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<document id="FCA2E35F862C4BA3499EB91CD73FED3D" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFB684558CFBFD4CDA9AFE0D" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes panamensis Kluge, 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="50" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="44" updateTime="1698734461607" updateUser="plazi">
<mods:mods id="D8EAC22367E4E6C204483705AAEF354E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F170FD856998AEF88E32052B8E0F9FE0">
<mods:title id="310A50AD057A9976DF245809B179CEC6">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
</mods:titleInfo>
<mods:name id="C2FCEBAB6A4E80A09FCF5B54ADBE4EBD" type="personal">
<mods:role id="0164D6105130645AFC7BA178886F0ED2">
<mods:roleTerm id="AD956802C34D48705933DE9E5426907C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D57EB85DD87407466FF39041EEFA510">Kluge, Nikita J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3B3449CBA53CF4D0C0F72EB4FBF4CD4C">text</mods:typeOfResource>
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<mods:title id="1D1A37522F8B28790E563367E8B2CDAC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="03775DD8B1D1333289C9877410B962E6">
<mods:date id="F58F30B83900CF5987D02068553F0022">2020</mods:date>
<mods:detail id="A62ABF44C3F00FD4A3B5E59D060CFEEE" type="pubDate">
<mods:number id="242875693939B3F8A691A4B754193A2B">2020-03-26</mods:number>
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<mods:number id="97F5D09547372E15A2F2E56BC9F5B1C6">4756</mods:number>
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<mods:number id="E43AAB2830127E243E67245B7B6E1B85">1</mods:number>
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<mods:classification id="FE149CD61FA4159F8BAC0AA895C355EB">journal article</mods:classification>
<mods:identifier id="F768697E0C3C31DB782330D6D6F89F97" type="DOI">10.11646/zootaxa.4756.1.1</mods:identifier>
<mods:identifier id="707BDF42702C7FCA029BE33087477D8B" type="GBIF-Dataset">4d48d52e-6505-4f62-909f-5da5c2cfb489</mods:identifier>
<mods:identifier id="4EEA9DA96FDD6EB9E87811F79EA066C5" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="EFB1E1866CBBF8300219A0DA33D1459A" type="Zenodo-Dep">3736895</mods:identifier>
<mods:identifier id="57AD93C1E3939DC483668A0E6EA10F1B" type="ZooBank">urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D</mods:identifier>
</mods:mods>
<treatment id="039787A6FFB684558CFBFD4CDA9AFE0D" ID-DOI="http://doi.org/10.5281/zenodo.3811773" ID-GBIF-Taxon="163420786" ID-Zenodo-Dep="3811773" LSID="urn:lsid:plazi:treatment:039787A6FFB684558CFBFD4CDA9AFE0D" httpUri="http://treatment.plazi.org/id/039787A6FFB684558CFBFD4CDA9AFE0D" lastPageId="49" lastPageNumber="50" pageId="43" pageNumber="44">
<subSubSection id="C324653BFFB6844F8CFBFD4CD921FD67" box="[151,543,682,709]" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8B8136B0FFB6844F8CFBFD4CD921FD67" blockId="43.[151,543,682,744]" box="[151,543,682,709]" pageId="43" pageNumber="44">
<heading id="D0C981DCFFB6844F8CFBFD4CD921FD67" bold="true" box="[151,543,682,709]" fontSize="10" level="2" pageId="43" pageNumber="44" reason="2">
<emphasis id="B94AEAA2FFB6844F8CFBFD4CD921FD67" bold="true" box="[151,543,682,709]" pageId="43" pageNumber="44">
4.
<taxonomicName id="4C3E4D33FFB6844F8CD9FD4CDAEBFD66" box="[181,469,682,709]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFB6844F8CD9FD4CDAEBFD66" bold="true" box="[181,469,682,709]" italics="true" pageId="43" pageNumber="44">Thraulodes panamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27957D9FFB6844F8DB1FD4DD921FD67" box="[477,543,683,709]" pageId="43" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFB6844F8CFBFD28DA7DFD4A" box="[151,323,717,744]" pageId="43" pageNumber="44" type="description">
<paragraph id="8B8136B0FFB6844F8CFBFD28DA7DFD4A" blockId="43.[151,543,682,744]" box="[151,323,717,744]" pageId="43" pageNumber="44">
(
<figureCitation id="13052A35FFB6844F8CF3FD28DA05FD4A" box="[159,315,717,744]" captionStart-0="FIGURES 122–132" captionStart-1="FIGURES 133–142" captionStart-2="FIGURES 143–148" captionStart-3="FIGURES 149–150" captionStartId-0="44.[151,264,1873,1898]" captionStartId-1="45.[151,264,1898,1923]" captionStartId-2="46.[151,264,1780,1805]" captionStartId-3="47.[151,264,1924,1949]" captionTargetBox-0="[161,1426,166,1853]" captionTargetBox-1="[151,1436,166,1878]" captionTargetBox-2="[151,1419,176,1748]" captionTargetBox-3="[151,1398,166,1905]" captionTargetId-0="figure@44.[161,1426,166,1853]" captionTargetId-1="figure@45.[151,1436,166,1878]" captionTargetId-2="figure@46.[151,1436,166,1760]" captionTargetId-3="figure@47.[151,1398,166,1905]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionTargetPageId-3="47" captionText-0="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." captionText-1="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." captionText-2="FIGURES 143–148. Thraulodes panamensis sp. n. 143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." captionText-3="FIGURES 149–150. Thraulodes panamensis sp. n., egg." figureDoi-0="http://doi.org/10.5281/zenodo.3736933" figureDoi-1="http://doi.org/10.5281/zenodo.3736935" figureDoi-2="http://doi.org/10.5281/zenodo.3736937" figureDoi-3="http://doi.org/10.5281/zenodo.3736939" httpUri-0="https://zenodo.org/record/3736933/files/figure.png" httpUri-1="https://zenodo.org/record/3736935/files/figure.png" httpUri-2="https://zenodo.org/record/3736937/files/figure.png" httpUri-3="https://zenodo.org/record/3736939/files/figure.png" pageId="43" pageNumber="44">Figs 122–150</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFB6844F8CFBFCF0D857FC93" box="[151,873,790,817]" pageId="43" pageNumber="44" type="etymology">
<paragraph id="8B8136B0FFB6844F8CFBFCF0D857FC93" blockId="43.[151,1437,790,2001]" box="[151,873,790,817]" pageId="43" pageNumber="44">
<emphasis id="B94AEAA2FFB6844F8CFBFCF0DA20FC92" bold="true" box="[151,286,790,816]" pageId="43" pageNumber="44">Etymology.</emphasis>
At present this species is known only from
<collectingCountry id="F3297620FFB6844F8F64FCF1D85AFC93" box="[776,868,791,817]" name="Panama" pageId="43" pageNumber="44">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFB6844F8CABFCDFD8A1FC47" pageId="43" pageNumber="44" type="materials_examined">
<paragraph id="8B8136B0FFB6844F8CABFCDFD8A1FC47" blockId="43.[151,1437,790,2001]" pageId="43" pageNumber="44">
<emphasis id="B94AEAA2FFB6844F8CABFCDFDA8AFCF6" bold="true" box="[199,436,825,852]" pageId="43" pageNumber="44">Material examined.</emphasis>
<materialsCitation id="3B563CEDFFB6844F8DD1FCDCDF6BFCDA" ID-GBIF-Occurrence="2594509923" collectingDate="2018-01-30" collectionCode="XVI" collectorName="N. Kluge &amp; L. Sheyko." country="Panama" latitude="8.916667" location="Rio Guabo" longLatPrecision="1300" longitude="-82.183334" municipality="Punta de Pena" pageId="43" pageNumber="44" specimenCount="1" specimenCount-male="1" stateProvince="Provincia Bocas del Toro" typeStatus="holotype">
<emphasis id="B94AEAA2FFB6844F8DD1FCDCD90AFCF6" bold="true" box="[445,564,826,852]" pageId="43" pageNumber="44">
<typeStatus id="54858812FFB6844F8DD1FCDCD910FCF6" box="[445,558,826,852]" pageId="43" pageNumber="44" type="holotype">Holotype</typeStatus>
:
</emphasis>
L-S-I
<specimenCount id="9D38FD39FFB6844F8E11FCDCD9AEFCF6" box="[637,656,826,852]" pageId="43" pageNumber="44" type="male">♂</specimenCount>
{specimen [
<collectionCode id="ED2FAE75FFB6844F8F4DFCDFD866FCF6" box="[801,856,825,852]" pageId="43" pageNumber="44">XVI</collectionCode>
] (7) 2018},
<collectingCountry id="F3297620FFB6844F8F86FCDCDF5AFCF6" box="[1002,1124,826,852]" name="Panama" pageId="43" pageNumber="44">PANAMA</collectingCountry>
,
<collectingRegion id="49FAF852FFB6844F881DFCDCDEA6FCF7" box="[1137,1432,826,853]" country="Panama" name="Bocas del Toro" pageId="43" pageNumber="44">Provincia Bocas del Toro</collectingRegion>
,
<collectingMunicipality id="6BE5ACCAFFB6844F8CFBFCB9DA08FCDB" box="[151,310,862,889]" pageId="43" pageNumber="44">Punta de Peña</collectingMunicipality>
,
<location id="8EE1606BFFB6844F8D2DFCB8DA84FCDA" LSID="urn:lsid:plazi:treatment:039787A6FFB684558CFBFD4CDA9AFE0D:8EE1606BFFB6844F8D2DFCB8DA84FCDA" box="[321,442,862,888]" country="Panama" latitude="8.916667" longLatPrecision="1300" longitude="-82.183334" municipality="Punta de Pena" name="Rio Guabo" pageId="43" pageNumber="44" stateProvince="Provincia Bocas del Toro">Rio Guabo</location>
(
<geoCoordinate id="EE0A5077FFB6844F8DA4FCBBD920FCDA" box="[456,542,861,888]" degrees="8" direction="north" minutes="55" orientation="latitude" pageId="43" pageNumber="44" precision="925" value="8.916667">8°55’N</geoCoordinate>
,
<geoCoordinate id="EE0A5077FFB6844F8E44FCB8D9AFFCDA" box="[552,657,862,888]" degrees="82" direction="west" minutes="11" orientation="longitude" pageId="43" pageNumber="44" precision="925" value="-82.183334">82°11’W</geoCoordinate>
),
<date id="FF801070FFB6844F8EC9FCB8D82CFCDA" box="[677,786,862,888]" pageId="43" pageNumber="44" value="2018-01-30">
<collectingDate id="EFC4E998FFB6844F8EC9FCB8D82CFCDA" box="[677,786,862,888]" pageId="43" pageNumber="44" value="2018-01-30">30.I.2018</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFB6844F8F38FCB9D887FCDA" box="[852,953,862,889]" pageId="43" pageNumber="44">N. Kluge</collectorName>
&amp;
<collectorName id="26CB5366FFB6844F8FB7FCB9DF6BFCDA" box="[987,1109,862,889]" pageId="43" pageNumber="44">L. Sheyko.</collectorName>
</materialsCitation>
<materialsCitation id="3B563CEDFFB6844F8836FCB8D81EFC3E" ID-GBIF-Occurrence="2594509909" collectingDate="2018-01-29" collectingDateMax="2018-01-31" collectingDateMin="2018-01-29" collectionCode="XVI" collectorName="N. Kluge &amp; L. Sheyko." country="Panama" latitude="8.916667" location="Rio Guabo" longLatPrecision="1300" longitude="-82.183334" municipality="Punta de Pena" pageId="43" pageNumber="44" specimenCount="5" specimenCount-female="1" specimenCount-larva="2" specimenCount-male="2" stateProvince="Provincia Bocas del Toro" typeStatus="paratype">
<emphasis id="B94AEAA2FFB6844F8836FCB8DFE5FCDA" bold="true" box="[1114,1243,862,888]" pageId="43" pageNumber="44">
<typeStatus id="54858812FFB6844F8836FCB8DFE8FCDA" box="[1114,1238,862,888]" pageId="43" pageNumber="44" type="paratype">Paratypes</typeStatus>
:
</emphasis>
the same locality and collectors,
<date id="FF801070FFB6844F8D53FC64DAE6FC3E" box="[319,472,898,924]" pageId="43" pageNumber="44" value="2018-01-29" valueMax="2018-01-31" valueMin="2018-01-29">
<collectingDate id="EFC4E998FFB6844F8D53FC64DAE6FC3E" box="[319,472,898,924]" pageId="43" pageNumber="44" value="2018-01-29" valueMax="2018-01-31" valueMin="2018-01-29">29–31.I.2018</collectingDate>
</date>
: 1 L-S-I
<specimenCount id="9D38FD39FFB6844F8E5BFC64D974FC3E" box="[567,586,898,924]" pageId="43" pageNumber="44" type="male">♂</specimenCount>
,
<specimenCount id="9D38FD39FFB6844F8E38FC64D945FC3E" box="[596,635,898,924]" pageId="43" pageNumber="44" type="larva">2 L</specimenCount>
/S
<specimenCount id="9D38FD39FFB6844F8EF8FC64D996FC3E" box="[660,680,898,924]" pageId="43" pageNumber="44" type="male">♂</specimenCount>
, 1 L-S-I
<specimenCount id="9D38FD39FFB6844F8F61FC64D825FC3E" box="[781,795,898,924]" pageId="43" pageNumber="44" type="female">♀</specimenCount>
;
</materialsCitation>
<materialsCitation id="3B563CEDFFB6844F8F4AFC64DE14FC62" ID-GBIF-Occurrence="2594509912" collectingDate="2018-02-10" collectorName="N. Kluge &amp; L. Sheyko" location="Rio" municipality="Parque Nacional Soberania" pageId="43" pageNumber="44" specimenCount="42" specimenCount-female="2" specimenCount-larva="39" specimenCount-male="1" stateProvince="Provincia de Colon">
<collectingRegion id="49FAF852FFB6844F8F4AFC64D8C0FC3E" box="[806,1022,898,924]" country="Panama" name="Colon" pageId="43" pageNumber="44">Provincia de Colon</collectingRegion>
,
<collectingMunicipality id="6BE5ACCAFFB6844F8865FC65DE04FC3E" box="[1033,1338,898,925]" pageId="43" pageNumber="44">Parque Nacional Soberania</collectingMunicipality>
,
<location id="8EE1606BFFB6844F8928FC64DE52FC3E" LSID="urn:lsid:plazi:treatment:039787A6FFB684558CFBFD4CDA9AFE0D:8EE1606BFFB6844F8928FC64DE52FC3E" box="[1348,1388,898,924]" municipality="Parque Nacional Soberania" name="Rio" pageId="43" pageNumber="44" stateProvince="Provincia de Colon">Rio</location>
Fri- joles, camina del oleoducto,
<date id="FF801070FFB6844F8DB9FC40D972FC62" box="[469,588,934,960]" pageId="43" pageNumber="44" value="2018-02-10">
<collectingDate id="EFC4E998FFB6844F8DB9FC40D972FC62" box="[469,588,934,960]" pageId="43" pageNumber="44" value="2018-02-10">10.II.2018</collectingDate>
</date>
, coll.
<collectorName id="26CB5366FFB6844F8EE2FC41D8B3FC62" box="[654,909,934,961]" pageId="43" pageNumber="44">N. Kluge &amp; L. Sheyko</collectorName>
: 2 L-S-I
<specimenCount id="9D38FD39FFB6844F8F9DFC40D8C1FC62" box="[1009,1023,934,960]" pageId="43" pageNumber="44" type="female">♀</specimenCount>
, 3 S-I
<specimenCount id="9D38FD39FFB6844F8828FC40DF66FC62" box="[1092,1112,934,960]" pageId="43" pageNumber="44" type="male">♂</specimenCount>
, 1 S-I
<specimenCount id="9D38FD39FFB6844F88CDFC40DF8FFC62" box="[1185,1201,934,960]" pageId="43" pageNumber="44" type="female">♀</specimenCount>
,
<specimenCount id="9D38FD39FFB6844F88D0FC40DE18FC62" box="[1212,1318,934,960]" pageId="43" pageNumber="44" type="larva">39 larvae</specimenCount>
;
</materialsCitation>
<materialsCitation id="3B563CEDFFB6844F895DFC40D8A1FC47" ID-GBIF-Occurrence="2594509929" collectingDate="1977-12-19" collectionCode="R" collectorName="E. of Caldera &amp; W. Flowers" location="Provincia Chiriqui" pageId="43" pageNumber="44" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Provincia Chiriqui">
<collectingRegion id="49FAF852FFB6844F895DFC40DBC9FC46" country="Panama" name="Chiriqui" pageId="43" pageNumber="44">Provincia Chiriqui</collectingRegion>
,
<collectorName id="26CB5366FFB6844F8D6FFC2DDAA4FC46" box="[259,410,970,997]" pageId="43" pageNumber="44">E. of Caldera</collectorName>
,
<date id="FF801070FFB6844F8DC9FC2CD90EFC46" box="[421,560,970,996]" pageId="43" pageNumber="44" value="1977-12-19">
<collectingDate id="EFC4E998FFB6844F8DC9FC2CD90EFC46" box="[421,560,970,996]" pageId="43" pageNumber="44" value="1977-12-19">19.XII.1977</collectingDate>
</date>
, coll.,
<collectionCode id="ED2FAE75FFB6844F8E17FC2DD9B1FC47" box="[635,655,971,997]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="43" pageNumber="44">R</collectionCode>
.
<collectorName id="26CB5366FFB6844F8EFFFC2DD827FC46" box="[659,793,970,997]" pageId="43" pageNumber="44">W. Flowers</collectorName>
:
<date id="FF801070FFB6844F8F4FFC2CD87CFC47" box="[803,834,970,997]" pageId="43" pageNumber="44">
<collectingDate id="EFC4E998FFB6844F8F4FFC2CD87CFC47" box="[803,834,970,997]" pageId="43" pageNumber="44">2 I</collectingDate>
</date>
<specimenCount id="9D38FD39FFB6844F8F2FFC2DD867FC47" box="[835,857,971,997]" pageId="43" pageNumber="44" type="male">♂</specimenCount>
,
<date id="FF801070FFB6844F8F09FC2CD8B9FC47" box="[869,903,970,997]" pageId="43" pageNumber="44">
<collectingDate id="EFC4E998FFB6844F8F09FC2CD8B9FC47" box="[869,903,970,997]" pageId="43" pageNumber="44">2 I</collectingDate>
</date>
<specimenCount id="9D38FD39FFB6844F8FEBFC2DD8A4FC47" box="[903,922,971,997]" pageId="43" pageNumber="44" type="female">♀</specimenCount>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324653BFFB684558CABFC0BDA9AFE0D" lastPageId="49" lastPageNumber="50" pageId="43" pageNumber="44" type="description">
<paragraph id="8B8136B0FFB6844F8CABFC0BDA4AFB3A" blockId="43.[151,1437,790,2001]" pageId="43" pageNumber="44">
<emphasis id="B94AEAA2FFB6844F8CABFC0BDA8BFBAB" bold="true" box="[199,437,1005,1033]" pageId="43" pageNumber="44">
Descriptions.
<emphasis id="B94AEAA2FFB6844F8D06FC09DA8BFBAB" bold="true" box="[362,437,1007,1033]" italics="true" pageId="43" pageNumber="44">Larva.</emphasis>
</emphasis>
CUTICULAR COLORATION (
<figureCitation id="13052A35FFB6844F8F4FFC08D8FCFBAA" box="[803,962,1006,1032]" captionStart="FIGURES 122–132" captionStartId="44.[151,264,1873,1898]" captionTargetBox="[161,1426,166,1853]" captionTargetId="figure@44.[161,1426,166,1853]" captionTargetPageId="44" captionText="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="43" pageNumber="44">Figs 122–127</figureCitation>
). Dorsal side of head, thorax and abdomen mostly brown with few blanks: pronotum bordered by light laterally, with or without other blanks; fore protoptera with convex and concave longitudinal veins lighter than background; abdominal terga with blanks on lateral parts, area mediad of tergalii attachments nearly unicolor brown. Femora mostly brown, with blanks; tibiae and tarsi uniformly light brown.
</paragraph>
<paragraph id="8B8136B0FFB6844F8CABFB44DA88FAA6" blockId="43.[151,1437,790,2001]" pageId="43" pageNumber="44">
HYPODERMAL COLORATION (
<figureCitation id="13052A35FFB6844F8E39FB44D987FB1E" box="[597,697,1186,1212]" captionStart="FIGURES 122–132" captionStartId="44.[151,264,1873,1898]" captionTargetBox="[161,1426,166,1853]" captionTargetId="figure@44.[161,1426,166,1853]" captionTargetPageId="44" captionText="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="43" pageNumber="44">Fig. 128</figureCitation>
). Abdominal terga, besides cuticular coloration (see above) often with paired transverse or round blackish spots corresponding to midway gray spots of imago. Tergalii brown or gray, tracheae black (
<figureCitation id="13052A35FFB6844F8D2BFB0CDA94FAA6" box="[327,426,1258,1284]" captionStart="FIGURES 122–132" captionStartId="44.[151,264,1873,1898]" captionTargetBox="[161,1426,166,1853]" captionTargetId="figure@44.[161,1426,166,1853]" captionTargetPageId="44" captionText="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="43" pageNumber="44">Fig. 129</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFB6844F8CABFAE8D90FFA37" blockId="43.[151,1437,790,2001]" pageId="43" pageNumber="44">
SHAPE AND SETATION. Clypeus widened distally; labrum 1.3–1.4 times wider than clypeus (
<figureCitation id="13052A35FFB6844F8946FAE8DEAEFA8A" box="[1322,1424,1294,1320]" captionStart="FIGURES 122–132" captionStartId="44.[151,264,1873,1898]" captionTargetBox="[161,1426,166,1853]" captionTargetId="figure@44.[161,1426,166,1853]" captionTargetPageId="44" captionText="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="43" pageNumber="44">Fig. 123</figureCitation>
). Labrum widest at 1/3 of length from base; initial fore margin (turned ventrally) without median emargination, with all 5 denticles wide; anterior transverse setal row regular, as wide as 3 denticles (as in
<figureCitation id="13052A35FFB6844F8835FAB0DF92FAD2" box="[1113,1196,1366,1392]" captionStart="FIGURES 86–93" captionStartId="37.[151,264,188,213]" captionTargetId="figure@36.[151,1436,342,2018]" captionTargetPageId="36" captionText="FIGURES 86–93. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." figureDoi="http://doi.org/10.5281/zenodo.3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="43" pageNumber="44">Fig. 88</figureCitation>
). Maxilla with 16–17 pectinate setae in apical-ventral row.
</paragraph>
<paragraph id="8B8136B0FFB6844F8CABFA79D88CFA7E" blockId="43.[151,1437,790,2001]" pageId="43" pageNumber="44">
Femora: Stout setae on anterior surface widened distally, apically rounded (
<figureCitation id="13052A35FFB6844F886BFA78DF56FA1A" box="[1031,1128,1438,1464]" captionStart="FIGURES 122–132" captionStartId="44.[151,264,1873,1898]" captionTargetBox="[161,1426,166,1853]" captionTargetId="figure@44.[161,1426,166,1853]" captionTargetPageId="44" captionText="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="43" pageNumber="44">Fig. 130</figureCitation>
). Irregular row of hairs near inner margin absent on fore femur, present on middle and hind femora.
</paragraph>
<paragraph id="8B8136B0FFB6844F8CABFA01DA88F9CF" blockId="43.[151,1437,790,2001]" pageId="43" pageNumber="44">
Fore tibia (
<figureCitation id="13052A35FFB6844F8D2AFA00DA94F9A2" box="[326,426,1510,1536]" captionStart="FIGURES 122–132" captionStartId="44.[151,264,1873,1898]" captionTargetBox="[161,1426,166,1853]" captionTargetId="figure@44.[161,1426,166,1853]" captionTargetPageId="44" captionText="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="43" pageNumber="44">Fig. 131</figureCitation>
): outer hairs form two irregular rows; inner-anterior row of recurved hairs absent; inneranterior row of stout setae absent; inner field of stout pointed setae moderately dense (setae lengths subequal to distances between them), its setae subequal in lengths, forming 2 rows (setae of anterior row partly dentate, setae of posterior row smooth).
</paragraph>
<paragraph id="8B8136B0FFB6844F8CABF990DAF9F8BD" blockId="43.[151,1437,790,2001]" pageId="43" pageNumber="44">
Hind tibia (
<figureCitation id="13052A35FFB6844F8D20F990DA8CF932" box="[332,434,1654,1680]" captionStart="FIGURES 122–132" captionStartId="44.[151,264,1873,1898]" captionTargetBox="[161,1426,166,1853]" captionTargetId="figure@44.[161,1426,166,1853]" captionTargetPageId="44" captionText="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="43" pageNumber="44">Fig. 132</figureCitation>
): outer-anterior row of stout setae consists of two clearly different
<typeStatus id="54858812FFB6844F88DBF991DFCCF933" box="[1207,1266,1655,1681]" pageId="43" pageNumber="44">types</typeStatus>
of stout setae: numerous very short truncated ones and several long spoon-like ones; outer-posterior row of stout setae consists of blunt stout setae of various lengths; posterior hairs located between these rows, numerous and form more than one row (besides row of hairs posteriad of outer-posterior row of stout setae); stout setae of inner-anterior row short, widened distally and blunt.
</paragraph>
<paragraph id="8B8136B0FFB6844F8CABF8CFDE23F8E1" blockId="43.[151,1437,790,2001]" box="[199,1309,1833,1859]" pageId="43" pageNumber="44">Claws with 6–9 denticles on rigid portion, with or without minute denticles on articulatory portion.</paragraph>
<paragraph id="8B8136B0FFB6844F8CABF8AAD8CBF8C4" blockId="43.[151,1437,790,2001]" box="[199,1013,1868,1894]" pageId="43" pageNumber="44">
Tergalii (
<figureCitation id="13052A35FFB6844F8D41F8AADAB1F8C4" box="[301,399,1868,1894]" captionStart="FIGURES 122–132" captionStartId="44.[151,264,1873,1898]" captionTargetBox="[161,1426,166,1853]" captionTargetId="figure@44.[161,1426,166,1853]" captionTargetPageId="44" captionText="FIGURES 122–132. Thraulodes panamensis sp. n. 122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="43" pageNumber="44">Fig. 129</figureCitation>
): each lamella slender, tracheae without side branches.
</paragraph>
<paragraph id="8B8136B0FFB6844F8CABF889DAC3F872" blockId="43.[151,1437,790,2001]" pageId="43" pageNumber="44">
Male genitalia in last larval instar (
<figureCitation id="13052A35FFB6844F8E3FF889D988F828" box="[595,694,1903,1930]" captionStart="FIGURES 143–148" captionStartId="46.[151,264,1780,1805]" captionTargetBox="[151,1419,176,1748]" captionTargetId="figure@46.[151,1436,166,1760]" captionTargetPageId="46" captionText="FIGURES 143–148. Thraulodes panamensis sp. n. 143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736937" httpUri="https://zenodo.org/record/3736937/files/figure.png" pageId="43" pageNumber="44">Fig. 147</figureCitation>
): protogonostyli short and separated one from another by shallow emargination. Each protopenis lobe with small, sharply projected gonopore-bearing process bent laterally; gonopores opened caudally-laterally.
</paragraph>
<caption id="DF416638FFB184488CFBF8B7D8DCF858" ID-DOI="http://doi.org/10.5281/zenodo.3736933" ID-Zenodo-Dep="3736933" httpUri="https://zenodo.org/record/3736933/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,264,1873,1898]" targetBox="[161,1426,166,1853]" targetPageId="44">
<paragraph id="8B8136B0FFB184488CFBF8B7D8DCF858" blockId="44.[151,1437,1873,2042]" pageId="44" pageNumber="45">
<emphasis id="B94AEAA2FFB184488CFBF8B7DA53F8C8" bold="true" box="[151,365,1873,1898]" pageId="44" pageNumber="45">FIGURES 122–132.</emphasis>
<taxonomicName id="4C3E4D33FFB184488D1FF8B4D95BF8C8" box="[371,613,1874,1898]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFB184488D1FF8B4D95BF8C8" box="[371,613,1874,1898]" italics="true" pageId="44" pageNumber="45">Thraulodes panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFB184488E06F8B4D99DF8C8" bold="true" box="[618,675,1874,1898]" pageId="44" pageNumber="45">
<taxonomicNameLabel id="A27957D9FFB184488E06F8B4D99DF8C8" box="[618,675,1874,1898]" pageId="44" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
122–127, larval exuviae at the same magnification: 122–124, parts of head with labrum and mandibles; 125, left half of pronotum and mesonotum; 126, part of sternum and left pleura with half of metanotum and hind protopteron, left fore, middle and hind legs; 127, abdominal terga and sterna; 128, immature larva with light cuticle and well visible blackish hypodermal markings on abdomen; 129, tergalius III; 130, anterior side of hind femur; 131, anterior side of fore tibia; 132, anterior side of hind tibia (122–127, 130, holotype).
</paragraph>
</caption>
<caption id="DF416638FFB084498CFBF88CDA5BF84D" ID-DOI="http://doi.org/10.5281/zenodo.3736935" ID-Zenodo-Dep="3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="45" pageNumber="46" startId="45.[151,264,1898,1923]" targetBox="[151,1436,166,1878]" targetPageId="45">
<paragraph id="8B8136B0FFB084498CFBF88CDA5BF84D" blockId="45.[151,1436,1898,2031]" pageId="45" pageNumber="46">
<emphasis id="B94AEAA2FFB084498CFBF88CDA4EF821" bold="true" box="[151,368,1898,1923]" pageId="45" pageNumber="46">FIGURES 133–142.</emphasis>
<taxonomicName id="4C3E4D33FFB084498D14F88CD953F820" box="[376,621,1898,1922]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFB084498D14F88CD953F820" box="[376,621,1898,1922]" italics="true" pageId="45" pageNumber="46">Thraulodes panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFB084498E19F88CD98FF820" bold="true" box="[629,689,1898,1922]" pageId="45" pageNumber="46">
<taxonomicNameLabel id="A27957D9FFB084498E19F88CD98FF820" box="[629,689,1898,1922]" pageId="45" pageNumber="46" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype).
</paragraph>
</caption>
<caption id="DF416638FFB3844A8CFBF912D990F8DB" ID-DOI="http://doi.org/10.5281/zenodo.3736937" ID-Zenodo-Dep="3736937" httpUri="https://zenodo.org/record/3736937/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,264,1780,1805]" targetBox="[151,1419,176,1748]" targetPageId="46">
<paragraph id="8B8136B0FFB3844A8CFBF912D990F8DB" blockId="46.[151,1436,1780,1913]" pageId="46" pageNumber="47">
<emphasis id="B94AEAA2FFB3844A8CFBF912DA53F8AF" bold="true" box="[151,365,1780,1805]" pageId="46" pageNumber="47">FIGURES 143–148.</emphasis>
<taxonomicName id="4C3E4D33FFB3844A8D1FF912D95AF8AE" box="[371,612,1780,1804]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFB3844A8D1FF912D95AF8AE" box="[371,612,1780,1804]" italics="true" pageId="46" pageNumber="47">Thraulodes panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFB3844A8E06F913D99DF8AF" bold="true" box="[618,675,1781,1805]" pageId="46" pageNumber="47">
<taxonomicNameLabel id="A27957D9FFB3844A8E06F913D99DF8AF" box="[618,675,1781,1805]" pageId="46" pageNumber="47" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis.
</paragraph>
</caption>
<caption id="DF416638FFB2844B8CFBF862D9E3F83F" ID-DOI="http://doi.org/10.5281/zenodo.3736939" ID-Zenodo-Dep="3736939" box="[151,733,1924,1949]" httpUri="https://zenodo.org/record/3736939/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,264,1924,1949]" targetBox="[151,1398,166,1905]" targetPageId="47">
<paragraph id="8B8136B0FFB2844B8CFBF862D9E3F83F" blockId="47.[151,733,1924,1949]" box="[151,733,1924,1949]" pageId="47" pageNumber="48">
<emphasis id="B94AEAA2FFB2844B8CFBF862DA50F83F" bold="true" box="[151,366,1924,1949]" pageId="47" pageNumber="48">FIGURES 149–150.</emphasis>
<taxonomicName id="4C3E4D33FFB2844B8D1FF863D95BF83F" box="[371,613,1925,1949]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFB2844B8D1FF863D95BF83F" box="[371,613,1925,1949]" italics="true" pageId="47" pageNumber="48">Thraulodes panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFB2844B8E07F863D99AF83F" bold="true" box="[619,676,1925,1949]" pageId="47" pageNumber="48">
<taxonomicNameLabel id="A27957D9FFB2844B8E07F863D99AF83F" box="[619,676,1925,1949]" pageId="47" pageNumber="48" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, egg.
</paragraph>
</caption>
<paragraph id="8B8136B0FFAD84548CABFF7ED99EFE2A" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
<emphasis id="B94AEAA2FFAD84548CABFF7EDA02FF11" bold="true" box="[199,316,152,179]" italics="true" pageId="48" pageNumber="49">Subimago</emphasis>
(
<figureCitation id="13052A35FFAD84548D21FF7FDAD3FF11" box="[333,493,153,179]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Figs 137–138</figureCitation>
,
<figureCitation id="13052A35FFAD84548D97FF7FD917FF16" box="[507,553,153,180]" captionStart="FIGURES 143–148" captionStartId="46.[151,264,1780,1805]" captionTargetBox="[151,1419,176,1748]" captionTargetId="figure@46.[151,1436,166,1760]" captionTargetPageId="46" captionText="FIGURES 143–148. Thraulodes panamensis sp. n. 143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736937" httpUri="https://zenodo.org/record/3736937/files/figure.png" pageId="48" pageNumber="49">146</figureCitation>
). CUTICULAR COLORATION. Cuticle mostly colorless, with brown areas. Flagellum of antenna brown. Pronotum colorless. Mesonotum mostly colorless, with contrasting dark brown antelateroparapsidal and lateroparapsidal sutures, brown anteronotal transverse impression and chromozone of medioscutum; chromozone of submedioscutum either nearly colorless (
<figureCitation id="13052A35FFAD84548FDEFEE5DF29FEBF" box="[946,1047,259,285]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Fig. 137</figureCitation>
), or light brownish, contrastingly lighter than chromozone of medioscutum (
<figureCitation id="13052A35FFAD84548E17FEC1D9E1FEE3" box="[635,735,295,321]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Fig. 138</figureCitation>
). Thoracic pleura and sterna colorless. On each leg femur entirely colorless, tibia and tarsus brown. Cuticle of wings colorless, microtrichia light brownish. Abdominal terga, sterna and gonostyli colorless, caudalii brown.
</paragraph>
<paragraph id="8B8136B0FFAD84548CABFE74DA1CFE50" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
TEXTURE. On tarsi of all legs, 1st tarsomere with microtrichia (as tibia), 2nd–5th tarsomeres coved by blunt microlepides; pointed microlepides present near apical margins of 2nd–4th tarsomeres of middle and hind legs (as in
<figureCitation id="13052A35FFAD84548CD8FE3EDA2BFE50" box="[180,277,472,498]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="48" pageNumber="49">Fig. 473</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFAD84548CABFE1DD894FD98" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
<emphasis id="B94AEAA2FFAD84548CABFE1DDA6BFDB4" bold="true" box="[199,341,507,534]" italics="true" pageId="48" pageNumber="49">Male imago</emphasis>
(
<figureCitation id="13052A35FFAD84548D0BFE1DD942FDB4" box="[359,636,507,534]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Figs 133–136, 140–142</figureCitation>
,
<figureCitation id="13052A35FFAD84548EE7FE1DD812FDB4" box="[651,812,507,534]" captionStart="FIGURES 143–148" captionStartId="46.[151,264,1780,1805]" captionTargetBox="[151,1419,176,1748]" captionTargetId="figure@46.[151,1436,166,1760]" captionTargetPageId="46" captionText="FIGURES 143–148. Thraulodes panamensis sp. n. 143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736937" httpUri="https://zenodo.org/record/3736937/files/figure.png" pageId="48" pageNumber="49">143–145, 148</figureCitation>
). Head brown. Antenna with scapus ocher, pedicellus brown, flagellum ocher. Dorsal eyes contiguous medially, dull orange.
</paragraph>
<paragraph id="8B8136B0FFAD84548CABFDA4DA56FD65" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
Prothorax whitish with pair of dark brown spots above coxa. Mesoscutum ocher with brown, scutellum whitish. Metanotum and lateral sides of pterothorax ocher with brown. Ventral side of pterothorax with mesothoracic episterna, furcasternal protuberances and furcasternal median impression dark brown, basisternum and other areas whitish (
<figureCitation id="13052A35FFAD84548C95FD4BDA65FD65" box="[249,347,684,711]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Fig. 135</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFAD84548CABFD36D812FBAA" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
Cuticle of legs mostly colorless; on fore leg apex of femur and base of tibia with cuticle light brownish. Legs with following hypodermal pigmentation on whitish background: Fore femur with triangular brown spot on anterior side at 1/4 from base and with multicolored apical band occupying distal 0.4 of femur; apical band orange, proximally and apically narrowly bordered by blackish-brown, on inner and outer sides with blackish-brown longitudinal stripes. Fore tibia with dark brown spots on base and on apex. Middle and hind femora with multicolored pre-apical band within apical 0.4 of femur; pre-apical band mainly orange-brown, proximally bordered by blackish-brown, on inner and outer sides with blackish-brown longitudinal stripes; apical 0.1 of femur non-pigmented. In
<typeStatus id="54858812FFAD8454895FFC40DEA6FC62" box="[1331,1432,934,960]" pageId="48" pageNumber="49" type="holotype">holotype</typeStatus>
, relation of fore femur to fore wing length 60:270; proportions femur/tibia/tarsomeres on fore leg 60:90:2:21:14:9:6; on middle leg 60:68:2:4:3:2:6; on hind leg 71:77:2:4:3:2:6.
</paragraph>
<paragraph id="8B8136B0FFAD84548CABFBF4D89EFB1E" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
Fore and hind wings with dark brown macula on costal brace and bases of Sc and R; longitudinal and cross veins colorless (proximal coloration of fore and hind wings appears during larval development and present both in subimago and imago). Costal cross veins proximad of bulla few, colorless and very thin, so that visible only under high magnification (
<figureCitation id="13052A35FFAD84548DECFB98DADAFB3A" box="[384,484,1150,1176]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Fig. 142</figureCitation>
). Pterostigmatic cross veins moderately dense, oblique, arched, non-branched (
<figureCitation id="13052A35FFAD84548903FB98DBFBFB1E" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Fig. 133</figureCitation>
). Hind wing with prominent or blunt costal projection (
<figureCitation id="13052A35FFAD84548F5DFB44D8AAFB1E" box="[817,916,1186,1212]" captionStart="FIGURES 143–148" captionStartId="46.[151,264,1780,1805]" captionTargetBox="[151,1419,176,1748]" captionTargetId="figure@46.[151,1436,166,1760]" captionTargetPageId="46" captionText="FIGURES 143–148. Thraulodes panamensis sp. n. 143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736937" httpUri="https://zenodo.org/record/3736937/files/figure.png" pageId="48" pageNumber="49">Fig. 148</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFAD84548CABFB20D82CFAEE" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">Abdominal hypodermal coloration: Tergum I brown. Terga II–VI whitish, translucent; terga III–VI with pair of small gray midway spots; pleura with blackish stigmatic dots. Tergum VII orange, tergum VIII anteriorly-medially orange and posteriorly-laterally white, terga XI–X orange. Sterna I–IX whitish. Caudalii with segments alternated as following: black; white; white with base black; white.</paragraph>
<paragraph id="8B8136B0FFAD84548CABFAB0DE3EFA7E" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
Genitalia (
<figureCitation id="13052A35FFAD84548D53FAB0DAE1FAD2" box="[319,479,1366,1392]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Figs 140–141</figureCitation>
,
<figureCitation id="13052A35FFAD84548D81FAB0D96AFAD3" box="[493,596,1366,1393]" captionStart="FIGURES 143–148" captionStartId="46.[151,264,1780,1805]" captionTargetBox="[151,1419,176,1748]" captionTargetId="figure@46.[151,1436,166,1760]" captionTargetPageId="46" captionText="FIGURES 143–148. Thraulodes panamensis sp. n. 143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736937" httpUri="https://zenodo.org/record/3736937/files/figure.png" pageId="48" pageNumber="49">143–146</figureCitation>
): Styliger and gonostyli whitish. Dorsal extension of styliger narrow and prominent. Penis lobes relatively long, moderately widened and divergent apically, with narrow lateral pouches. Apico-lateral area forming ear, projected laterally-caudally. Telopenes in form of spear-like rolls, attached on dorsal side and directed medially-dorsally, nearly straight, with groove opened dorsally-laterally (
<figureCitation id="13052A35FFAD845488FDFA24DFCDFA7E" box="[1169,1267,1474,1500]" captionStart="FIGURES 143–148" captionStartId="46.[151,264,1780,1805]" captionTargetBox="[151,1419,176,1748]" captionTargetId="figure@46.[151,1436,166,1760]" captionTargetPageId="46" captionText="FIGURES 143–148. Thraulodes panamensis sp. n. 143, genitalia of male imago in lateral view; 144, apex of penis with spearlike roll in dorsal view; 145, genitalia of the same specimen in ventral view; 146, subimaginal exuviae of genitalia at the same magnification; 147, larval exuviae of protogonostyli and protopenis at the same magnification; 148, hind wing (146, 147, holotype). Abbreviations: lp, pateral pouch; tp, telopenis." figureDoi="http://doi.org/10.5281/zenodo.3736937" httpUri="https://zenodo.org/record/3736937/files/figure.png" pageId="48" pageNumber="49">Fig. 144</figureCitation>
).
</paragraph>
<paragraph id="8B8136B0FFAD84548CABFA03D8FCF986" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
<emphasis id="B94AEAA2FFAD84548CABFA03DA4CF9A2" bold="true" box="[199,370,1509,1536]" italics="true" pageId="48" pageNumber="49">Female imago</emphasis>
(
<figureCitation id="13052A35FFAD84548DE9FA00DAD2F9A2" box="[389,492,1510,1536]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Fig. 139</figureCitation>
). Pronotum medially brown with ocher, laterally whitish; abdominal terga II–VI brownish with ocher; in other respects coloration similar to that of male.
</paragraph>
<paragraph id="8B8136B0FFAD84548CABF9C8DAC0F9CE" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
<emphasis id="B94AEAA2FFAD84548CABF9C8DA3FF9EA" bold="true" box="[199,257,1582,1608]" italics="true" pageId="48" pageNumber="49">Eggs</emphasis>
(
<figureCitation id="13052A35FFAD84548D63F9C8DA90F9EA" box="[271,430,1581,1608]" captionStart="FIGURES 149–150" captionStartId="47.[151,264,1924,1949]" captionTargetBox="[151,1398,166,1905]" captionTargetId="figure@47.[151,1398,166,1905]" captionTargetPageId="47" captionText="FIGURES 149–150. Thraulodes panamensis sp. n., egg." figureDoi="http://doi.org/10.5281/zenodo.3736939" httpUri="https://zenodo.org/record/3736939/files/figure.png" pageId="48" pageNumber="49">Figs 149–150</figureCitation>
). Mostly barrel-shaped. Each KTC surrounded by thick ring-like cove. Other chorion with evenly dispersed protuberances.
</paragraph>
<paragraph id="8B8136B0FFAD84548CABF993D8CBF932" blockId="48.[151,1437,152,2040]" box="[199,1013,1653,1681]" pageId="48" pageNumber="49">
<emphasis id="B94AEAA2FFAD84548CABF993DA70F932" bold="true" box="[199,334,1653,1680]" pageId="48" pageNumber="49">Dimension.</emphasis>
Fore wing length (and approximate body length)
<quantity id="4CC69B55FFAD84548F10F993D8CFF932" box="[892,1009,1653,1680]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="48" pageNumber="49" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.5–7 mm</quantity>
.
</paragraph>
<paragraph id="8B8136B0FFAD84548CABF97FD8A6F95E" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
<emphasis id="B94AEAA2FFAD84548CABF97FDA5CF916" bold="true" box="[199,354,1689,1716]" pageId="48" pageNumber="49">Comparison.</emphasis>
Male imago of
<taxonomicName id="4C3E4D33FFAD84548E7EF97CD9FFF916" box="[530,705,1690,1716]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFAD84548E7EF97CD9FFF916" box="[530,705,1690,1716]" italics="true" pageId="48" pageNumber="49">Th. panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFAD84548EA4F97CD839F916" bold="true" box="[712,775,1690,1716]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A27957D9FFAD84548EA4F97CD839F916" box="[712,775,1690,1716]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has genital structure, coloration of thorax, wings and abdomen the same as in
<taxonomicName id="4C3E4D33FFAD84548D1FF958D9E5F97A" authority="Dominguez 1986" authorityName="Dominguez" authorityYear="1986" box="[371,731,1726,1752]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bolivianus">
<emphasis id="B94AEAA2FFAD84548D1FF958D92DF97A" box="[371,531,1726,1752]" italics="true" pageId="48" pageNumber="49">Th. bolivianus</emphasis>
<bibRefCitation id="EFAF4B41FFAD84548E76F958D9E5F97A" author="Dominguez, E." box="[538,731,1726,1752]" pageId="48" pageNumber="49" pagination="149 - 153" refId="ref69183" refString="Dominguez, E. (1986) Thraulodes bolivianus, una nueva especie de la familia Leptophlebiidae (Insecta: Ephemeroptera) de Bolivia. Acta Zoologica Lilloana, 38, 149 - 153." type="journal article" year="1986">Dominguez 1986</bibRefCitation>
</taxonomicName>
described from
<collectingCountry id="F3297620FFAD84548FFEF958D8DAF97A" box="[914,996,1726,1752]" name="Bolivia" pageId="48" pageNumber="49">Bolivia</collectingCountry>
and
<taxonomicName id="4C3E4D33FFAD84548876F958DEA2F97A" authority="Dominguez et al. 2015" authorityName="Dominguez" authorityYear="2015" box="[1050,1436,1725,1752]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="insular">
<emphasis id="B94AEAA2FFAD84548876F958DFABF97A" box="[1050,1173,1726,1752]" italics="true" pageId="48" pageNumber="49">Th. insular</emphasis>
Domínguez
<emphasis id="B94AEAA2FFAD84548949F959DE69F97A" box="[1317,1367,1726,1752]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2015
</taxonomicName>
described by
<bibRefCitation id="EFAF4B41FFAD84548D40F904D933F95E" author="Zuniga, M. C. &amp; Molineri, C. &amp; Dominguez, E. &amp; Cardona, W." box="[300,525,1761,1788]" pageId="48" pageNumber="49" pagination="281 - 296" refId="ref72771" refString="Zuniga, M. C., Molineri, C., Dominguez, E. &amp; Cardona, W. (2015) Leptophlebiidae (Insecta: Ephemeroptera) from Gorgona Island National Natural Park (Tropical Eastern Pacific, Colombia) with the description of two new species. Annales de Limnologie-International Journal of Limnology, 51, 281 - 296. https: // doi. org / 10.1051 / limn / 2015026" type="journal article" year="2015">
Zúñiga
<emphasis id="B94AEAA2FFAD84548DEFF905DA8AF95E" box="[387,436,1762,1788]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. (2015)
</bibRefCitation>
from Gorgona Island in
<collectingCountry id="F3297620FFAD84548F4EF904D8AAF95E" box="[802,916,1762,1788]" name="Colombia" pageId="48" pageNumber="49">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="8B8136B0FFAD84548CABF8E0D918F82E" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
<taxonomicName id="4C3E4D33FFAD84548CABF8E0DA48F882" box="[199,374,1798,1824]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFAD84548CABF8E0DA48F882" box="[199,374,1798,1824]" italics="true" pageId="48" pageNumber="49">Th. panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFAD84548D10F8E0DA85F882" bold="true" box="[380,443,1798,1824]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A27957D9FFAD84548D10F8E0DA85F882" box="[380,443,1798,1824]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C3E4D33FFAD84548E20F8E0D9D2F882" authorityName="Dominguez" authorityYear="1986" box="[588,748,1798,1824]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bolivianus">
<emphasis id="B94AEAA2FFAD84548E20F8E0D9D2F882" box="[588,748,1798,1824]" italics="true" pageId="48" pageNumber="49">Th. bolivianus</emphasis>
</taxonomicName>
by the coloration of the fore femur: proximad of the multicolored apical band,
<taxonomicName id="4C3E4D33FFAD84548D33F8CCD930F8E6" box="[351,526,1834,1860]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFAD84548D33F8CCD930F8E6" box="[351,526,1834,1860]" italics="true" pageId="48" pageNumber="49">Th. panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFAD84548E7AF8CCD96BF8E6" bold="true" box="[534,597,1834,1860]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A27957D9FFAD84548E7AF8CCD96BF8E6" box="[534,597,1834,1860]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a small spot (
<figureCitation id="13052A35FFAD84548F72F8CCD841F8E6" box="[798,895,1834,1860]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="48" pageNumber="49">Fig. 134</figureCitation>
), while
<taxonomicName id="4C3E4D33FFAD84548FB5F8CCDF44F8E6" authorityName="Dominguez" authorityYear="1986" box="[985,1146,1834,1860]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="bolivianus">
<emphasis id="B94AEAA2FFAD84548FB5F8CCDF44F8E6" box="[985,1146,1834,1860]" italics="true" pageId="48" pageNumber="49">Th. bolivianus</emphasis>
</taxonomicName>
has a wide band reaching the multicolored apical band: «una banda amplia, que ocupa el 1/3 medio y que toca la banda apical, también grisnegruzca» (
<bibRefCitation id="EFAF4B41FFAD84548D75F894D927F82E" author="Dominguez, E." box="[281,537,1905,1932]" pageId="48" pageNumber="49" pagination="149 - 153" refId="ref69183" refString="Dominguez, E. (1986) Thraulodes bolivianus, una nueva especie de la familia Leptophlebiidae (Insecta: Ephemeroptera) de Bolivia. Acta Zoologica Lilloana, 38, 149 - 153." type="journal article" year="1986">Domínguez 1986: 150</bibRefCitation>
).
</paragraph>
<paragraph id="8B8136B0FFAD84548CABF870D971F85A" blockId="48.[151,1437,152,2040]" pageId="48" pageNumber="49">
<taxonomicName id="4C3E4D33FFAD84548CABF870DA46F812" box="[199,376,1942,1968]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFAD84548CABF870DA46F812" box="[199,376,1942,1968]" italics="true" pageId="48" pageNumber="49">Th. panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFAD84548DECF870DAFFF812" bold="true" box="[384,449,1942,1968]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A27957D9FFAD84548DECF870DAFFF812" box="[384,449,1942,1968]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C3E4D33FFAD84548E3BF870D9EDF812" box="[599,723,1942,1968]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="insular">
<emphasis id="B94AEAA2FFAD84548E3BF870D9EDF812" box="[599,723,1942,1968]" italics="true" pageId="48" pageNumber="49">Th. insular</emphasis>
</taxonomicName>
by the absence of a black spot on proximal part of middle and hind femora (
<taxonomicName id="4C3E4D33FFAD84548D41F85CDAE5F876" box="[301,475,1978,2004]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFAD84548D41F85CDAE5F876" box="[301,475,1978,2004]" italics="true" pageId="48" pageNumber="49">Th. panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFAD84548D8DF85CD921F876" bold="true" box="[481,543,1978,2004]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="A27957D9FFAD84548D8DF85CD921F876" box="[481,543,1978,2004]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has such a spot on fore femur only) and by orange color of the dorsal eyes (while in
<taxonomicName id="4C3E4D33FFAD84548CD8F838DA11F85A" box="[180,303,2014,2040]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="insular">
<emphasis id="B94AEAA2FFAD84548CD8F838DA11F85A" box="[180,303,2014,2040]" italics="true" pageId="48" pageNumber="49">Th. insular</emphasis>
</taxonomicName>
the dorsal eyes are gray).
</paragraph>
<paragraph id="8B8136B0FFAC84558CABFF7FD98FFE82" blockId="49.[151,1437,153,431]" pageId="49" pageNumber="50">
<taxonomicName id="4C3E4D33FFAC84558CABFF7FDA46FF11" box="[199,376,153,179]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFAC84558CABFF7FDA46FF11" box="[199,376,153,179]" italics="true" pageId="49" pageNumber="50">Th. panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFAC84558DEDFF7FDAFCFF11" bold="true" box="[385,450,153,179]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A27957D9FFAC84558DEDFF7FDAFCFF11" box="[385,450,153,179]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C3E4D33FFAC84558E37FF7FD842FF11" authority="Traver 1946" authorityName="Traver" authorityYear="1946" box="[603,892,153,179]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="B94AEAA2FFAC84558E37FF7FD9D4FF11" box="[603,746,153,179]" italics="true" pageId="49" pageNumber="50">Th. centralis</emphasis>
<bibRefCitation id="EFAF4B41FFAC84558E9EFF7FD842FF11" author="Traver, J. R." box="[754,892,153,179]" pageId="49" pageNumber="50" pagination="418 - 436" refId="ref72450" refString="Traver, J. R. (1946) Notes on Neotropical mayflies. Part I. Family Baetidae, subfamily Leptophlebiinae. Revista de Entomologia, 17, 418 - 436." type="journal article" year="1946">Traver 1946</bibRefCitation>
</taxonomicName>
by structure of genitalia; it differs by the white background of legs in imago and subimago (instead of yellowish orange in subimago of
<taxonomicName id="4C3E4D33FFAC845588EFFF5BDE2CFF75" authorityName="Traver" authorityYear="1946" box="[1155,1298,189,215]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="B94AEAA2FFAC845588EFFF5BDE2CFF75" box="[1155,1298,189,215]" italics="true" pageId="49" pageNumber="50">Th. centralis</emphasis>
</taxonomicName>
), by the abdominal tergum VII darker than VIII (instead of lighter in
<taxonomicName id="4C3E4D33FFAC84558F5DFF07D8FEFF59" authorityName="Traver" authorityYear="1946" box="[817,960,225,251]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="B94AEAA2FFAC84558F5DFF07D8FEFF59" box="[817,960,225,251]" italics="true" pageId="49" pageNumber="50">Th. centralis</emphasis>
</taxonomicName>
) and by white abdominal sterna (while in
<taxonomicName id="4C3E4D33FFAC84558CFBFEE3DA1BFEBD" authorityName="Traver" authorityYear="1946" box="[151,293,261,287]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="B94AEAA2FFAC84558CFBFEE3DA1BFEBD" box="[151,293,261,287]" italics="true" pageId="49" pageNumber="50">Th. centralis</emphasis>
</taxonomicName>
sterna VII-IX are dark red-brown).
</paragraph>
<paragraph id="8B8136B0FFAC84558CABFECFDF85FEE1" blockId="49.[151,1437,153,431]" box="[199,1211,297,323]" pageId="49" pageNumber="50">
Imagines of
<taxonomicName id="4C3E4D33FFAC84558D3FFECFD93DFEE1" box="[339,515,297,323]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="panamensis" status="sp. nov.">
<emphasis id="B94AEAA2FFAC84558D3FFECFD93DFEE1" box="[339,515,297,323]" italics="true" pageId="49" pageNumber="50">Th. panamensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFAC84558E66FECFD977FEE1" bold="true" box="[522,585,297,323]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A27957D9FFAC84558E66FECFD977FEE1" box="[522,585,297,323]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differ from other examined species by white scutellum.
</paragraph>
<paragraph id="8B8136B0FFAC84558CABFEABDA9AFE0D" blockId="49.[151,1437,153,431]" pageId="49" pageNumber="50">
Egg structure of
<taxonomicName id="4C3E4D33FFAC84558DEDFEABD90EFEC5" box="[385,560,333,359]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B94AEAA2FFAC84558DEDFEABD90EFEC5" box="[385,560,333,359]" italics="true" pageId="49" pageNumber="50">Th. panamensis</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C3E4D33FFAC84558F62FEABDFA9FEC5" authority="Dominguez 1987" authorityName="Dominguez" authorityYear="1987" box="[782,1175,333,359]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cochunaensis">
<emphasis id="B94AEAA2FFAC84558F62FEABD8EEFEC5" box="[782,976,333,359]" italics="true" pageId="49" pageNumber="50">Th. cochunaensis</emphasis>
<bibRefCitation id="EFAF4B41FFAC84558FBAFEABDFA9FEC5" author="Dominguez, E." box="[982,1175,333,359]" pageId="49" pageNumber="50" pagination="47 - 65" refId="ref69218" refString="Dominguez, E. (1987) El genero Thraulodes (Ephemeroptera: Leptophlebiidae) en la Republica Argentina. Acta Zoologica Lilloana, 39, 47 - 65." type="journal article" year="1987">Domínguez 1987</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFAF4B41FFAC845588C9FEABDE52FEC5" author="Dominguez, E." box="[1189,1388,333,359]" pageId="49" pageNumber="50" pagination="47 - 65" refId="ref69218" refString="Dominguez, E. (1987) El genero Thraulodes (Ephemeroptera: Leptophlebiidae) en la Republica Argentina. Acta Zoologica Lilloana, 39, 47 - 65." type="journal article" year="1987">Domínguez 1987</bibRefCitation>
: fig. 5.2), while these species have quite different structure of male imagines (genitalia of
<taxonomicName id="4C3E4D33FFAC84558806FE97DE10FE29" authorityName="Dominguez" authorityYear="1987" box="[1130,1326,369,395]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="cochunaensis">
<emphasis id="B94AEAA2FFAC84558806FE97DE10FE29" box="[1130,1326,369,395]" italics="true" pageId="49" pageNumber="50">Th. cochunaensis</emphasis>
</taxonomicName>
resemble that of
<taxonomicName id="4C3E4D33FFAC84558C8AFE73DA73FE0D" box="[230,333,405,431]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
<emphasis id="B94AEAA2FFAC84558C8AFE73DA73FE0D" box="[230,333,405,431]" italics="true" pageId="49" pageNumber="50">Th. niger</emphasis>
</taxonomicName>
<emphasis id="B94AEAA2FFAC84558D38FE73DAAAFE0D" bold="true" box="[340,404,405,431]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A27957D9FFAC84558D38FE73DAAAFE0D" box="[340,404,405,431]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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