27FD3DFADFFC348D9B" box="[741,973,1382,1408]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B96EEADEFFB71F27FD3DFADFFC348D9B" box="[741,973,1382,1408]" italics="true" pageId="72" pageNumber="73">Coccothrinax rigida</emphasis>
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</taxonomicName>
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has a unique combination of qualitative character states and is recognized as a phylogenetic species. It is notable for its leaf segments without indumentum, scales, or wax abaxially; otherwise it is similar to
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<taxonomicName id="4C1A4D4FFFB71F27FD44FA16FCC28DD3" authorityName="Beccari" authorityYear="1907" baseAuthorityName="Kunth" box="[668,827,1455,1481]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="miraguama">
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<emphasis id="B96EEADEFFB71F27FD44FA16FCC28DD3" box="[668,827,1455,1481]" italics="true" pageId="72" pageNumber="73">C. miraguama</emphasis>
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</taxonomicName>
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.
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<bibRefCitation id="EF8B4B3DFFB71F27FC9EFA17FC378DD3" author="Leon" box="[838,974,1454,1480]" pageId="72" pageNumber="73" pagination="107 - 156" refId="ref62742" refString="Leon (1939) Contribucion al estudio de las palmas de Cuba. III. Memorias de la Sociedad Cubana de Historia Natural " Felipe Poey " 13: 107 - 156." type="journal article" year="1939">
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<collectingRegion id="49DEF82EFFB71F27FC9EFA17FC798DD3" box="[838,896,1454,1480]" country="Nicaragua" name="Leon" pageId="72" pageNumber="73">León</collectingRegion>
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(1939)
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</bibRefCitation>
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placed
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<taxonomicName id="4C1A4D4FFFB71F27FBC7FA16FB798DD3" box="[1055,1152,1454,1481]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="rigida">
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<emphasis id="B96EEADEFFB71F27FBC7FA16FB798DD3" box="[1055,1152,1454,1481]" italics="true" pageId="72" pageNumber="73">C. rigida</emphasis>
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</taxonomicName>
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in his series
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<taxonomicName id="4C1A4D4FFFB71F27FADEFA17FE798DF7" genus="Longispadicae" pageId="72" pageNumber="73" rank="section" section="Pauciramosae">
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<emphasis id="B96EEADEFFB71F27FADEFA17FA528DD3" box="[1286,1451,1454,1480]" italics="true" pageId="72" pageNumber="73">Longispadicae</emphasis>
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section
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<emphasis id="B96EEADEFFB71F27FF06FA6AFE798DF7" box="[222,384,1491,1516]" italics="true" pageId="72" pageNumber="73">Pauciramosae</emphasis>
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</taxonomicName>
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. However it does not appear to have elongate inflorescences, as do other species in the section.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |