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<document id="BD2F51586C3F50A3CB50E941729293E3" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFAC1F3CFF50FF27FF158E43" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax spissa Bailey 1939" docType="treatment" docVersion="2" lastPageNumber="84" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="84" updateTime="1695980191673" updateUser="ExternalLinkService">
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<mods:title id="6D1FB2B6B92CF58FDCA3E422E08F8350">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<subSubSection id="C3006547FFAC1F3CFF50FF27FACA88C7" pageId="83" pageNumber="84" type="nomenclature">
<paragraph id="8BA536CCFFAC1F3CFF50FF27FACA88C7" blockId="83.[136,1451,158,221]" pageId="83" pageNumber="84">
<emphasis id="B96EEADEFFAC1F3CFF50FF27FD5488A3" bold="true" box="[136,685,158,185]" pageId="83" pageNumber="84">
1.37.
<taxonomicName id="4C1A4D4FFFAC1F3CFF11FF27FD5E88A3" ID-CoL="WLZM" ID-ENA="480904" authority="Bailey (1939 b: 253)" authorityName="Bailey" authorityPageNumber="253" authorityYear="1939" box="[201,679,158,185]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFAC1F3CFF11FF27FE4A88A3" bold="true" box="[201,435,158,185]" italics="true" pageId="83" pageNumber="84">Coccothrinax spissa</emphasis>
<bibRefCitation id="EF8B4B3DFFAC1F3CFE65FF27FD5E88A3" author="Bailey, L." box="[445,679,158,184]" pageId="83" pageNumber="84" pagination="247 - 259" refId="ref60742" refString="Bailey, L. (1939 b) Coccothrinax in the southern Greater Antilles. Gentes Herbarum 4: 247 - 259." type="journal article" year="1939">Bailey (1939b: 253)</bibRefCitation>
</taxonomicName>
.
</emphasis>
<materialsCitation id="3B723C91FFAC1F3CFD6FFF26FACA88C7" collectingDate="1939-07-27" collectionCode="L, BH" country="Dominican Republic" location="Bani" municipality="Hispaniola" pageId="83" pageNumber="84" specimenCount="2" stateProvince="Santo Domingo" typeStatus="lectotype">
<typeStatus id="54A1886EFFAC1F3CFD6FFF26FCDE88A2" box="[695,807,159,185]" pageId="83" pageNumber="84">Lectotype</typeStatus>
(designated by
<bibRefCitation id="EF8B4B3DFFAC1F3CFC07FF27FB2288A3" author="Reveal, J. &amp; Nixon, K." box="[991,1243,158,184]" pageId="83" pageNumber="84" pagination="1 - 46" refId="ref64245" refString="Reveal, J. &amp; Nixon, K. (2013) The palm type collection in the Liberty Hyde Bailey Hortorium (BH). Phytoneuron 79: 1 - 46." type="journal article" year="2013">Reveal &amp; Nixon 2013</bibRefCitation>
):—
<collectingCountry id="F30D765CFFAC1F3CFADAFF27FEEA88C7" name="Dominican Republic" pageId="83" pageNumber="84">DOMINICAN REPUBLIC</collectingCountry>
.
<collectingMunicipality id="6BC1ACB6FFAC1F3CFEC5FF7BFE6088C7" box="[285,409,194,220]" pageId="83" pageNumber="84">Hispaniola</collectingMunicipality>
,
<collectingRegion id="49DEF82EFFAC1F3CFE7DFF7BFDA088C7" box="[421,601,194,220]" country="Dominican Republic" name="Santo Domingo" pageId="83" pageNumber="84">Santo Domingo</collectingRegion>
, near
<location id="8EC56017FFAC1F3CFD42FF7BFD2B88C7" LSID="urn:lsid:plazi:treatment:03B387DAFFAC1F3CFF50FF27FF158E43:8EC56017FFAC1F3CFD42FF7BFD2B88C7" box="[666,722,194,220]" country="Dominican Republic" municipality="Hispaniola" name="Bani" pageId="83" pageNumber="84" stateProvince="Santo Domingo">Bani</location>
,
<date id="FFA4100CFFAC1F3CFD05FF7BFC8888C7" box="[733,881,194,220]" pageId="83" pageNumber="84" value="1939-07-27">
<collectingDate id="EFE0E9E4FFAC1F3CFD05FF7BFC8888C7" box="[733,881,194,220]" pageId="83" pageNumber="84" value="1939-07-27">27 July 1939</collectingDate>
</date>
,
<emphasis id="B96EEADEFFAC1F3CFCA4FF7AFBEB88C7" box="[892,1042,194,220]" italics="true" pageId="83" pageNumber="84">
<collectionCode id="ED0BAE09FFAC1F3CFCA4FF7AFC7688C7" box="[892,911,195,220]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="83" pageNumber="84" type="Herbarium">L</collectionCode>
. Bailey 271
</emphasis>
(
<typeStatus id="54A1886EFFAC1F3CFBF9FF7BFB7388C7" box="[1057,1162,194,220]" pageId="83" pageNumber="84">lectotype</typeStatus>
<collectionCode id="ED0BAE09FFAC1F3CFB48FF7BFB4688C7" box="[1168,1215,194,220]" country="USA" lsid="urn:lsid:biocol.org:col:15518" name="L. H. Bailey Hortorium, Cornell University" pageId="83" pageNumber="84" type="Herbarium">BH</collectionCode>
!).
<figureCitation id="13212A49FFAC1F3CFB00FF7BFACA88C7" box="[1240,1331,194,220]" captionStart="PLATE 26" captionStartId="84.[136,214,1917,1938]" captionTargetBox="[151,1433,194,1891]" captionTargetId="figure-27@84.[150,1434,193,1893]" captionTargetPageId="84" captionText="PLATE 26. Coccothrinax spissa, Dominican Republic, with swollen stems. (Image by Eladio Fernández)." figureDoi="http://doi.org/10.5281/zenodo.8389419" httpUri="https://zenodo.org/record/8389419/files/figure.png" pageId="83" pageNumber="84">Plate 26</figureCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFAC1F3CFF50FEB3FC1F8B5B" pageId="83" pageNumber="84" type="description">
<paragraph id="8BA536CCFFAC1F3CFF50FEB3FC1F8B5B" blockId="83.[136,1452,266,1624]" pageId="83" pageNumber="84">
<emphasis id="B96EEADEFFAC1F3CFF50FEB3FF29893F" bold="true" box="[136,208,266,292]" pageId="83" pageNumber="84">Stems</emphasis>
4.9(2.56.5) m long and 24.2(11.050.0) cm diameter, solitary, often swollen.
<emphasis id="B96EEADEFFAC1F3CFB95FEB3FB66893F" bold="true" box="[1101,1183,266,292]" pageId="83" pageNumber="84">Leaves</emphasis>
more or less deciduous or only leaf bases persisting on stem; leaf sheath fibers 0.4(0.10.7) mm diameter, thin, closely woven, not forming persistent ligules and soon disintegrating at the apices; petioles 16.6(11.926.4) mm diameter just below the apex; palmans 22.4(14.031.5) cm long, relatively long, without prominent adaxial veins; leaf blades not wedge-shaped; segments 60(4771) per leaf, the middle ones 74.5(51.599.5) cm long and 3.3(2.54.4) cm wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively long and narrow, tapering from base to apex, scarcely folded, flexible and not leathery, a shoulder or constriction absent or poorly developed, the apices thin, deeply splitting and breaking off; middle leaf segment apices attenuate; leaf segments not waxy or sometimes with a deciduous, thin layer of wax adaxially, densely indumentose abaxially, with irregularly shaped, semi-persistent, interlocking, fimbriate hairs without an obvious center, with well-developed transverse veinlets.
<emphasis id="B96EEADEFFAC1F3CFF50FDCBFEC88A97" bold="true" box="[136,305,626,652]" pageId="83" pageNumber="84">Inflorescences</emphasis>
curving, arching, or pendulous amongst the leaves, with few to numerous partial inflorescences; rachis bracts swollen, woody, not or sparsely tomentose; partial inflorescences 5(36); proximalmost rachillae straight, 13.6(7.522.0) cm long and 1.6(0.92.4) mm diameter in fruit; rachillae glabrous at or near anthesis; stamens 10(9 13); fruit pedicels 1.3(0.52.7) mm long;
<emphasis id="B96EEADEFFAC1F3CFDBDFD67FD5F8AE3" bold="true" box="[613,678,734,760]" pageId="83" pageNumber="84">fruits</emphasis>
8.6(6.710.1) mm long and 8.6(6.59.9) mm diameter, purple, red-purple, white to purple, magenta, magenta to blackish; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFAC1F3CFF65FCF3FF158E43" pageId="83" pageNumber="84" type="distribution">
<paragraph id="8BA536CCFFAC1F3CFF65FCF3FD068B93" blockId="83.[136,1452,266,1624]" pageId="83" pageNumber="84">
<emphasis id="B96EEADEFFAC1F3CFF65FCF3FDF78B7F" bold="true" box="[189,526,842,868]" pageId="83" pageNumber="84">Distribution and habitat:—</emphasis>
<collectingCountry id="F30D765CFFAC1F3CFDD7FCF3FDBE8B7F" box="[527,583,842,868]" name="Haiti" pageId="83" pageNumber="84">Haiti</collectingCountry>
and
<collectingCountry id="F30D765CFFAC1F3CFD5EFCF3FC8A8B7F" box="[646,883,842,868]" name="Dominican Republic" pageId="83" pageNumber="84">Dominican Republic</collectingCountry>
(
<figureCitation id="13212A49FFAC1F3CFC5FFCF3FC258B7F" box="[903,988,842,868]" captionStart="FIGURE 18" captionStartId="87.[136,229,2053,2075]" captionTargetBox="[155,1429,746,2024]" captionTargetId="figure-268@87.[150,1434,741,2029]" captionTargetPageId="87" captionText="FIGURE 18. Distribution of C. spissa, C. torrida, and C. yunquensis." figureDoi="http://doi.org/10.5281/zenodo.8389425" httpUri="https://zenodo.org/record/8389425/files/figure.png" pageId="83" pageNumber="84">Fig. 18</figureCitation>
) in broad leaf forest or dry forest, and persisting in disturbed areas, at 243(7675) m elevation.
</paragraph>
<paragraph id="8BA536CCFFAC1F3CFF65FC2BFB048C7A" blockId="83.[136,1452,266,1624]" pageId="83" pageNumber="84">
<emphasis id="B96EEADEFFAC1F3CFF65FC2BFE508BB7" bold="true" box="[189,425,914,940]" pageId="83" pageNumber="84">Taxonomic notes:—</emphasis>
Two preliminary species (
<emphasis id="B96EEADEFFAC1F3CFD1EFC2AFC4D8BB7" box="[710,948,914,941]" italics="true" pageId="83" pageNumber="84">
<taxonomicName id="4C1A4D4FFFAC1F3CFD1EFC2AFCD58BB7" authorityName="Bailey" authorityYear="1939" box="[710,812,915,941]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="spissa">C. spissa</taxonomicName>
,
<taxonomicName id="4C1A4D4FFFAC1F3CFCEDFC2AFC4D8BB7" authorityName="Burret" authorityYear="1929" box="[821,948,914,941]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="fragrans">C. fragrans</taxonomicName>
</emphasis>
from Hispaniola) share a unique combination of qualitative character states and are recognized as a phylogenetic species, for which the earliest name is
<taxonomicName id="4C1A4D4FFFAC1F3CFAE6FC0EFA5D8BCB" authorityName="Bailey" authorityYear="1939" box="[1342,1444,951,977]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFAC1F3CFAE6FC0EFA5D8BCB" box="[1342,1444,951,977]" italics="true" pageId="83" pageNumber="84">C. spissa</emphasis>
</taxonomicName>
. Note that the preliminary species
<taxonomicName id="4C1A4D4FFFAC1F3CFDDCFC62FD7F8BEF" authorityName="Burret" authorityYear="1929" box="[516,646,986,1013]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="fragrans">
<emphasis id="B96EEADEFFAC1F3CFDDCFC62FD7F8BEF" box="[516,646,986,1013]" italics="true" pageId="83" pageNumber="84">C. fragrans</emphasis>
</taxonomicName>
refers here only to specimens from Hispaniola, which were mistakenly referred to by
<bibRefCitation id="EF8B4B3DFFAC1F3CFEF0FC47FE288C03" author="Bailey, L." box="[296,465,1022,1048]" pageId="83" pageNumber="84" pagination="247 - 259" refId="ref60742" refString="Bailey, L. (1939 b) Coccothrinax in the southern Greater Antilles. Gentes Herbarum 4: 247 - 259." type="journal article" year="1939">Bailey (1939b)</bibRefCitation>
as
<taxonomicName id="4C1A4D4FFFAC1F3CFE2FFC46FD818C03" authorityName="Burret" authorityYear="1929" box="[503,632,1022,1049]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="fragrans">
<emphasis id="B96EEADEFFAC1F3CFE2FFC46FD818C03" box="[503,632,1022,1049]" italics="true" pageId="83" pageNumber="84">C. fragrans</emphasis>
</taxonomicName>
. Stems, leaves, and inflorescences of
<taxonomicName id="4C1A4D4FFFAC1F3CFBFBFC47FAFE8C03" authorityName="Bailey" authorityYear="1939" box="[1059,1287,1022,1048]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFAC1F3CFBFBFC47FAFE8C03" box="[1059,1287,1022,1048]" italics="true" pageId="83" pageNumber="84">Coccothrinax spissa</emphasis>
</taxonomicName>
are the largest of any species in the genus, and stems may be swollen or columnar. Sometimes
<taxonomicName id="4C1A4D4FFFAC1F3CFC22FB9AFBA78C27" authorityName="Bailey" authorityYear="1939" box="[1018,1118,1059,1085]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFAC1F3CFC22FB9AFBA78C27" box="[1018,1118,1059,1085]" italics="true" pageId="83" pageNumber="84">C. spissa</emphasis>
</taxonomicName>
is sympatric with
<taxonomicName id="4C1A4D4FFFAC1F3CFAF1FB9AFA558C27" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[1321,1452,1059,1085]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFAC1F3CFAF1FB9AFA558C27" box="[1321,1452,1059,1085]" italics="true" pageId="83" pageNumber="84">C. argentea</emphasis>
</taxonomicName>
and it can be difficult to distinguish specimens. It seems likely that there are hybrids between the two.
</paragraph>
<paragraph id="8BA536CCFFAC1F3CFF65FBD3FB748D9A" blockId="83.[136,1452,266,1624]" pageId="83" pageNumber="84">
<emphasis id="B96EEADEFFAC1F3CFF65FBD3FE448C9F" bold="true" box="[189,445,1130,1156]" pageId="83" pageNumber="84">Subspecific variation</emphasis>
:—
<taxonomicName id="4C1A4D4FFFAC1F3CFE38FBD3FD318C9F" authorityName="Bailey" authorityYear="1939" box="[480,712,1130,1156]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFAC1F3CFE38FBD3FD318C9F" box="[480,712,1130,1156]" italics="true" pageId="83" pageNumber="84">Coccothrinax spissa</emphasis>
</taxonomicName>
is widely distributed throughout Hispaniola. The population in the southern part of
<collectingCountry id="F30D765CFFAC1F3CFEB2FB37FDE98CB3" box="[362,528,1166,1192]" name="Dominican Republic" pageId="83" pageNumber="84">the Dominican</collectingCountry>
Republic, which includes the
<typeStatus id="54A1886EFFAC1F3CFCBCFB36FC6D8CB2" box="[868,916,1167,1193]" pageId="83" pageNumber="84">type</typeStatus>
locality, occurs in a dryer habitat. Stems of this population are usually considered to be consistently swollen. However, according to the label of
<specimenCount id="9D1CFD45FFAC1F3CFB62FB0AFAAB8CD7" box="[1210,1362,1202,1229]" pageId="83" pageNumber="84" type="generic">one specimen</specimenCount>
(
<emphasis id="B96EEADEFFAC1F3CFABAFB0AFF398CEB" italics="true" pageId="83" pageNumber="84">Moore 7592</emphasis>
) there is considerable variation in the amount of swelling. As shown in the image in
<bibRefCitation id="EF8B4B3DFFAC1F3CFBB7FB6FFAFD8CEB" author="Sayers, J." box="[1135,1284,1238,1264]" pageId="83" pageNumber="84" pagination="20 - 27" refId="ref64690" refString="Sayers, J. (2012) Coccothrinax in Los Angeles. The Palm Journal 202: 20 - 27." type="journal article" year="2012">Sayers (2012</bibRefCitation>
, pages 2627), some stems are swollen and some columnar. Nevertheless, specimens from this southern population seem quite similar to one another, especially in their rachillae that dry a distinctive light brown color. One specimen (
<emphasis id="B96EEADEFFAC1F3CFB12FAA6FA9C8D23" box="[1226,1381,1310,1336]" italics="true" pageId="83" pageNumber="84">Zanoni 31093</emphasis>
) from the southern population was determined by the collector as
<taxonomicName id="4C1A4D4FFFAC1F3CFCF0FAFAFC558D47" authorityName="Sargent ex Schumann" authorityYear="1901" baseAuthorityName="Loddiges ex Schultes &amp; Schultes" box="[808,940,1347,1373]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="argentea">
<emphasis id="B96EEADEFFAC1F3CFCF0FAFAFC558D47" box="[808,940,1347,1373]" italics="true" pageId="83" pageNumber="84">C. argentea</emphasis>
</taxonomicName>
and said to occur in a mixed population with
<taxonomicName id="4C1A4D4FFFAC1F3CFF50FADEFF148D9B" authorityName="Bailey" authorityYear="1939" box="[136,237,1383,1409]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFAC1F3CFF50FADEFF148D9B" box="[136,237,1383,1409]" italics="true" pageId="83" pageNumber="84">C. spissa</emphasis>
</taxonomicName>
. However,
<emphasis id="B96EEADEFFAC1F3CFEB7FADEFDF28D9B" box="[367,523,1382,1408]" italics="true" pageId="83" pageNumber="84">Zanoni 31093</emphasis>
is here considered to be a depauperate plant of
<taxonomicName id="4C1A4D4FFFAC1F3CFBF8FADEFB7C8D9B" authorityName="Bailey" authorityYear="1939" box="[1056,1157,1383,1409]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="83" pageNumber="84" phylum="Tracheophyta" rank="species" species="spissa">
<emphasis id="B96EEADEFFAC1F3CFBF8FADEFB7C8D9B" box="[1056,1157,1383,1409]" italics="true" pageId="83" pageNumber="84">C. spissa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA536CCFFAC1F3CFF65FA32FF158E43" blockId="83.[136,1452,266,1624]" pageId="83" pageNumber="84">
In other areas, specimens appear much more variable. Stems within a population may be either swollen or columnar, as illustrated in
<bibRefCitation id="EF8B4B3DFFAC1F3CFEE2FA17FE338DD3" author="Hodel, D." box="[314,458,1454,1480]" pageId="83" pageNumber="84" pagination="4 - 21" refId="ref62267" refString="Hodel, D. (2013) Coccothrinax. Always late but worth the wait. The Palm Journal 203: 4 - 21." type="journal article" year="2013">Hodel (2013</bibRefCitation>
, fig. 43) and
<bibRefCitation id="EF8B4B3DFFAC1F3CFD86FA17FC428DD3" author="Fernandez, E. &amp; Gottschalk, A." box="[606,955,1454,1480]" pageId="83" pageNumber="84" refId="ref61896" refString="Fernandez, E. &amp; Gottschalk, A. (2017) Palmas de Espanola. Grupo Sid, 159 pp." type="book" year="2017">Fernández &amp; Gottschalk (2017</bibRefCitation>
, page 130). There is one unusual population in northeastern
<collectingCountry id="F30D765CFFAC1F3CFEEFFA6BFDDD8DF7" box="[311,548,1490,1516]" name="Dominican Republic" pageId="83" pageNumber="84">Dominican Republic</collectingCountry>
. This comprises
<specimenCount id="9D1CFD45FFAC1F3CFD07FA6AFC7F8DF7" box="[735,902,1490,1517]" pageId="83" pageNumber="84" type="generic">two specimens</specimenCount>
from near Cabrera. One of these (
<emphasis id="B96EEADEFFAC1F3CFADFFA6AFA5B8DF7" box="[1287,1442,1490,1516]" italics="true" pageId="83" pageNumber="84">Montrero 433</emphasis>
) has a slightly costapalmate leaf and shorter pedicels, and fruits mature from white to reddish-purple. The second specimen (
<emphasis id="B96EEADEFFAC1F3CFEDBF9A2FE598E2F" box="[259,416,1562,1588]" italics="true" pageId="83" pageNumber="84">Zanoni 35835</emphasis>
) has longer pedicels. A few other specimens, from scattered localities, have unusually long pedicels.
</paragraph>
</subSubSection>
</treatment>
</document>