192 lines
19 KiB
XML
192 lines
19 KiB
XML
<document id="B212FEE88116D08A85F52E82735C7027" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. & Liang, Hongbin" docDate="2015" docId="039787812E46FFCA94FD4BBFEDDCEB10" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) roninus Bates 1873" docType="treatment" docVersion="7" lastPageNumber="12" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="12" updateTime="1698562432667" updateUser="plazi">
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<mods:titleInfo id="A21F9A1C3B2D3785F9D8AC90FF565962">
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<mods:title id="E02BE8F1C267758B280B05A7D0EA67E5">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
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<mods:name id="2A9705DB41C063C7C0B60D414E16E213" type="personal">
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<mods:namePart id="9154F754B50FF78EC16FE75CC5BC4EE6">Kataev, Boris M.</mods:namePart>
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<mods:roleTerm id="51ABC5F127E9C2529086492E5E850E0E">Author</mods:roleTerm>
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<mods:namePart id="D84F15D03FD19A1ADC4FEC145B4CDA7B">Liang, Hongbin</mods:namePart>
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<mods:typeOfResource id="5261713BF96A54F88EC0141676FFC6F0">text</mods:typeOfResource>
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<mods:title id="706563B309A0B4554D7F2E5EFA06DD00">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B798FC1C7B8005434421C3D73881AFC7">
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<mods:date id="C6F345B62EE876214AEA47AB53244523">2015</mods:date>
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<mods:detail id="3DD4C1BD7F2FB82F115E463704E37EEF" type="volume">
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<mods:number id="D9E352BF42921C7F859AA7037ABB8EA2">3920</mods:number>
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<mods:detail id="C05C9BA6304257B339059CE163966AE7" type="issue">
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<mods:number id="31E9952A68891E278A49CBF0993261A7">1</mods:number>
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<mods:identifier id="A03D11965116E523B76BA42AB91F41C3" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
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<mods:identifier id="80891675039743907F1E319B3E72675A" type="GBIF-Dataset">7289c86d-0ed1-4e93-b7b3-bc596042d854</mods:identifier>
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<treatment id="039787812E46FFCA94FD4BBFEDDCEB10" ID-DOI="http://doi.org/10.5281/zenodo.6114252" ID-GBIF-Taxon="127671658" ID-Zenodo-Dep="6114252" LSID="urn:lsid:plazi:treatment:039787812E46FFCA94FD4BBFEDDCEB10" httpUri="http://treatment.plazi.org/id/039787812E46FFCA94FD4BBFEDDCEB10" lastPageNumber="12" pageId="11" pageNumber="12">
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<subSubSection id="C324651C2E46FFCA94FD4BBFE800EED5" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8B8136972E46FFCA94FD4BBFEBF8EEB4" blockId="11.[151,735,404,463]" box="[151,735,404,430]" pageId="11" pageNumber="12">
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<heading id="D0C981FB2E46FFCA94FD4BBFEBF8EEB4" bold="true" box="[151,735,404,430]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
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<taxonomicName id="4C3E4D142E46FFCA94FD4BBFEBF8EEB4" ID-CoL="6LDTM" authority="Bates, 1873" authorityName="Bates" authorityYear="1873" box="[151,735,404,430]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="roninus" subGenus="Pseudoophonus">
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||
<emphasis id="B94AEA852E46FFCA94FD4BBFEBF8EEB4" bold="true" box="[151,735,404,430]" pageId="11" pageNumber="12">
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<emphasis id="B94AEA852E46FFCA94FD4BBFE82EEEB4" bold="true" box="[151,265,404,430]" italics="true" pageId="11" pageNumber="12">Harpalus</emphasis>
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||
(
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<emphasis id="B94AEA852E46FFCA95704BBFE8FDEEB4" bold="true" box="[282,474,404,430]" italics="true" pageId="11" pageNumber="12">Pseudoophonus</emphasis>
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)
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<emphasis id="B94AEA852E46FFCA95814BBFEB6EEEB4" bold="true" box="[491,585,404,430]" italics="true" pageId="11" pageNumber="12">roninus</emphasis>
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<bibRefCitation id="EFAF4B662E46FFCA963B4BBFEBF8EEB4" author="Bates" box="[593,735,404,430]" pageId="11" pageNumber="12" refString="Bates, H. W. (1873) On the Geodephagous Coleoptera of Japan. The Transactions of the Royal Entomological Society of London, 1873, 219 - 322." type="journal article" year="1873">Bates, 1873</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B8136972E46FFCA94FD4B9DE800EED5" blockId="11.[151,735,404,463]" box="[151,295,438,463]" pageId="11" pageNumber="12">
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(
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<figureCitation id="13052A122E46FFCA94F54B9DE839EED5" box="[159,286,438,463]" captionStart="FIGURES 19 – 23" captionStartId="10.[151,263,1623,1646]" captionTargetBox="[188,1401,420,1593]" captionTargetId="figure@10.[176,1411,409,1602]" captionTargetPageId="10" captionText="FIGURES 19 – 23. Harpalus, median lobe of aedeagus (19, apical portion). 19 – 21, H. ussuriensis (19, Maritime Terr.; 20 – 21, Qinghai); 22 – 23, H. roninus (Heilongjiang). 19 – 20, 22, view from the left side; 21, 23, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287889/files/figure.png" pageId="11" pageNumber="12">Figs 22–23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C324651C2E46FFCA94FD4BD4EA7EED0E" box="[151,857,510,532]" pageId="11" pageNumber="12" type="reference_group">
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<paragraph id="8B8136972E46FFCA94FD4BD4EA7EED0E" blockId="11.[151,857,510,532]" box="[151,857,510,532]" pageId="11" pageNumber="12">
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<treatmentCitationGroup id="AB2E11B92E46FFCA94FD4BD4EA7EED0E" box="[151,857,510,532]" pageId="11" pageNumber="12">
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<taxonomicName id="4C3E4D142E46FFCA94FD4BD4E8DAED0E" ID-CoL="6LDTM" authority="Bates, 1873: 260" authorityName="Bates" authorityPageNumber="260" authorityYear="1873" box="[151,509,510,532]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="roninus">
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<emphasis id="B94AEA852E46FFCA94FD4BD4E86DED0E" box="[151,330,511,532]" italics="true" pageId="11" pageNumber="12">Harpalus roninus</emphasis>
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<treatmentCitation id="0A9F10862E46FFCA95254BD4E8DAED0E" author="Bates" box="[335,509,510,532]" page="260" pageId="11" pageNumber="12" year="1873">
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<bibRefCitation id="EFAF4B662E46FFCA95254BD4E8EDED0E" author="Bates" box="[335,458,510,532]" pageId="11" pageNumber="12" refString="Bates, H. W. (1873) On the Geodephagous Coleoptera of Japan. The Transactions of the Royal Entomological Society of London, 1873, 219 - 322." type="journal article" year="1873">Bates, 1873</bibRefCitation>
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: 260
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</treatmentCitation>
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||
</taxonomicName>
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.
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<typeStatus id="548588352E46FFCA966D4BD4EB1EED0E" box="[519,569,511,532]" pageId="11" pageNumber="12">Type</typeStatus>
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locality: "Nagasaki",
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<collectingCountry id="F32976072E46FFCA97724BD4EA72ED0E" box="[792,853,511,532]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C324651C2E46FFCA94FD486AEA11EDDC" pageId="11" pageNumber="12" type="materials_examined">
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<paragraph id="8B8136972E46FFCA94FD486AEA58ED43" blockId="11.[151,1437,577,1034]" box="[151,895,577,602]" pageId="11" pageNumber="12">
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<emphasis id="B94AEA852E46FFCA94FD486AE81AED40" bold="true" box="[151,317,577,602]" pageId="11" pageNumber="12">
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<typeStatus id="548588352E46FFCA94FD486AE9F5ED40" box="[151,210,577,602]" pageId="11" pageNumber="12">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="548588352E46FFCA9526486AE890ED43" box="[332,439,577,601]" pageId="11" pageNumber="12" type="syntype">Syntypes</typeStatus>
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:
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<specimenCount id="9D38FD1E2E46FFCA95AB486AE8CAED43" box="[449,493,577,601]" pageId="11" pageNumber="12" type="generic">1 ♂</specimenCount>
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, 1 ♀, labeled "Nagasaki" (
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<collectionCode id="ED2FAE522E46FFCA977D486AEA54ED43" LSID="urn:lsid:biocol.org:col:34988" box="[791,883,577,601]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="12">MNHN</collectionCode>
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).
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</paragraph>
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<paragraph id="8B8136972E46FFCA94AD484EEA11EDDC" blockId="11.[151,1437,577,1034]" pageId="11" pageNumber="12">
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<emphasis id="B94AEA852E46FFCA94AD484EEB1AED64" bold="true" box="[199,573,613,638]" pageId="11" pageNumber="12">Additional material examined</emphasis>
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. More than
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<specimenCount id="9D38FD1E2E46FFCA96B4484DEAABED64" box="[734,908,613,638]" pageId="11" pageNumber="12" type="generic">170 specimens</specimenCount>
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(133 from
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<collectingCountry id="F32976072E46FFCA9048484EED4EED64" box="[1058,1129,613,638]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
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), including the following specimens from Jilin and Hubei.
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<collectingCountry id="F32976072E46FFCA966C48A2EB68EDB8" box="[518,591,649,674]" name="China" pageId="11" pageNumber="12">
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||
<emphasis id="B94AEA852E46FFCA966C48A2EB68EDB8" bold="true" box="[518,591,649,674]" pageId="11" pageNumber="12">China</emphasis>
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</collectingCountry>
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.
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<collectionCode id="ED2FAE522E46FFCA963648A1EB84EDBB" box="[604,675,650,673]" pageId="11" pageNumber="12">JILIN</collectionCode>
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:
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<specimenCount id="9D38FD1E2E46FFCA96C748A1EBFEEDBB" box="[685,729,650,674]" pageId="11" pageNumber="12" type="generic">1 ♂</specimenCount>
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, Changchun Co.,
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<date id="FF8010572E46FFCA97F548A1ED34EDB8" box="[927,1043,650,674]" pageId="11" pageNumber="12" value="1982-09-09">9.IX.1982</date>
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, Mao Zenghua leg. (
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<collectionCode id="ED2FAE522E46FFCA909248A1EC0BEDB8" box="[1272,1324,650,674]" pageId="11" pageNumber="12">IOZ</collectionCode>
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).
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<collectionCode id="ED2FAE522E46FFCA915548A1ECBEEDBB" box="[1343,1433,650,673]" pageId="11" pageNumber="12">HUBEI</collectionCode>
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: 1 ♀, Xinshan,
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<quantity id="4CC69B722E46FFCA95504885E8B6EDDF" box="[314,401,686,710]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="12" unit="m" value="1300.0">1300 m</quantity>
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,
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<date id="FF8010572E46FFCA95F74885EB01EDDC" box="[413,550,686,710]" pageId="11" pageNumber="12" value="1980-07-29">
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29.VII.
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<quantity id="4CC69B722E46FFCA95814885EB01EDDC" box="[491,550,686,710]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="11" pageNumber="12" unit="m" value="1980.0">1980</quantity>
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</date>
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m, Yu Peiyu leg. (
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<collectionCode id="ED2FAE522E46FFCA969C4885EA0DEDDC" box="[758,810,686,710]" pageId="11" pageNumber="12">IOZ</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C324651C2E46FFCA94AD48FAED72EC2B" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="8B8136972E46FFCA94AD48FAED72EC2B" blockId="11.[151,1437,577,1034]" pageId="11" pageNumber="12">
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<emphasis id="B94AEA852E46FFCA94AD48FAE81DEDF0" bold="true" box="[199,314,721,746]" pageId="11" pageNumber="12">Diagnosis</emphasis>
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. This species is similar in external characters to
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<taxonomicName id="4C3E4D142E46FFCA970B48F9ED27EDF3" box="[865,1024,722,745]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ussuriensis">
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<emphasis id="B94AEA852E46FFCA970B48F9ED27EDF3" box="[865,1024,722,745]" italics="true" pageId="11" pageNumber="12">H. ussuriensis</emphasis>
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</taxonomicName>
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, but distinctly differs from the latter in black legs. Internal sac of aedeagus (
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<figureCitation id="13052A122E46FFCA962548DEEBE8EC14" box="[591,719,757,782]" captionStart="FIGURES 19 – 23" captionStartId="10.[151,263,1623,1646]" captionTargetBox="[188,1401,420,1593]" captionTargetId="figure@10.[176,1411,409,1602]" captionTargetPageId="10" captionText="FIGURES 19 – 23. Harpalus, median lobe of aedeagus (19, apical portion). 19 – 21, H. ussuriensis (19, Maritime Terr.; 20 – 21, Qinghai); 22 – 23, H. roninus (Heilongjiang). 19 – 20, 22, view from the left side; 21, 23, dorsal view. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287889/files/figure.png" pageId="11" pageNumber="12">Figs 22–23</figureCitation>
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) is armed with a small medial spiny patch on the left side of the median lobe and a separate medium-sized spine apically. Body length
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<quantity id="4CC69B722E46FFCA97C14932ED76EC2B" box="[939,1105,793,818]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.01" metricValueMin="1.69" pageId="11" pageNumber="12" unit="mm" value="18.5" valueMax="20.1" valueMin="16.9">16.9–20.1 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C324651C2E46FFCA94AD4916EDDCEB10" pageId="11" pageNumber="12" type="distribution">
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||
<paragraph id="8B8136972E46FFCA94AD4916EA80EC84" blockId="11.[151,1437,577,1034]" pageId="11" pageNumber="12">
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||
<emphasis id="B94AEA852E46FFCA94AD4916E87EEC4C" bold="true" box="[199,345,829,854]" pageId="11" pageNumber="12">Distribution</emphasis>
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||
.
