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<document id="9324E44B62A34B51396E57569650B71C" ID-DOI="10.5281/zenodo.199230" ID-GBIF-Dataset="401871e9-df9f-48d2-bf20-554170b12052" ID-ISSN="1175-5326" ID-Zenodo-Dep="199230" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460332324328" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2010" docId="039705178029041EFF02FE89FE91C476" docLanguage="en" docName="zt02677p048.pdf" docOrigin="Zootaxa 2677" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cavoportunus dubius Laurie 1906, comb. nov." docType="treatment" docVersion="9" lastPageNumber="45" masterDocId="FFAE7D6F802A0416FF95FFDBFC30C554" masterDocTitle="A new genus of the family Portunidae (Crustacea: Decapoda: Brachyura) and the identity of Portunus (Cycloachelous) yoronensis Sakai, 1974" masterLastPageNumber="48" masterPageNumber="38" pageNumber="41" updateTime="1698258926458" updateUser="plazi">
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<mods:title id="D2393EEFC25290530627A51AF83D55AA">A new genus of the family Portunidae (Crustacea: Decapoda: Brachyura) and the identity of Portunus (Cycloachelous) yoronensis Sakai, 1974</mods:title>
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<paragraph id="8B81B40180290415FF02FE89FED0C438" blockId="3.[151,736,338,399]" box="[151,736,338,364]" pageId="3" pageNumber="41">
<heading id="D0C9036D80290415FF02FE89FED0C438" bold="true" box="[151,736,338,364]" fontSize="11" level="1" pageId="3" pageNumber="41" reason="1">
<emphasis id="B94A681380290415FF02FE89FED0C438" bold="true" box="[151,736,338,364]" pageId="3" pageNumber="41">
<taxonomicName id="4C3ECF8280290415FF02FE89FE63C438" ID-CoL="RYDG" authority="Laurie, 1906" authorityName="Laurie" authorityYear="1906" box="[151,595,338,364]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="dubius" status="comb. nov.">
<emphasis id="B94A681380290415FF02FE89FDABC438" bold="true" box="[151,411,338,364]" italics="true" pageId="3" pageNumber="41">Cavoportunus dubius</emphasis>
(
<bibRefCitation id="EFAFC9F080290415FE3FFE89FE7AC438" author="Laurie" box="[426,586,338,364]" pageId="3" pageNumber="41" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie, 1906</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A279D56880290415FDCFFE89FED0C438" box="[602,736,338,364]" pageId="3" pageNumber="41">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B81B40180290415FF02FEAEFD27C4DB" blockId="3.[151,736,338,399]" box="[151,279,373,399]" pageId="3" pageNumber="41">
(
<figureCitation id="1305A88480290415FF0AFEAEFD3FC4DB" box="[159,271,373,399]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,257,1484,1508]" captionStartId-1="5.[151,255,1926,1950]" captionStartId-2="6.[151,256,1604,1628]" captionStartId-3="7.[151,255,1387,1411]" captionTargetBox-0="[151,1436,714,1462]" captionTargetBox-1="[228,1359,194,1913]" captionTargetBox-2="[151,1436,194,1575]" captionTargetBox-3="[187,1406,238,1346]" captionTargetId-0="figure@3.[151,1436,714,1463]" captionTargetId-1="figure@5.[228,1359,194,1915]" captionTargetId-2="figure@6.[151,1436,194,1583]" captionTargetId-3="figure@7.[170,1417,194,1365]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Live colour. Cavoportunus dubius (Laurie, 1906). Female (13.0 × 10.2 mm) (NSMT-KY- 08 - 15), dorsal overall view. Scales = 1.0 mm. (Photograph: H. Komatsu)." captionText-1="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." captionText-2="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." captionText-3="FIGURE 4. Cavoportunus dubius (Laurie, 1906). A, lectotype male, (10.2 × 8.1 mm) (NHM 1907.5.22.320); B-F. male, (18.5 × 14.6 mm) (ZRC 2001.0627). A, carapace; B, male abdomen; C, male thoracic sternum with intact right G 1; D, different views of left G 1; E, right G 2; F, distal part of right G 2. Scales: A-E = 1.0 mm, F = 0.2 mm." httpUri-0="https://zenodo.org/record/199231/files/figure.png" httpUri-1="https://zenodo.org/record/199232/files/figure.png" httpUri-2="https://zenodo.org/record/199233/files/figure.png" httpUri-3="https://zenodo.org/record/199234/files/figure.png" pageId="3" pageNumber="41">Figs. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324E78A80290415FF02FE1BFD1EC7C3" pageId="3" pageNumber="41" type="reference_group">
<paragraph id="8B81B40180290415FF02FE1BFEFAC483" blockId="3.[151,1436,448,663]" box="[151,714,448,471]" pageId="3" pageNumber="41">
<treatmentCitationGroup id="AB2E932F80290415FF02FE1BFEFAC483" box="[151,714,448,471]" pageId="3" pageNumber="41">
<taxonomicName id="4C3ECF8280290415FF02FE1BFEB1C483" ID-CoL="47475" authority="Laurie, 1906: 416" authorityName="Laurie" authorityPageNumber="416" authorityYear="1906" box="[151,641,448,471]" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="dubia" subGenus="Achelous">
<emphasis id="B94A681380290415FF02FE1BFCCBC483" box="[151,251,448,471]" italics="true" pageId="3" pageNumber="41">Neptunus</emphasis>
(
<emphasis id="B94A681380290415FE9EFE1BFD5DC483" box="[267,365,448,471]" italics="true" pageId="3" pageNumber="41">Achelous</emphasis>
)
<emphasis id="B94A681380290415FEE9FE1BFD89C483" box="[380,441,448,471]" italics="true" pageId="3" pageNumber="41">dubia</emphasis>
<treatmentCitation id="0A9F921080290415FE55FE1BFEB1C483" author="Laurie" box="[448,641,448,471]" page="416" pageId="3" pageNumber="41" year="1906">
<bibRefCitation id="EFAFC9F080290415FE55FE1BFE7CC483" author="Laurie" box="[448,588,448,471]" pageId="3" pageNumber="41" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie, 1906</bibRefCitation>
: 416
</treatmentCitation>
</taxonomicName>
, fig. 9.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B81B40180290415FF02FE3BFD09C703" blockId="3.[151,1436,448,663]" pageId="3" pageNumber="41">
<treatmentCitationGroup id="AB2E932F80290415FF02FE3BFD09C703" pageId="3" pageNumber="41">
<taxonomicName id="4C3ECF8280290415FF02FE3BFDF0C4A3" ID-CoL="4M2JG" box="[151,448,480,503]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="dubius" subGenus="Achelous">
<emphasis id="B94A681380290415FF02FE3BFCC9C4A3" box="[151,249,480,503]" italics="true" pageId="3" pageNumber="41">Portunus</emphasis>
(
<emphasis id="B94A681380290415FE9DFE3BFD5AC4A3" box="[264,362,480,503]" italics="true" pageId="3" pageNumber="41">Achelous</emphasis>
)
<emphasis id="B94A681380290415FEECFE3BFDF0C4A3" box="[377,448,480,503]" italics="true" pageId="3" pageNumber="41">dubius</emphasis>
</taxonomicName>
<treatmentCitation id="0A9F921080290415FE74FE3BFF28C4A3" author="Stephenson" box="[481,792,480,503]" page="20" pageId="3" pageNumber="41" year="1967">
<bibRefCitation id="EFAFC9F080290415FE74FE3BFEC1C4A3" author="Stephenson" box="[481,753,480,503]" pageId="3" pageNumber="41" refString="Stephenson, W. &amp; Rees, M. (1967) Some portunid crabs from the Pacific and Indian Oceans in the collections of the Smithsonian Institution. Proceedings of the United States National Museum, 120, 1 - 114." type="journal article" year="1967">Stephenson &amp; Rees 1967</bibRefCitation>
: 20
</treatmentCitation>
, fig 3. —
<treatmentCitation id="0A9F921080290415FC17FE3BF84EC4A3" author="Stephenson" box="[898,1150,480,503]" page="135" pageId="3" pageNumber="41" year="1972">
<bibRefCitation id="EFAFC9F080290415FC17FE3BF878C4A3" author="Stephenson" box="[898,1096,480,503]" pageId="3" pageNumber="41" refString="Stephenson, W. (1972 a) An annotated check list and key to the Indo-West-Pacific swimming crabs (Crustacea: Decapoda: Portunidae). Bulletin of the Royal Society of New Zealand, 10, 1 - 64." type="journal article" year="1972" yearSuffix="a">Stephenson 1972a</bibRefCitation>
: 135
</treatmentCitation>
. —
<treatmentCitation id="0A9F921080290415FB3CFE3BF9A7C4A3" author="Stephenson" box="[1193,1431,480,503]" page="15" pageId="3" pageNumber="41" year="1972">
<bibRefCitation id="EFAFC9F080290415FB3CFE3BF941C4A3" author="Stephenson" box="[1193,1393,480,503]" pageId="3" pageNumber="41" refString="Stephenson, W. (1972 b) Portunid crabs from the Indo-West-Pacific and Western America in the Zoological Museum, Copenhagen (Decapoda, Brachyura, Portunidae). Steenstrupia, 2 (9), 127 - 156." type="journal article" year="1972" yearSuffix="b">Stephenson 1972b</bibRefCitation>
: 15
</treatmentCitation>
, 38.
<treatmentCitation id="0A9F921080290415FE90FDDBFDD9C743" author="Stephenson" box="[261,489,512,535]" page="16" pageId="3" pageNumber="41" year="1976">
<bibRefCitation id="EFAFC9F080290415FE90FDDBFDF0C743" author="Stephenson" box="[261,448,512,535]" pageId="3" pageNumber="41" refString="Stephenson, W. (1976) Notes on Indo-West-Pacific portunids (Decapoda, Portunidae) in the Smithsonian Institution. Crustaceana, 31 (1), 11 - 26." type="journal article" year="1976">Stephenson 1976</bibRefCitation>
: 16
</treatmentCitation>
.
<treatmentCitation id="0A9F921080290415FD9CFDDBFEF8C743" author="Moosa" box="[521,712,512,535]" page="143" pageId="3" pageNumber="41" year="1981">
<bibRefCitation id="EFAFC9F080290415FD9CFDDBFEA3C743" author="Moosa" box="[521,659,512,535]" pageId="3" pageNumber="41" refString="Moosa, M. K. (1981) Portunus (Lupocycloporus) wilsoni, a new portunid from the Indo-Australian region (Brachyura, Portunidae). Crustaceana, 40 (1), 26 - 30." type="journal article" year="1981">Moosa 1981</bibRefCitation>
: 143
</treatmentCitation>
, 147. —
<treatmentCitation id="0A9F921080290415FCB8FDDBF83AC743" author="Crosnier" box="[813,1034,512,535]" page="401" pageId="3" pageNumber="41" year="1984">
<bibRefCitation id="EFAFC9F080290415FCB8FDDBFFE7C743" author="Crosnier" box="[813,983,512,535]" pageId="3" pageNumber="41" refString="Crosnier, A. (1984) Sur quelques Portunidae (Crustacea Decapoda Brachyura) des iles Seychelles. Bulletin du Museum national d'Histoire naturelle, Section A, Zoologie, Biologie et Ecologie Animales, Paris, 4 e serie, 6 (2), 397 - 419." type="journal article" year="1984" yearSuffix="b">Crosnier 1984b</bibRefCitation>
: 401
</treatmentCitation>
; 1985: 33, figs. 1a-e; 2002: 404. —
<treatmentCitation id="0A9F921080290415FF52FDFBFD91C763" author="Zarenkov" box="[199,417,544,567]" page="115" pageId="3" pageNumber="41" year="1994">
<bibRefCitation id="EFAFC9F080290415FF52FDFBFD5DC763" author="Zarenkov" box="[199,365,544,567]" pageId="3" pageNumber="41" refString="Zarenkov, N. A. (1994) Crabs from seamounts of the western part of the Indian Ocean [in Russian]. Trudy Instituta Okeanologii imeni P. P. Shirshova, 129, 97 - 125." type="journal article" year="1994">Zarenkov 1994</bibRefCitation>
: 115
</treatmentCitation>
. —
<treatmentCitation id="0A9F921080290415FE59FDFBFEBAC763" author="Moosa" box="[460,650,544,567]" page="521" pageId="3" pageNumber="41" year="1996">
<bibRefCitation id="EFAFC9F080290415FE59FDFBFE64C763" author="Moosa" box="[460,596,544,567]" pageId="3" pageNumber="41" refString="Moosa, M. K. (1996) Crustacea Decapoda: deep-water swimming crabs from the south-west Pacific, particularly New Caledonia (Brachyura, Portunidae). In: Crosnier, A. (Ed.) Resultats des Campagnes MUSORSTOM, vol. 15. Memoires du Museum national d'Histoire naturelle (Paris), 168., pp. 503 - 530." type="book chapter" year="1996">Moosa 1996</bibRefCitation>
: 521
</treatmentCitation>
. —
<treatmentCitation id="0A9F921080290415FD20FDFBFF44C763" author="Poupin" box="[693,884,544,567]" page="31" pageId="3" pageNumber="41" year="1996">
<bibRefCitation id="EFAFC9F080290415FD20FDFBFF7CC763" author="Poupin" box="[693,844,544,567]" pageId="3" pageNumber="41" refString="Poupin, J. (1996) Crustacea Decapoda of French Polynesia (Astacidea, Palinuridea, Anomura, Brachyura). Atoll Research Bulletin, 442, 1 - 114." type="journal article" year="1996" yearSuffix="a">Poupin 1996a</bibRefCitation>
: 31
</treatmentCitation>
. —
<treatmentCitation id="0A9F921080290415FC35FDFBF937C763" author="Neumann" box="[928,1287,544,567]" page="19" pageId="3" pageNumber="41" year="1999">
<bibRefCitation id="EFAFC9F080290415FC35FDFBF8EFC763" author="Neumann" box="[928,1247,544,567]" pageId="3" pageNumber="41" refString="Neumann, V. &amp; Spiridonov, V. A. (1999) Shallow water crabs from the Western Indian Ocean: Portunoidea and Xanthoidea excluding Pilumnidae (Crustacea, Decapoda, Brachyura). Tropical Zoology, 12, 9 - 66." type="journal article" year="1999">Neumann &amp; Spiridonov 1999</bibRefCitation>
: 19
</treatmentCitation>
. —– Ng
<emphasis id="B94A681380290415FAF2FDFBF9A5C763" box="[1383,1429,544,567]" italics="true" pageId="3" pageNumber="41">et al</emphasis>
. 2008: 151.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B81B40180290415FF02FDBBFD1EC7C3" blockId="3.[151,1436,448,663]" pageId="3" pageNumber="41">
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<taxonomicName id="4C3ECF8280290415FF02FDBBFED2C723" ID-CoL="4M2K3" authority="Sakai, 1974: 90" authorityName="Sakai" authorityPageNumber="90" authorityYear="1974" box="[151,738,608,631]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="yoronensis" subGenus="Cycloachelous">
<emphasis id="B94A681380290415FF02FDBBFCC9C723" box="[151,249,608,631]" italics="true" pageId="3" pageNumber="41">Portunus</emphasis>
(
<emphasis id="B94A681380290415FE9FFDBBFD97C723" box="[266,423,608,631]" italics="true" pageId="3" pageNumber="41">Cycloachelous</emphasis>
)
<emphasis id="B94A681380290415FE2CFDBBFE1BC723" box="[441,555,608,631]" italics="true" pageId="3" pageNumber="41">yoronensis</emphasis>
<treatmentCitation id="0A9F921080290415FDA1FDBBFED2C723" author="Sakai" box="[564,738,608,631]" page="90" pageId="3" pageNumber="41" year="1974">
<bibRefCitation id="EFAFC9F080290415FDA1FDBBFE87C723" author="Sakai" box="[564,695,608,631]" pageId="3" pageNumber="41" refString="Sakai, T. (1974) Notes from the carcinological fauna of Japan (V). Researches on Crustacea, 6, 86 - 102." type="journal article" year="1974">Sakai, 1974</bibRefCitation>
: 90
</treatmentCitation>
</taxonomicName>
. —
<treatmentCitation id="0A9F921080290415FC84FDBBFFF5C723" author="Sakai" box="[785,965,608,631]" page="349" pageId="3" pageNumber="41" year="1976">
<bibRefCitation id="EFAFC9F080290415FC84FDBBFFBEC723" author="Sakai" box="[785,910,608,631]" pageId="3" pageNumber="41" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxxix + 773 pp.; vol. 2 [Japanese text], 461 pp.; vol. 3 [plates], 16 pp., pls. 1 - 251." type="book chapter" year="1976">Sakai 1976</bibRefCitation>
: 349
</treatmentCitation>
, figs. 189a-d. — Ng
<emphasis id="B94A681380290415FB25FDBBF8D1C723" box="[1200,1249,608,631]" italics="true" pageId="3" pageNumber="41">et al</emphasis>
. 2008: 151. (See Remarks)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF41E48980290415FF02FA17FF39C357" httpUri="https://zenodo.org/record/199231/files/figure.png" pageId="3" pageNumber="41" targetBox="[151,1436,714,1462]" targetPageId="3">
<paragraph id="8B81B40180290415FF02FA17FF39C357" blockId="3.[151,1436,1484,1539]" pageId="3" pageNumber="41">
<emphasis id="B94A681380290415FF02FA17FD2DC0B0" bold="true" box="[151,285,1484,1508]" pageId="3" pageNumber="41">FIGURE 1.</emphasis>
Live colour.
<taxonomicName id="4C3ECF8280290415FE21FA16FF7EC0B7" authority="Laurie, 1906" authorityName="Laurie" authorityYear="1906" box="[436,846,1484,1508]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A681380290415FE21FA16FE92C0B0" box="[436,674,1485,1508]" italics="true" pageId="3" pageNumber="41">Cavoportunus dubius</emphasis>
(Laurie, 1906)
</taxonomicName>
. Female (13.0 ×10.2 mm) (NSMT-KY-08-15), dorsal overall view. Scales = 1.0 mm. (Photograph: H. Komatsu).
</paragraph>
</caption>
<subSubSection id="C324E78A80290412FF02F9EAF858C476" lastPageId="4" lastPageNumber="42" pageId="3" pageNumber="41" type="materials_examined">
<paragraph id="8B81B40180290412FF02F9EAF858C476" blockId="3.[151,1437,1585,2022]" lastBlockId="4.[151,1437,151,2010]" lastPageId="4" lastPageNumber="42" pageId="3" pageNumber="41">
<emphasis id="B94A681380290415FF02F9EAFDADC31F" bold="true" box="[151,413,1585,1611]" pageId="3" pageNumber="41">Material examined.</emphasis>
<typeStatus id="54850AA380290415FE32F9E9FE15C318" box="[423,549,1586,1612]" pageId="3" pageNumber="41" type="lectotype">Lectotype</typeStatus>
(here designated): male (10.2 ×
<quantity id="4CC619E480290415FC5CF9E9F81BC318" box="[969,1067,1586,1612]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="3" pageNumber="41" unit="mm" value="8.1">8.1 mm</quantity>
) (NHM 1907.5.22.320a), off Negombo,
<collectingCountry id="F329F49180290415FE8FF98CFDBDC325" box="[282,397,1623,1649]" name="Sri Lanka" pageId="3" pageNumber="41">Sri Lanka</collectingCountry>
or Gulf of Manaar, coral reef;
<specimenCount id="9D387F8880290415FD6EF98CFF7AC325" box="[763,842,1623,1649]" pageId="3" pageNumber="41" type="male">1 male</specimenCount>
<typeStatus id="54850AA380290415FCC7F98CFFC3C325" box="[850,1011,1623,1649]" pageId="3" pageNumber="41" type="paralectotype">paralectotype</typeStatus>
, broken up (NHM 1907.5.22.320b), same data as
<typeStatus id="54850AA380290415FEA5F9A7FDAFC3C2" box="[304,415,1660,1686]" pageId="3" pageNumber="41" type="lectotype">lectotype</typeStatus>
. Other material:
<specimenCount id="9D387F8880290415FDF7F9A7FE8BC3C2" box="[610,699,1660,1686]" pageId="3" pageNumber="41" type="male">2 males</specimenCount>
(18.5 ×
<quantity id="4CC619E480290415FC88F9A7FFB8C3C2" box="[797,904,1660,1686]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="3" pageNumber="41" unit="mm" value="14.6">14.6 mm</quantity>
, 17.4 ×
<quantity id="4CC619E480290415FC71F9A7F860C3C2" box="[996,1104,1660,1686]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="3" pageNumber="41" unit="mm" value="13.3">13.3 mm</quantity>
),
<specimenCount id="9D387F8880290415FBF1F9A7F8FAC3C2" box="[1124,1226,1660,1686]" pageId="3" pageNumber="41" type="female">1 female</specimenCount>
(18.5 ×
<quantity id="4CC619E480290415FABEF9A7F9A5C3C2" box="[1323,1429,1660,1686]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="3" pageNumber="41" unit="mm" value="14.3">14.3 mm</quantity>
) (
<collectionCode id="ED2F2CC480290415FF0AF979FCEAC3E8" box="[159,218,1698,1724]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="41">ZRC</collectionCode>
2001.0627),
<collectingCountry id="F329F49180290415FEE0F979FDCCC3E8" box="[373,508,1698,1724]" name="Philippines" pageId="3" pageNumber="41">Philippines</collectingCountry>
, Bohol, Panglao Is., Balicasag Is.,
<quantity id="4CC619E480290415FC1AF979F837C3E8" box="[911,1031,1698,1724]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="5.0" metricValueMin="0.5" pageId="3" pageNumber="41" unit="m" value="275.0" valueMax="500.0" valueMin="50.0">50500 m</quantity>
, tangle nets,
<date id="FF8092C180290415FB0BF979F9B2C3E8" box="[1182,1410,1698,1724]" pageId="3" pageNumber="41" value="2001-11-28">28 November 2001</date>
;
<specimenCount id="9D387F8880290415FA18F979FCE0C3B5" pageId="3" pageNumber="41" type="male">1 male</specimenCount>
(14.8 ×
<quantity id="4CC619E480290415FEA6F91CFDAEC3B5" box="[307,414,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="3" pageNumber="41" unit="mm" value="11.4">11.4 mm</quantity>
),
<specimenCount id="9D387F8880290415FE27F91CFDC3C3B5" box="[434,499,1735,1761]" pageId="3" pageNumber="41" type="generic">1 juv.</specimenCount>
male (9.7 ×
<quantity id="4CC619E480290415FD12F91CFED5C3B5" box="[647,741,1735,1761]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="41" unit="mm" value="7.3">7.3 mm</quantity>
),
<specimenCount id="9D387F8880290415FD6CF91CFF6FC3B5" box="[761,863,1735,1761]" pageId="3" pageNumber="41" type="female">1 female</specimenCount>
(13.4 ×
<quantity id="4CC619E480290415FC54F91CF81CC3B5" box="[961,1068,1735,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="41" unit="mm" value="10.2">10.2 mm</quantity>
) (WMNH-Na-Cr 0539),
<collectingCountry id="F329F49180290415FAC7F91CF9A8C3B5" box="[1362,1432,1735,1761]" name="Japan" pageId="3" pageNumber="41">Japan</collectingCountry>
, Wakayama, Kushimoto, Kamiura,
<quantity id="4CC619E480290415FDB9F937FEADC252" box="[556,669,1772,1798]" metricMagnitude="1" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="41" unit="m" value="25.0" valueMax="30.0" valueMin="20.0">20 30 m</quantity>
,
<date id="FF8092C180290415FD3DF937FF58C252" box="[680,872,1772,1798]" pageId="3" pageNumber="41" value="1979-11">November 1979</date>
;
<specimenCount id="9D387F8880290415FCE1F937FFE9C252" box="[884,985,1772,1798]" pageId="3" pageNumber="41" type="female">1 female</specimenCount>
(13.0 ×
<quantity id="4CC619E480290415FBAEF937F894C252" box="[1083,1188,1772,1798]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="41" unit="mm" value="10.2">10.2 mm</quantity>
) (NSMT-KY-08-15),
<collectingCountry id="F329F49180290415FF02F8C9FCEDC278" box="[151,221,1810,1836]" name="Japan" pageId="3" pageNumber="41">Japan</collectingCountry>
, Ogasawara Is., south of Chichi-jima Is., 27°03.846΄N 142°07.644΄E 27°03.663΄N 142°07.869΄E, 109-
<quantity id="4CC619E480290415FF02F8ECFCEAC205" box="[151,218,1847,1873]" metricMagnitude="1" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="41" unit="m" value="97.0">97 m</quantity>
, dredge, coll. RV
<emphasis id="B94A681380290415FE2BF8E3FDCFC205" box="[446,511,1848,1873]" italics="true" pageId="3" pageNumber="41">Koyo</emphasis>
,
<date id="FF8092C180290415FD85F8ECFED6C205" box="[528,742,1847,1873]" pageId="3" pageNumber="41" value="2008-10-28">28 October 2008</date>
;
<specimenCount id="9D387F8880290415FD61F8ECFF0BC205" box="[756,827,1847,1873]" pageId="3" pageNumber="41" type="generic">1 juv.</specimenCount>
male (8.2 ×
<quantity id="4CC619E480290415FC48F8ECF80FC205" box="[989,1087,1847,1873]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="41" unit="mm" value="6.5">6.5 mm</quantity>
) (NSMT-KY-08-18),
<collectingCountry id="F329F49180290415FADBF8ECF9A8C205" box="[1358,1432,1847,1873]" name="Japan" pageId="3" pageNumber="41">Japan</collectingCountry>
, Ogasawara Is., east of Chichi-jima Is., 27°05.014΄N 142°14.894΄E 27°04.881΄N 142°14.865΄E, 49
<quantity id="4CC619E480290415FAC2F887F9A7C222" box="[1367,1431,1884,1910]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="41" unit="m" value="48.0">48 m</quantity>
, dredge, coll. RV
<emphasis id="B94A681380290415FEF5F859FDAEC2CF" box="[352,414,1922,1947]" italics="true" pageId="3" pageNumber="41">Koyo</emphasis>
,
<date id="FF8092C180290415FE38F859FE47C2C8" box="[429,631,1922,1948]" pageId="3" pageNumber="41" value="2008-10-29">29 October 2008</date>
;
<specimenCount id="9D387F8880290415FD16F859FEF6C2C8" box="[643,710,1922,1948]" pageId="3" pageNumber="41" type="generic">1 juv.</specimenCount>
male (8.4 ×
<quantity id="4CC619E480290415FCC1F859FF80C2C8" box="[852,944,1922,1948]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="3" pageNumber="41" unit="mm" value="6.7">6.7 mm</quantity>
) (NSMT-KY-08-26), west of Chichi-jima Is., Ogasawara Is,,
<collectingCountry id="F329F49180290415FEE2F87CFD8CC295" box="[375,444,1959,1985]" name="Japan" pageId="3" pageNumber="41">Japan</collectingCountry>
, 27°04.840΄N 142°08.934΄E - 27°04.980΄N 142°09.151΄E,
<quantity id="4CC619E480290415FBFEF87CF8E0C295" box="[1131,1232,1959,1985]" metricMagnitude="1" metricUnit="m" metricValue="8.55" metricValueMax="8.7" metricValueMin="8.4" pageId="3" pageNumber="41" unit="m" value="85.5" valueMax="87.0" valueMin="84.0">84-87 m</quantity>
, dredge, coll. RV Koyo,
<date id="FF8092C180290415FF78F817FDF4C2B2" box="[237,452,1996,2022]" pageId="3" pageNumber="41" value="2008-10-30">30 October 2008</date>
;
<specimenCount id="9D387F8880290415FE47F817FE29C2B2" box="[466,537,1996,2022]" pageId="3" pageNumber="41" type="generic">1 juv.</specimenCount>
male (8.9 ×
<quantity id="4CC619E480290415FD29F817FF2EC2B2" box="[700,798,1996,2022]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="41" unit="mm" value="6.8">6.8 mm</quantity>
) (NSMT-KY-09-27),
<collectingCountry id="F329F49180290415FBBBF817F848C2B2" box="[1070,1144,1996,2022]" name="Japan" pageId="3" pageNumber="41">Japan</collectingCountry>
, Ogasawara Is., east of Anijima-Seto Is.,, 27°06.292΄N 142°13.877΄E 27°06.284΄N 142°14.012΄E,
<quantity id="4CC619E4802E0412FB8DFF43F8A4C5E6" box="[1048,1172,152,178]" metricMagnitude="1" metricUnit="m" metricValue="8.22" metricValueMax="8.34" metricValueMin="8.1" pageId="4" pageNumber="42" unit="m" value="82.2" valueMax="83.4" valueMin="81.0">81-83.4 m</quantity>
,
<quantity id="4CC619E4802E0412FB0AFF43F8D7C5E6" box="[1183,1255,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="42" unit="cm" value="50.0">50 cm</quantity>
<collectionCode id="ED2F2CC4802E0412FB7BFF43F910C5E6" box="[1262,1312,152,178]" httpUri="http://grbio.org/cool/z2jx-p0kp" name="Ocean Research Institute" pageId="4" pageNumber="42">ORI</collectionCode>
biological dredge, coll. RV
<emphasis id="B94A6813802E0412FEF1FF66FD93C582" box="[356,419,189,214]" italics="true" pageId="4" pageNumber="42">Koyo</emphasis>
,
<date id="FF8092C1802E0412FE27FF66FE61C583" box="[434,593,189,215]" pageId="4" pageNumber="42" value="2009-07-15">15 July 2009</date>
;
<specimenCount id="9D387F88802E0412FDCBFF66FE80C583" box="[606,688,189,215]" pageId="4" pageNumber="42" type="male">1 male</specimenCount>
(12.3 ×
<quantity id="4CC619E4802E0412FC82FF66FF48C583" box="[791,888,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="4" pageNumber="42" unit="mm" value="9.8">9.8 mm</quantity>
),
<specimenCount id="9D387F88802E0412FC18FF66FFE0C583" box="[909,976,189,215]" pageId="4" pageNumber="42" type="generic">2 juv.</specimenCount>
males (9.8 ×
<quantity id="4CC619E4802E0412FBE3FF66F8E5C583" box="[1142,1237,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="4" pageNumber="42" unit="mm" value="7.9">7.9 mm</quantity>
, 11.2 × 9.0 mm) (NSMT-SY-09-18),
<collectingCountry id="F329F491802E0412FE1CFF39FDE1C5A8" box="[393,465,226,252]" name="Japan" pageId="4" pageNumber="42">Japan</collectingCountry>
, Ogasawara Is., west off Chichi-jima Is., 27°06.11΄N 142°08.89΄E
<geoCoordinate id="EE0AD2C6802E0412FA86FF39F9ACC5A8" box="[1299,1436,226,252]" direction="north" orientation="latitude" pageId="4" pageNumber="42" precision="9" value="27.101166">27°06.07'N</geoCoordinate>
<geoCoordinate id="EE0AD2C6802E0412FF02FED3FD1AC476" box="[151,298,264,290]" direction="east" orientation="longitude" pageId="4" pageNumber="42" precision="9" value="142.151">142°09.06'E</geoCoordinate>
, 101
<quantity id="4CC619E4802E0412FEE5FED3FD98C476" box="[368,424,264,290]" metricMagnitude="1" metricUnit="m" metricValue="9.8" pageId="4" pageNumber="42" unit="m" value="98.0">98m</quantity>
,
<quantity id="4CC619E4802E0412FE26FED3FDC4C476" box="[435,500,264,290]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="42" unit="cm" value="50.0">50cm</quantity>
dredge, coll. TRV
<emphasis id="B94A6813802E0412FD41FED3FF5FC475" box="[724,879,264,289]" italics="true" pageId="4" pageNumber="42">Shinyo-Maru</emphasis>
,
<date id="FF8092C1802E0412FCEBFED3F853C476" box="[894,1123,264,290]" pageId="4" pageNumber="42" value="2009-11-18">18 November 2009</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C324E78A802E0412FF50FEF6FECAC1BB" pageId="4" pageNumber="42" type="diagnosis">
<paragraph id="8B81B401802E0412FF50FEF6FECAC1BB" blockId="4.[151,1437,151,2010]" pageId="4" pageNumber="42">
<emphasis id="B94A6813802E0412FF50FEF6FD78C413" bold="true" box="[197,328,301,327]" pageId="4" pageNumber="42">Diagnosis.</emphasis>
Carapace hexagonal in shape, relatively long, width approximately 1.3 times length, dorsal surface glabrous, microscopically granular with distinct regions; only metagastric ridge distinctly visible, other ridges are indistinct as granules diffused into large areas (
<figureCitation id="1305A884802E0412FC17FEA3FFE4C4C6" box="[898,980,376,402]" captionStart="FIGURE 1" captionStartId="3.[151,257,1484,1508]" captionTargetBox="[151,1436,714,1462]" captionTargetId="figure@3.[151,1436,714,1463]" captionTargetPageId="3" captionText="FIGURE 1. Live colour. Cavoportunus dubius (Laurie, 1906). Female (13.0 × 10.2 mm) (NSMT-KY- 08 - 15), dorsal overall view. Scales = 1.0 mm. (Photograph: H. Komatsu)." httpUri="https://zenodo.org/record/199231/files/figure.png" pageId="4" pageNumber="42">Figs. 1</figureCitation>
,
<figureCitation id="1305A884802E0412FC76FEA3FFC2C4C6" box="[995,1010,376,402]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">2</figureCitation>
A, 4A). Frontal margin with 4 sharp teeth, protruding beyond inner supraorbital angles, directed dorsally; lateral teeth stronger, longer than median ones; inner supraobital angles rounded (
<figureCitation id="1305A884802E0412FDE5FE19FEF3C488" box="[624,707,450,476]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">Figs. 2</figureCitation>
A, 4A). Upper orbital margin with single fissure (
<figureCitation id="1305A884802E0412FA9AFE19F951C488" box="[1295,1377,450,476]" captionStart="FIGURE 1" captionStartId="3.[151,257,1484,1508]" captionTargetBox="[151,1436,714,1462]" captionTargetId="figure@3.[151,1436,714,1463]" captionTargetPageId="3" captionText="FIGURE 1. Live colour. Cavoportunus dubius (Laurie, 1906). Female (13.0 × 10.2 mm) (NSMT-KY- 08 - 15), dorsal overall view. Scales = 1.0 mm. (Photograph: H. Komatsu)." httpUri="https://zenodo.org/record/199231/files/figure.png" pageId="4" pageNumber="42">Figs. 1</figureCitation>
,
<figureCitation id="1305A884802E0412FAE4FE19F9B0C488" box="[1393,1408,450,476]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">2</figureCitation>
A, 4A), lower margin with acute forward-directed tooth (
<figureCitation id="1305A884802E0412FC8CFE33FF52C756" box="[793,866,488,514]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">Fig. 2</figureCitation>
B). Anterolateral margin with 9 teeth, first tooth stoutest, size of second to ninth teeth subequal (
<figureCitation id="1305A884802E0412FC92FDCBFF52C77E" box="[775,866,528,554]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">Figs. 2</figureCitation>
A, 4A). Merus of third maxilliped produced anterolaterally, length to width ratio approximately 1.27; length to width ratio of ischium 1.57 (
<figureCitation id="1305A884802E0412FB60FDEEF976C71B" box="[1269,1350,565,591]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">Figs. 2</figureCitation>
C, 3C). Cheliped covered with fine granules; ventral margin with squamiform markings; merus with 3 or 4 spines on anterior margin, a single spine at distal end of posterior margin; carpus with sharp spine on internal angle, outer surface with carina leading to spine; manus with 3 spines, one at wrist articulation, 2 on inner, outer margins of upper surface at distal end; outer surface with 3 granular costae at lower half; the first costa highly raised; movable fingers slender, as long as manus (
<figureCitation id="1305A884802E0412FD66FD2BFF76C65E" box="[755,838,752,778]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">Figs. 2</figureCitation>
A, B, 3A, B). Fifth walking legs with short merus, posterior margin with fine denticles; propodus with fine, short, dense setae; without spines (
<figureCitation id="1305A884802E0412FB74FCCEF903C67B" box="[1249,1331,789,815]" captionStart="FIGURE 1" captionStartId="3.[151,257,1484,1508]" captionTargetBox="[151,1436,714,1462]" captionTargetId="figure@3.[151,1436,714,1463]" captionTargetPageId="3" captionText="FIGURE 1. Live colour. Cavoportunus dubius (Laurie, 1906). Female (13.0 × 10.2 mm) (NSMT-KY- 08 - 15), dorsal overall view. Scales = 1.0 mm. (Photograph: H. Komatsu)." httpUri="https://zenodo.org/record/199231/files/figure.png" pageId="4" pageNumber="42">Figs. 1</figureCitation>
,
<figureCitation id="1305A884802E0412FAD6FCCEF962C67B" box="[1347,1362,789,815]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">2</figureCitation>
A, B). Male abdomen triangular, with tapering lateral margins, converging towards posteror; posterior margin of somite 3 convex with median cleft; lateral margins of somite 4 evenly convex; lateral margins of somite 5 slightly convex (
<figureCitation id="1305A884802E0412FEC9FC5EFD9FC6CB" box="[348,431,901,927]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">Figs. 2</figureCitation>
B, 4B). Median line present on thoracic sternites 5 to 8 of both sexes. Male adominal holding system with prominent spur-like press-button on thoracic sternite 5 next to the sternal suture 5/6 (
<figureCitation id="1305A884802E0412FF0AFC0BFCC0C6BE" box="[159,240,976,1002]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Figs. 3</figureCitation>
D, 4C), present also in female and juveniles but relatively weaker (
<figureCitation id="1305A884802E0412FB92FC0BF867C6BE" box="[1031,1111,976,1002]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Figs. 3</figureCitation>
G, H); sternite 8 of thoracic sternum of adult male with median groove, with 2 deep depressions (anterior larger) on each side to accommodate G1 (
<figureCitation id="1305A884802E0412FEEDFBC1FDFAC160" box="[376,458,1050,1076]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Figs. 3</figureCitation>
D, 4C). G1 very short, relatively stout, S-shaped; basal part relatively slender, not dilated; subdistal part distinctly dilated, forming large lobe-like process; distal part rounded, with distinct flattened folds on tip (
<figureCitation id="1305A884802E0412FE0EFBBEFDDCC12B" box="[411,492,1125,1151]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Figs. 3</figureCitation>
E, 4D). G2 elongated, tapering, about four-fifths length of G1, bilobed tip (
<figureCitation id="1305A884802E0412FAF4FBBEFC97C1F0" captionStart="FIGURE 4" captionStartId="7.[151,255,1387,1411]" captionTargetBox="[187,1406,238,1346]" captionTargetId="figure@7.[170,1417,194,1365]" captionTargetPageId="7" captionText="FIGURE 4. Cavoportunus dubius (Laurie, 1906). A, lectotype male, (10.2 × 8.1 mm) (NHM 1907.5.22.320); B-F. male, (18.5 × 14.6 mm) (ZRC 2001.0627). A, carapace; B, male abdomen; C, male thoracic sternum with intact right G 1; D, different views of left G 1; E, right G 2; F, distal part of right G 2. Scales: A-E = 1.0 mm, F = 0.2 mm." httpUri="https://zenodo.org/record/199234/files/figure.png" pageId="4" pageNumber="42">Figs. 4</figureCitation>
E, F). Penis coxal, long, slender, proximally lodged between 2 raised processes of sternite 8 (
<figureCitation id="1305A884802E0412FABFFB51F944C1F0" box="[1322,1396,1162,1188]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Fig. 3</figureCitation>
D). Vulvae mostly surrounded by thick, whitish, crescent-shaped rim; salient, short, whitish sternal cover on outer margin; separate, small, round operculum (
<figureCitation id="1305A884802E0412FD1BFB0EFEE3C1BB" box="[654,723,1237,1263]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Fig. 3</figureCitation>
G).
</paragraph>
</subSubSection>
<subSubSection id="C324E78A802E0412FF50FB21FFAEC0A0" pageId="4" pageNumber="42" type="description">
<paragraph id="8B81B401802E0412FF50FB21FD3BC00B" blockId="4.[151,1437,151,2010]" pageId="4" pageNumber="42">
<emphasis id="B94A6813802E0412FF50FB21FD70C040" bold="true" box="[197,320,1274,1300]" pageId="4" pageNumber="42">Variation.</emphasis>
The adult female abdomen is ovate and covers much of the sternum, with the telson triangular (
<figureCitation id="1305A884802E0412FF35FAFBFCD6C06E" box="[160,230,1312,1338]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Fig. 3</figureCitation>
F). In juvenile males, the G1s relatively smaller, with depressions on thoracic sternum less developed (
<figureCitation id="1305A884802E0412FF0AFA9EFCD4C00B" box="[159,228,1349,1375]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Fig. 3</figureCitation>
H).
</paragraph>
<paragraph id="8B81B401802E0412FF50FAB1FFAEC0A0" blockId="4.[151,1437,151,2010]" pageId="4" pageNumber="42">
<emphasis id="B94A6813802E0412FF50FAB1FD25C0D0" bold="true" box="[197,277,1386,1412]" pageId="4" pageNumber="42">Color.</emphasis>
White to transparent, some areas with scattered orange dots, especially on the carapace and chelipeds (
<figureCitation id="1305A884802E0412FEB7FA4BFD5EC0FE" box="[290,366,1424,1450]" captionStart="FIGURE 1" captionStartId="3.[151,257,1484,1508]" captionTargetBox="[151,1436,714,1462]" captionTargetId="figure@3.[151,1436,714,1463]" captionTargetPageId="3" captionText="FIGURE 1. Live colour. Cavoportunus dubius (Laurie, 1906). Female (13.0 × 10.2 mm) (NSMT-KY- 08 - 15), dorsal overall view. Scales = 1.0 mm. (Photograph: H. Komatsu)." httpUri="https://zenodo.org/record/199231/files/figure.png" pageId="4" pageNumber="42">Fig. 1</figureCitation>
). Silvery iridescence is very distinct on the posterolateral junction of the carapace, protruding edges of the first two abdominal somites, upper surface of the movable finger, carina on the outer side and upper spines of the palm and the spine on the outer wrist.
</paragraph>
</subSubSection>
<subSubSection id="C324E78A802E0412FF50F9DBFF62C34E" box="[197,850,1536,1562]" pageId="4" pageNumber="42" type="distribution">
<paragraph id="8B81B401802E0412FF50F9DBFF62C34E" blockId="4.[151,1437,151,2010]" box="[197,850,1536,1562]" pageId="4" pageNumber="42">
<emphasis id="B94A6813802E0412FF50F9DBFD55C34E" bold="true" box="[197,357,1536,1562]" pageId="4" pageNumber="42">Distribution.</emphasis>
Indian Ocean,
<collectingCountry id="F329F491802E0412FD8DF9DBFED3C34E" box="[536,739,1536,1562]" name="French Polynesia" pageId="4" pageNumber="42">French Polynesia</collectingCountry>
to
<collectingCountry id="F329F491802E0412FC9DF9DBFF7EC34E" box="[776,846,1536,1562]" name="Japan" pageId="4" pageNumber="42">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C324E78A802E0410FF50F9FEFD87C2ED" lastPageId="6" lastPageNumber="44" pageId="4" pageNumber="42" type="discussion">
<paragraph id="8B81B401802E0412FF50F9FEFDA5C28E" blockId="4.[151,1437,151,2010]" pageId="4" pageNumber="42">
<emphasis id="B94A6813802E0412FF50F9FEFD0FC36B" bold="true" box="[197,319,1573,1599]" pageId="4" pageNumber="42">Remarks.</emphasis>
<bibRefCitation id="EFAFC9F0802E0412FED2F9FEFDDDC36B" author="Laurie" box="[327,493,1573,1599]" pageId="4" pageNumber="42" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie (1906)</bibRefCitation>
listed two specimens in his material examined of his description, one from the Gulf of Manaar and one from off Negombo in
<collectingCountry id="F329F491802E0412FD2FF991FF01C330" box="[698,817,1610,1636]" name="Sri Lanka" pageId="4" pageNumber="42">Sri Lanka</collectingCountry>
. No
<typeStatus id="54850AA3802E0412FCF2F991FFFEC330" box="[871,974,1610,1636]" pageId="4" pageNumber="42" type="holotype">holotype</typeStatus>
was indicated so both are
<typeStatus id="54850AA3802E0412FA91F991F95EC330" box="[1284,1390,1610,1636]" pageId="4" pageNumber="42" type="syntype">syntypes</typeStatus>
. He provided measurements of one specimen, with a carapace length of
<quantity id="4CC619E4802E0412FC5BF9ABF813C3DE" box="[974,1059,1648,1674]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="42" unit="mm" value="13.0">13 mm</quantity>
(
<bibRefCitation id="EFAFC9F0802E0412FBA6F9ABF938C3DE" author="Laurie" box="[1075,1288,1648,1674]" pageId="4" pageNumber="42" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie 1906: 416</bibRefCitation>
) but did not indicate where the specimen was from. He also provided figures of the left cheliped, both third maxillipeds, male abdomen and two G1s of one specimen (
<bibRefCitation id="EFAFC9F0802E0412FD2CF961FFA9C380" author="Laurie" box="[697,921,1722,1748]" pageId="4" pageNumber="42" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie 1906: fig. 9</bibRefCitation>
) but did not indicate its size or locality. We have checked the
<typeStatus id="54850AA3802E0412FEFFF93BFDADC3AE" box="[362,413,1760,1786]" pageId="4" pageNumber="42">type</typeStatus>
material but only one specimen is still intact, a male measuring 10.2 ×
<quantity id="4CC619E4802E0412FB7BF93BF97DC3AE" box="[1262,1357,1760,1786]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="42" unit="mm" value="8.1">8.1 mm</quantity>
. There are parts of another specimen in the same specimen jar that corresponds exactly to the structures figured by Laurie (
<figureCitation id="1305A884802E0412FF62F8F1FD0EC210" box="[247,318,1834,1860]" captionStart="FIGURE 3" captionStartId="6.[151,256,1604,1628]" captionTargetBox="[151,1436,194,1575]" captionTargetId="figure@6.[151,1436,194,1583]" captionTargetPageId="6" captionText="FIGURE 3. Cavoportunus dubius (Laurie, 1906). A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G 1; E, different views of G 1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm." httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="4" pageNumber="42">Fig. 3</figureCitation>
A, B, C;
<bibRefCitation id="EFAFC9F0802E0412FE38F8F1FE75C210" author="Laurie" box="[429,581,1834,1860]" pageId="4" pageNumber="42" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie 1906</bibRefCitation>
: fig. 9). Although the carapace and all other parts of this specimen are missing, we believe the
<quantity id="4CC619E4802E0412FE22F88BFE3BC23E" box="[439,523,1872,1898]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="42" unit="mm" value="13.0">13 mm</quantity>
carapace length specimen listed by
<bibRefCitation id="EFAFC9F0802E0412FC29F88BF851C23E" author="Laurie" box="[956,1121,1872,1898]" pageId="4" pageNumber="42" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie (1906)</bibRefCitation>
is probably the broken-up one. The intact specimen measures
<quantity id="4CC619E4802E0412FDCBF8AEFE8FC2DB" box="[606,703,1909,1935]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="4" pageNumber="42" unit="mm" value="8.1">8.1 mm</quantity>
in carapace length and is clearly smaller than the broken specimen. We hereby designate the complete specimen as the
<typeStatus id="54850AA3802E0412FC07F841F832C2E0" box="[914,1026,1946,1972]" pageId="4" pageNumber="42" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C3ECF82802E0412FBB9F840FCEBC28E" authority="Laurie, 1906" authorityName="Laurie" authorityYear="1906" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A6813802E0412FBB9F840F902C2E0" box="[1068,1330,1947,1972]" italics="true" pageId="4" pageNumber="42">Cavoportunus dubius</emphasis>
(
<bibRefCitation id="EFAFC9F0802E0412FAD6F841FCE3C28E" author="Laurie" pageId="4" pageNumber="42" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie, 1906</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="1305A884802E0412FF7EF81BFD0CC28E" box="[235,316,1984,2010]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="4" pageNumber="42">Figs. 2</figureCitation>
A, 4A).
</paragraph>
<caption id="DF41E489802F0413FF02F85DFD63C289" httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="5" pageNumber="43" targetBox="[228,1359,194,1913]" targetPageId="5">
<paragraph id="8B81B401802F0413FF02F85DFD63C289" blockId="5.[151,1436,1926,2013]" pageId="5" pageNumber="43">
<emphasis id="B94A6813802F0413FF02F85DFD29C2CA" bold="true" box="[151,281,1926,1950]" pageId="5" pageNumber="43">FIGURE 2.</emphasis>
<taxonomicName id="4C3ECF82802F0413FEB5F85CFE99C2C9" authority="Laurie, 1906" authorityName="Laurie" authorityYear="1906" box="[288,681,1926,1950]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A6813802F0413FEB5F85CFE35C2CA" box="[288,517,1927,1950]" italics="true" pageId="5" pageNumber="43">Cavoportunus dubius</emphasis>
(Laurie, 1906)
</taxonomicName>
. A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace.
</paragraph>
</caption>
<caption id="DF41E489802C0410FF02F99FFCF8C38F" httpUri="https://zenodo.org/record/199233/files/figure.png" pageId="6" pageNumber="44" targetBox="[151,1436,194,1575]" targetPageId="6">
<paragraph id="8B81B401802C0410FF02F99FFCF8C38F" blockId="6.[151,1437,1604,1755]" pageId="6" pageNumber="44">
<emphasis id="B94A6813802C0410FF02F99FFD2CC308" bold="true" box="[151,284,1604,1628]" pageId="6" pageNumber="44">FIGURE 3.</emphasis>
<taxonomicName id="4C3ECF82802C0410FEB6F99EFE84C30F" authority="Laurie, 1906" authorityName="Laurie" authorityYear="1906" box="[291,692,1604,1628]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A6813802C0410FEB6F99EFE3CC308" box="[291,524,1605,1628]" italics="true" pageId="6" pageNumber="44">Cavoportunus dubius</emphasis>
(Laurie, 1906)
</taxonomicName>
. A-C, parts of paralectotype, male (NHM 1907.5.22.320). A, B, left cheliped; C, right third maxiliped; D, E, male (18.5 × 14.6 mm) (ZRC 2001.0627). D, thoracic sternum with intact right G1; E, different views of G1; F, G, female (18.5 × 14.3 mm (ZRC 2001.0627). F, abdomen; G, thoracic sternum; H, juv. male (9.7 × 7.3 mm) (WMNH-Na-Cr 0539) thoracic sternum. Abbreviations: p = penis; pb = press button. Scales = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B81B401802C0410FF53F8D1FD87C2ED" blockId="6.[151,1437,1802,1977]" pageId="6" pageNumber="44">
The provenance of the
<typeStatus id="54850AA3802C0410FE40F8D1FE70C270" box="[469,576,1802,1828]" pageId="6" pageNumber="44" type="lectotype">lectotype</typeStatus>
cannot be ascertained.
<bibRefCitation id="EFAFC9F0802C0410FCC7F8D1FFC4C270" author="Laurie" box="[850,1012,1802,1828]" pageId="6" pageNumber="44" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie (1906)</bibRefCitation>
did not indicate precisely where his two specimens came from, so the
<typeStatus id="54850AA3802C0410FDBFF8F4FEA6C21D" box="[554,662,1839,1865]" pageId="6" pageNumber="44" type="lectotype">lectotype</typeStatus>
was from either the Gulf of Manaar or Negombo. The label with the
<typeStatus id="54850AA3802C0410FF56F88FFD31C23A" box="[195,257,1876,1902]" pageId="6" pageNumber="44">types</typeStatus>
mentions only the Gulf of Manaar and its catalogue number suggests only one specimen, although there are clearly two specimens. In any case, the two
<typeStatus id="54850AA3802C0410FCA7F8A1FF57C2C0" box="[818,871,1914,1940]" pageId="6" pageNumber="44">type</typeStatus>
localities are very close to each other so it is probably not significant.
</paragraph>
</subSubSection>
<caption id="DF41E489802D0411FF02FAB0F8F5C096" httpUri="https://zenodo.org/record/199234/files/figure.png" pageId="7" pageNumber="45" targetBox="[187,1406,238,1346]" targetPageId="7">
<paragraph id="8B81B401802D0411FF02FAB0F8F5C096" blockId="7.[151,1436,1387,1474]" pageId="7" pageNumber="45">
<emphasis id="B94A6813802D0411FF02FAB0FD29C0D7" bold="true" box="[151,281,1387,1411]" pageId="7" pageNumber="45">FIGURE 4.</emphasis>
<taxonomicName id="4C3ECF82802D0411FEB5FAB0FE9AC0D6" authority="Laurie, 1906" authorityName="Laurie" authorityYear="1906" box="[288,682,1387,1410]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A6813802D0411FEB5FAB0FE35C0D6" box="[288,517,1387,1410]" italics="true" pageId="7" pageNumber="45">Cavoportunus dubius</emphasis>
(Laurie, 1906)
</taxonomicName>
. A, lectotype male, (10.2 ×8.1 mm) (NHM 1907.5.22.320); B-F. male, (18.5 ×14.6 mm) (ZRC 2001.0627). A, carapace; B, male abdomen; C, male thoracic sternum with intact right G1; D, different views of left G1; E, right G2; F, distal part of right G2. Scales: A-E = 1.0 mm, F = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C324E78A802D041EFF50FA2AFE91C476" lastPageId="8" lastPageNumber="46" pageId="7" pageNumber="45" type="reference_group">
<paragraph id="8B81B401802D0411FF50FA2AF841C244" blockId="7.[151,1437,1520,2032]" pageId="7" pageNumber="45">
<taxonomicName id="4C3ECF82802D0411FF50FA2AFD8CC35E" ID-CoL="4M2K3" box="[197,444,1521,1546]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="yoronensis">
<emphasis id="B94A6813802D0411FF50FA2AFD8CC35E" box="[197,444,1521,1546]" italics="true" pageId="7" pageNumber="45">Portunus yoronensis</emphasis>
</taxonomicName>
was originally described from one male specimen collected from Yoron Is.,
<collectingCountry id="F329F491802D0411FACDFA2BF9ACC35E" box="[1368,1436,1520,1546]" name="Japan" pageId="7" pageNumber="45">Japan</collectingCountry>
(
<bibRefCitation id="EFAFC9F0802D0411FF0AF9CDFD6AC364" author="Sakai" box="[159,346,1558,1584]" pageId="7" pageNumber="45" refString="Sakai, T. (1974) Notes from the carcinological fauna of Japan (V). Researches on Crustacea, 6, 86 - 102." type="journal article" year="1974">Sakai 1974: 91</bibRefCitation>
) but the
<typeStatus id="54850AA3802D0411FE5FF9CDFDCEC364" box="[458,510,1558,1584]" pageId="7" pageNumber="45">type</typeStatus>
specimen could not be found in the Japanese repositories that still hold his material or in the Senckenberg Museum, Frankfurt, where some of his own material is now kept. It is, however, possible it is mixed with material in other jars or has not been catalogued. On the basis of the descriptions and figures by
<bibRefCitation id="EFAFC9F0802D0411FE7DF95DFE4CC3F4" author="Sakai" box="[488,636,1670,1696]" pageId="7" pageNumber="45" refString="Sakai, T. (1974) Notes from the carcinological fauna of Japan (V). Researches on Crustacea, 6, 86 - 102." type="journal article" year="1974">Sakai (1974</bibRefCitation>
,
<bibRefCitation id="EFAFC9F0802D0411FD1FF95DFEF8C3F4" author="Sakai" box="[650,712,1670,1696]" pageId="7" pageNumber="45" refString="Sakai, T. (1976) Crabs of Japan and the Adjacent Seas. Kodansha, Tokyo, vol. 1 [English text], xxxix + 773 pp.; vol. 2 [Japanese text], 461 pp.; vol. 3 [plates], 16 pp., pls. 1 - 251." type="book chapter" year="1976">1976</bibRefCitation>
) as well as examination of recent specimens of this species from
<collectingCountry id="F329F491802D0411FF4EF970FD14C391" box="[219,292,1707,1733]" name="Japan" pageId="7" pageNumber="45">Japan</collectingCountry>
,
<taxonomicName id="4C3ECF82802D0411FEA4F970FFAEC391" ID-CoL="4M2K3" authority="Sakai, 1974" authorityName="Sakai" authorityYear="1974" box="[305,926,1707,1733]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="yoronensis" subGenus="Cycloachelous">
<emphasis id="B94A6813802D0411FEA4F970FD92C390" box="[305,418,1707,1732]" italics="true" pageId="7" pageNumber="45">Portunus</emphasis>
(
<emphasis id="B94A6813802D0411FE21F970FE58C390" box="[436,616,1707,1732]" italics="true" pageId="7" pageNumber="45">Cycloachelous</emphasis>
)
<emphasis id="B94A6813802D0411FDE9F970FF30C390" box="[636,768,1707,1732]" italics="true" pageId="7" pageNumber="45">yoronensis</emphasis>
<bibRefCitation id="EFAFC9F0802D0411FC9CF970FFAEC391" author="Sakai" box="[777,926,1707,1733]" pageId="7" pageNumber="45" refString="Sakai, T. (1974) Notes from the carcinological fauna of Japan (V). Researches on Crustacea, 6, 86 - 102." type="journal article" year="1974">Sakai, 1974</bibRefCitation>
</taxonomicName>
, is clearly a junior synonym of
<taxonomicName id="4C3ECF82802D0411FABCF970FE3AC3BE" ID-CoL="47475" authority="Laurie, 1906" authorityName="Laurie" authorityYear="1906" class="Malacostraca" family="Portunidae" genus="Neptunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="dubia" subGenus="Achelous">
<emphasis id="B94A6813802D0411FABCF970F9ACC390" box="[1321,1436,1707,1732]" italics="true" pageId="7" pageNumber="45">Neptunus</emphasis>
(
<emphasis id="B94A6813802D0411FF34F90AFD3EC3BE" box="[161,270,1745,1770]" italics="true" pageId="7" pageNumber="45">Achelous</emphasis>
)
<emphasis id="B94A6813802D0411FEB5F90AFD54C3BE" box="[288,356,1745,1770]" italics="true" pageId="7" pageNumber="45">dubia</emphasis>
<bibRefCitation id="EFAFC9F0802D0411FEF8F90BFE3AC3BE" author="Laurie" box="[365,522,1744,1770]" pageId="7" pageNumber="45" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie, 1906</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFAFC9F0802D0411FD83F90BFED4C3BE" author="Crosnier" box="[534,740,1744,1770]" pageId="7" pageNumber="45" refString="Crosnier, A. (1984) Sur quelques Portunidae (Crustacea Decapoda Brachyura) des iles Seychelles. Bulletin du Museum national d'Histoire naturelle, Section A, Zoologie, Biologie et Ecologie Animales, Paris, 4 e serie, 6 (2), 397 - 419." type="journal article" year="1984" yearSuffix="b">Crosnier (1984b)</bibRefCitation>
had in fact synonymized
<taxonomicName id="4C3ECF82802D0411FB89F90AF88AC3BE" ID-CoL="4M2K3" box="[1052,1210,1745,1770]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="yoronensis">
<emphasis id="B94A6813802D0411FB89F90AF88AC3BE" box="[1052,1210,1745,1770]" italics="true" pageId="7" pageNumber="45">P. yoronensis</emphasis>
</taxonomicName>
with
<taxonomicName id="4C3ECF82802D0411FA95F90AF95EC3BE" ID-CoL="RYDG" box="[1280,1390,1745,1770]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A6813802D0411FA95F90AF95EC3BE" box="[1280,1390,1745,1770]" italics="true" pageId="7" pageNumber="45">P. dubius</emphasis>
</taxonomicName>
but this has not been widely recognized by other authors. We agree with his assessment.
</paragraph>
<paragraph id="8B81B401802D041EFF50F8C0FE91C476" blockId="7.[151,1437,1520,2032]" lastBlockId="8.[151,1436,152,290]" lastPageId="8" lastPageNumber="46" pageId="7" pageNumber="45">
<bibRefCitation id="EFAFC9F0802D0411FF50F8C0FE33C261" author="Stephenson" box="[197,515,1819,1845]" pageId="7" pageNumber="45" refString="Stephenson, W. &amp; Rees, M. (1967) Some portunid crabs from the Pacific and Indian Oceans in the collections of the Smithsonian Institution. Proceedings of the United States National Museum, 120, 1 - 114." type="journal article" year="1967">Stephenson &amp; Rees (1967)</bibRefCitation>
reported a male and a female of
<taxonomicName id="4C3ECF82802D0411FC1FF8C0F8B7C260" ID-CoL="RYDG" box="[906,1159,1819,1844]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A6813802D0411FC1FF8C0F8B7C260" box="[906,1159,1819,1844]" italics="true" pageId="7" pageNumber="45">Cavoportunus dubius</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C3ECF82802D0411FB2CF8C0F915C260" ID-CoL="9CLRM" box="[1209,1317,1819,1844]" class="Malacostraca" family="Portunidae" genus="Portunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B94A6813802D0411FB2CF8C0F915C260" box="[1209,1317,1819,1844]" italics="true" pageId="7" pageNumber="45">Portunus</emphasis>
</taxonomicName>
) from the
<collectingCountry id="F329F491802D0411FF02F89BFD2FC20E" box="[151,287,1856,1882]" name="Philippines" pageId="7" pageNumber="45">Philippines</collectingCountry>
, and noted that the &quot;… abdomen of the single male in the present collection is narrower, and the ultimate segment is longer than figured by
<bibRefCitation id="EFAFC9F0802D0411FD06F8BDFFB8C2D4" author="Laurie" box="[659,904,1894,1920]" pageId="7" pageNumber="45" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie (1906, fig. 9)</bibRefCitation>
&quot;. We have re-examined the
<typeStatus id="54850AA3802D0411FB46F8BDF935C2D4" box="[1235,1285,1894,1920]" pageId="7" pageNumber="45">type</typeStatus>
specimen of
<taxonomicName id="4C3ECF82802D0411FF02F850FD90C2F0" ID-CoL="RYDG" box="[151,416,1931,1956]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A6813802D0411FF02F850FD90C2F0" box="[151,416,1931,1956]" italics="true" pageId="7" pageNumber="45">Cavoportunus dubius</emphasis>
</taxonomicName>
, which was used for the photographs in
<bibRefCitation id="EFAFC9F0802D0411FC09F850F8ADC2F1" author="Laurie" box="[924,1181,1931,1957]" pageId="7" pageNumber="45" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie (1906: fig. 9)</bibRefCitation>
, and somite 6 of the abdomen is actually not as short as figured by
<bibRefCitation id="EFAFC9F0802D0411FD28F86BFF82C29E" author="Laurie" box="[701,946,1968,1994]" pageId="7" pageNumber="45" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.) Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists, Part 5, Supplementary Reports, 40, pp. 349 - 432." type="journal article" year="1906">Laurie (1906: fig. 9)</bibRefCitation>
. Its actual shape actually closely matches the figures of
<bibRefCitation id="EFAFC9F0802D0411FED0F80DFEC6C2A4" author="Stephenson" box="[325,758,2006,2032]" pageId="7" pageNumber="45" refString="Stephenson, W. &amp; Rees, M. (1967) Some portunid crabs from the Pacific and Indian Oceans in the collections of the Smithsonian Institution. Proceedings of the United States National Museum, 120, 1 - 114." type="journal article" year="1967" yearSuffix="b">Stephenson &amp; Rees (1967: fig. 3b)</bibRefCitation>
. However, the figure by Stephenson &amp; Rees shows the anterior margin of the third male abdominal somite (
<bibRefCitation id="EFAFC9F08022041EFD6BFF43F8B4C5E6" author="Stephenson" box="[766,1156,152,178]" pageId="8" pageNumber="46" refString="Stephenson, W. &amp; Rees, M. (1967) Some portunid crabs from the Pacific and Indian Oceans in the collections of the Smithsonian Institution. Proceedings of the United States National Museum, 120, 1 - 114." type="journal article" year="1967" yearSuffix="b">Stephenson &amp; Rees 1967: fig. 3b</bibRefCitation>
) as almost straight, and not sinuous as those observed in other
<taxonomicName id="4C3ECF828022041EFDCEFF66FF66C582" ID-CoL="RYDG" box="[603,854,189,214]" class="Malacostraca" family="Portunidae" genus="Cavoportunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="46" phylum="Arthropoda" rank="species" species="dubius">
<emphasis id="B94A68138022041EFDCEFF66FF66C582" box="[603,854,189,214]" italics="true" pageId="8" pageNumber="46">Cavoportunus dubius</emphasis>
</taxonomicName>
specimens or figures or photographs provided by other authors (
<figureCitation id="1305A8848022041EFED5FF39FDA2C5A8" box="[320,402,226,252]" captionStart="FIGURE 2" captionStartId="5.[151,255,1926,1950]" captionTargetBox="[228,1359,194,1913]" captionTargetId="figure@5.[228,1359,194,1915]" captionTargetPageId="5" captionText="FIGURE 2. Cavoportunus dubius (Laurie, 1906). A, lectotype male (10.2 × 8.1 mm) (NHM 1907.5.22.320); B, C, male (18.5 × 14.6 mm) (ZRC 2001.0627). A, overall dorsal view; B, ventral view of thoracic sternum and abdomen; C, frontal view of carapace." httpUri="https://zenodo.org/record/199232/files/figure.png" pageId="8" pageNumber="46">Figs. 2</figureCitation>
B, 4B). The observed differences are possibly due to the rather schematic and inaccurate figure of
<bibRefCitation id="EFAFC9F08022041EFE91FED3FEADC476" author="Stephenson" box="[260,669,264,290]" pageId="8" pageNumber="46" refString="Stephenson, W. &amp; Rees, M. (1967) Some portunid crabs from the Pacific and Indian Oceans in the collections of the Smithsonian Institution. Proceedings of the United States National Museum, 120, 1 - 114." type="journal article" year="1967" yearSuffix="b">Stephenson &amp; Rees (1967: fig. 3b)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>