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<document id="FC8B9764D58BC03AF49712BA7DF612EF" ID-CLB-Dataset="28042" ID-DOI="10.11646/zootaxa.4508.3.2" ID-GBIF-Dataset="1a35d13d-57fb-4944-80b7-c73bb58274e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607143" ID-ZooBank="4C0CAE39-B5C3-419C-9A4D-493806280141" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586026139" checkinUser="plazi" docAuthor="Lee, Chi-Feng &amp; Bezdĕk, Jan" docDate="2018" docId="03928782A84AFFC01E897350FA72FCFE" docLanguage="en" docName="zootaxa.4508.3.2.pdf" docOrigin="Zootaxa 4508 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Theopea Baly 1864" docType="key" docVersion="9" lastPageNumber="337" masterDocId="FFABFFFAA848FFC31E1E7544FFE9FFEA" masterDocTitle="Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) of East Asia: species lacking modified clypeus in males and the T. sauteri species group" masterLastPageNumber="376" masterPageNumber="334" pageNumber="336" updateTime="1698713057748" updateUser="ExternalLinkService">
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<mods:title id="9FDEE3C78557D6B8324F9290F2E42A81">Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) of East Asia: species lacking modified clypeus in males and the T. sauteri species group</mods:title>
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<mods:namePart id="B73207C2707E3FEDCA62D38E1D2DB9C0">Lee, Chi-Feng</mods:namePart>
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<mods:namePart id="DBDEC13D3BE81A0ABAF7BB8B09D9E607">Bezdĕk, Jan</mods:namePart>
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<treatment id="03928782A84AFFC01E897350FA72FCFE" ID-DOI="http://doi.org/10.5281/zenodo.5957157" ID-GBIF-Taxon="154495864" ID-Zenodo-Dep="5957157" LSID="urn:lsid:plazi:treatment:03928782A84AFFC01E897350FA72FCFE" httpUri="http://treatment.plazi.org/id/03928782A84AFFC01E897350FA72FCFE" lastPageId="3" lastPageNumber="337" pageId="2" pageNumber="336">
<subSubSection id="C321651FA84AFFC11E897350FAE4F9C4" box="[151,1293,1556,1582]" pageId="2" pageNumber="336" type="nomenclature">
<paragraph id="8B843694A84AFFC11E897350FAE4F9C4" blockId="2.[151,1293,1556,1582]" box="[151,1293,1556,1582]" pageId="2" pageNumber="336">
<heading id="D0CC81F8A84AFFC11E897350FAE4F9C4" bold="true" box="[151,1293,1556,1582]" fontSize="11" level="1" pageId="2" pageNumber="336" reason="1">
<emphasis id="B94FEA86A84AFFC11E897350FAE4F9C4" bold="true" box="[151,1293,1556,1582]" pageId="2" pageNumber="336">
Key to the species of
<taxonomicName id="4C3B4D17A84AFFC11F827350FDEDF9C4" authorityName="Baly" authorityYear="1864" box="[412,516,1556,1582]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B94FEA86A84AFFC11F827350FDEDF9C4" bold="true" box="[412,516,1556,1582]" italics="true" pageId="2" pageNumber="336">Theopea</emphasis>
</taxonomicName>
in
<collectingCountry id="F32C7604A84AFFC11C357350FD4AF9C4" box="[555,675,1556,1582]" name="China" pageId="2" pageNumber="336">East Asia</collectingCountry>
, excluding species with modified clypeus in males
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C321651FA84AFFC01E89731EFA72FCFE" lastPageId="3" lastPageNumber="337" pageId="2" pageNumber="336" type="key">
<keyStep id="30CF2E31A84AFFC11E89731EFA72F966" pageId="2" pageNumber="336">
<paragraph id="8B843694A84AFFC11E89731EFA72F985" blockId="2.[151,1436,1626,2028]" box="[151,1435,1626,1647]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E89731EFA72F985" box="[151,1435,1626,1647]" pageId="2" pageNumber="336">1 Longitudinal ridges on elytra distinct...................................................................... 2</keyLead>
</paragraph>
<paragraph id="8B843694A84AFFC11E897333FA72F966" blockId="2.[151,1436,1626,2028]" box="[151,1435,1655,1676]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E897333FA72F966" box="[151,1435,1655,1676]" pageId="2" pageNumber="336">- Longitudinal ridges on elytra indistinct or reduced........................................................... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84AFFC11E8973D1FA72F8E8" pageId="2" pageNumber="336">
<paragraph id="8B843694A84AFFC11E8973D1FA75F92D" blockId="2.[151,1436,1626,2028]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E8973D1FA75F92D" pageId="2" pageNumber="336">
2 Elytra with extremely coarse punctures, space between punctures narrower than diameters of punctures; head and prothorax yellow except vertex and pronotum;
<collectingCountry id="F32C7604A84AFFC11C5F73F6FD6EF92D" box="[577,647,1714,1735]" name="Taiwan" pageId="2" pageNumber="336">Taiwan</collectingCountry>
(
<figureCitation id="13002A11A84AFFC11C8A73F6FD39F92D" box="[660,720,1714,1735]" captionStart="FIGURES 2" captionStartId="4.[151,264,1892,1915]" captionTargetBox="[224,1387,200,1873]" captionTargetId="figure@4.[174,1418,193,1876]" captionTargetPageId="4" captionText="FIGURES 2. Habitus of Theopea aureoviridis Chûjô and T. collaris Kimoto. 2A. T. aureoviridis, male, dorsal view; 2B. Same, ventral view; 2C. T. aureoviridis, female, dorsal view; 2D. T. collaris, male, Tahanshan (大漢山), dorsal view; 2E. Same, ventral view; 2F. T. collaris, female, Tahanshan (大漢山), dorsal view; 2G. T. collaris, male, Kenting (墾丁), dorsal view; 2H. T. collaris, female, Nanjenshan (南仁山), dorsal view." httpUri="https://zenodo.org/record/2607151/files/figure.png" pageId="2" pageNumber="336">Figs 2</figureCitation>
D2H).........................................
<taxonomicName id="4C3B4D17A84AFFC11B1C73F6FA75F92D" authority="Kimoto" authorityName="Kimoto. Solid" authorityYear="1989" box="[1282,1436,1714,1735]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="B94FEA86A84AFFC11B1C73F6FAA2F92D" box="[1282,1355,1714,1735]" italics="true" pageId="2" pageNumber="336">collaris</emphasis>
Kimoto
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B843694A84AFFC11E89738BFA72F8E8" blockId="2.[151,1436,1626,2028]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E89738BFA72F8E8" pageId="2" pageNumber="336">- Elytra with coarse punctures, space between punctures broader than diameters of punctures; head and prothorax metallic blue or green except mouth parts............................................................................. 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84AFFC11E89724EFA75F89D" pageId="2" pageNumber="336">
<paragraph id="8B843694A84AFFC11E89724EFA75F8D7" blockId="2.[151,1436,1626,2028]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E89724EFA75F8D7" pageId="2" pageNumber="336">
3 General color metallic green; longitudinal ridges on elytra apically abbreviated from apical 1/3 (
<figureCitation id="13002A11A84AFFC11A97724EFB2EF8F5" box="[1161,1223,1802,1823]" captionStart="FIGURES 4" captionStartId="7.[151,264,1905,1928]" captionTargetBox="[172,1331,184,1883]" captionTargetId="figure@7.[151,1435,180,1892]" captionTargetPageId="7" captionText="FIGURES 4. Habitus of Theopea cheni sp. nov. and T. sauteri Chûjô. 4A. T. cheni sp. nov., male, dorsal view; 4B. Same, ventral view; 4C. T. cheni sp. nov., female, dorsal view; 4D. T. sauteri, male, Mutanchihshan (ËṚḾƜ), dorsal view; 4E. Same, ventral view; 4F. T. sauteri, male, Liyuan (Mȁ), dorsal view; 4G. T. sauteri, female, Liyuan (Mȁ), dorsal view; 4H. T. sauteri, male, Lilungshan (BÊƜ), dorsal view; 4I. Same, ventral view." httpUri="https://zenodo.org/record/2607157/files/figure.png" pageId="2" pageNumber="336">Figs 4</figureCitation>
A4C); antennomeres IIIVII straight in males (
<figureCitation id="13002A11A84AFFC11FC97263FDCDF8D6" box="[471,548,1831,1852]" captionStart="FIGURES 5" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[166,1393,217,1920]" captionTargetId="figure@8.[151,1436,193,1932]" captionTargetPageId="8" captionText="FIGURES 5. Diagnostic characters of Theopea cheni sp. nov. 5A. Antenna, male; 5B. Antenna, female; 5C. Penis, dorsal view; 5D. Penis, lateral view; 5E. Penis, ventral view; 5F. Gonocoxae; 5G. Abdominal ventrite VIII; 5H. Apical margin of abdominal ventrite V, female; 5I. Tarsi of front leg, male; 5J. Tarsi of front leg, female; 5K. Spermatheca." httpUri="https://zenodo.org/record/2607159/files/figure.png" pageId="2" pageNumber="336">Fig. 5A</figureCitation>
);
<collectingCountry id="F32C7604A84AFFC11C287263FD95F8D6" box="[566,636,1831,1852]" name="Taiwan" pageId="2" pageNumber="336">Taiwan</collectingCountry>
.....................................................
<taxonomicName id="4C3B4D17A84AFFC11AE67263FAA1F8D6" authority="Lee &amp; Bezdĕk, 2018" authorityName="Lee &amp; Bezdĕk" authorityYear="2018" box="[1272,1352,1831,1852]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="cheni" status="sp. nov.">
<emphasis id="B94FEA86A84AFFC11AE67263FAA1F8D6" box="[1272,1352,1831,1852]" italics="true" pageId="2" pageNumber="336">T. cheni</emphasis>
</taxonomicName>
<emphasis id="B94FEA86A84AFFC11B517263FA75F8D7" bold="true" box="[1359,1436,1831,1853]" pageId="2" pageNumber="336">
<taxonomicNameLabel id="A27C57FDA84AFFC11B517263FA75F8D7" box="[1359,1436,1831,1853]" pageId="2" pageNumber="336" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B843694A84AFFC11E897201FA75F89D" blockId="2.[151,1436,1626,2028]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E897201FA75F89D" pageId="2" pageNumber="336">
- General color metallic blue; longitudinal ridges on elytra not apically abbreviated; antennomeres IIIVII more or less curved................................................................................4. (
<taxonomicName id="4C3B4D17A84AFFC11AAE7226FAE0F89D" authorityName="Chujo. Solid" authorityYear="1935" box="[1200,1289,1890,1911]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="336" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B94FEA86A84AFFC11AAE7226FAE0F89D" box="[1200,1289,1890,1911]" italics="true" pageId="2" pageNumber="336">T. sauteri</emphasis>
</taxonomicName>
species group)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84AFFC11E89723BFA75F858" pageId="2" pageNumber="336">
<paragraph id="8B843694A84AFFC11E89723BFA72F87E" blockId="2.[151,1436,1626,2028]" box="[151,1435,1919,1940]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E89723BFA72F87E" box="[151,1435,1919,1940]" pageId="2" pageNumber="336">
<specimenCount id="9D3DFD1DA84AFFC11E89723BFEF5F87E" box="[151,284,1919,1940]" pageId="2" pageNumber="336" type="male">4 Males</specimenCount>
with longitudinal ridges on the elytra more or less reduced................................................ 5
</keyLead>
</paragraph>
<paragraph id="8B843694A84AFFC11E8972D9FA75F858" blockId="2.[151,1436,1626,2028]" box="[151,1436,1949,1970]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E8972D9FA75F858" box="[151,1436,1949,1970]" pageId="2" pageNumber="336">- Males with longitudinal ridges on the elytra prominent...................................................... w8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84AFFC11E8972FEFA72F806" pageId="2" pageNumber="336">
<paragraph id="8B843694A84AFFC11E8972FEFA72F825" blockId="2.[151,1436,1626,2028]" box="[151,1435,1978,1999]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E8972FEFA72F825" box="[151,1435,1978,1999]" pageId="2" pageNumber="336">
5 Penis asymmetric, bent to the right (
<figureCitation id="13002A11A84AFFC11C2272FEFD6AF825" box="[572,643,1978,1999]" captionStart="FIGURES 14" captionStartId="25.[151,264,1855,1878]" captionTargetBox="[155,1372,203,1830]" captionTargetId="figure@25.[151,1436,193,1835]" captionTargetPageId="25" captionText="FIGURES 14. Diagnostic characters of Theopea coerulea Gressitt &amp; Kimoto. 14A. Antenna, male; 14B. Antenna, female; 14C. Penis, dorsal view; 14D. Penis, lateral view; 14E. Penis, ventral view; 14F. Gonocoxae; 14G. Abdominal ventrite VIII; 14H. Apical margin of abdominal ventrite V, female; 14I. Tarsi of front leg, male; 14J. Tarsi of front leg, female; 14K. Spermatheca." httpUri="https://zenodo.org/record/2607181/files/figure.png" pageId="2" pageNumber="336">Figs 14</figureCitation>
C14E, 20C20E)................................................. 6
</keyLead>
</paragraph>
<paragraph id="8B843694A84AFFC11E897293FA72F806" blockId="2.[151,1436,1626,2028]" box="[151,1435,2007,2028]" pageId="2" pageNumber="336">
<keyLead id="30CA95A1A84AFFC11E897293FA72F806" box="[151,1435,2007,2028]" pageId="2" pageNumber="336">
- Penis symmetrical (
<figureCitation id="13002A11A84AFFC11F807293FE0CF806" box="[414,485,2007,2028]" captionStart="FIGURES 15" captionStartId="26.[151,264,1952,1975]" captionTargetBox="[151,1387,200,1927]" captionTargetId="figure@26.[151,1436,193,1932]" captionTargetPageId="26" captionText="FIGURES 15. Diagnostic characters of Theopea geiseri sp. nov. 15A. Antenna, male; 15B. Antenna, female; 15C. Penis, dorsal view; 15D. Penis, lateral view; 15E. Penis, ventral view; 15F. Gonocoxae; 15G. Abdominal ventrite VIII; 15H. Apical margin of abdominal ventrite V, female; 15I. Tarsi of front leg, male; 15J. Tarsi of front leg, female; 15K. Spermatheca." httpUri="https://zenodo.org/record/2607183/files/figure.png" pageId="2" pageNumber="336">Figs 15</figureCitation>
C15E, 18C18F)............................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84BFFC01E8975D3FA75FEEE" pageId="3" pageNumber="337">
<paragraph id="8B843694A84BFFC01E8975D3FA75FF20" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E8975D3FA75FF20" pageId="3" pageNumber="337">
6 Penis relatively slender, 10.0x longer than wide; dorsal sclerite of endophallus extremely elongate, 3.6x longer than basal piece;
<collectingCountry id="F32C7604A84BFFC01F3E75F1FEB9FF20" box="[288,336,181,202]" name="Laos" pageId="3" pageNumber="337">Laos</collectingCountry>
(
<figureCitation id="13002A11A84BFFC01F4875F1FE77FF20" box="[342,414,181,202]" captionStart="FIGURES 20" captionStartId="39.[151,264,1940,1963]" captionTargetBox="[170,1414,211,1923]" captionTargetId="figure@39.[151,1436,188,1927]" captionTargetPageId="39" captionText="FIGURES 20. Diagnostic characters of Theopea sekerkai sp. nov. 20A. Antenna, male; 20B. Antenna, female; 20C. Penis, dorsal view; 20D. Penis, lateral view; 20E. Penis, ventral view; 20F. Gonocoxae; 20G. Abdominal ventrite VIII; 20H. Apical margin of abdominal ventrite V, female; 20I. Tarsi of front leg, male; 20J. Tarsi of front leg, female; 20K. Spermatheca." httpUri="https://zenodo.org/record/2607197/files/figure.png" pageId="3" pageNumber="337">Figs 20</figureCitation>
C20E)...............................................................
<taxonomicName id="4C3B4D17A84BFFC01AFE75F1FAA0FF20" authority="Lee &amp; Bezdĕk, 2018" authorityName="Lee &amp; Bezdĕk" authorityYear="2018" box="[1248,1353,181,202]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="sekerkai" status="sp. nov.">
<emphasis id="B94FEA86A84BFFC01AFE75F1FAA0FF20" box="[1248,1353,181,202]" italics="true" pageId="3" pageNumber="337">T. sekerkai</emphasis>
</taxonomicName>
<emphasis id="B94FEA86A84BFFC01B5175F0FA75FF20" bold="true" box="[1359,1436,180,202]" pageId="3" pageNumber="337">
<taxonomicNameLabel id="A27C57FDA84BFFC01B5175F0FA75FF20" box="[1359,1436,180,202]" pageId="3" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B843694A84BFFC01E897596FA75FEEE" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E897596FA75FEEE" pageId="3" pageNumber="337">
- Penis relatively broad, 9.0x longer than wide; dorsal sclerite of endophallus less elongate, 1.6x longer than basal piece;
<collectingCountry id="F32C7604A84BFFC01B7C7596FA75FF0D" box="[1378,1436,210,231]" name="China" pageId="3" pageNumber="337">China</collectingCountry>
(
<figureCitation id="13002A11A84BFFC01EF975ABFEC6FEEE" box="[231,303,239,260]" captionStart="FIGURES 14" captionStartId="25.[151,264,1855,1878]" captionTargetBox="[155,1372,203,1830]" captionTargetId="figure@25.[151,1436,193,1835]" captionTargetPageId="25" captionText="FIGURES 14. Diagnostic characters of Theopea coerulea Gressitt &amp; Kimoto. 14A. Antenna, male; 14B. Antenna, female; 14C. Penis, dorsal view; 14D. Penis, lateral view; 14E. Penis, ventral view; 14F. Gonocoxae; 14G. Abdominal ventrite VIII; 14H. Apical margin of abdominal ventrite V, female; 14I. Tarsi of front leg, male; 14J. Tarsi of front leg, female; 14K. Spermatheca." httpUri="https://zenodo.org/record/2607181/files/figure.png" pageId="3" pageNumber="337">Figs 14</figureCitation>
C14E)...............................................................
<taxonomicName id="4C3B4D17A84BFFC01A6875ABFA75FEEE" authority="Gressitt &amp; Kimoto" authorityName="Gressitt &amp; Kimoto" box="[1142,1436,239,260]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="coerulea">
<emphasis id="B94FEA86A84BFFC01A6875ABFB0AFEEE" box="[1142,1251,239,260]" italics="true" pageId="3" pageNumber="337">T. coerulea</emphasis>
Gressitt &amp; Kimoto
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84BFFC01E897449FA75FE90" pageId="3" pageNumber="337">
<paragraph id="8B843694A84BFFC01E897449FA75FEAA" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E897449FA75FEAA" pageId="3" pageNumber="337">
7 Triangular sclerites of endophallus elongate; ventral sclerites absent; basal piece longer than apical piece, with longitudinal row of tiny teeth along lateral margin;
<collectingCountry id="F32C7604A84BFFC01C4D746EFD6DFED5" box="[595,644,298,319]" name="India" pageId="3" pageNumber="337">India</collectingCountry>
(
<figureCitation id="13002A11A84BFFC01C8F746EFD3EFED5" box="[657,727,298,319]" captionStart="FIGURES 15" captionStartId="26.[151,264,1952,1975]" captionTargetBox="[151,1387,200,1927]" captionTargetId="figure@26.[151,1436,193,1932]" captionTargetPageId="26" captionText="FIGURES 15. Diagnostic characters of Theopea geiseri sp. nov. 15A. Antenna, male; 15B. Antenna, female; 15C. Penis, dorsal view; 15D. Penis, lateral view; 15E. Penis, ventral view; 15F. Gonocoxae; 15G. Abdominal ventrite VIII; 15H. Apical margin of abdominal ventrite V, female; 15I. Tarsi of front leg, male; 15J. Tarsi of front leg, female; 15K. Spermatheca." httpUri="https://zenodo.org/record/2607183/files/figure.png" pageId="3" pageNumber="337">Figs 15</figureCitation>
C15E)......................................
<taxonomicName id="4C3B4D17A84BFFC01AF2746EFAA0FED5" authority="Lee &amp; Bezdĕk, 2018" authorityName="Lee &amp; Bezdĕk" authorityYear="2018" box="[1260,1353,298,319]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="geiseri" status="sp. nov.">
<emphasis id="B94FEA86A84BFFC01AF2746EFAA0FED5" box="[1260,1353,298,319]" italics="true" pageId="3" pageNumber="337">T. geiseri</emphasis>
</taxonomicName>
<emphasis id="B94FEA86A84BFFC01B51746EFA75FEAA" bold="true" box="[1359,1436,298,320]" pageId="3" pageNumber="337">
<taxonomicNameLabel id="A27C57FDA84BFFC01B51746EFA75FEAA" box="[1359,1436,298,320]" pageId="3" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B843694A84BFFC01E897403FA75FE90" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E897403FA75FE90" pageId="3" pageNumber="337">
- Triangular sclerites of endophallus small; ventral sclerites present; basal piece shorter than apical piece, without tiny teeth;
<collectingCountry id="F32C7604A84BFFC01EFF7421FEF4FE90" box="[225,285,357,378]" name="China" pageId="3" pageNumber="337">China</collectingCountry>
,
<collectingCountry id="F32C7604A84BFFC01F387421FEBEFE90" box="[294,343,357,378]" name="Laos" pageId="3" pageNumber="337">Laos</collectingCountry>
,
<collectingCountry id="F32C7604A84BFFC01F7F7421FE5BFE90" box="[353,434,357,378]" name="Vietnam" pageId="3" pageNumber="337">Vietnam</collectingCountry>
(
<figureCitation id="13002A11A84BFFC01FA07421FDEFFE90" box="[446,518,357,378]" captionStart="FIGURES 18" captionStartId="31.[151,264,1855,1878]" captionTargetBox="[155,1432,217,1835]" captionTargetId="figure@31.[151,1436,193,1835]" captionTargetPageId="31" captionText="FIGURES 18. Diagnostic characters of Theopea laosensis sp. nov. 18A. Antenna, male; 18B. Antenna, female; 18C. Penis, dorsal view; 18D. Endophallic sclerites, Vietnam; 18E. Penis, lateral view; 18F. Penis, ventral view; 18G. Gonocoxae; 18H. Abdominal ventrite VIII; 18I. Apical margin of abdominal ventrite V, female; 18J. Tarsi of front leg, male; 18K. Tarsi of front leg, female; 18L. Spermatheca." httpUri="https://zenodo.org/record/2607189/files/figure.png" pageId="3" pageNumber="337">Figs 18</figureCitation>
C18F)......................................................
<emphasis id="B94FEA86A84BFFC01AC37421FAA1FE90" box="[1245,1352,357,378]" italics="true" pageId="3" pageNumber="337">
..
<taxonomicName id="4C3B4D17A84BFFC01AEE7421FAA1FE90" authority="Lee &amp; Bezdĕk, 2018" authorityName="Lee &amp; Bezdĕk" authorityYear="2018" box="[1264,1352,357,378]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="laosensis" status="sp. nov.">laosensis</taxonomicName>
</emphasis>
<emphasis id="B94FEA86A84BFFC01B517420FA75FE90" bold="true" box="[1359,1436,356,378]" pageId="3" pageNumber="337">
<taxonomicNameLabel id="A27C57FDA84BFFC01B517420FA75FE90" box="[1359,1436,356,378]" pageId="3" pageNumber="337" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84BFFC01E8974C6FA75FE1A" pageId="3" pageNumber="337">
<paragraph id="8B843694A84BFFC01E8974C6FA75FE5E" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<materialsCitation id="3B533CC9A84BFFC01E8974C6FA75FE5E" ID-GBIF-Occurrence="2028139097" collectingDate="1919-01" country="Taiwan" location="Taiwan" pageId="3" pageNumber="337" specimenCount="8" specimenCount-male="8">
<keyLead id="30CA95A1A84BFFC01E8974C6FA75FE5E" pageId="3" pageNumber="337">
<specimenCount id="9D3DFD1DA84BFFC01E8974C6FEF5FE7D" box="[151,284,386,407]" pageId="3" pageNumber="337" type="male">8 Males</specimenCount>
with antennomeres IIIX moderately curved (
<figureCitation id="13002A11A84BFFC01CC374C6FCDEFE7D" box="[733,823,386,407]" captionStart="FIGURES 19" captionStartId="37.[151,264,1952,1975]" captionTargetBox="[158,1408,204,1924]" captionTargetId="figure@37.[151,1436,193,1932]" captionTargetPageId="37" captionText="FIGURES 19. Diagnostic characters of Theopea sauteri Chûjô. 19A. Antenna, male; 19B. Antenna, female; 19C. Penis, dorsal view; 19D. Penis, lateral view; 19E. Penis, ventral view; 19F. Gonocoxae; 19G. Abdominal ventrite VIII; 19H. Apical margin of abdominal ventrite V, female; 19I. Tarsi of front leg, male; 19J. Tarsi of front leg, female; 19K. Spermatheca." httpUri="https://zenodo.org/record/2607195/files/figure.png" pageId="3" pageNumber="337">Fig. 19A</figureCitation>
); swollen tarsomeres I of front legs not apically narrowed (
<figureCitation id="13002A11A84BFFC01B6B74C6FEEAFE5E" captionStart="FIGURES 19" captionStartId="37.[151,264,1952,1975]" captionTargetBox="[158,1408,204,1924]" captionTargetId="figure@37.[151,1436,193,1932]" captionTargetPageId="37" captionText="FIGURES 19. Diagnostic characters of Theopea sauteri Chûjô. 19A. Antenna, male; 19B. Antenna, female; 19C. Penis, dorsal view; 19D. Penis, lateral view; 19E. Penis, ventral view; 19F. Gonocoxae; 19G. Abdominal ventrite VIII; 19H. Apical margin of abdominal ventrite V, female; 19I. Tarsi of front leg, male; 19J. Tarsi of front leg, female; 19K. Spermatheca." httpUri="https://zenodo.org/record/2607195/files/figure.png" pageId="3" pageNumber="337">
Fig.
<date id="FF851054A84BFFC01EFF74DBFEEAFE5E" box="[225,259,415,436]" pageId="3" pageNumber="337" value="1919-01">
<collectingDate id="EFC1E9BCA84BFFC01EFF74DBFEEAFE5E" box="[225,259,415,436]" pageId="3" pageNumber="337" value="1919-01">19I</collectingDate>
</date>
</figureCitation>
);
<collectingCountry id="F32C7604A84BFFC01F0B74DBFEB2FE5E" box="[277,347,415,436]" name="Taiwan" pageId="3" pageNumber="337">Taiwan</collectingCountry>
.............................................................................
<taxonomicName id="4C3B4D17A84BFFC01B1F74DBFA75FE5E" authority="Chujo" authorityName="Chujo. Solid" authorityYear="1935" box="[1281,1436,415,436]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B94FEA86A84BFFC01B1F74DBFAB2FE5E" box="[1281,1371,415,436]" italics="true" pageId="3" pageNumber="337">T. sauteri</emphasis>
Chûjô
</taxonomicName>
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="8B843694A84BFFC01E8974F9FA75FE1A" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E8974F9FA75FE1A" pageId="3" pageNumber="337">
- Males with antennomeres IIIX straight (
<figureCitation id="13002A11A84BFFC01C7374F9FD2EFE38" box="[621,711,445,466]" captionStart="FIGURES 16" captionStartId="28.[151,264,1943,1966]" captionTargetBox="[221,1347,609,1899]" captionTargetId="figure@28.[218,1353,589,1924]" captionTargetPageId="28" captionText="FIGURES 16. Diagnostic characters of Theopea hainanensis sp. nov. 16A. Antenna, male; 16B. Penis, dorsal view; 16C. Penis, lateral view; 16D. Penis, ventral view; 16E. Tarsi of front leg, male; AP: apical piece, BP: basal piece, DS: dorsal sclerite, TS: triangular sclerite, VS: ventral sclerite." httpUri="https://zenodo.org/record/2607185/files/figure.png" pageId="3" pageNumber="337">Fig. 16A</figureCitation>
); swollen tarsomeres I of front legs apically narrowed (
<figureCitation id="13002A11A84BFFC01AC774F9FAC6FE38" box="[1241,1327,445,466]" captionStart="FIGURES 16" captionStartId="28.[151,264,1943,1966]" captionTargetBox="[221,1347,609,1899]" captionTargetId="figure@28.[218,1353,589,1924]" captionTargetPageId="28" captionText="FIGURES 16. Diagnostic characters of Theopea hainanensis sp. nov. 16A. Antenna, male; 16B. Penis, dorsal view; 16C. Penis, lateral view; 16D. Penis, ventral view; 16E. Tarsi of front leg, male; AP: apical piece, BP: basal piece, DS: dorsal sclerite, TS: triangular sclerite, VS: ventral sclerite." httpUri="https://zenodo.org/record/2607185/files/figure.png" pageId="3" pageNumber="337">Fig. 16E</figureCitation>
);
<collectingCountry id="F32C7604A84BFFC01B5F74F9FA6EFE38" box="[1345,1415,445,466]" name="Taiwan" pageId="3" pageNumber="337">Taiwan</collectingCountry>
...................................................................................
<taxonomicName id="4C3B4D17A84BFFC01AA7749EFAABFE05" authorityName="Lee &amp; Bezdĕk" authorityYear="2018" box="[1209,1346,474,495]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="hainanensis">
<emphasis id="B94FEA86A84BFFC01AA7749EFAABFE05" box="[1209,1346,474,495]" italics="true" pageId="3" pageNumber="337">T. hainanensis</emphasis>
</taxonomicName>
.
<emphasis id="B94FEA86A84BFFC01B51749EFA75FE1A" bold="true" box="[1359,1436,474,496]" pageId="3" pageNumber="337">sp. nov.</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84BFFC01E8974B3FA75FDAD" pageId="3" pageNumber="337">
<paragraph id="8B843694A84BFFC01E8974B3FA75FDC0" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E8974B3FA75FDC0" pageId="3" pageNumber="337">
9 Elytra with extremely coarse punctures, space between punctures narrower than diameters of punctures (
<figureCitation id="13002A11A84BFFC01AD574B3FAEEFDE6" box="[1227,1287,503,524]" captionStart="FIGURES 2" captionStartId="4.[151,264,1892,1915]" captionTargetBox="[224,1387,200,1873]" captionTargetId="figure@4.[174,1418,193,1876]" captionTargetPageId="4" captionText="FIGURES 2. Habitus of Theopea aureoviridis Chûjô and T. collaris Kimoto. 2A. T. aureoviridis, male, dorsal view; 2B. Same, ventral view; 2C. T. aureoviridis, female, dorsal view; 2D. T. collaris, male, Tahanshan (大漢山), dorsal view; 2E. Same, ventral view; 2F. T. collaris, female, Tahanshan (大漢山), dorsal view; 2G. T. collaris, male, Kenting (墾丁), dorsal view; 2H. T. collaris, female, Nanjenshan (南仁山), dorsal view." httpUri="https://zenodo.org/record/2607151/files/figure.png" pageId="3" pageNumber="337">Figs 2</figureCitation>
A2C);
<collectingCountry id="F32C7604A84BFFC01B4B74B3FA65FDE6" box="[1365,1420,503,524]" name="Japan" pageId="3" pageNumber="337">Japan</collectingCountry>
.....................................................................................
<taxonomicName id="4C3B4D17A84BFFC01AD17751FA75FDC0" authority="Chujo" authorityName="Chujo" box="[1231,1436,533,554]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="aureoviridis">
<emphasis id="B94FEA86A84BFFC01AD17751FAB2FDC0" box="[1231,1371,533,554]" italics="true" pageId="3" pageNumber="337">T. aureoviridis</emphasis>
Chûjô
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B843694A84BFFC01E897776FA75FDAD" blockId="3.[151,1436,151,788]" box="[151,1436,562,583]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E897776FA75FDAD" box="[151,1436,562,583]" pageId="3" pageNumber="337">
- Elytra with moderately coarse punctures, space between punctures broader than diameters of punctures (
<figureCitation id="13002A11A84BFFC01AD77776FAEEFDAD" box="[1225,1287,562,583]" captionStart="FIGURES 8" captionStartId="13.[151,264,1911,1934]" captionTargetBox="[240,1371,200,1893]" captionTargetId="figure@13.[196,1401,193,1894]" captionTargetPageId="13" captionText="FIGURES 8. Habitus of Theopea irregularis Takizawa. 8A. Male, Taipeingshan (‡ ṬƜ), dorsal view; 8B. Same, ventral view; 8C. Male, Motien (±), dorsal view; 8D. Male, Tsuifeng (Ṝẘ), dorsal view; 8E. Female, Anmashan (ḎḄƜ), dorsal view; 8F. Same, ventral view; 8G. Female, Mingchi (ŖḾ), dorsal view; 8H. Male, Tatachia (ḆḆffi), dorsal view; 8I. Female, Tatachia (ḆḆffi), dorsal view." httpUri="https://zenodo.org/record/2607165/files/figure.png" pageId="3" pageNumber="337">Figs 8</figureCitation>
,
<figureCitation id="13002A11A84BFFC01B0C7776FAC2FDAD" box="[1298,1323,562,583]" captionStart="FIGURES 11" captionStartId="19.[151,264,1905,1928]" captionTargetBox="[181,1387,185,1905]" captionTargetId="figure@19.[151,1435,181,1893]" captionTargetPageId="19" captionText="FIGURES 11. Habitus of Theopea kanmiyai (Kimoto). 11A. Male, Alishan (ȐBƜ), dorsal view; 11B. Same, ventral view; 11C. Male, Motien (±), dorsal view; 11D. Male, Tahanshan (XÃƜ), dorsal view; 11E. Female, Paling (ƂD), dorsal view; 11F. Female, Chungchihkuan (¢ŽṞ), dorsal view; 11G. Female, Tahanshan (XÃƜ), dorsal view; 11H. Female, Tahanshan (XÃƜ), dorsal view." httpUri="https://zenodo.org/record/2607173/files/figure.png" pageId="3" pageNumber="337">11</figureCitation>
)....... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CF2E31A84BFFC01E89770BFA72FCFE" pageId="3" pageNumber="337">
<paragraph id="8B843694A84BFFC01E89770BFA75FD56" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E89770BFA75FD56" pageId="3" pageNumber="337">
10 General color sexually dimorphic, elytra yellowish brown with metallic green sides, pronotum and vertex metallic green in males; elytra entirely metallic green, prothorax and head yellow in females; hypomeron yellowish brown (
<figureCitation id="13002A11A84BFFC01AC17729FACCFD68" box="[1247,1317,621,642]" captionStart="FIGURES 11" captionStartId="19.[151,264,1905,1928]" captionTargetBox="[181,1387,185,1905]" captionTargetId="figure@19.[151,1435,181,1893]" captionTargetPageId="19" captionText="FIGURES 11. Habitus of Theopea kanmiyai (Kimoto). 11A. Male, Alishan (ȐBƜ), dorsal view; 11B. Same, ventral view; 11C. Male, Motien (±), dorsal view; 11D. Male, Tahanshan (XÃƜ), dorsal view; 11E. Female, Paling (ƂD), dorsal view; 11F. Female, Chungchihkuan (¢ŽṞ), dorsal view; 11G. Female, Tahanshan (XÃƜ), dorsal view; 11H. Female, Tahanshan (XÃƜ), dorsal view." httpUri="https://zenodo.org/record/2607173/files/figure.png" pageId="3" pageNumber="337">Fig. 11</figureCitation>
); antenna in males relatively shorter, antennomeres VIX less than six times longer than wide (
<figureCitation id="13002A11A84BFFC01DE977CEFBB8FD75" box="[1015,1105,650,671]" captionStart="FIGURES 12" captionStartId="20.[151,264,1855,1878]" captionTargetBox="[161,1395,219,1829]" captionTargetId="figure@20.[151,1436,193,1835]" captionTargetPageId="20" captionText="FIGURES 12. Diagnostic characters of Theopea kanmiyai (Kimoto). 11A. Antenna, male; 11B. Antenna, female; 11C. Penis, dorsal view; 11D. Penis, lateral view; 11E. Penis, ventral view; 11F. Gonocoxae; 11G. Abdominal ventrite VIII; 11H. Apical margin of abdominal ventrite V, female; 11I. Tarsi of front leg, male; 11J. Tarsi of front leg, female; 11K. Spermatheca." httpUri="https://zenodo.org/record/2607177/files/figure.png" pageId="3" pageNumber="337">Fig. 12A</figureCitation>
);
<collectingCountry id="F32C7604A84BFFC01A7C77CEFB41FD75" box="[1122,1192,650,671]" name="Taiwan" pageId="3" pageNumber="337">Taiwan</collectingCountry>
........................................................................................................
<taxonomicName id="4C3B4D17A84BFFC01AD777E3FA75FD56" authority="(Kimoto)" baseAuthorityName="Kimoto" box="[1225,1436,679,700]" class="Insecta" family="Chrysomelidae" genus="Hoplosaenidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="kanmiyai">
<emphasis id="B94FEA86A84BFFC01AD777E3FAD5FD56" box="[1225,1340,679,700]" italics="true" pageId="3" pageNumber="337">T. kanmiyai</emphasis>
(Kimoto)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B843694A84BFFC01E897781FA72FCFE" blockId="3.[151,1436,151,788]" pageId="3" pageNumber="337">
<keyLead id="30CA95A1A84BFFC01E897781FA72FCFE" pageId="3" pageNumber="337">
- General color not sexually dimorphic, elytra, pronotum, and vertex metallic green or blue in both sexes; hypomeron dark or blackish brown (
<figureCitation id="13002A11A84BFFC01F9C77A6FE54FD1D" box="[386,445,738,759]" captionStart="FIGURES 8" captionStartId="13.[151,264,1911,1934]" captionTargetBox="[240,1371,200,1893]" captionTargetId="figure@13.[196,1401,193,1894]" captionTargetPageId="13" captionText="FIGURES 8. Habitus of Theopea irregularis Takizawa. 8A. Male, Taipeingshan (‡ ṬƜ), dorsal view; 8B. Same, ventral view; 8C. Male, Motien (±), dorsal view; 8D. Male, Tsuifeng (Ṝẘ), dorsal view; 8E. Female, Anmashan (ḎḄƜ), dorsal view; 8F. Same, ventral view; 8G. Female, Mingchi (ŖḾ), dorsal view; 8H. Male, Tatachia (ḆḆffi), dorsal view; 8I. Female, Tatachia (ḆḆffi), dorsal view." httpUri="https://zenodo.org/record/2607165/files/figure.png" pageId="3" pageNumber="337">Fig. 8</figureCitation>
); antenna in males more slender, antennomeres VIX more than six times longer than wide (
<figureCitation id="13002A11A84BFFC01B5F77A6FEE9FCFE" captionStart="FIGURES 9" captionStartId="14.[151,264,1952,1975]" captionTargetBox="[157,1430,215,1928]" captionTargetId="figure@14.[151,1436,193,1932]" captionTargetPageId="14" captionText="FIGURES 9. Diagnostic characters of Theopea irregularis Takizawa. 9A. Antenna, male; 9B. Antenna, male; 9C. Antenna, female; 9E. Penis, dorsal view; 9E. Penis, lateral view; 9F. Penis, ventral view; 9G. Gonocoxae; 9H. Abdominal ventrite VIII; 9I. Apical margin of abdominal ventrite V, female; 9J. Tarsi of front leg, male; 9K. Tarsi of front leg, female; 9L. Spermatheca." httpUri="https://zenodo.org/record/2607169/files/figure.png" pageId="3" pageNumber="337">Figs. 9A, 9B</figureCitation>
);
<collectingCountry id="F32C7604A84BFFC01F0F77BBFEBEFCFE" box="[273,343,767,788]" name="Taiwan" pageId="3" pageNumber="337">Taiwan</collectingCountry>
........................................................................
<taxonomicName id="4C3B4D17A84BFFC01AA577BBFA72FCFE" authority="Takizawa" authorityName="Takizawa" authorityYear="1978" box="[1211,1435,767,788]" class="Insecta" family="Chrysomelidae" genus="Hoplosaenidea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="337" phylum="Arthropoda" rank="species" species="irregularis">
<emphasis id="B94FEA86A84BFFC01AA577BBFAD3FCFE" box="[1211,1338,767,788]" italics="true" pageId="3" pageNumber="337">T. irregularis</emphasis>
Takizawa
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>