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<collectingCountry id="F32976072E46FFCA95034916E897EC4C" box="[361,432,829,854]" name="China" pageId="11" pageNumber="12">China</collectingCountry>
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: mainly in the north-eastern part. It was recorded from Heilongjiang, Liaoning, Jiangsu, Shanghai, Shanxi, Shaanxi, Sichuan (
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<bibRefCitation id="EFAF4B662E46FFCA962F4949EA07EC60" author="Kataev" box="[581,800,865,890]" pageId="11" pageNumber="12" refString="Kataev, B. M., Wrase, D. W. & Ito, N. (2003) Harpalina. In: Lobl, L. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
|
||
Kataev
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<emphasis id="B94AEA852E46FFCA96CA4949EBF3EC63" box="[672,724,865,889]" italics="true" pageId="11" pageNumber="12">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
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||
), Jilin, Hebei, Henan, Hubei, Hunan, and
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<collectingCountry id="F32976072E46FFCA9162494AEC7DEC60" box="[1288,1370,865,890]" name="Taiwan" pageId="11" pageNumber="12">Taiwan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFAF4B662E46FFCA91064949E9F3EC84" author="Hua" pageId="11" pageNumber="12" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua 2002</bibRefCitation>
|
||
). We provided here the additional material from Jilin and Hubei.
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||
</paragraph>
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<paragraph id="8B8136972E46FFCA94AD4982EDDCEB10" blockId="11.[151,1437,577,1034]" pageId="11" pageNumber="12">
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The species is also known from Eastern
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<collectingCountry id="F32976072E46FFCA96A94982EA29ECD8" box="[707,782,937,962]" name="Russia" pageId="11" pageNumber="12">Russia</collectingCountry>
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||
(Buryatia, Amur Province, Khabarovsk and Maritime Territories), North and
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<collectingCountry id="F32976072E46FFCA95F549E6EB09ECFF" box="[415,558,973,998]" name="South Korea" pageId="11" pageNumber="12">South Korea</collectingCountry>
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including Jejudo, and
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<collectingCountry id="F32976072E46FFCA975B49E5EA56ECFF" box="[817,881,974,997]" name="Japan" pageId="11" pageNumber="12">Japan</collectingCountry>
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(Honshu, Kyushu, and Tsushima Islands) (
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<bibRefCitation id="EFAF4B662E46FFCA913449E6E9F5EB10" author="Habu" pageId="11" pageNumber="12" refString="Habu, A. (1973) Carabidae: Harpalini (Insecta, Coleoptera). Fauna Japonica. Keigaku Publishing Co., Tokio, xiii + 430 pp." type="book" year="1973">Habu 1973</bibRefCitation>
|
||
;
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<bibRefCitation id="EFAF4B662E46FFCA94B749DAE87BEB10" author="Lafer" box="[221,348,1009,1034]" pageId="11" pageNumber="12" refString="Lafer, G. Sh. (1989) Fam. Carabidae-The ground-beetles. In: Lehr, P. A. (Ed.), Opredelitel' nasekomykh Dal'nego Vostoka SSSR v shesti tomakh. Tom III. Zhestkokrylye, ili zhuki. Chast' 1. Nauka, Leningrad, pp. 70 - 222. [in Russian]" type="journal article" year="1989">Lafer 1989</bibRefCitation>
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;
|
||
<bibRefCitation id="EFAF4B662E46FFCA950249DAEBA9EB10" author="Kryzhanovskij" box="[360,654,1009,1034]" pageId="11" pageNumber="12" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the groundbeetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers, Sofia - Moscow, 272 pp." type="book" year="1995">
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Kryzhanovskij
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;
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<bibRefCitation id="EFAF4B662E46FFCA96F049D9EA0CEB10" author="Kataev" box="[666,811,1010,1034]" pageId="11" pageNumber="12" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
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;
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<bibRefCitation id="EFAF4B662E46FFCA975D49D9ED20EB10" author="Kataev" box="[823,1031,1009,1034]" pageId="11" pageNumber="12" refString="Kataev, B. M., Wrase, D. W. & Ito, N. (2003) Harpalina. In: Lobl, L. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
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<bibRefCitation id="EFAF4B662E46FFCA901349DAEDC8EB10" author="Paik" box="[1145,1263,1009,1034]" pageId="11" pageNumber="12" refString="Moon, Ch. - S. & Paik, J. - Ch. (2006) Notes on Harpalini (Coleoptera, Carabidae) from Korea (1 - 3). Korean Journal of Soil Zoology, 11 (1 - 2), 30 - 84." type="journal article" year="2006">Paik 2006</bibRefCitation>
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).
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||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |