treatments-xml/data/03/92/5B/03925B12ED6BFFB6FF5FFF7EFDBDFE95.xml

<document id="03D3DE37A73E701624327426BD93F861" ID-CLB-Dataset="9278" ID-DOI="10.11646/zootaxa.4895.1.6" ID-GBIF-Dataset="277eca87-67c5-42bf-b494-397887598bb9" ID-ISSN="1175-5326" ID-Zenodo-Dep="4321548" ID-ZooBank="988D35CE-5652-485F-A1F4-01E55D3F20C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607972527210" checkinUser="plazi" docAuthor="Medeiros, Lucas Silva De, Moreira, Cristiano Rangel, Pinna, Mário De &amp; Lima, Sergio Maia Queiroz" docDate="2020" docId="03925B12ED6BFFB6FF5FFF7EFDBDFE95" docLanguage="en" docName="zootaxa.4895.1.6.pdf" docOrigin="Zootaxa 4895 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microcambeva bendego Medeiros &amp; Moreira &amp; Pinna &amp; Lima 2020, new species" docType="treatment" docVersion="10" lastPageNumber="120" masterDocId="FFAB236AED69FFBCFFC8FFE9FFC5FFF0" masterDocTitle="A new catfish species of Microcambeva Costa &amp; Bockmann 1994 (Siluriformes Trichomycteridae) from a coastal basin in Rio de Janeiro State, southeastern Brazil" masterLastPageNumber="123" masterPageNumber="111" pageNumber="113" updateTime="1698847546314" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="41DC19B8485477C9F28F8F3B3F2B2BC7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6C5588E22CDD81C36DF57797FC14C24A">
<mods:title id="10905ED172B384D791DFD64D77ADADE5">A new catfish species of Microcambeva Costa &amp; Bockmann 1994 (Siluriformes Trichomycteridae) from a coastal basin in Rio de Janeiro State, southeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="DC5625939CEFDAE0ABB96222B0F38FC9" type="personal">
<mods:role id="B1153587CBF558EE43A529A4306344B9">
<mods:roleTerm id="AC072C6CCE3A85EB50032C5E7B22E26E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4DB2E18146277E10F8ACD6C2535C079C">Medeiros, Lucas Silva De</mods:namePart>
<mods:nameIdentifier id="2A4CC29292AEC9B6220206B973205C2C" type="ORCID">0000-0001-9859-2128</mods:nameIdentifier>
<mods:affiliation id="73B57BF18163EE5A134B58EC1C2963A8">Laboratório de Ictiologia Sistemática e Evolutiva, Departamento de Botânica e Zoologia, Centro de Biociências, Universidade Federal do Rio Grande do Norte, 59978 - 970 Natal, RN, Brazil. &amp; lucasmedeiros 0 @ hotmail. com; https: // orcid. org / 0000 - 0001 - 9859 - 2128</mods:affiliation>
<mods:nameIdentifier id="3D344B8B1243C8E6CEF4A12FEC4F5013" type="email">lucasmedeiros0@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="902265B2BE72D0871EAB9AB6E39F578D" type="personal">
<mods:role id="4820604DC49B39B5E8840EE56E532B87">
<mods:roleTerm id="3499AF7CF6BB912C1D7CEBF19220946C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C9FDCE96682CEF4751E06A61394E0FC">Moreira, Cristiano Rangel</mods:namePart>
<mods:nameIdentifier id="9968494B169011C435228CF98586CCBB" type="ORCID">0000-0003-4599-7214</mods:nameIdentifier>
<mods:affiliation id="7AFCF5E0883F2B567733814AB9C5BD3C">Museu Nacional, Universidade Federal do Rio de Janeiro, Departamento de Vertebrados, Quinta da Boa Vista, 20940 - 040 Rio de Janeiro, RJ, Brazil. moreira. c. r @ gmail. com; https: // orcid. org / 0000 - 0003 - 4599 - 7214</mods:affiliation>
<mods:nameIdentifier id="11D709DE96C2E678BA53F06EF792BA24" type="email">moreira.c.r@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2789861991B977F91070F6F8B1885029" type="personal">
<mods:role id="67A8DF64F296DBAEE42C14483202A36F">
<mods:roleTerm id="598F0676596CF73A25EEBC56DA275207">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="52C5BA43B7013C5AAAB793581D276B6D">Pinna, Mário De</mods:namePart>
<mods:nameIdentifier id="2900E01AAB3B2CCAAA5340975657AE4B" type="ORCID">0000-0003-1711-4816</mods:nameIdentifier>
<mods:affiliation id="822A030A2847921A95E6502AF62271E6">Museu de Zoologia da Universidade de São Paulo, Universidade de São Paulo, Av. Nazaré, 481, 04263 - 000 São Paulo, SP, Brazil. pinna @ ib. usp. br; https: // orcid. org / 0000 - 0003 - 1711 - 4816</mods:affiliation>
<mods:nameIdentifier id="5FD60A435C0E7402F2E62D6A471E1CEB" type="email">pinna@ib.usp.br</mods:nameIdentifier>
</mods:name>
<mods:name id="EB01AC87BADDD8088203DA4639A5CF8F" type="personal">
<mods:role id="942524D8C7471D85BC99A9D739BEF335">
<mods:roleTerm id="52E832FEDCB159619E95EEDC104FB602">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2FE8808CD30C6D1CC60740F535BF1527">Lima, Sergio Maia Queiroz</mods:namePart>
<mods:nameIdentifier id="4A7E1982CBB3B4E32CB3B5CFC9A1FCB6" type="ORCID">0000-0001-9365-4879</mods:nameIdentifier>
<mods:affiliation id="3B78C5B0D86210DE23652D8D9806304E">Laboratório de Ictiologia Sistemática e Evolutiva, Departamento de Botânica e Zoologia, Centro de Biociências, Universidade Federal do Rio Grande do Norte, 59978 - 970 Natal, RN, Brazil. &amp; smaialima @ gmail. com; https: // orcid. org / 0000 - 0001 - 9365 - 4879</mods:affiliation>
<mods:nameIdentifier id="0D12B67A76C5BDF2C1D59C28327F2400" type="email">smaialima@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="FFC02AA40C20EA97B1660B992805A927">text</mods:typeOfResource>
<mods:relatedItem id="FA8F6041A5F13489140CECECD444571B" type="host">
<mods:titleInfo id="D5206AEE45B044744C1630AF13FBE404">
<mods:title id="A09B4AF2E6F58CE194BD895069753627">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B406EFF240ADC9F6AEA72C4CC14DD276">
<mods:date id="87EBA84FFB2F3FE95EC4615C14FC713E">2020</mods:date>
<mods:detail id="5C39DC53B64EDE23213230FD9DEAEE3C" type="pubDate">
<mods:number id="3AC5DAD5C3C85D793D0F8DEA28FB7D7B">2020-12-14</mods:number>
</mods:detail>
<mods:detail id="27DAEAF2EBA22800596B677C5FD00AB1" type="volume">
<mods:number id="39BECCF1E94B5951CEB12D078A3AED14">4895</mods:number>
</mods:detail>
<mods:detail id="DF1A529591AD6E52334C6C9C5F98A5C7" type="issue">
<mods:number id="D3A7BD52BB794BD30D5E61C4FD19336D">1</mods:number>
</mods:detail>
<mods:extent id="219D1F33144A1B9526B47B2EFB1DA179" unit="page">
<mods:start id="15AF116C06821F86BB6D185CE92BDE39">111</mods:start>
<mods:end id="3A9FB1E0B4ED034A9EF402BB2D3B8ED1">123</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="48BAFF15DA8D64617E3A6FC860508C92">journal article</mods:classification>
<mods:identifier id="74BC58610E2E417B9957AC25C793F362" type="CLB-Dataset">9278</mods:identifier>
<mods:identifier id="B0EB1E48828BC1D0AA08E9B576732D24" type="DOI">10.11646/zootaxa.4895.1.6</mods:identifier>
<mods:identifier id="42772D5DDD334D872F9B0D34FB783254" type="GBIF-Dataset">277eca87-67c5-42bf-b494-397887598bb9</mods:identifier>
<mods:identifier id="8462340F277040860456C1DA36D44C4F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D077B23E2DE79936F35F8BFC1EDB01E9" type="Zenodo-Dep">4321548</mods:identifier>
<mods:identifier id="4E2EC78DDCAFE50F01A80DBAC3E77B57" type="ZooBank">988D35CE-5652-485F-A1F4-01E55D3F20C9</mods:identifier>
</mods:mods>
<treatment id="03925B12ED6BFFB6FF5FFF7EFDBDFE95" ID-DOI="http://doi.org/10.5281/zenodo.4323864" ID-GBIF-Taxon="176092295" ID-Zenodo-Dep="4323864" LSID="urn:lsid:plazi:treatment:03925B12ED6BFFB6FF5FFF7EFDBDFE95" httpUri="http://treatment.plazi.org/id/03925B12ED6BFFB6FF5FFF7EFDBDFE95" lastPageId="10" lastPageNumber="120" pageId="2" pageNumber="113">
<subSubSection id="C321B98FED6BFFBEFF5FFF7EFD8BFF41" box="[151,590,151,178]" pageId="2" pageNumber="113" type="nomenclature">
<paragraph id="8B84EA04ED6BFFBEFF5FFF7EFD8BFF41" blockId="2.[151,590,151,213]" box="[151,590,151,178]" pageId="2" pageNumber="113">
<heading id="D0CC5D68ED6BFFBEFF5FFF7EFD8BFF41" bold="true" box="[151,590,151,178]" fontSize="11" level="1" pageId="2" pageNumber="113" reason="1">
<emphasis id="B94F3616ED6BFFBEFF5FFF7EFD8BFF41" bold="true" box="[151,590,151,178]" pageId="2" pageNumber="113">
<taxonomicName id="4C3B9187ED6BFFBEFF5FFF7EFE72FF42" authority="Medeiros &amp; Moreira &amp; Pinna &amp; Lima, 2020" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[151,439,151,178]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="bendego" status="sp. nov.">
<emphasis id="B94F3616ED6BFFBEFF5FFF7EFE72FF42" bold="true" box="[151,439,151,178]" italics="true" pageId="2" pageNumber="113">Microcambeva bendego</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27C8B6DED6BFFBEFE76FF7EFD8BFF41" box="[446,590,151,177]" pageId="2" pageNumber="113" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C321B98FED6BFFBEFF5FFF52FE8EFF25" box="[151,331,187,213]" pageId="2" pageNumber="113" type="description">
<paragraph id="8B84EA04ED6BFFBEFF5FFF52FE8EFF25" blockId="2.[151,590,151,213]" box="[151,331,187,213]" pageId="2" pageNumber="113">
(
<figureCitation id="1300F681ED6BFFBEFF68FF52FF27FF25" box="[160,226,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1945,1969]" captionTargetBox="[151,1435,993,1921]" captionTargetId="figure-362@2.[151,1435,993,1921]" captionTargetPageId="2" captionText="FIGURE 1. Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Rio Guapiaçu, near Cachoeiras de Macacu, rio Guapi-Macacu basin, Guapimirim Municipality, Rio de Janeiro State, southeastern Brazil. a. lateral view; b. dorsal view; c. ventral view. Scale: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4321550" httpUri="https://zenodo.org/record/4321550/files/figure.png" pageId="2" pageNumber="113">Fig. 1</figureCitation>
, 
<tableCitation id="C6B9DFBFED6BFFBEFF27FF52FE85FF25" box="[239,320,187,213]" captionStart="TABLE 1" captionStartId="3.[151,239,151,175]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data of holotype (MNRJ 52042, 28.1 mm SL) and two paratypes (MNRJ 48616, 26.94 mm SL and MZUSP 125789, 27.8 mm SL) of Microcambeva bendego." httpUri="http://table.plazi.org/id/DF44BA8CED6AFFBFFF5FFF7EFCFDFF23" pageId="2" pageNumber="113" tableUuid="DF44BA8CED6AFFBFFF5FFF7EFCFDFF23">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C321B98FED6BFFBEFF5FFEEAFEFFFE5D" pageId="2" pageNumber="113" type="materials_examined">
<paragraph id="8B84EA04ED6BFFBEFF5FFEEAFEB2FE95" blockId="2.[151,1437,259,970]" pageId="2" pageNumber="113">
<materialsCitation id="3B53E059ED6BFFBEFF5FFEEAFEB2FE95" ID-GBIF-Occurrence="2995635302" collectingDate="2016-08-31" collectionCode="V" collectorName="P. A. Buckup &amp; D. F. Moraes-Jr &amp; V. Brito" country="Brazil" county="Guapimirim Municipality" latitude="-22.5925" location="Guapimirim Municipality" longLatPrecision="21" longitude="-42.88889" municipality="rio Guapiacu" pageId="2" pageNumber="113" specimenCode="MNRJ 52042" specimenCount="1" stateProvince="Rio de Janeiro State" typeStatus="holotype">
<emphasis id="B94F3616ED6BFFBEFF5FFEEAFECEFEED" bold="true" box="[151,267,259,285]" pageId="2" pageNumber="113">
<typeStatus id="548054A6ED6BFFBEFF5FFEEAFEC3FEED" box="[151,262,259,285]" pageId="2" pageNumber="113" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB9D427FED6BFFBEFEDBFEEDFE6AFEEE" box="[275,431,259,286]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="113">MNRJ 52042</specimenCode>
, 
<quantity id="4CC347E1ED6BFFBEFE73FEEAFDE5FEEE" box="[443,544,259,286]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="2" pageNumber="113" unit="mm" value="28.1">28.1 mm</quantity>
SL; 
<collectingCountry id="F32CAA94ED6BFFBEFD90FEEAFD64FEED" box="[600,673,259,285]" name="Brazil" pageId="2" pageNumber="113">Brazil</collectingCountry>
: 
<collectingRegion id="49FF24E6ED6BFFBEFD65FEEAFC51FEED" box="[685,916,259,285]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="113">Rio de Janeiro State</collectingRegion>
, 
<collectingCounty id="62E59288ED6BFFBEFC68FEEAFB00FEED" box="[928,1221,259,285]" pageId="2" pageNumber="113">Guapimirim Municipality</collectingCounty>
; rio 
<locationDeviation id="9A36F326ED6BFFBEFB31FEEAFD78FEB1" pageId="2" pageNumber="113">Guapiaçu near Cachoeiras de Macacu, rio Guapi-Macacu basin</locationDeviation>
, 
<geoCoordinate id="EE0F8CC3ED6BFFBEFD01FECEFC8BFEB2" box="[713,846,295,322]" degrees="22" direction="south" minutes="35" orientation="latitude" pageId="2" pageNumber="113" precision="15" seconds="33" value="-22.5925">22°35’33”S</geoCoordinate>
<geoCoordinate id="EE0F8CC3ED6BFFBEFC9FFECEFC23FEB2" box="[855,998,295,322]" degrees="42" direction="west" minutes="53" orientation="longitude" pageId="2" pageNumber="113" precision="15" seconds="20" value="-42.88889">42°53’20”W</geoCoordinate>
, 
<collectorName id="26CE8FD2ED6BFFBEFC3BFEC1FB49FEB1" box="[1011,1164,295,322]" pageId="2" pageNumber="113">P. A. Buckup</collectorName>
, 
<collectorName id="26CE8FD2ED6BFFBEFB51FEC1FA89FEB2" box="[1177,1356,296,322]" pageId="2" pageNumber="113">D. F. Moraes-Jr</collectorName>
and 
<collectorName id="26CE8FD2ED6BFFBEFA4DFEC1FF16FE95" pageId="2" pageNumber="113">V. Brito</collectorName>
, 
<date id="FF85CCC4ED6BFFBEFF17FEA2FEB6FE95" box="[223,371,331,357]" pageId="2" pageNumber="113" value="2016-08-31">
<collectingDate id="EFC1352CED6BFFBEFF17FEA2FEB6FE95" box="[223,371,331,357]" pageId="2" pageNumber="113" value="2016-08-31">31 Aug 2016</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B84EA04ED6BFFBEFF0FFE86FEFFFE5D" blockId="2.[151,1437,259,970]" pageId="2" pageNumber="113">
<materialsCitation id="3B53E059ED6BFFBEFF0FFE86FC76FE79" ID-GBIF-Occurrence="2995635303" box="[199,947,367,394]" collectingDate="2016-08-31" collectionCode="MNRJ" collectorName="de Macacu &amp; P. A. Buckup &amp; Brito" country="Brazil" county="Guapimirim Municipality" latitude="-22.5925" location="Cachoeiras de Macacu" longLatPrecision="21" longitude="-42.88889" municipality="rio Guapiacu" pageId="2" pageNumber="113" specimenCode="MNRJ 48616" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<emphasis id="B94F3616ED6BFFBEFF0FFE86FE83FE79" bold="true" box="[199,326,367,393]" pageId="2" pageNumber="113">
<typeStatus id="548054A6ED6BFFBEFF0FFE86FE84FE79" box="[199,321,367,393]" pageId="2" pageNumber="113" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DB9D427FED6BFFBEFE85FE99FE2DFE79" box="[333,488,367,394]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="113">MNRJ 48616</specimenCode>
, 1, 
<quantity id="4CC347E1ED6BFFBEFDC7FE86FDB6FE7A" box="[527,627,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="2" pageNumber="113" unit="mm" value="26.9">26.9 mm</quantity>
SL, collected with holotype
</materialsCitation>
. 
<materialsCitation id="3B53E059ED6BFFBEFC75FE86FEF3FE5D" ID-GBIF-Occurrence="2995635301" collectingDate="2016-08-31" collectionCode="MZUSP" collectorName="de Macacu &amp; P. A. Buckup &amp; Brito" country="Brazil" county="Guapimirim Municipality" latitude="-22.5925" location="Cachoeiras de Macacu" longLatPrecision="21" longitude="-42.88889" municipality="rio Guapiacu" pageId="2" pageNumber="113" specimenCode="MZUSP 125789" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="paratype">
<specimenCode id="DB9D427FED6BFFBEFC75FE86FBB2FE7A" box="[957,1143,367,394]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="113">MZUSP 125789</specimenCode>
, 1, 
<quantity id="4CC347E1ED6BFFBEFB56FE86FAC7FE7A" box="[1182,1282,367,394]" metricMagnitude="-2" metricUnit="m" metricValue="2.78" pageId="2" pageNumber="113" unit="mm" value="27.8">27.8 mm</quantity>
SL, collected with holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C321B98FED6BFFBEFF0FFE5EFDBEFCCA" pageId="2" pageNumber="113" type="diagnosis">
<paragraph id="8B84EA04ED6BFFBEFF0FFE5EFDBEFCCA" blockId="2.[151,1437,259,970]" pageId="2" pageNumber="113">
<emphasis id="B94F3616ED6BFFBEFF0FFE5EFE87FE21" bold="true" box="[199,322,439,465]" pageId="2" pageNumber="113">Diagnosis.</emphasis>
<taxonomicName id="4C3B9187ED6BFFBEFE81FE5EFD92FE21" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[329,599,439,465]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED6BFFBEFE81FE5EFD92FE21" box="[329,599,439,465]" italics="true" pageId="2" pageNumber="113">Microcambeva bendego</emphasis>
</taxonomicName>
is distinguished from all congeners by the long finger-like projections, as long as orbital diameter (
<emphasis id="B94F3616ED6BFFBEFE67FE35FE08FE05" box="[431,461,476,501]" italics="true" pageId="2" pageNumber="113">vs.</emphasis>
projections absent in 
<taxonomicName id="4C3B9187ED6BFFBEFD09FE35FCACFE05" authorityName="Costa, Katz &amp; Vilardo" authorityYear="2020" box="[705,873,475,501]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="filamentosa">
<emphasis id="B94F3616ED6BFFBEFD09FE35FCACFE05" box="[705,873,475,501]" italics="true" pageId="2" pageNumber="113">M. filamentosa</emphasis>
</taxonomicName>
and smaller than orbital diameter in the remaining congeners), and the more numerous opercular odontodes (19 
<emphasis id="B94F3616ED6BFFBEFC82FDE9FCA4FDE9" box="[842,865,512,537]" italics="true" pageId="2" pageNumber="113">vs</emphasis>
. 
<quantity id="4CC347E1ED6BFFBEFCB8FE16FC72FDE9" box="[880,951,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="2" pageNumber="113" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
<taxonomicName id="4C3B9187ED6BFFBEFC77FDE9FBF8FDE9" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[959,1085,511,537]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="barbata">
<emphasis id="B94F3616ED6BFFBEFC77FDE9FBF8FDE9" box="[959,1085,511,537]" italics="true" pageId="2" pageNumber="113">M. barbata</emphasis>
</taxonomicName>
, 
<quantity id="4CC347E1ED6BFFBEFB84FE16FB64FDE9" box="[1100,1185,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="2" pageNumber="113" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">9–12 in</quantity>
<emphasis id="B94F3616ED6BFFBEFB61FDE9FA99FDE9" box="[1193,1372,512,537]" italics="true" pageId="2" pageNumber="113">
<taxonomicName id="4C3B9187ED6BFFBEFB61FDE9FA9DFDE9" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[1193,1368,512,537]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="mucuriensis">M. mucuriensis</taxonomicName>
,
</emphasis>
<quantity id="4CC347E1ED6BFFBEFAACFE16FF68FDCD" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.556" metricValueMin="2.286" pageId="2" pageNumber="113" unit="in" value="11.5" valueMax="14.0" valueMin="9.0">9–14 in</quantity>
<taxonomicName id="4C3B9187ED6BFFBEFF7EFDCDFEFDFDCD" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[182,312,547,573]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED6BFFBEFF7EFDCDFEFDFDCD" box="[182,312,547,573]" italics="true" pageId="2" pageNumber="113">M. ribeirae</emphasis>
</taxonomicName>
, 
<quantity id="4CC347E1ED6BFFBEFE80FDCAFE69FDCD" box="[328,428,547,573]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.048" metricValueMin="2.794" pageId="2" pageNumber="113" unit="in" value="11.5" valueMax="12.0" valueMin="11.0">11–12 in</quantity>
<taxonomicName id="4C3B9187ED6BFFBEFE7DFDCDFDD8FDCD" authorityName="Mattos &amp; Lima" authorityYear="2010" box="[437,541,547,573]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="draco">
<emphasis id="B94F3616ED6BFFBEFE7DFDCDFDD8FDCD" box="[437,541,547,573]" italics="true" pageId="2" pageNumber="113">M. draco</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED6BFFBEFD90FDCDFCC6FDCD" authorityName="Costa, Katz &amp; Vilardo" authorityYear="2020" box="[600,771,547,573]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="filamentosa">
<emphasis id="B94F3616ED6BFFBEFD90FDCDFCC6FDCD" box="[600,771,547,573]" italics="true" pageId="2" pageNumber="113">M. filamentosa</emphasis>
</taxonomicName>
, and 
<quantity id="4CC347E1ED6BFFBEFC8CFDCAFC6CFDCD" box="[836,937,547,574]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="2" pageNumber="113" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
<taxonomicName id="4C3B9187ED6BFFBEFC7AFDCDFB84FDCD" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[946,1089,548,573]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="jucuensis">
<emphasis id="B94F3616ED6BFFBEFC7AFDCDFB84FDCD" box="[946,1089,548,573]" italics="true" pageId="2" pageNumber="113">M. jucuensis</emphasis>
</taxonomicName>
). It is also distinguished from all congeners, except 
<taxonomicName id="4C3B9187ED6BFFBEFE43FDA1FDCFFD91" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[395,522,583,609]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED6BFFBEFE43FDA1FDCFFD91" box="[395,522,583,609]" italics="true" pageId="2" pageNumber="113">M. ribeirae</emphasis>
</taxonomicName>
, by the unmodified first pectoral-fin ray (
<emphasis id="B94F3616ED6BFFBEFC28FDA1FC32FD91" box="[992,1015,584,609]" italics="true" pageId="2" pageNumber="113">vs</emphasis>
. first pectoral-fin ray filamentous). It differs from all species of the genus, except 
<taxonomicName id="4C3B9187ED6BFFBEFD59FD85FCD4FD75" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[657,785,619,645]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED6BFFBEFD59FD85FCD4FD75" box="[657,785,619,645]" italics="true" pageId="2" pageNumber="113">M. ribeirae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED6BFFBEFC82FD85FC31FD75" authorityName="Costa, Katz &amp; Vilardo" authorityYear="2020" box="[842,1012,619,645]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="filamentosa">
<emphasis id="B94F3616ED6BFFBEFC82FD85FC31FD75" box="[842,1012,619,645]" italics="true" pageId="2" pageNumber="113">M. filamentosa</emphasis>
</taxonomicName>
, by the absence of ossification in the anterior cartilage of the autopalatine (
<emphasis id="B94F3616ED6BFFBEFD8DFD79FD99FD59" box="[581,604,656,681]" italics="true" pageId="2" pageNumber="113">vs</emphasis>
. presence), and by the supraorbital pores (s6) fused into a single median pore, positioned on the middle of head (
<emphasis id="B94F3616ED6BFFBEFD9FFD5DFDB0FD3D" box="[599,629,692,717]" italics="true" pageId="2" pageNumber="113">vs.</emphasis>
paired s6 pore). The new species differs further from 
<taxonomicName id="4C3B9187ED6BFFBEFB19FD5DFABFFD3D" authorityName="Costa, Katz &amp; Vilardo" authorityYear="2020" box="[1233,1402,691,717]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="filamentosa">
<emphasis id="B94F3616ED6BFFBEFB19FD5DFABFFD3D" box="[1233,1402,691,717]" italics="true" pageId="2" pageNumber="113">M. filamentosa</emphasis>
</taxonomicName>
by having 8 interopercular odontodes (
<emphasis id="B94F3616ED6BFFBEFDECFD31FD87FD01" box="[548,578,728,753]" italics="true" pageId="2" pageNumber="113">vs.</emphasis>
6) and 33 vertebrae (
<emphasis id="B94F3616ED6BFFBEFCE6FD31FC89FD01" box="[814,844,728,753]" italics="true" pageId="2" pageNumber="113">vs.</emphasis>
36), and from 
<taxonomicName id="4C3B9187ED6BFFBEFC38FD31FBAAFD01" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[1008,1135,727,753]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED6BFFBEFC38FD31FBAAFD01" box="[1008,1135,727,753]" italics="true" pageId="2" pageNumber="113">M. ribeirae</emphasis>
</taxonomicName>
by the rictal barbels reaching the anterior portion of the interopercular patch of odontodes (
<emphasis id="B94F3616ED6BFFBEFC48FD15FC52FCE5" box="[896,919,764,789]" italics="true" pageId="2" pageNumber="113">vs</emphasis>
. reaching middle of orbit), and the three first rays of the dorsal-fin unbranched (
<emphasis id="B94F3616ED6BFFBEFDD3FCC9FDF7FCC9" box="[539,562,800,825]" italics="true" pageId="2" pageNumber="113">vs</emphasis>
. two).
</paragraph>
</subSubSection>
<subSubSection id="C321B98FED6BFFBBFF0FFCAAFBC0FE53" lastPageId="7" lastPageNumber="118" pageId="2" pageNumber="113" type="description">
<paragraph id="8B84EA04ED6BFFBEFF0FFCAAFB0DFC39" blockId="2.[151,1437,259,970]" pageId="2" pageNumber="113">
<emphasis id="B94F3616ED6BFFBEFF0FFCAAFE9CFCAD" bold="true" box="[199,345,835,861]" pageId="2" pageNumber="113">Description.</emphasis>
Morphometric data of 
<typeStatus id="548054A6ED6BFFBEFDAAFCAAFD01FCAD" box="[610,708,835,861]" pageId="2" pageNumber="113" type="holotype">holotype</typeStatus>
and two 
<typeStatus id="548054A6ED6BFFBEFCF8FCADFC5EFCAE" box="[816,923,836,862]" pageId="2" pageNumber="113" type="paratype">paratypes</typeStatus>
presented in 
<tableCitation id="C6B9DFBFED6BFFBEFBFEFCAAFB4EFCAD" box="[1078,1163,835,861]" captionStart="TABLE 1" captionStartId="3.[151,239,151,175]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data of holotype (MNRJ 52042, 28.1 mm SL) and two paratypes (MNRJ 48616, 26.94 mm SL and MZUSP 125789, 27.8 mm SL) of Microcambeva bendego." httpUri="http://table.plazi.org/id/DF44BA8CED6AFFBFFF5FFF7EFCFDFF23" pageId="2" pageNumber="113" tableUuid="DF44BA8CED6AFFBFFF5FFF7EFCFDFF23">Table 1</tableCitation>
. Body elongated, cylindrical immediately posterior to head to pelvic-fin origin, gradually compressed towards caudal peduncle (
<figureCitation id="1300F681ED6BFFBEFAF5FC8EFA48FC71" box="[1341,1421,871,897]" captionStart="FIGURE 1" captionStartId="2.[151,250,1945,1969]" captionTargetBox="[151,1435,993,1921]" captionTargetId="figure-362@2.[151,1435,993,1921]" captionTargetPageId="2" captionText="FIGURE 1. Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Rio Guapiaçu, near Cachoeiras de Macacu, rio Guapi-Macacu basin, Guapimirim Municipality, Rio de Janeiro State, southeastern Brazil. a. lateral view; b. dorsal view; c. ventral view. Scale: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4321550" httpUri="https://zenodo.org/record/4321550/files/figure.png" pageId="2" pageNumber="113">Fig. 1a</figureCitation>
). Dorsal profile gently convex from snout to dorsal-fin origin, and straight along caudal peduncle. Ventral profile convex from jaw to pelvic-fin insertion, straight from that point to terminus of caudal peduncle.
</paragraph>
<caption id="DF44BA8CED6BFFBEFF5FF870FDCCF808" ID-DOI="http://doi.org/10.5281/zenodo.4321550" ID-Zenodo-Dep="4321550" httpUri="https://zenodo.org/record/4321550/files/figure.png" pageId="2" pageNumber="113" startId="2.[151,250,1945,1969]" targetBox="[151,1435,993,1921]" targetPageId="2">
<paragraph id="8B84EA04ED6BFFBEFF5FF870FDCCF808" blockId="2.[151,1436,1945,2041]" pageId="2" pageNumber="113">
<emphasis id="B94F3616ED6BFFBEFF5FF870FED0F841" bold="true" box="[151,277,1945,1969]" pageId="2" pageNumber="113">FIGURE 1.</emphasis>
<taxonomicName id="4C3B9187ED6BFFBEFED5F870FDD7F841" authority="Medeiros &amp; Moreira &amp; Pinna &amp; Lima, 2020" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[285,530,1945,1969]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="113" phylum="Chordata" rank="species" species="bendego" status="sp. nov.">
<emphasis id="B94F3616ED6BFFBEFED5F870FDD7F841" box="[285,530,1945,1969]" italics="true" pageId="2" pageNumber="113">Microcambeva bendego</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A27C8B6DED6BFFBEFDE9F870FD5AF841" box="[545,671,1945,1969]" pageId="2" pageNumber="113" rank="species">new species</taxonomicNameLabel>
, holotype, MNRJ 52042, 28.1 mm SL. Rio Guapiaçu, near Cachoeiras de Macacu, rio Guapi-Macacu basin, Guapimirim Municipality, Rio de Janeiro State, southeastern Brazil. 
<emphasis id="B94F3616ED6BFFBEFB60F854FB70F825" bold="true" box="[1192,1205,1981,2005]" pageId="2" pageNumber="113">a</emphasis>
. lateral view; 
<emphasis id="B94F3616ED6BFFBEFA8CF854FA97F825" bold="true" box="[1348,1362,1981,2005]" pageId="2" pageNumber="113">b</emphasis>
. dorsal view; 
<emphasis id="B94F3616ED6BFFBEFF1EF808FF24F809" bold="true" box="[214,225,2017,2041]" pageId="2" pageNumber="113">c</emphasis>
. ventral view. Scale: 10 mm.
</paragraph>
</caption>
<caption id="DF44BA8CED6AFFBFFF5FFF7EFCFDFF23" ID-Table-UUID="DF44BA8CED6AFFBFFF5FFF7EFCFDFF23" httpUri="http://table.plazi.org/id/DF44BA8CED6AFFBFFF5FFF7EFCFDFF23" pageId="3" pageNumber="114" targetBox="[166,1407,227,1213]" targetIsTable="true" targetPageId="3">
<paragraph id="8B84EA04ED6AFFBFFF5FFF7EFCFDFF23" blockId="3.[151,1436,151,212]" pageId="3" pageNumber="114">
<emphasis id="B94F3616ED6AFFBFFF5FFF7EFECFFF5F" bold="true" box="[151,266,151,175]" pageId="3" pageNumber="114">TABLE 1.</emphasis>
Morphometric data of holotype (MNRJ 52042, 28.1 mm SL) and two paratypes (MNRJ 48616, 26.94 mm SL and MZUSP 125789, 27.8 mm SL) of 
<taxonomicName id="4C3B9187ED6AFFBFFDF9FF52FCF4FF23" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[561,817,187,211]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="114" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED6AFFBFFDF9FF52FCF4FF23" box="[561,817,187,211]" italics="true" pageId="3" pageNumber="114">Microcambeva bendego</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B84EA04ED6AFFBFFDAAFF0DFAB5FB4D" pageId="3" pageNumber="114">
<table id="F93B18A4ED6A0043FF6EFF0AFABAFB4D" box="[166,1407,227,1213]" gridcols="5" gridrows="24" pageId="3" pageNumber="114">
<tr id="350BE846ED6A0043FF6EFF0AFABAFED0" box="[166,1407,227,288]" gridrow="0" pageId="3" pageNumber="114" rowspan-0="1">
<th id="76DA813AED6A0043FDAAFF0AFCCAFED0" box="[610,783,227,288]" gridcol="1" gridrow="0" pageId="3" pageNumber="114">
<emphasis id="B94F3616ED6AFFBFFDAAFF0DFD00FF0C" bold="true" box="[610,709,228,252]" pageId="3" pageNumber="114">Holotype</emphasis>
MNRJ 52042
</th>
<th id="76DA813AED6A0043FC87FF0AFC39FED0" box="[847,1020,227,288]" gridcol="2" gridrow="0" pageId="3" pageNumber="114">
<emphasis id="B94F3616ED6AFFBFFC87FF0DFC74FF0C" bold="true" box="[847,945,228,252]" pageId="3" pageNumber="114">Paratype</emphasis>
MNRJ 48616
</th>
<th id="76DA813AED6A0043FBF7FF0AFB39FED0" box="[1087,1276,227,288]" gridcol="3" gridrow="0" pageId="3" pageNumber="114">
<emphasis id="B94F3616ED6AFFBFFBF7FF0DFB64FF0C" bold="true" box="[1087,1185,228,252]" pageId="3" pageNumber="114">Paratype</emphasis>
MZUSP 125789
</th>
<th id="76DA813AED6A0043FA89FF0AFABAFED0" box="[1345,1407,227,288]" gridcol="4" gridrow="0" pageId="3" pageNumber="114">
<emphasis id="B94F3616ED6AFFBFFA89FF0AFABAFF0B" bold="true" box="[1345,1407,227,251]" pageId="3" pageNumber="114">Mean</emphasis>
</th>
</tr>
<tr id="350BE846ED6A0043FF6EFEC6FABAFEB8" box="[166,1407,303,328]" gridrow="1" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFEC6FE3AFEB8" box="[166,511,303,328]" gridcol="0" gridrow="1" pageId="3" pageNumber="114">
<emphasis id="B94F3616ED6AFFBFFF6EFEC6FE48FEB7" bold="true" box="[166,397,303,327]" pageId="3" pageNumber="114">Standard length—SL</emphasis>
</th>
<td id="76DA813AED6A0043FDAAFEC6FCCAFEB8" box="[610,783,303,328]" gridcol="1" gridrow="1" pageId="3" pageNumber="114">28.1</td>
<td id="76DA813AED6A0043FC87FEC6FC39FEB8" box="[847,1020,303,328]" gridcol="2" gridrow="1" pageId="3" pageNumber="114">26.9</td>
<td id="76DA813AED6A0043FBF7FEC6FB39FEB8" box="[1087,1276,303,328]" gridcol="3" gridrow="1" pageId="3" pageNumber="114">27.8</td>
<td id="76DA813AED6A0043FA89FEC6FABAFEB8" box="[1345,1407,303,328]" gridcol="4" gridrow="1" pageId="3" pageNumber="114">27.5</td>
</tr>
<tr id="350BE846ED6A0043FF6EFEB1FABAFE81" box="[166,1407,344,369]" gridrow="2" pageId="3" pageNumber="114" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76DA813AED6A0043FF6EFEB1FE3AFE81" box="[166,511,344,369]" gridcol="0" gridrow="2" pageId="3" pageNumber="114">Percentage of SL</th>
</tr>
<tr id="350BE846ED6A0043FF6EFE69FABAFE69" box="[166,1407,384,409]" gridrow="3" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFE69FE3AFE69" box="[166,511,384,409]" gridcol="0" gridrow="3" pageId="3" pageNumber="114">Body depth</th>
<td id="76DA813AED6A0043FDAAFE69FCCAFE69" box="[610,783,384,409]" gridcol="1" gridrow="3" pageId="3" pageNumber="114">3.7</td>
<td id="76DA813AED6A0043FC87FE69FC39FE69" box="[847,1020,384,409]" gridcol="2" gridrow="3" pageId="3" pageNumber="114">3.2</td>
<td id="76DA813AED6A0043FBF7FE69FB39FE69" box="[1087,1276,384,409]" gridcol="3" gridrow="3" pageId="3" pageNumber="114">3.5</td>
<td id="76DA813AED6A0043FA89FE69FABAFE69" box="[1345,1407,384,409]" gridcol="4" gridrow="3" pageId="3" pageNumber="114">3.5</td>
</tr>
<tr id="350BE846ED6A0043FF6EFE40FABAFE31" box="[166,1407,425,449]" gridrow="4" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFE40FE3AFE31" box="[166,511,425,449]" gridcol="0" gridrow="4" pageId="3" pageNumber="114">Caudal peduncle depth</th>
<td id="76DA813AED6A0043FDAAFE40FCCAFE31" box="[610,783,425,449]" gridcol="1" gridrow="4" pageId="3" pageNumber="114">2.6</td>
<td id="76DA813AED6A0043FC87FE40FC39FE31" box="[847,1020,425,449]" gridcol="2" gridrow="4" pageId="3" pageNumber="114">2.1</td>
<td id="76DA813AED6A0043FBF7FE40FB39FE31" box="[1087,1276,425,449]" gridcol="3" gridrow="4" pageId="3" pageNumber="114">2.1</td>
<td id="76DA813AED6A0043FA89FE40FABAFE31" box="[1345,1407,425,449]" gridcol="4" gridrow="4" pageId="3" pageNumber="114">2.3</td>
</tr>
<tr id="350BE846ED6A0043FF6EFE38FABAFE19" box="[166,1407,465,489]" gridrow="5" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFE38FE3AFE19" box="[166,511,465,489]" gridcol="0" gridrow="5" pageId="3" pageNumber="114">Body width</th>
<td id="76DA813AED6A0043FDAAFE38FCCAFE19" box="[610,783,465,489]" gridcol="1" gridrow="5" pageId="3" pageNumber="114">2.5</td>
<td id="76DA813AED6A0043FC87FE38FC39FE19" box="[847,1020,465,489]" gridcol="2" gridrow="5" pageId="3" pageNumber="114">1.0</td>
<td id="76DA813AED6A0043FBF7FE38FB39FE19" box="[1087,1276,465,489]" gridcol="3" gridrow="5" pageId="3" pageNumber="114">1.0</td>
<td id="76DA813AED6A0043FA89FE38FABAFE19" box="[1345,1407,465,489]" gridcol="4" gridrow="5" pageId="3" pageNumber="114">1.5</td>
</tr>
<tr id="350BE846ED6A0043FF6EFE10FABAFDE1" box="[166,1407,505,529]" gridrow="6" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFE10FE3AFDE1" box="[166,511,505,529]" gridcol="0" gridrow="6" pageId="3" pageNumber="114">Caudal penduncle width</th>
<td id="76DA813AED6A0043FDAAFE10FCCAFDE1" box="[610,783,505,529]" gridcol="1" gridrow="6" pageId="3" pageNumber="114">2.2</td>
<td id="76DA813AED6A0043FC87FE10FC39FDE1" box="[847,1020,505,529]" gridcol="2" gridrow="6" pageId="3" pageNumber="114">0.6</td>
<td id="76DA813AED6A0043FBF7FE10FB39FDE1" box="[1087,1276,505,529]" gridcol="3" gridrow="6" pageId="3" pageNumber="114">0.6</td>
<td id="76DA813AED6A0043FA89FE10FABAFDE1" box="[1345,1407,505,529]" gridcol="4" gridrow="6" pageId="3" pageNumber="114">1.1</td>
</tr>
<tr id="350BE846ED6A0043FF6EFDC8FABAFDCA" box="[166,1407,545,570]" gridrow="7" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFDC8FE3AFDCA" box="[166,511,545,570]" gridcol="0" gridrow="7" pageId="3" pageNumber="114">Caudal penduncle length</th>
<td id="76DA813AED6A0043FDAAFDC8FCCAFDCA" box="[610,783,545,570]" gridcol="1" gridrow="7" pageId="3" pageNumber="114">8.8</td>
<td id="76DA813AED6A0043FC87FDC8FC39FDCA" box="[847,1020,545,570]" gridcol="2" gridrow="7" pageId="3" pageNumber="114">5.0</td>
<td id="76DA813AED6A0043FBF7FDC8FB39FDCA" box="[1087,1276,545,570]" gridcol="3" gridrow="7" pageId="3" pageNumber="114">5.7</td>
<td id="76DA813AED6A0043FA89FDC8FABAFDCA" box="[1345,1407,545,570]" gridcol="4" gridrow="7" pageId="3" pageNumber="114">6.5</td>
</tr>
<tr id="350BE846ED6A0043FF6EFDA3FABAFD92" box="[166,1407,586,610]" gridrow="8" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFDA3FE3AFD92" box="[166,511,586,610]" gridcol="0" gridrow="8" pageId="3" pageNumber="114">Dorsal-fin base length</th>
<td id="76DA813AED6A0043FDAAFDA3FCCAFD92" box="[610,783,586,610]" gridcol="1" gridrow="8" pageId="3" pageNumber="114">3.3</td>
<td id="76DA813AED6A0043FC87FDA3FC39FD92" box="[847,1020,586,610]" gridcol="2" gridrow="8" pageId="3" pageNumber="114">2.8</td>
<td id="76DA813AED6A0043FBF7FDA3FB39FD92" box="[1087,1276,586,610]" gridcol="3" gridrow="8" pageId="3" pageNumber="114">2.6</td>
<td id="76DA813AED6A0043FA89FDA3FABAFD92" box="[1345,1407,586,610]" gridcol="4" gridrow="8" pageId="3" pageNumber="114">2.9</td>
</tr>
<tr id="350BE846ED6A0043FF6EFD9BFABAFD7A" box="[166,1407,626,650]" gridrow="9" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFD9BFE3AFD7A" box="[166,511,626,650]" gridcol="0" gridrow="9" pageId="3" pageNumber="114">Anal-fin base length</th>
<td id="76DA813AED6A0043FDAAFD9BFCCAFD7A" box="[610,783,626,650]" gridcol="1" gridrow="9" pageId="3" pageNumber="114">2.1</td>
<td id="76DA813AED6A0043FC87FD9BFC39FD7A" box="[847,1020,626,650]" gridcol="2" gridrow="9" pageId="3" pageNumber="114">2.3</td>
<td id="76DA813AED6A0043FBF7FD9BFB39FD7A" box="[1087,1276,626,650]" gridcol="3" gridrow="9" pageId="3" pageNumber="114">2.5</td>
<td id="76DA813AED6A0043FA89FD9BFABAFD7A" box="[1345,1407,626,650]" gridcol="4" gridrow="9" pageId="3" pageNumber="114">2.3</td>
</tr>
<tr id="350BE846ED6A0043FF6EFD73FABAFD42" box="[166,1407,666,690]" gridrow="10" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFD73FE3AFD42" box="[166,511,666,690]" gridcol="0" gridrow="10" pageId="3" pageNumber="114">Pelvic-fin length</th>
<td id="76DA813AED6A0043FDAAFD73FCCAFD42" box="[610,783,666,690]" gridcol="1" gridrow="10" pageId="3" pageNumber="114">4.7</td>
<td id="76DA813AED6A0043FC87FD73FC39FD42" box="[847,1020,666,690]" gridcol="2" gridrow="10" pageId="3" pageNumber="114">4.1</td>
<td id="76DA813AED6A0043FBF7FD73FB39FD42" box="[1087,1276,666,690]" gridcol="3" gridrow="10" pageId="3" pageNumber="114">4.3</td>
<td id="76DA813AED6A0043FA89FD73FABAFD42" box="[1345,1407,666,690]" gridcol="4" gridrow="10" pageId="3" pageNumber="114">4.4</td>
</tr>
<tr id="350BE846ED6A0043FF6EFD2BFABAFD2B" box="[166,1407,706,731]" gridrow="11" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFD2BFE3AFD2B" box="[166,511,706,731]" gridcol="0" gridrow="11" pageId="3" pageNumber="114">Distance between pelvic-fin bases</th>
<td id="76DA813AED6A0043FDAAFD2BFCCAFD2B" box="[610,783,706,731]" gridcol="1" gridrow="11" pageId="3" pageNumber="114">1.1</td>
<td id="76DA813AED6A0043FC87FD2BFC39FD2B" box="[847,1020,706,731]" gridcol="2" gridrow="11" pageId="3" pageNumber="114">0.6</td>
<td id="76DA813AED6A0043FBF7FD2BFB39FD2B" box="[1087,1276,706,731]" gridcol="3" gridrow="11" pageId="3" pageNumber="114">1.0</td>
<td id="76DA813AED6A0043FA89FD2BFABAFD2B" box="[1345,1407,706,731]" gridcol="4" gridrow="11" pageId="3" pageNumber="114">0.9</td>
</tr>
<tr id="350BE846ED6A0043FF6EFD03FABAFCF3" box="[166,1407,746,771]" gridrow="12" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFD03FE3AFCF3" box="[166,511,746,771]" gridcol="0" gridrow="12" pageId="3" pageNumber="114">Pectoral-fin length</th>
<td id="76DA813AED6A0043FDAAFD03FCCAFCF3" box="[610,783,746,771]" gridcol="1" gridrow="12" pageId="3" pageNumber="114">5.6</td>
<td id="76DA813AED6A0043FC87FD03FC39FCF3" box="[847,1020,746,771]" gridcol="2" gridrow="12" pageId="3" pageNumber="114">3.8</td>
<td id="76DA813AED6A0043FBF7FD03FB39FCF3" box="[1087,1276,746,771]" gridcol="3" gridrow="12" pageId="3" pageNumber="114">4.3</td>
<td id="76DA813AED6A0043FA89FD03FABAFCF3" box="[1345,1407,746,771]" gridcol="4" gridrow="12" pageId="3" pageNumber="114">4.5</td>
</tr>
<tr id="350BE846ED6A0043FF6EFCFAFABAFCDB" box="[166,1407,787,811]" gridrow="13" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFCFAFE3AFCDB" box="[166,511,787,811]" gridcol="0" gridrow="13" pageId="3" pageNumber="114">Predorsal length</th>
<td id="76DA813AED6A0043FDAAFCFAFCCAFCDB" box="[610,783,787,811]" gridcol="1" gridrow="13" pageId="3" pageNumber="114">17.0</td>
<td id="76DA813AED6A0043FC87FCFAFC39FCDB" box="[847,1020,787,811]" gridcol="2" gridrow="13" pageId="3" pageNumber="114">13.9</td>
<td id="76DA813AED6A0043FBF7FCFAFB39FCDB" box="[1087,1276,787,811]" gridcol="3" gridrow="13" pageId="3" pageNumber="114">14.8</td>
<td id="76DA813AED6A0043FA89FCFAFABAFCDB" box="[1345,1407,787,811]" gridcol="4" gridrow="13" pageId="3" pageNumber="114">15.2</td>
</tr>
<tr id="350BE846ED6A0043FF6EFCD2FABAFCA3" box="[166,1407,827,851]" gridrow="14" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFCD2FE3AFCA3" box="[166,511,827,851]" gridcol="0" gridrow="14" pageId="3" pageNumber="114">Prepelvic length</th>
<td id="76DA813AED6A0043FDAAFCD2FCCAFCA3" box="[610,783,827,851]" gridcol="1" gridrow="14" pageId="3" pageNumber="114">14.9</td>
<td id="76DA813AED6A0043FC87FCD2FC39FCA3" box="[847,1020,827,851]" gridcol="2" gridrow="14" pageId="3" pageNumber="114">12.5</td>
<td id="76DA813AED6A0043FBF7FCD2FB39FCA3" box="[1087,1276,827,851]" gridcol="3" gridrow="14" pageId="3" pageNumber="114">12.3</td>
<td id="76DA813AED6A0043FA89FCD2FABAFCA3" box="[1345,1407,827,851]" gridcol="4" gridrow="14" pageId="3" pageNumber="114">13.2</td>
</tr>
<tr id="350BE846ED6A0043FF6EFC8AFABAFC8B" box="[166,1407,867,891]" gridrow="15" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFC8AFE3AFC8B" box="[166,511,867,891]" gridcol="0" gridrow="15" pageId="3" pageNumber="114">
<emphasis id="B94F3616ED6AFFBFFF6EFC8AFEA0FC8B" bold="true" box="[166,357,867,891]" pageId="3" pageNumber="114">Head length (HL)</emphasis>
</th>
<td id="76DA813AED6A0043FDAAFC8AFCCAFC8B" box="[610,783,867,891]" gridcol="1" gridrow="15" pageId="3" pageNumber="114">6.0</td>
<td id="76DA813AED6A0043FC87FC8AFC39FC8B" box="[847,1020,867,891]" gridcol="2" gridrow="15" pageId="3" pageNumber="114">5.1</td>
<td id="76DA813AED6A0043FBF7FC8AFB39FC8B" box="[1087,1276,867,891]" gridcol="3" gridrow="15" pageId="3" pageNumber="114">5.8</td>
<td id="76DA813AED6A0043FA89FC8AFABAFC8B" box="[1345,1407,867,891]" gridcol="4" gridrow="15" pageId="3" pageNumber="114">5.7</td>
</tr>
<tr id="350BE846ED6A0043FF6EFC65FABAFC54" box="[166,1407,908,932]" gridrow="16" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFC65FABAFC54" box="[166,1407,908,932]" colspan="5" colspanRight="4" gridcol="0" gridrow="16" pageId="3" pageNumber="114">Percentage of HL</th>
</tr>
<tr id="350BE846ED6A0043FF6EFC5DFABAFC3C" box="[166,1407,948,972]" gridrow="17" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFC5DFE3AFC3C" box="[166,511,948,972]" gridcol="0" gridrow="17" pageId="3" pageNumber="114">Head depth</th>
<td id="76DA813AED6A0043FDAAFC5DFCCAFC3C" box="[610,783,948,972]" gridcol="1" gridrow="17" pageId="3" pageNumber="114">3.0</td>
<td id="76DA813AED6A0043FC87FC5DFC39FC3C" box="[847,1020,948,972]" gridcol="2" gridrow="17" pageId="3" pageNumber="114">2.3</td>
<td id="76DA813AED6A0043FBF7FC5DFB39FC3C" box="[1087,1276,948,972]" gridcol="3" gridrow="17" pageId="3" pageNumber="114">2.4</td>
<td id="76DA813AED6A0043FA89FC5DFABAFC3C" box="[1345,1407,948,972]" gridcol="4" gridrow="17" pageId="3" pageNumber="114">2.6</td>
</tr>
<tr id="350BE846ED6A0043FF6EFC35FABAFC04" box="[166,1407,988,1012]" gridrow="18" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFC35FE3AFC04" box="[166,511,988,1012]" gridcol="0" gridrow="18" pageId="3" pageNumber="114">Head width</th>
<td id="76DA813AED6A0043FDAAFC35FCCAFC04" box="[610,783,988,1012]" gridcol="1" gridrow="18" pageId="3" pageNumber="114">4.9</td>
<td id="76DA813AED6A0043FC87FC35FC39FC04" box="[847,1020,988,1012]" gridcol="2" gridrow="18" pageId="3" pageNumber="114">4.1</td>
<td id="76DA813AED6A0043FBF7FC35FB39FC04" box="[1087,1276,988,1012]" gridcol="3" gridrow="18" pageId="3" pageNumber="114">4.3</td>
<td id="76DA813AED6A0043FA89FC35FABAFC04" box="[1345,1407,988,1012]" gridcol="4" gridrow="18" pageId="3" pageNumber="114">4.4</td>
</tr>
<tr id="350BE846ED6A0043FF6EFBEDFABAFBEC" box="[166,1407,1028,1052]" gridrow="19" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFBEDFE3AFBEC" box="[166,511,1028,1052]" gridcol="0" gridrow="19" pageId="3" pageNumber="114">Interorbital width</th>
<td id="76DA813AED6A0043FDAAFBEDFCCAFBEC" box="[610,783,1028,1052]" gridcol="1" gridrow="19" pageId="3" pageNumber="114">0.8</td>
<td id="76DA813AED6A0043FC87FBEDFC39FBEC" box="[847,1020,1028,1052]" gridcol="2" gridrow="19" pageId="3" pageNumber="114">0.9</td>
<td id="76DA813AED6A0043FBF7FBEDFB39FBEC" box="[1087,1276,1028,1052]" gridcol="3" gridrow="19" pageId="3" pageNumber="114">1.2</td>
<td id="76DA813AED6A0043FA89FBEDFABAFBEC" box="[1345,1407,1028,1052]" gridcol="4" gridrow="19" pageId="3" pageNumber="114">1.0</td>
</tr>
<tr id="350BE846ED6A0043FF6EFBC4FABAFBB5" box="[166,1407,1069,1093]" gridrow="20" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFBC4FE3AFBB5" box="[166,511,1069,1093]" gridcol="0" gridrow="20" pageId="3" pageNumber="114">Preorbital length</th>
<td id="76DA813AED6A0043FDAAFBC4FCCAFBB5" box="[610,783,1069,1093]" gridcol="1" gridrow="20" pageId="3" pageNumber="114">2.7</td>
<td id="76DA813AED6A0043FC87FBC4FC39FBB5" box="[847,1020,1069,1093]" gridcol="2" gridrow="20" pageId="3" pageNumber="114">2.4</td>
<td id="76DA813AED6A0043FBF7FBC4FB39FBB5" box="[1087,1276,1069,1093]" gridcol="3" gridrow="20" pageId="3" pageNumber="114">2.6</td>
<td id="76DA813AED6A0043FA89FBC4FABAFBB5" box="[1345,1407,1069,1093]" gridcol="4" gridrow="20" pageId="3" pageNumber="114">2.6</td>
</tr>
<tr id="350BE846ED6A0043FF6EFBBCFABAFB9D" box="[166,1407,1109,1133]" gridrow="21" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFBBCFE3AFB9D" box="[166,511,1109,1133]" gridcol="0" gridrow="21" pageId="3" pageNumber="114">Eye diameter</th>
<td id="76DA813AED6A0043FDAAFBBCFCCAFB9D" box="[610,783,1109,1133]" gridcol="1" gridrow="21" pageId="3" pageNumber="114">1.2</td>
<td id="76DA813AED6A0043FC87FBBCFC39FB9D" box="[847,1020,1109,1133]" gridcol="2" gridrow="21" pageId="3" pageNumber="114">1.1</td>
<td id="76DA813AED6A0043FBF7FBBCFB39FB9D" box="[1087,1276,1109,1133]" gridcol="3" gridrow="21" pageId="3" pageNumber="114">0.7</td>
<td id="76DA813AED6A0043FA89FBBCFABAFB9D" box="[1345,1407,1109,1133]" gridcol="4" gridrow="21" pageId="3" pageNumber="114">1.0</td>
</tr>
<tr id="350BE846ED6A0043FF6EFB94FABAFB65" box="[166,1407,1149,1173]" gridrow="22" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFB94FE3AFB65" box="[166,511,1149,1173]" gridcol="0" gridrow="22" pageId="3" pageNumber="114">Mouth width</th>
<td id="76DA813AED6A0043FDAAFB94FCCAFB65" box="[610,783,1149,1173]" gridcol="1" gridrow="22" pageId="3" pageNumber="114">1.3</td>
<td id="76DA813AED6A0043FC87FB94FC39FB65" box="[847,1020,1149,1173]" gridcol="2" gridrow="22" pageId="3" pageNumber="114">1.5</td>
<td id="76DA813AED6A0043FBF7FB94FB39FB65" box="[1087,1276,1149,1173]" gridcol="3" gridrow="22" pageId="3" pageNumber="114">1.2</td>
<td id="76DA813AED6A0043FA89FB94FABAFB65" box="[1345,1407,1149,1173]" gridcol="4" gridrow="22" pageId="3" pageNumber="114">1.3</td>
</tr>
<tr id="350BE846ED6A0043FF6EFB4CFABAFB4D" box="[166,1407,1189,1213]" gridrow="23" pageId="3" pageNumber="114">
<th id="76DA813AED6A0043FF6EFB4CFE3AFB4D" box="[166,511,1189,1213]" gridcol="0" gridrow="23" pageId="3" pageNumber="114">Internarial width</th>
<td id="76DA813AED6A0043FDAAFB4CFCCAFB4D" box="[610,783,1189,1213]" gridcol="1" gridrow="23" pageId="3" pageNumber="114">0.1</td>
<td id="76DA813AED6A0043FC87FB4CFC39FB4D" box="[847,1020,1189,1213]" gridcol="2" gridrow="23" pageId="3" pageNumber="114">0.4</td>
<td id="76DA813AED6A0043FBF7FB4CFB39FB4D" box="[1087,1276,1189,1213]" gridcol="3" gridrow="23" pageId="3" pageNumber="114">0.4</td>
<td id="76DA813AED6A0043FA89FB4CFABAFB4D" box="[1345,1407,1189,1213]" gridcol="4" gridrow="23" pageId="3" pageNumber="114">0.3</td>
</tr>
</table>
</paragraph>
<paragraph id="8B84EA04ED6AFFBFFF0FFB04FEB1F9BB" blockId="3.[151,1437,1261,2043]" pageId="3" pageNumber="114">
Head triangular in dorsal view, moderately depressed, longer than wide (
<figureCitation id="1300F681ED6AFFBFFBC0FB04FB9EFAF8" box="[1032,1115,1261,1288]" captionStart="FIGURE 1" captionStartId="2.[151,250,1945,1969]" captionTargetBox="[151,1435,993,1921]" captionTargetId="figure-362@2.[151,1435,993,1921]" captionTargetPageId="2" captionText="FIGURE 1. Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Rio Guapiaçu, near Cachoeiras de Macacu, rio Guapi-Macacu basin, Guapimirim Municipality, Rio de Janeiro State, southeastern Brazil. a. lateral view; b. dorsal view; c. ventral view. Scale: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4321550" httpUri="https://zenodo.org/record/4321550/files/figure.png" pageId="3" pageNumber="114">Fig. 1b</figureCitation>
). Mouth subterminal. Snout rectangular, with slight lateral compression after origin of maxillary barbel. Nostrils circular, posterior ones larger than anterior ones. Anterior nostril surrounded by skin continuous with nasal barbel, posterior nostril with small half-moon skin fold, approximately as large as nasal opening. Posterior nostril closer to anterior nostril than to margin of eye, anterior nostril closer to upper lip than posterior nostril. Barbels tapering distally. Nasal barbel originating on median portion of anterior nostril, reaching posterior portion of posterior nostril. Maxillary barbel reaching posterior half of interopercular patch of odontodes. Rictal barbel reaching anterior half of interopercular patch of odontodes. Pair of finger-like projections approximately as long as eye diameter, inserted anterior to branchial isthmus (
<figureCitation id="1300F681ED6AFFBFFF1EF9E4FED9F9D7" box="[214,284,1549,1575]" captionStart="FIGURE 2" captionStartId="4.[151,250,1592,1616]" captionTargetBox="[151,1435,417,1568]" captionTargetId="figure-124@4.[151,1435,417,1568]" captionTargetPageId="4" captionText="FIGURE 2. Ventral view of holotype of Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL, white arrow indicates the finger-like projections. Scale bar: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4321552" httpUri="https://zenodo.org/record/4321552/files/figure.png" pageId="3" pageNumber="114">Fig. 2</figureCitation>
). Eyes round, positioned dorsally at middle distance between snout and posterior portion of opercular patch of odontodes.
</paragraph>
<paragraph id="8B84EA04ED6AFFBFFF0FF9BCFAE0F89B" blockId="3.[151,1437,1261,2043]" pageId="3" pageNumber="114">
Pectoral fin subtriangular, with seven rays (i+6). First unbranched ray not filamentous, approximately 60% as long as first branched ray, last branched ray shortest. Tips of rays extending beyond interradial membrane. Adipose organ round, translucent, dorsal to pectoral fin. Dorsal fin semicircular, nine rays (iii+6), origin at vertical through 14 
<superScript id="7C4E474CED6AFFBFFF7BF928FF05F93F" attach="left" box="[179,192,1729,1743]" fontSize="6" pageId="3" pageNumber="114">th</superScript>
vertebrae. Pelvic fins subtriangular, with five rays (i+4) extending beyond interradial membrane; pelvic-fin origin at vertical through 10 
<superScript id="7C4E474CED6AFFBFFE7BF90CFE05F903" attach="left" box="[435,448,1765,1779]" fontSize="6" pageId="3" pageNumber="114">th</superScript>
vertebrae, its tip not reaching urogenital papilla. Urogenital papilla conic. Anal fin with seven rays (iii+4) and semicircular distal margin, originating at vertical through 19 
<superScript id="7C4E474CED6AFFBFFBE3F8E0FBFDF8E7" attach="left" box="[1067,1080,1801,1815]" fontSize="6" pageId="3" pageNumber="114">th</superScript>
vertebrae. Caudal fin truncated, with 13 rays (i+11+i), six in dorsal plate (i+5) and seven in ventral plate (6+i). Six procurrent caudal-fin rays dorsally and ventrally. Vertebrae 33, 27 caudal and six precaudal. Five pleural ribs. Six branchiostegal rays.
</paragraph>
<paragraph id="8B84EA04ED6AFFB8FF0FF89CFAFFFE79" blockId="3.[151,1437,1261,2043]" lastBlockId="4.[151,1437,150,393]" lastPageId="4" lastPageNumber="115" pageId="3" pageNumber="114">
Mesethmoid long, cornua short, straight and pointed; slight depression at anterior portion of the mesethmoid; main body of bone compressed at posterior portion, and pointed. Lateral margin straight along mid-length (
<figureCitation id="1300F681ED6AFFBFFA99F870FA55F844" box="[1361,1424,1945,1972]" captionStart="FIGURE 3" captionStartId="5.[151,250,1708,1732]" captionTargetBox="[177,1410,813,1685]" captionTargetId="figure-281@5.[177,1410,813,1685]" captionTargetPageId="5" captionText="FIGURE 3. Skull, Hyoid Arch, Jaws, Opercular Apparatus, pectoral fin and girdle, Suspensorium, anterior vertebrae and Weberian complex of Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Dorsal view (A) and Ventral view (B). Abbreviations: AAR, Anguloarticular; ACH: Anterior ceratohyal; BAR: Barbular; BAS+EXO: Basioccipital-exoccipital bone; BRR: Branchiostegal rays; CLE: Cleithrum; DEN: Dentary; EPO: Epioccipital; HYO: Hyomandibula; IOP: Interopercle; LAN: Lacrimal-antorbital; LAT: Lateral ethmoid; MAX: Maxilla; MET: Mesethmoid; MPT: Metapterygoid; OPE: Opercle; ORB: Orbitosphenoid; PAT: Parietal; PAL: Autoplatine; PCH: Posterior ceratohyal; PMX: Premaxilla; POP, Preopercle; PSC: Posttemporo-supracleithrum; PSO: Parieto-supraoccipital; PTE: Pterotic; PUH: Parurohyal; QUA: Quadrate; SCO, Scapulocoracoid; SPH+POT+PSF: Sphenotic + Prootic + Pterosphenoid complex; VHH: Ventral hypohyal; VOM: Vomer; WEB: Capsule of Weberian apparatus; psp S6: Posterior supraorbital pore S6. Scale bar: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4321554" httpUri="https://zenodo.org/record/4321554/files/figure.png" pageId="3" pageNumber="114">Fig.3</figureCitation>
). Frontals slender; progressively more spaced from each other posteriorly. Two pores of supraorbital canal present, s3 pore opening at anterior portion of frontal, and median s6 pore at center of neurocranium, near anterior edge of cranial fontanel. Cranial fontanel extending for approximately 75% of neurocranium length, with rounded anterior and posterior ends. Sphenotic+ pterosphenoid+ prootic trapezoid, prominent, with elongated and pointed lateral process at middle portion, bearing part of laterosensory canal, opening as single pore i11. Pterotic square-like, without lateral process, bearing preopercle pore i11. Epioccipital rectangular. Posttemporo-supracleithrum rectangular, with small lateral process near its proximal portion. Vomer long, arrow-shaped, with lateral constriction at anterior portion and forked anteriorly; thin lateral projections anterior to constriction. Lateral ethmoid rectangular, without lateral projections. Basioccipital fused with exoccipital anteriorly and with Weberian complex posteriorly.
</paragraph>
<caption id="DF44BA8CED6DFFB8FF5FF9D1FD37F984" ID-DOI="http://doi.org/10.5281/zenodo.4321552" ID-Zenodo-Dep="4321552" httpUri="https://zenodo.org/record/4321552/files/figure.png" pageId="4" pageNumber="115" startId="4.[151,250,1592,1616]" targetBox="[151,1435,417,1568]" targetPageId="4">
<paragraph id="8B84EA04ED6DFFB8FF5FF9D1FD37F984" blockId="4.[151,1437,1592,1652]" pageId="4" pageNumber="115">
<emphasis id="B94F3616ED6DFFB8FF5FF9D1FED6F9A0" bold="true" box="[151,275,1592,1616]" pageId="4" pageNumber="115">FIGURE 2.</emphasis>
Ventral view of holotype of 
<taxonomicName id="4C3B9187ED6DFFB8FDFEF9D1FCECF9A0" authority="Medeiros &amp; Moreira &amp; Pinna &amp; Lima, 2020" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[566,809,1592,1616]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="115" phylum="Chordata" rank="species" species="bendego" status="sp. nov.">
<emphasis id="B94F3616ED6DFFB8FDFEF9D1FCECF9A0" box="[566,809,1592,1616]" italics="true" pageId="4" pageNumber="115">Microcambeva bendego</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A27C8B6DED6DFFB8FCFEF9D1FC77F9A0" box="[822,946,1592,1616]" pageId="4" pageNumber="115" rank="species">new species</taxonomicNameLabel>
, holotype, MNRJ 52042, 28.1 mm SL, white arrow indicates the finger-like projections. Scale bar: 1.0mm.
</paragraph>
</caption>
<paragraph id="8B84EA04ED6DFFB8FF0FF94BFAF4F880" blockId="4.[151,1437,1698,2013]" pageId="4" pageNumber="115">
Premaxilla with two tooth rows. Eight conical teeth in labial row and 
<quantity id="4CC347E1ED6DFFB8FC10F94BFBD4F94C" box="[984,1041,1698,1724]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="4" pageNumber="115" unit="in" value="12.0">12 in</quantity>
lingual; premaxillary dorsal protuberance present and expanded near palatine cartilage (
<figureCitation id="1300F681ED6DFFB8FD35F92FFC86F910" box="[765,835,1734,1760]" captionStart="FIGURE 3" captionStartId="5.[151,250,1708,1732]" captionTargetBox="[177,1410,813,1685]" captionTargetId="figure-281@5.[177,1410,813,1685]" captionTargetPageId="5" captionText="FIGURE 3. Skull, Hyoid Arch, Jaws, Opercular Apparatus, pectoral fin and girdle, Suspensorium, anterior vertebrae and Weberian complex of Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Dorsal view (A) and Ventral view (B). Abbreviations: AAR, Anguloarticular; ACH: Anterior ceratohyal; BAR: Barbular; BAS+EXO: Basioccipital-exoccipital bone; BRR: Branchiostegal rays; CLE: Cleithrum; DEN: Dentary; EPO: Epioccipital; HYO: Hyomandibula; IOP: Interopercle; LAN: Lacrimal-antorbital; LAT: Lateral ethmoid; MAX: Maxilla; MET: Mesethmoid; MPT: Metapterygoid; OPE: Opercle; ORB: Orbitosphenoid; PAT: Parietal; PAL: Autoplatine; PCH: Posterior ceratohyal; PMX: Premaxilla; POP, Preopercle; PSC: Posttemporo-supracleithrum; PSO: Parieto-supraoccipital; PTE: Pterotic; PUH: Parurohyal; QUA: Quadrate; SCO, Scapulocoracoid; SPH+POT+PSF: Sphenotic + Prootic + Pterosphenoid complex; VHH: Ventral hypohyal; VOM: Vomer; WEB: Capsule of Weberian apparatus; psp S6: Posterior supraorbital pore S6. Scale bar: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4321554" httpUri="https://zenodo.org/record/4321554/files/figure.png" pageId="4" pageNumber="115">Fig. 3</figureCitation>
). Maxilla long, 50% larger than premaxilla, its distal tip pointed and triangular expansion on its ventrolateral portion. Autopalatine long with moderately concave lateral margin, and straight mesial margin. Posterior process of autopalatine long and pointed. Lacrimal-antorbital elongated and cylindrical approximately 75% of barbular length. Barbular bone long and cylindrical; lacrimal-antorbital and barbular disposed in line, separated by gap approximately equivalent to length of lacrimal-antorbital.
</paragraph>
<paragraph id="8B84EA04ED6DFFB9FF0FF893FDB4FE05" blockId="4.[151,1437,1698,2013]" lastBlockId="5.[151,1437,151,789]" lastPageId="5" lastPageNumber="116" pageId="4" pageNumber="115">
Metapterygoid small and elliptical. Quadrate elongated and concave, with small dorsally-directed process forming distal synchondrosis with metapterygoid (
<figureCitation id="1300F681ED6DFFB8FD5CF877FD1CF848" box="[660,729,1950,1976]" captionStart="FIGURE 4" captionStartId="6.[151,250,1189,1213]" captionTargetBox="[151,1435,583,1165]" captionTargetId="figure-200@6.[151,1435,583,1165]" captionTargetPageId="6" captionText="FIGURE 4. Suspensorium, opercular apparatus and jaws of Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Left lateral view. Abbreviations: ACH: Anterior ceratohyal; BRR: Branchiostegal rays; DEN: Dentary; HYO: Hyomandibula; IOP: Interopercle; LAT: Lateral ethmoid; MAX: Maxilla; MET: Mesethmoid; OPE: Opercle; ORB: Orbitosphenoid; PMX: Premaxilla; POP: Preopercle; PUH: Parurohyal; QUA: Quadrate; VHH: Ventral hypohyal. Scale bar: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4321556" httpUri="https://zenodo.org/record/4321556/files/figure.png" pageId="4" pageNumber="115">Fig. 4</figureCitation>
). Quadrate-hyomandibula fenestra lacking defined shape. Hyomandibula narrow, long, with pointed anterior process with slightly ventrally-curved anteriorly. Ventroposterior margin of hyomandibula with lateral condyle articulating with opercle. Preopercle narrow, pointed anteriorly and round posteriorly, its anteromedial region concave with lateral extension articulating with interopercle, its central portion with pointed process. Opercle narrow, with 19 conical, posteriorly-directed odontodes arranged obliquely in five irregular rows on posterior region. Opercle with pronounced dorsomedial concavity and small pointed process anteriorly. Opercle articulating dorsally with hyomandibula via small condyle on anterior medial process. Interopercle narrow with eight conical, posteriorly-oriented odontodes arranged in two oblique irregular rows on posterior region. Interopercle with deep concavity in dorsomedial region, with small pointed process on anterior portion. Odontodes progressively larger and more strongly curved posteriorly, on both opercle and interopercle. Dentary straight, with two rows of small conical teeth, seven teeth in inner row and nine in outer one. Prominent coronoid process and associated Meckel’s cartilage.
</paragraph>
<paragraph id="8B84EA04ED6CFFB9FF0FFE16FC7BFCE5" blockId="5.[151,1437,151,789]" pageId="5" pageNumber="116">
Parurohyal with two long wings tapering gradually from base to tip, with slight concavity between them. Posterior median process straight and pointed, shorter than lateral wings. Hypobranchial foramen small and circular. Anterior portion of parurohyal with two small condyles articulating with ventral hypohyals (
<figureCitation id="1300F681ED6CFFB9FB51FDAEFB35FD91" box="[1177,1264,583,609]" captionStart="FIGURE 3" captionStartId="5.[151,250,1708,1732]" captionTargetBox="[177,1410,813,1685]" captionTargetId="figure-281@5.[177,1410,813,1685]" captionTargetPageId="5" captionText="FIGURE 3. Skull, Hyoid Arch, Jaws, Opercular Apparatus, pectoral fin and girdle, Suspensorium, anterior vertebrae and Weberian complex of Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Dorsal view (A) and Ventral view (B). Abbreviations: AAR, Anguloarticular; ACH: Anterior ceratohyal; BAR: Barbular; BAS+EXO: Basioccipital-exoccipital bone; BRR: Branchiostegal rays; CLE: Cleithrum; DEN: Dentary; EPO: Epioccipital; HYO: Hyomandibula; IOP: Interopercle; LAN: Lacrimal-antorbital; LAT: Lateral ethmoid; MAX: Maxilla; MET: Mesethmoid; MPT: Metapterygoid; OPE: Opercle; ORB: Orbitosphenoid; PAT: Parietal; PAL: Autoplatine; PCH: Posterior ceratohyal; PMX: Premaxilla; POP, Preopercle; PSC: Posttemporo-supracleithrum; PSO: Parieto-supraoccipital; PTE: Pterotic; PUH: Parurohyal; QUA: Quadrate; SCO, Scapulocoracoid; SPH+POT+PSF: Sphenotic + Prootic + Pterosphenoid complex; VHH: Ventral hypohyal; VOM: Vomer; WEB: Capsule of Weberian apparatus; psp S6: Posterior supraorbital pore S6. Scale bar: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4321554" httpUri="https://zenodo.org/record/4321554/files/figure.png" pageId="5" pageNumber="116">Fig. 3B</figureCitation>
). Ventral hypohyal triangular, with gentle concavity on dorsal portion and slightly depression on ventral portion, accommodating parurohyal condyles. Anterior ceratohyal cylindrical, with constriction at middle; its anterior margin nearly straight, its posterior margin concave dorsally and convex ventrally with both ends lined with cartilage. Posterior ceratohyal roughly triangular, with irregular margins. Six branchiostegal rays, cylindrical and pointed, three on posterior ceratohyal, one on interceratohyal cartilage and two on posterior ceratohyal.
</paragraph>
<caption id="DF44BA8CED6CFFB9FF5FF945FC0BF814" ID-DOI="http://doi.org/10.5281/zenodo.4321554" ID-Zenodo-Dep="4321554" httpUri="https://zenodo.org/record/4321554/files/figure.png" pageId="5" pageNumber="116" startId="5.[151,250,1708,1732]" targetBox="[177,1410,813,1685]" targetPageId="5">
<paragraph id="8B84EA04ED6CFFB9FF5FF945FC0BF814" blockId="5.[151,1437,1708,2020]" pageId="5" pageNumber="116">
<emphasis id="B94F3616ED6CFFB9FF5FF945FED7F934" bold="true" box="[151,274,1708,1732]" pageId="5" pageNumber="116">FIGURE 3.</emphasis>
Skull, Hyoid Arch, Jaws, Opercular Apparatus, pectoral fin and girdle, Suspensorium, anterior vertebrae and Weberian complex of 
<taxonomicName id="4C3B9187ED6CFFB9FEF5F939FDF5F918" authority="Medeiros &amp; Moreira &amp; Pinna &amp; Lima, 2020" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[317,560,1744,1768]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="116" phylum="Chordata" rank="species" species="bendego" status="sp. nov.">
<emphasis id="B94F3616ED6CFFB9FEF5F939FDF5F918" box="[317,560,1744,1768]" italics="true" pageId="5" pageNumber="116">Microcambeva bendego</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A27C8B6DED6CFFB9FDF5F938FD7CF918" box="[573,697,1744,1768]" pageId="5" pageNumber="116" rank="species">new species</taxonomicNameLabel>
, holotype, MNRJ 52042, 28.1 mm SL. Dorsal view (A) and Ventral view (B). Abbreviations: AAR, Anguloarticular; ACH: Anterior ceratohyal; BAR: Barbular; BAS+EXO: Basioccipital-exoccipital bone; BRR: Branchiostegal rays; CLE: Cleithrum; DEN: Dentary; EPO: Epioccipital; HYO: Hyomandibula; IOP: Interopercle; LAN: Lacrimal-antorbital; LAT: Lateral ethmoid; MAX: Maxilla; MET: Mesethmoid; MPT: Metapterygoid; OPE: Opercle; ORB: Orbitosphenoid; PAT: Parietal; PAL: Autoplatine; PCH: Posterior ceratohyal; PMX: Premaxilla; POP, Preopercle; PSC: Posttemporo-supracleithrum; PSO: Parieto-supraoccipital; PTE: Pterotic; PUH: Parurohyal; QUA: Quadrate; SCO, Scapulocoracoid; SPH+POT+PSF: Sphenotic + Prootic + Pterosphenoid complex; VHH: Ventral hypohyal; VOM: Vomer; WEB: Capsule of Weberian apparatus; psp S6: Posterior supraorbital pore S6. Scale bar: 1.0mm.
</paragraph>
</caption>
<paragraph id="8B84EA04ED6FFFBAFF0FFF7EFD9AFDDF" blockId="6.[151,1437,151,560]" pageId="6" pageNumber="117">
Basibranchial 1 absent. Basibranchial 2–3 rod-like with anterior and posterior tips cartilaginous; basibranchial- 4 fully cartilaginous, circular and flattened, with slight constriction at middle (
<figureCitation id="1300F681ED6FFFBAFBCEFF50FB9BFF23" box="[1030,1118,185,211]" captionStart="FIGURE 3" captionStartId="5.[151,250,1708,1732]" captionTargetBox="[177,1410,813,1685]" captionTargetId="figure-281@5.[177,1410,813,1685]" captionTargetPageId="5" captionText="FIGURE 3. Skull, Hyoid Arch, Jaws, Opercular Apparatus, pectoral fin and girdle, Suspensorium, anterior vertebrae and Weberian complex of Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Dorsal view (A) and Ventral view (B). Abbreviations: AAR, Anguloarticular; ACH: Anterior ceratohyal; BAR: Barbular; BAS+EXO: Basioccipital-exoccipital bone; BRR: Branchiostegal rays; CLE: Cleithrum; DEN: Dentary; EPO: Epioccipital; HYO: Hyomandibula; IOP: Interopercle; LAN: Lacrimal-antorbital; LAT: Lateral ethmoid; MAX: Maxilla; MET: Mesethmoid; MPT: Metapterygoid; OPE: Opercle; ORB: Orbitosphenoid; PAT: Parietal; PAL: Autoplatine; PCH: Posterior ceratohyal; PMX: Premaxilla; POP, Preopercle; PSC: Posttemporo-supracleithrum; PSO: Parieto-supraoccipital; PTE: Pterotic; PUH: Parurohyal; QUA: Quadrate; SCO, Scapulocoracoid; SPH+POT+PSF: Sphenotic + Prootic + Pterosphenoid complex; VHH: Ventral hypohyal; VOM: Vomer; WEB: Capsule of Weberian apparatus; psp S6: Posterior supraorbital pore S6. Scale bar: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4321554" httpUri="https://zenodo.org/record/4321554/files/figure.png" pageId="6" pageNumber="117">Fig. 3B</figureCitation>
). Hypobranchial 1, rod-like, anterior and posterior ends with globose cartilages; hypobranchial 2–3 conical, with constriction at middle. Ceratobranchial 1–4 cylindrical, with irregular dorsal margins. Ceratobranchial 1–2 with small laminar expansion at ventromedial region; ceratobranchial 3 with small round process at ventromedial region; ceratobranchial 4 with pointed process near anterodorsal margin. Five conical pharyngeal teeth irregularly arranged on ceratobranchial-5. Epibranchial 1 Y-shaped, with pronounced concavity at distal portion. Epibranchial 2 rod-like, with two lateral expansions, a mid-dorsal one directed anteriorly, and an anteroventral one directed posteriorly. Epibranchial 3 laminar with expansion distally at anteroventral portion; epibranchial 4 rectangular. Pharyngobranchials 1–2 absent; pharyngobranchial 3 rod-like, with thin roundish cartilages at tips; pharyngobranchial 4 entirely cartilaginous, inserted at anterior portion of upper pharyngeal tooth plate. Tooth plate large, trapezoid in shape, with 12 conical teeth distributed in single row on ventral surface.
</paragraph>
<caption id="DF44BA8CED6FFFBAFF5FFB4CFAEBFAD9" ID-DOI="http://doi.org/10.5281/zenodo.4321556" ID-Zenodo-Dep="4321556" httpUri="https://zenodo.org/record/4321556/files/figure.png" pageId="6" pageNumber="117" startId="6.[151,250,1189,1213]" targetBox="[151,1435,583,1165]" targetPageId="6">
<paragraph id="8B84EA04ED6FFFBAFF5FFB4CFAEBFAD9" blockId="6.[151,1437,1189,1321]" pageId="6" pageNumber="117">
<emphasis id="B94F3616ED6FFFBAFF5FFB4CFED7FB4D" bold="true" box="[151,274,1189,1213]" pageId="6" pageNumber="117">FIGURE 4.</emphasis>
Suspensorium, opercular apparatus and jaws of 
<taxonomicName id="4C3B9187ED6FFFBAFD3EFB4CFC2DFB4D" authority="Medeiros &amp; Moreira &amp; Pinna &amp; Lima, 2020" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[758,1000,1189,1213]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="117" phylum="Chordata" rank="species" species="bendego" status="sp. nov.">
<emphasis id="B94F3616ED6FFFBAFD3EFB4CFC2DFB4D" box="[758,1000,1189,1213]" italics="true" pageId="6" pageNumber="117">Microcambeva bendego</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A27C8B6DED6FFFBAFC3CFB4CFBAAFB4D" box="[1012,1135,1189,1213]" pageId="6" pageNumber="117" rank="species">new species</taxonomicNameLabel>
, holotype, MNRJ 52042, 28.1 mm SL. Left lateral view. Abbreviations: ACH: Anterior ceratohyal; BRR: Branchiostegal rays; DEN: Dentary; HYO: Hyomandibula; IOP: Interopercle; LAT: Lateral ethmoid; MAX: Maxilla; MET: Mesethmoid; OPE: Opercle; ORB: Orbitosphenoid; PMX: Premaxilla; POP: Preopercle; PUH: Parurohyal; QUA: Quadrate; VHH: Ventral hypohyal. Scale bar: 1.0mm.
</paragraph>
</caption>
<paragraph id="8B84EA04ED6FFFBAFF0FFABEFD6DF977" blockId="6.[151,1437,1367,2017]" pageId="6" pageNumber="117">
Cleithrum triangular and flat; scapulacoracoid ossified, roughly triangular. Basipterygium square, excluding anterior processes; internal and external processes long and slender with pointed tips; external process larger than internal; posterior process rudimentary; pelvic splint pointed and small. Hypural complex composed of two plates (
<figureCitation id="1300F681ED6FFFBAFF68FA56FF21FA2A" box="[160,228,1471,1498]" captionStart="FIGURE 5" captionStartId="7.[151,250,1252,1276]" captionTargetBox="[151,1435,721,1229]" captionTargetId="figure-298@7.[151,1435,721,1229]" captionTargetPageId="7" captionText="FIGURE 5. Caudal skeleton of Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Left lateral view. Abbreviations: HES: Hemal spine; HYP 1–2: Hypural plate 1–2 plus parhypural, fused; HYP 3–5: Hypural plate 3–5 fused; NES: Neural spine; URO: Uroneural. Scale bar: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4321558" httpUri="https://zenodo.org/record/4321558/files/figure.png" pageId="6" pageNumber="117">Fig. 5</figureCitation>
). Lower plate rectangular, supposedly corresponding to fused parhypural plus hypurals 1–2, and upper plate corresponding to hypurals 3–5. Uroneural long and equal in size to adjacent neural spine, in contact with, but not fused to, upper hypural plate. Two procurrent rays associated with uroneural. Neural spine long and pointed, reaching approximately half of upper hypural plate. Hemal spine larger than neural one, not contacting lower hypural plate, and reaching to distal portion of lower hypural plate; two procurrent rays associated with hemal spine. Ventral procurrent rays larger than dorsal ones (
<figureCitation id="1300F681ED6FFFBAFD9EF985FD5FF977" box="[598,666,1644,1671]" captionStart="FIGURE 5" captionStartId="7.[151,250,1252,1276]" captionTargetBox="[151,1435,721,1229]" captionTargetId="figure-298@7.[151,1435,721,1229]" captionTargetPageId="7" captionText="FIGURE 5. Caudal skeleton of Microcambeva bendego, new species, holotype, MNRJ 52042, 28.1 mm SL. Left lateral view. Abbreviations: HES: Hemal spine; HYP 1–2: Hypural plate 1–2 plus parhypural, fused; HYP 3–5: Hypural plate 3–5 fused; NES: Neural spine; URO: Uroneural. Scale bar: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.4321558" httpUri="https://zenodo.org/record/4321558/files/figure.png" pageId="6" pageNumber="117">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B84EA04ED6FFFBBFF0FF966FBC0FE53" blockId="6.[151,1437,1367,2017]" lastBlockId="7.[151,1437,151,697]" lastPageId="7" lastPageNumber="118" pageId="6" pageNumber="117">
<emphasis id="B94F3616ED6FFFBAFF0FF966FE4BF959" bold="true" box="[199,398,1679,1705]" pageId="6" pageNumber="117">Color in alcohol.</emphasis>
Dorsum, lateral and ventrum of body, almost yellowish pale. Four rows of large dark spots, two along sides, one middorsal and one midventral. Lower lateral row midlateral, with nine unevenly-spaced spots extending along longitudinal skeletogenous septum. First spot dorsoposteriorly to base of pectoral fin. Last spot of midlateral series largest, covering hypural plate and central portion of base of caudal fin. Upper lateral row extending along dorsal part of flanks, with three well-defined, bilaterally-aligned anterior spots (third one at transverse line through origin of dorsal fin), plus three or four poorly-defined and bilaterly misaligned ones. Middorsal series with three spots evenly spaced between occiput and origin of dorsal fin (with third one right at fin origin), plus fourth one at middle of dorsal-fin base. Anterior two middorsal spots intercalated with those of upper lateral row. Isolated final middorsal spot on terminus of edge of caudal peduncle. Midventral row with three spots, aligned respectively with distal portion of pelvic fins, middle of anal-fin base and middle of caudal peduncle. Faint files of dark chromato-phores along limits of myotomes, more pronouncedly on epaxial series. Head with large dark field extending over middle of posterior part of neurocranium, from slightly anteriorly to eyes to limit of epaxial musculature, forming two dark arms at that limit. Cephalic color pattern poorly-defined. Eyes and iris black. Narrow dark field extending laterally between anterior and posterior nostrils. Remainder of dorsal surface of head with light uniform covering of dark chromatophores. Nostrils outlined in abrupt white, in stark contrast with remainder of dorsal surface of head. Dark concentration on opercular patch of odontodes. Ventral part of head entirely white. Maxillary and nasal barbels with dark chromatophores concentrated at base. Rictal barbel white. Dark concentrations on proximal third of pectoral fin. Remaining fins hyaline. Pale-gray vertical bar on caudal-fin base.
</paragraph>
</subSubSection>
<subSubSection id="C321B98FED6EFFBBFF0FFE45FBFEFD49" pageId="7" pageNumber="118" type="etymology">
<paragraph id="8B84EA04ED6EFFBBFF0FFE45FBFEFD49" blockId="7.[151,1437,151,697]" pageId="7" pageNumber="118">
<emphasis id="B94F3616ED6EFFBBFF0FFE45FE8BFE36" bold="true" box="[199,334,428,454]" pageId="7" pageNumber="118">Etymology.</emphasis>
The specific epithet is derived from the second-largest meteorite discovered in 
<collectingCountry id="F32CAA94ED6EFFBBFB12FE45FAE4FE36" box="[1242,1313,428,454]" name="Brazil" pageId="7" pageNumber="118">Brazil</collectingCountry>
, the Bendegó. Found in 
<quantity id="4CC347E1ED6EFFBBFEF1FE26FE48FE19" box="[313,397,463,489]" metricMagnitude="1" metricUnit="m" metricValue="4.55676" pageId="7" pageNumber="118" unit="in" value="1794.0">1794 in</quantity>
Northeastern 
<collectingCountry id="F32CAA94ED6EFFBBFDE4FE26FDB6FE19" box="[556,627,463,489]" name="Brazil" pageId="7" pageNumber="118">Brazil</collectingCountry>
, it was transported to the Museu Nacional in 1888, where it became part of its exhibit ever since. In 2018, a fire of huge proportions destroyed part of the Museu Nacional, Universidade Federal do 
<collectingRegion id="49FF24E6ED6EFFBBFEDEFDFDFE7DFDDE" box="[278,440,532,558]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="118">Rio de Janeiro</collectingRegion>
(MNRJ) including some of its bicentennial collections. Even though part of the building collapsed, the Bendegó remained intact at the main entrance of the museum, where it was seen by the crowd that gathered the day after the fire, becoming a symbol of the resistance of the institution. This is not only an homage to the MNRJ, its employees and students, but also an allusion to the resilience of the species herein described in Atlantic Forest basin severely impacted by anthropic actions. A noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF44BA8CED6EFFBBFF5FFB0DFD17FAB4" ID-DOI="http://doi.org/10.5281/zenodo.4321558" ID-Zenodo-Dep="4321558" httpUri="https://zenodo.org/record/4321558/files/figure.png" pageId="7" pageNumber="118" startId="7.[151,250,1252,1276]" targetBox="[151,1435,721,1229]" targetPageId="7">
<paragraph id="8B84EA04ED6EFFBBFF5FFB0DFD17FAB4" blockId="7.[151,1437,1252,1348]" pageId="7" pageNumber="118">
<emphasis id="B94F3616ED6EFFBBFF5FFB0DFED6FB0C" bold="true" box="[151,275,1252,1276]" pageId="7" pageNumber="118">FIGURE 5.</emphasis>
Caudal skeleton of 
<taxonomicName id="4C3B9187ED6EFFBBFE28FB0DFD16FB0C" authority="Medeiros &amp; Moreira &amp; Pinna &amp; Lima, 2020" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[480,723,1252,1276]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="species" species="bendego" status="sp. nov.">
<emphasis id="B94F3616ED6EFFBBFE28FB0DFD16FB0C" box="[480,723,1252,1276]" italics="true" pageId="7" pageNumber="118">Microcambeva bendego</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A27C8B6DED6EFFBBFD28FB0DFC98FB0C" box="[736,861,1252,1276]" pageId="7" pageNumber="118" rank="species">new species</taxonomicNameLabel>
, holotype, MNRJ 52042, 28.1 mm SL. Left lateral view. Abbreviations: HES: Hemal spine; HYP 1–2: Hypural plate 1–2 plus parhypural, fused; HYP 3–5: Hypural plate 3–5 fused; NES: Neural spine; URO: Uroneural. Scale bar: 1.0mm.
</paragraph>
</caption>
<caption id="DF44BA8CED6EFFBBFF5FF8B6FD4AF813" ID-DOI="http://doi.org/10.5281/zenodo.4321560" ID-Zenodo-Dep="4321560" httpUri="https://zenodo.org/record/4321560/files/figure.png" pageId="7" pageNumber="118" startId="7.[151,250,1887,1911]" targetBox="[157,1429,1377,1858]" targetPageId="7">
<paragraph id="8B84EA04ED6EFFBBFF5FF8B6FD4AF813" blockId="7.[151,1437,1887,2019]" pageId="7" pageNumber="118">
<emphasis id="B94F3616ED6EFFBBFF5FF8B6FED6F887" bold="true" box="[151,275,1887,1911]" pageId="7" pageNumber="118">FIGURE 6.</emphasis>
Geographic distribution 
<taxonomicName id="4C3B9187ED6EFFBBFDDAF8B6FD6CF887" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[530,681,1887,1911]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED6EFFBBFDDAF8B6FD6CF887" box="[530,681,1887,1911]" italics="true" pageId="7" pageNumber="118">Microcambeva</emphasis>
</taxonomicName>
catfishes in the Atlantic Forest costal basins of southeastern Brazil. White star: type locality new species 
<taxonomicName id="4C3B9187ED6EFFBBFE11F86DFD96F86B" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[473,595,1923,1947]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED6EFFBBFE11F86DFD96F86B" box="[473,595,1923,1947]" italics="true" pageId="7" pageNumber="118">M. bendego</emphasis>
</taxonomicName>
; Black square: type locality of 
<taxonomicName id="4C3B9187ED6EFFBBFC56F86DFBD7F86B" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[926,1042,1923,1947]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="species" species="barbata">
<emphasis id="B94F3616ED6EFFBBFC56F86DFBD7F86B" box="[926,1042,1923,1947]" italics="true" pageId="7" pageNumber="118">M. barbata</emphasis>
</taxonomicName>
; Blue hexagon: 
<taxonomicName id="4C3B9187ED6EFFBBFB08F86DFA9FF86B" authorityName="Costa, Katz &amp; Vilardo" authorityYear="2020" box="[1216,1370,1923,1947]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="species" species="filamentosa">
<emphasis id="B94F3616ED6EFFBBFB08F86DFA9FF86B" box="[1216,1370,1923,1947]" italics="true" pageId="7" pageNumber="118">M. filamentosa</emphasis>
</taxonomicName>
; Gray diamond: 
<taxonomicName id="4C3B9187ED6EFFBBFF35F841FEB4F84F" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[253,369,1959,1983]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED6EFFBBFF35F841FEB4F84F" box="[253,369,1959,1983]" italics="true" pageId="7" pageNumber="118">M. ribeirae</emphasis>
</taxonomicName>
; Red triangle: 
<taxonomicName id="4C3B9187ED6EFFBBFDC0F841FD4DF84F" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[520,648,1960,1983]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="species" species="jucuensis">
<emphasis id="B94F3616ED6EFFBBFDC0F841FD4DF84F" box="[520,648,1960,1983]" italics="true" pageId="7" pageNumber="118">M. jucuensis</emphasis>
</taxonomicName>
; Yellow circle: 
<taxonomicName id="4C3B9187ED6EFFBBFCE0F841FC01F84F" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[808,964,1960,1983]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="species" species="mucuriensis">
<emphasis id="B94F3616ED6EFFBBFCE0F841FC01F84F" box="[808,964,1960,1983]" italics="true" pageId="7" pageNumber="118">M. mucuriensis</emphasis>
</taxonomicName>
; Pentagon orange: 
<taxonomicName id="4C3B9187ED6EFFBBFB41F841FB23F84F" authorityName="Mattos &amp; Lima" authorityYear="2010" box="[1161,1254,1959,1983]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="118" phylum="Chordata" rank="species" species="draco">
<emphasis id="B94F3616ED6EFFBBFB41F841FB23F84F" box="[1161,1254,1959,1983]" italics="true" pageId="7" pageNumber="118">M. draco</emphasis>
</taxonomicName>
. White line: Guanabara Bay region; Black line: River basin limits.
</paragraph>
</caption>
<subSubSection id="C321B98FED61FFB4FF0FFF7FFBEBFE21" pageId="8" pageNumber="119" type="distribution">
<paragraph id="8B84EA04ED61FFB4FF0FFF7FFB0BFF26" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="119">
<emphasis id="B94F3616ED61FFB4FF0FFF7FFEA4FF40" bold="true" box="[199,353,150,176]" pageId="8" pageNumber="119">Distribution.</emphasis>
Known only from its 
<typeStatus id="548054A6ED61FFB4FD96FF7EFD4BFF41" box="[606,654,151,177]" pageId="8" pageNumber="119">type</typeStatus>
locality, in the middle section of the rio Guapiaçu, in the Guapi-Macacu system, a basin that drains directly into the northeastern portion of Guanabara Bay (
<figureCitation id="1300F681ED61FFB4FBB4FF52FB05FF26" box="[1148,1216,187,214]" captionStart="FIGURE 6" captionStartId="7.[151,250,1887,1911]" captionTargetBox="[157,1429,1377,1858]" captionTargetId="figure-355@7.[151,1435,1372,1862]" captionTargetPageId="7" captionText="FIGURE 6. Geographic distribution Microcambeva catfishes in the Atlantic Forest costal basins of southeastern Brazil. White star: type locality new species M. bendego; Black square: type locality of M. barbata; Blue hexagon: M. filamentosa; Gray diamond: M. ribeirae; Red triangle: M. jucuensis; Yellow circle: M. mucuriensis; Pentagon orange: M. draco. White line: Guanabara Bay region; Black line: River basin limits." figureDoi="http://doi.org/10.5281/zenodo.4321560" httpUri="https://zenodo.org/record/4321560/files/figure.png" pageId="8" pageNumber="119">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8B84EA04ED61FFB4FF0FFF37FBEBFE21" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="119">
<emphasis id="B94F3616ED61FFB4FF0FFF37FDE2FF09" bold="true" box="[199,551,222,249]" pageId="8" pageNumber="119">Habitat and ecological notes.</emphasis>
<taxonomicName id="4C3B9187ED61FFB4FDF9FF36FC84FF09" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[561,833,223,249]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED61FFB4FDF9FF36FC84FF09" box="[561,833,223,249]" italics="true" pageId="8" pageNumber="119">Microcambeva bendego</emphasis>
</taxonomicName>
was found in the middle course of rio Guapiaçu, in a stream approximately 
<quantity id="4CC347E1ED61FFB4FE78FEEAFE29FEED" box="[432,492,259,285]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="119" unit="m" value="10.0">10 m</quantity>
wide and 
<quantity id="4CC347E1ED61FFB4FDA3FEEAFD5CFEED" box="[619,665,259,285]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="119" unit="m" value="1.0">1 m</quantity>
deep, with sandy bottom and riparian vegetation composed mostly of grass-like 
<taxonomicName id="4C3B9187ED61FFB4FE85FECEFE16FEB1" authorityName="de Jussieu" authorityYear="1789" box="[333,467,295,321]" class="Liliopsida" family="Cyperaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="119" phylum="Tracheophyta" rank="family">Cyperaceae</taxonomicName>
. The new species was collected along with 
<taxonomicName id="4C3B9187ED61FFB4FC04FECEFA52FEB1" authority="(Steindachner 1877)" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[972,1431,295,321]" class="Actinopterygii" family="Loricariidae" genus="Hypostomus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="affinis">
<emphasis id="B94F3616ED61FFB4FC04FECEFB63FEB1" box="[972,1190,295,321]" italics="true" pageId="8" pageNumber="119">Hypostomus affinis</emphasis>
(Steindachner 1877)
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FF5FFEA2FD7CFE95" authority="(Steindachner 1877)" baseAuthorityName="Steindachner" baseAuthorityYear="1877" box="[151,697,331,357]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="maculicauda">
<emphasis id="B94F3616ED61FFB4FF5FFEA2FE09FE95" box="[151,460,331,357]" italics="true" pageId="8" pageNumber="119">Parotocinclus maculicauda</emphasis>
(Steindachner 1877)
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FD3AFEA2FC4CFE95" authorityName="Bleeker in Bleeker" authorityYear="1862" box="[754,905,331,357]" class="Actinopterygii" family="Loricariidae" genus="Rineloricaria" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED61FFB4FD3AFEA2FC4CFE95" box="[754,905,331,357]" italics="true" pageId="8" pageNumber="119">Rineloricaria</emphasis>
</taxonomicName>
sp., plus two introduced aquarium species, the three-spot gourami 
<taxonomicName id="4C3B9187ED61FFB4FEB8FE86FCE1FE79" authority="(Pallas 1770)" baseAuthorityName="Pallas" baseAuthorityYear="1770" box="[368,804,367,393]" class="Actinopterygii" family="Osphronemidae" genus="Trichopodus" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="trichopterus">
<emphasis id="B94F3616ED61FFB4FEB8FE86FD4EFE79" box="[368,651,367,393]" italics="true" pageId="8" pageNumber="119">Trichopodus trichopterus</emphasis>
(Pallas 1770)
</taxonomicName>
and the jewel tetra 
<taxonomicName id="4C3B9187ED61FFB4FC33FE86FF19FE5D" authority="(Steindachner 1882)" baseAuthorityName="Steindachner" baseAuthorityYear="1882" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="eques">
<emphasis id="B94F3616ED61FFB4FC33FE86FB3EFE78" box="[1019,1275,367,393]" italics="true" pageId="8" pageNumber="119">Hyphessobrycon eques</emphasis>
(Steindachner 1882)
</taxonomicName>
. A recent visit to the 
<typeStatus id="548054A6ED61FFB4FE0AFE7AFE37FE5D" box="[450,498,403,429]" pageId="8" pageNumber="119">type</typeStatus>
locality did not secure any 
<taxonomicName id="4C3B9187ED61FFB4FCEBFE7AFC0FFE5D" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[803,970,403,429]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED61FFB4FCEBFE7AFC0FFE5D" box="[803,970,403,429]" italics="true" pageId="8" pageNumber="119">Microcambeva</emphasis>
</taxonomicName>
specimens. According to local fishermen, the African catfish 
<taxonomicName id="4C3B9187ED61FFB4FEA5FE5EFD3CFE21" authority="(Burchell 1822)" baseAuthorityName="Burchell" baseAuthorityYear="1822" box="[365,761,439,465]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="gariepinus">
<emphasis id="B94F3616ED61FFB4FEA5FE5EFD85FE20" box="[365,576,439,465]" italics="true" pageId="8" pageNumber="119">Clarias gariepinus</emphasis>
(Burchell 1822)
</taxonomicName>
is also found in the stream.
</paragraph>
</subSubSection>
<subSubSection id="C321B98FED61FFB6FF0FFE33FDBDFE95" lastPageId="10" lastPageNumber="121" pageId="8" pageNumber="119" type="discussion">
<paragraph id="8B84EA04ED61FFB4FF0FFE33FD90FD5A" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="119">
<emphasis id="B94F3616ED61FFB4FF0FFE33FE89FE04" bold="true" box="[199,332,474,500]" pageId="8" pageNumber="119">Discussion.</emphasis>
<taxonomicName id="4C3B9187ED61FFB4FE9CFE32FDA7FE05" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[340,610,475,501]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED61FFB4FE9CFE32FDA7FE05" box="[340,610,475,501]" italics="true" pageId="8" pageNumber="119">Microcambeva bendego</emphasis>
</taxonomicName>
has all four putative synapomorphies so far proposed for the genus, viz., a reduction in the quadrate process, a rectangular ventral hypural plate; and a reduced number of opercular and interopercular odontodes (
<bibRefCitation id="EFAA97F5ED61FFB4FE6EFDCAFD03FDCD" author="Costa, W. J. E. M. &amp; Bockmann, F. A." box="[422,710,547,573]" pageId="8" pageNumber="119" pagination="715 - 730" refId="ref7972" refString="Costa, W. J. E. M. &amp; Bockmann, F. A. (1994) A new genus and species of Sarcoglanidinae (Siluriformes: Trichomycteridae) from southeastern Brazil, with a re-examination of subfamilial phylogeny. Journal of Natural History, 28, 715 - 730. https: // doi. org / 10.1080 / 00222939400770331" type="journal article" year="1994">Costa &amp; Bockmann 1994</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FD19FDCAFC58FDCD" author="Costa, W. J. E. M. &amp; Henschel, E. &amp; Katz, A. M." box="[721,925,547,573]" pageId="8" pageNumber="119" pagination="159 - 173" refId="ref8282" refString="Costa, W. J. E. M., Henschel, E. &amp; Katz, A. M. (2020 a) Multigene phylogeny reveals convergent evolution in small interstitial catfishes from the Amazon and Atlantic forests (Siluriformes: Trichomycteridae). Zoologica Scripta, 49, 159 - 173. https: // doi. org / 10.1111 / zsc. 12403" type="journal article" year="2020">
Costa 
<emphasis id="B94F3616ED61FFB4FCDFFDCAFC8DFDCD" box="[791,840,547,573]" italics="true" pageId="8" pageNumber="119">et al</emphasis>
. 2020a
</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FC60FDCAFC37FDCE" author="Costa, W. J. E. M. &amp; Vilardo, P. &amp; Katz, A. M." box="[936,1010,547,574]" pageId="8" pageNumber="119" pagination="12 - 17" refId="ref8350" refString="Costa, W. J. E. M., Vilardo, P. &amp; Katz, A. M. (2020 b) Sympatric sister species with divergent morphological features of psammophilic catfishes of the south-eastern Brazilian genus Microcambeva (Siluriformes: Trichomycteridae). Zoologischer Anzeiger, 285, 12 - 17. https: // doi. org / 10.1016 / j. jcz. 2020.01.001" type="journal article" year="2020">2020b</bibRefCitation>
), although polarity of those characters are debatable. The new species has the largest number of opercular odontodes (19) in the genus (
<quantity id="4CC347E1ED61FFB4FB02FDAEFAD4FD91" box="[1226,1297,583,609]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="8" pageNumber="119" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
<taxonomicName id="4C3B9187ED61FFB4FAD0FDAEFA53FD91" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[1304,1430,583,609]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="barbata">
<emphasis id="B94F3616ED61FFB4FAD0FDAEFA53FD91" box="[1304,1430,583,609]" italics="true" pageId="8" pageNumber="119">M. barbata</emphasis>
</taxonomicName>
, 
<quantity id="4CC347E1ED61FFB4FF5FFD82FF2EFD75" box="[151,235,619,645]" metricMagnitude="-1" metricUnit="m" metricValue="2.921" metricValueMax="3.556" metricValueMin="2.286" pageId="8" pageNumber="119" unit="in" value="11.5" valueMax="14.0" valueMin="9.0">9–14 in</quantity>
<taxonomicName id="4C3B9187ED61FFB4FF39FD82FE58FD74" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[241,413,619,644]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="mucuriensis">
<emphasis id="B94F3616ED61FFB4FF39FD82FE58FD74" box="[241,413,619,644]" italics="true" pageId="8" pageNumber="119">M. mucuriensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FE62FD82FDECFD75" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[426,553,619,645]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED61FFB4FE62FD82FDECFD75" box="[426,553,619,645]" italics="true" pageId="8" pageNumber="119">M. ribeirae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FDFEFD82FD59FD75" authorityName="Mattos &amp; Lima" authorityYear="2010" box="[566,668,619,645]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="draco">
<emphasis id="B94F3616ED61FFB4FDFEFD82FD59FD75" box="[566,668,619,645]" italics="true" pageId="8" pageNumber="119">M. draco</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FD19FD82FCBDFD75" authorityName="Costa, Katz &amp; Vilardo" authorityYear="2020" box="[721,888,619,645]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="filamentosa">
<emphasis id="B94F3616ED61FFB4FD19FD82FCBDFD75" box="[721,888,619,645]" italics="true" pageId="8" pageNumber="119">M. filamentosa</emphasis>
</taxonomicName>
, and 
<quantity id="4CC347E1ED61FFB4FC7BFD83FBD3FD75" box="[947,1046,618,645]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="8" pageNumber="119" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
<taxonomicName id="4C3B9187ED61FFB4FBD4FD82FB6DFD74" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[1052,1192,619,644]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="jucuensis">
<emphasis id="B94F3616ED61FFB4FBD4FD82FB6DFD74" box="[1052,1192,619,644]" italics="true" pageId="8" pageNumber="119">M. jucuensis</emphasis>
</taxonomicName>
) (
<bibRefCitation id="EFAA97F5ED61FFB4FB77FD82FF17FD59" author="Costa, W. J. E. M. &amp; Bockmann, F. A." pageId="8" pageNumber="119" pagination="715 - 730" refId="ref7972" refString="Costa, W. J. E. M. &amp; Bockmann, F. A. (1994) A new genus and species of Sarcoglanidinae (Siluriformes: Trichomycteridae) from southeastern Brazil, with a re-examination of subfamilial phylogeny. Journal of Natural History, 28, 715 - 730. https: // doi. org / 10.1080 / 00222939400770331" type="journal article" year="1994">Costa &amp; Bockmann 1994</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FF16FD66FE65FD59" author="Costa, W. J. E. &amp; Lima, S. M. Q. &amp; Bizerril, C. R. S. F." box="[222,416,655,681]" pageId="8" pageNumber="119" pagination="1 - 10" refId="ref8036" refString="Costa, W. J. E. M, Lima, S. M. Q. &amp; Bizerril, C. R. S. F. (2004) Microcambeva ribeirae sp. n. (Teleostei: Siluriformes: Trichomycteridae): a new sarcoglanidine catfish from the Rio Ribeira do Iguape basin, southeastern Brazil. Zootaxa, 563 (1), 1 - 10. https: // doi. org / 10.11646 / zootaxa. 563.1.1" type="journal article" year="2004">
Costa 
<emphasis id="B94F3616ED61FFB4FEEDFD66FE98FD59" box="[293,349,655,681]" italics="true" pageId="8" pageNumber="119">et al.</emphasis>
2004
</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FE63FD66FE37FD59" author="Costa, W. J. E. M. &amp; Katz, A. M. &amp; Mattos, J. L. O. &amp; Rangel-Pereira, F. S." box="[427,498,655,681]" pageId="8" pageNumber="119" pagination="1837 - 1851" refId="ref8120" refString="Costa, W. J. E. M., Katz, A. M., Mattos, J. L. O. &amp; Rangel-Pereira, F. S. (2019 a) Two new species of miniature psammophilic sarcoglanidine catfishes of the genus Microcambeva from the Atlantic Forest of eastern Brazil (Siluriformes: Trichomycteridae). Journal of Natural History, 53, 1837 - 1851. https: // doi. org / 10.1080 / 00222933.2019.1669729" type="journal article" year="2019">2019a</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FE36FD66FD8DFD5A" author="Costa, W. J. E. M. &amp; Vilardo, P. &amp; Katz, A. M." box="[510,584,655,682]" pageId="8" pageNumber="119" pagination="12 - 17" refId="ref8350" refString="Costa, W. J. E. M., Vilardo, P. &amp; Katz, A. M. (2020 b) Sympatric sister species with divergent morphological features of psammophilic catfishes of the south-eastern Brazilian genus Microcambeva (Siluriformes: Trichomycteridae). Zoologischer Anzeiger, 285, 12 - 17. https: // doi. org / 10.1016 / j. jcz. 2020.01.001" type="journal article" year="2020">2020b</bibRefCitation>
).
</paragraph>
<paragraph id="8B84EA04ED61FFB4FF0FFD5AFCE2FC39" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="119">
An additional putative synapomorphy of the genus is the presence of a distinct pair of finger-like projections, similar in position to a mentonian barbel. This barbel of unknown function occurs in all species of the genus, except 
<taxonomicName id="4C3B9187ED61FFB4FF5FFD12FDECFCE6" authority="(Costa et al. 2020 b)" baseAuthorityName="Costa" baseAuthorityYear="2020" box="[151,553,763,790]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="filamentosa">
<emphasis id="B94F3616ED61FFB4FF5FFD12FE85FCE5" box="[151,320,763,789]" italics="true" pageId="8" pageNumber="119">M. filamentosa</emphasis>
(
<bibRefCitation id="EFAA97F5ED61FFB4FE86FD12FDE5FCE6" author="Costa, W. J. E. M. &amp; Vilardo, P. &amp; Katz, A. M." box="[334,544,763,790]" pageId="8" pageNumber="119" pagination="12 - 17" refId="ref8350" refString="Costa, W. J. E. M., Vilardo, P. &amp; Katz, A. M. (2020 b) Sympatric sister species with divergent morphological features of psammophilic catfishes of the south-eastern Brazilian genus Microcambeva (Siluriformes: Trichomycteridae). Zoologischer Anzeiger, 285, 12 - 17. https: // doi. org / 10.1016 / j. jcz. 2020.01.001" type="journal article" year="2020">
Costa 
<emphasis id="B94F3616ED61FFB4FE5FFD12FE15FCE5" box="[407,464,763,789]" italics="true" pageId="8" pageNumber="119">et al.</emphasis>
2020b
</bibRefCitation>
)
</taxonomicName>
, and at least five unrelated trichomycterid species, 
<taxonomicName id="4C3B9187ED61FFB4FBAFFD12FE62FCC9" authority="(Steindachner 1879)" baseAuthorityName="Steindachner" baseAuthorityYear="1879" class="Actinopterygii" family="Trichomycteridae" genus="Pseudostegophilus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="maculatus">
<emphasis id="B94F3616ED61FFB4FBAFFD12FF7DFCC8" italics="true" pageId="8" pageNumber="119">Pseudostegophilus maculatus</emphasis>
(Steindachner 1879)
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FE7CFCF6FBC5FCC9" authority="Myers &amp; Weitzman 1966" authorityName="Myers &amp; Weitzman" authorityYear="1966" box="[436,1024,799,825]" class="Actinopterygii" family="Trichomycteridae" genus="Malacoglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="gelatinosus">
<emphasis id="B94F3616ED61FFB4FE7CFCF6FD12FCC9" box="[436,727,799,825]" italics="true" pageId="8" pageNumber="119">Malacoglanis gelatinosus</emphasis>
<bibRefCitation id="EFAA97F5ED61FFB4FD16FCF6FBC5FCC9" author="Myers, G. S. &amp; Weitzman, S. H." box="[734,1024,799,825]" pageId="8" pageNumber="119" pagination="277 - 287" refId="ref9021" refString="Myers, G. S. &amp; Weitzman, S. H. (1966) Two remarkable new trichomycterid catfishes from the Amazon basin in Brazil and Colombia. Journal of Zoology, 149 (3), 277 - 287. https: // doi. org / 10.1111 / j. 1469 - 7998.1966. tb 04049. x" type="journal article" year="1966">Myers &amp; Weitzman 1966</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FBC4FCF6FEEEFCAD" authority="de Pinna &amp; Starnes 1990" authorityName="de Pinna &amp; Starnes" authorityYear="1990" class="Actinopterygii" family="Trichomycteridae" genus="Stenolicmus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="sarmientoi">
<emphasis id="B94F3616ED61FFB4FBC4FCF6FAD1FCC8" box="[1036,1300,799,825]" italics="true" pageId="8" pageNumber="119">Stenolicmus sarmientoi</emphasis>
de Pinna &amp; Starnes 1990
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FEFDFCAAFC8EFCAD" authority="de Pinna &amp; Winemiller 2000" authorityName="de Pinna &amp; Winemiller" authorityYear="2000" box="[309,843,835,861]" class="Actinopterygii" family="Trichomycteridae" genus="Ammoglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="pulex">
<emphasis id="B94F3616ED61FFB4FEFDFCAAFDC1FCAD" box="[309,516,835,861]" italics="true" pageId="8" pageNumber="119">Ammoglanis pulex</emphasis>
<bibRefCitation id="EFAA97F5ED61FFB4FDC2FCAAFC8EFCAD" author="de Pinna, M. C. C. &amp; Winemiller, K. O." box="[522,843,835,861]" pageId="8" pageNumber="119" pagination="255 - 264" refId="ref9311" refString="de Pinna, M. C. C. &amp; Winemiller, K. O. (2000) A new species of Ammoglanis (Siluriformes: Trichomycteridae) from Venezuela. Ichthyological Exploration of Freshwaters, 11 (3), 255 - 264." type="journal article" year="2000">de Pinna &amp; Winemiller 2000</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C3B9187ED61FFB4FC4BFCAAFEF4FC71" authority="Henschel, Braganca, Rangel-Pereira &amp; Costa 2019" authorityName="Henschel, Braganca, Rangel-Pereira &amp; Costa" authorityYear="2019" class="Actinopterygii" family="Trichomycteridae" genus="Ammoglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="obliquus">
<emphasis id="B94F3616ED61FFB4FC4BFCAAFBC7FCAD" box="[899,1026,835,861]" italics="true" pageId="8" pageNumber="119">A. obliquus</emphasis>
Henschel, Bragança, Rangel-Pereira &amp; Costa 2019
</taxonomicName>
(
<bibRefCitation id="EFAA97F5ED61FFB4FEF7FC8EFD4CFC71" author="de Pinna, M. C. C. &amp; Winemiller, K. O." box="[319,649,871,897]" pageId="8" pageNumber="119" pagination="255 - 264" refId="ref9311" refString="de Pinna, M. C. C. &amp; Winemiller, K. O. (2000) A new species of Ammoglanis (Siluriformes: Trichomycteridae) from Venezuela. Ichthyological Exploration of Freshwaters, 11 (3), 255 - 264." type="journal article" year="2000">de Pinna &amp; Winemiller, 2000</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FD5BFC8EFCB9FC71" author="Henschel, E. &amp; Braganca, P. H. N. &amp; Rangel-Pereira, F. &amp; Costa, W. J. E. M." box="[659,892,871,897]" pageId="8" pageNumber="119" pagination="67 - 72" refId="ref8487" refString="Henschel, E., Braganca, P. H. N., Rangel-Pereira, F. &amp; Costa, W. J. E. M. (2020) A new psammophilic species of the catfish genus Ammoglanis (Siluriformes, Trichomycteridae) from the Amazon River basin, northern Brazil. Zoosystematics and Evolution, 96 (1), 67 - 72. https: // doi. org / 10.3897 / zse. 96.48952" type="journal article" year="2020">
Henschel 
<emphasis id="B94F3616ED61FFB4FCCAFC8EFCFCFC71" box="[770,825,871,897]" italics="true" pageId="8" pageNumber="119">et al.</emphasis>
2020
</bibRefCitation>
). The structure in 
<taxonomicName id="4C3B9187ED61FFB4FB8BFC8EFB03FC71" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[1091,1222,871,897]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED61FFB4FB8BFC8EFB03FC71" box="[1091,1222,871,897]" italics="true" pageId="8" pageNumber="119">M. bendego</emphasis>
</taxonomicName>
is the largest so far recorded for the genus, in stark contrast to the minute barbels seen in 
<taxonomicName id="4C3B9187ED61FFB4FC64FC62FBD6FC55" authorityName="Mattos &amp; Lima" authorityYear="2010" box="[940,1043,907,933]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="draco">
<emphasis id="B94F3616ED61FFB4FC64FC62FBD6FC55" box="[940,1043,907,933]" italics="true" pageId="8" pageNumber="119">M. draco</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FBEAFC62FB75FC54" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[1058,1200,907,932]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="jucuensis">
<emphasis id="B94F3616ED61FFB4FBEAFC62FB75FC54" box="[1058,1200,907,932]" italics="true" pageId="8" pageNumber="119">M. jucuensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C3B9187ED61FFB4FB27FC62FA59FC54" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[1263,1436,907,932]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="mucuriensis">
<emphasis id="B94F3616ED61FFB4FB27FC62FA59FC54" box="[1263,1436,907,932]" italics="true" pageId="8" pageNumber="119">M. mucuriensis</emphasis>
</taxonomicName>
and the medium-sized ones in 
<taxonomicName id="4C3B9187ED61FFB4FE25FC46FDAEFC39" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[493,619,943,969]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="barbata">
<emphasis id="B94F3616ED61FFB4FE25FC46FDAEFC39" box="[493,619,943,969]" italics="true" pageId="8" pageNumber="119">M. barbata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FD69FC46FCE5FC39" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[673,800,943,969]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED61FFB4FD69FC46FCE5FC39" box="[673,800,943,969]" italics="true" pageId="8" pageNumber="119">M. ribeirae</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84EA04ED61FFB4FF0FFC3AFE30FA89" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="119">
<emphasis id="B94F3616ED61FFB4FF0FFC3AFD54FC1D" box="[199,657,979,1005]" italics="true" pageId="8" pageNumber="119">
<taxonomicName id="4C3B9187ED61FFB4FF0FFC3AFE13FC1D" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[199,470,979,1005]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="ribeirae">Microcambeva ribeirae</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FE2DFC3AFD54FC1D" box="[485,657,979,1005]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="filamentosa">M. filamentosa</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C3B9187ED61FFB4FD07FC3AFC92FC1D" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[719,855,979,1005]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED61FFB4FD07FC3AFC92FC1D" box="[719,855,979,1005]" italics="true" pageId="8" pageNumber="119">M. bendego</emphasis>
</taxonomicName>
share three character-states not observed in other congeners: median fusion of the third pore (s6) of the supraorbital canal, absence of the anterior autopalatine ossification, and a deep concavity between the base of the ascending process of opercle and the patch of odontodes. Among 
<emphasis id="B94F3616ED61FFB4FF3DFBD6FE67FBA9" box="[245,418,1087,1113]" italics="true" pageId="8" pageNumber="119">
<taxonomicName id="4C3B9187ED61FFB4FF3DFBD6FE5AFBA9" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[245,415,1087,1113]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">Microcambeva</taxonomicName>
,
</emphasis>
the new species additionally shares exclusively with 
<taxonomicName id="4C3B9187ED61FFB4FBC4FBD6FB48FBA9" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[1036,1165,1087,1113]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED61FFB4FBC4FBD6FB48FBA9" box="[1036,1165,1087,1113]" italics="true" pageId="8" pageNumber="119">M. ribeirae</emphasis>
</taxonomicName>
two distinct features: a non-filamentous first pectoral-fin ray and conspicuous series of dark spots along the body. In other congeners, the first pectoral-fin ray is modified as a filament, as the long filament in 
<taxonomicName id="4C3B9187ED61FFB4FC65FB6EFB93FB51" box="[941,1110,1159,1185]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="filamentosa">
<emphasis id="B94F3616ED61FFB4FC65FB6EFB93FB51" box="[941,1110,1159,1185]" italics="true" pageId="8" pageNumber="119">M. filamentosa</emphasis>
</taxonomicName>
(70–80% pectoral-fin length without filament), medium sized in 
<taxonomicName id="4C3B9187ED61FFB4FDE5FB42FD69FB35" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[557,684,1195,1221]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="barbata">
<emphasis id="B94F3616ED61FFB4FDE5FB42FD69FB35" box="[557,684,1195,1221]" italics="true" pageId="8" pageNumber="119">M. barbata</emphasis>
</taxonomicName>
(40%) and 
<taxonomicName id="4C3B9187ED61FFB4FCFAFB42FC5CFB35" authorityName="Mattos &amp; Lima" authorityYear="2010" box="[818,921,1195,1221]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="draco">
<emphasis id="B94F3616ED61FFB4FCFAFB42FC5CFB35" box="[818,921,1195,1221]" italics="true" pageId="8" pageNumber="119">M. draco</emphasis>
</taxonomicName>
(50%), and short in 
<taxonomicName id="4C3B9187ED61FFB4FB4CFB42FAD7FB34" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" box="[1156,1298,1195,1220]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="jucuensis">
<emphasis id="B94F3616ED61FFB4FB4CFB42FAD7FB34" box="[1156,1298,1195,1220]" italics="true" pageId="8" pageNumber="119">M. jucuensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FA82FB42FF39FB18" authorityName="Costa, Katz, Mattos &amp; Rangel-Pereira" authorityYear="2019" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="mucuriensis">
<emphasis id="B94F3616ED61FFB4FA82FB42FF39FB18" italics="true" pageId="8" pageNumber="119">M. mucuriensis</emphasis>
</taxonomicName>
(5% to 10%). In the sarcoglanidines 
<taxonomicName id="4C3B9187ED61FFB4FD56FB26FB64FB19" authority="Myers &amp; Weitzman 1966" authorityName="Myers &amp; Weitzman" authorityYear="1966" box="[670,1185,1231,1257]" class="Actinopterygii" family="Trichomycteridae" genus="Sarcoglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="simplex">
<emphasis id="B94F3616ED61FFB4FD56FB26FCBAFB19" box="[670,895,1231,1257]" italics="true" pageId="8" pageNumber="119">Sarcoglanis simplex</emphasis>
<bibRefCitation id="EFAA97F5ED61FFB4FC4EFB26FB64FB19" author="Myers, G. S. &amp; Weitzman, S. H." box="[902,1185,1231,1257]" pageId="8" pageNumber="119" pagination="277 - 287" refId="ref9021" refString="Myers, G. S. &amp; Weitzman, S. H. (1966) Two remarkable new trichomycterid catfishes from the Amazon basin in Brazil and Colombia. Journal of Zoology, 149 (3), 277 - 287. https: // doi. org / 10.1111 / j. 1469 - 7998.1966. tb 04049. x" type="journal article" year="1966">Myers &amp; Weitzman 1966</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FB1FFB26FF39FAFD" authorityName="Myers &amp; Weitzman" authorityYear="1966" class="Actinopterygii" family="Trichomycteridae" genus="Malacoglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="gelatinosus">
<emphasis id="B94F3616ED61FFB4FB1FFB26FF39FAFD" italics="true" pageId="8" pageNumber="119">Malacoglanis gelatinosus</emphasis>
</taxonomicName>
, the trichomycterine 
<taxonomicName id="4C3B9187ED61FFB4FE23FB1AFCF3FAFD" authority="Eigenmann 1917" authorityName="Eigenmann" authorityYear="1917" box="[491,822,1267,1293]" class="Actinopterygii" family="Trichomycteridae" genus="Scleronema" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED61FFB4FE23FB1AFDABFAFD" box="[491,622,1267,1293]" italics="true" pageId="8" pageNumber="119">Scleronema</emphasis>
Eigenmann 1917
</taxonomicName>
, and in the stegophiline 
<taxonomicName id="4C3B9187ED61FFB4FB8DFB1AFA59FAFD" baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1912" box="[1093,1436,1267,1293]" class="Actinopterygii" family="Trichomycteridae" genus="Ochmacanthus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="batrachostoma">
<emphasis id="B94F3616ED61FFB4FB8DFB1AFA59FAFD" box="[1093,1436,1267,1293]" italics="true" pageId="8" pageNumber="119">Ochmacanthus batrachostoma</emphasis>
</taxonomicName>
(
<collectingRegion id="49FF24E6ED61FFB4FF57FAFEFEC5FAC1" box="[159,256,1303,1329]" country="Venezuela" name="Miranda" pageId="8" pageNumber="119">Miranda</collectingRegion>
Ribeiro 1912) a small reduction in the size of the first pectoral filament is observed (
<bibRefCitation id="EFAA97F5ED61FFB4FB76FAFEFF17FAA5" author="Myers, G. S. &amp; Weitzman, S. H." pageId="8" pageNumber="119" pagination="277 - 287" refId="ref9021" refString="Myers, G. S. &amp; Weitzman, S. H. (1966) Two remarkable new trichomycterid catfishes from the Amazon basin in Brazil and Colombia. Journal of Zoology, 149 (3), 277 - 287. https: // doi. org / 10.1111 / j. 1469 - 7998.1966. tb 04049. x" type="journal article" year="1966">Myers &amp; Weitzman 1966</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FF15FAD2FE1CFAA5" author="Neto, S. C. &amp; de Pinna, M. C. C." box="[221,473,1339,1365]" pageId="8" pageNumber="119" pagination="1" refId="ref9083" refString="Neto, S. C. &amp; de Pinna, M. C. C. (2016) Redescription of Ochmacanthus batrachostoma (Miranda-Ribeiro, 1912) (Siluriformes: Trichomycteridae): a possible case of incipient paedomorphism. Neotropical Ichthyology, 14 (1). [published online] https: // doi. org / 10.1590 / 1982 - 0224 - 20150030" type="journal article" year="2016">Neto &amp; de Pinna 2016</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FE2CFAD2FCC3FAA5" author="Ferrer, J. &amp; Malabarba, L. R." box="[484,774,1339,1365]" pageId="8" pageNumber="119" pagination="2" refId="ref8421" refString="Ferrer, J. &amp; Malabarba, L. R. (2020) Systematic revision of the Neotropical catfish genus Scleronema (Siluriformes: Trichomycteridae), with descriptions of six new species from Pampa grasslands. Neotropical Ichthyology, 18 (2). [published online] https: // doi. org / 10.1590 / 1982 - 0224 - 2019 - 0081" type="journal article" year="2020">Ferrer &amp; Malabarba 2020</bibRefCitation>
); however this reduction is not comparable to that observed in 
<taxonomicName id="4C3B9187ED61FFB4FF7CFAB6FEF6FA89" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[180,307,1375,1401]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED61FFB4FF7CFAB6FEF6FA89" box="[180,307,1375,1401]" italics="true" pageId="8" pageNumber="119">M. ribeirae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FEA2FAB6FE2BFA89" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[362,494,1375,1401]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED61FFB4FEA2FAB6FE2BFA89" box="[362,494,1375,1401]" italics="true" pageId="8" pageNumber="119">M. bendego</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84EA04ED61FFB4FF0FFA6AFCEBF94D" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="119">
Several psammophilic species display a reduction in integumentary pigmentation, as observed in the sarcoglanidines 
<taxonomicName id="4C3B9187ED61FFB4FF2BFA4EFEBDFA30" authorityName="de Pinna &amp; Starnes" authorityYear="1990" box="[227,376,1447,1473]" class="Actinopterygii" family="Trichomycteridae" genus="Stenolicmus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="sarmientoi">
<emphasis id="B94F3616ED61FFB4FF2BFA4EFEBDFA30" box="[227,376,1447,1473]" italics="true" pageId="8" pageNumber="119">S. sarmientoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FE40FA4EFCFCFA31" authority="de Pinna 1989" authorityName="de Pinna" authorityYear="1989" box="[392,825,1447,1473]" class="Actinopterygii" family="Trichomycteridae" genus="Stauroglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="gouldingi">
<emphasis id="B94F3616ED61FFB4FE40FA4EFD4FFA31" box="[392,650,1447,1473]" italics="true" pageId="8" pageNumber="119">Stauroglanis gouldingi</emphasis>
<bibRefCitation id="EFAA97F5ED61FFB4FD5BFA4EFCFCFA31" author="de Pinna, M. C. C." box="[659,825,1447,1473]" pageId="8" pageNumber="119" pagination="1 - 39" refId="ref9263" refString="de Pinna, M. C. C. (1989) A new sarcoglanidine catfish, phylogeny of its subfamily, and an appraisal of the phyletic status of the Trichomycterinae (Teleostei, Trichomycteridae). American Museum novitates, 2950, 1 - 39." type="journal article" year="1989">de Pinna 1989</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED61FFB4FC8EFA4EFC28FA31" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[838,1005,1447,1473]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED61FFB4FC8EFA4EFC28FA31" box="[838,1005,1447,1473]" italics="true" pageId="8" pageNumber="119">Microcambeva</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FBEFFA4EFAF9FA31" authority="Costa 1994" authorityName="Costa" authorityYear="1994" box="[1063,1340,1447,1473]" class="Actinopterygii" family="Trichomycteridae" genus="Ammoglanis" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED61FFB4FBEFFA4EFB76FA31" box="[1063,1203,1447,1473]" italics="true" pageId="8" pageNumber="119">Ammoglanis</emphasis>
<bibRefCitation id="EFAA97F5ED61FFB4FB73FA4EFAF9FA31" author="Costa, W. J. E. M. &amp; Bockmann, F. A." box="[1211,1340,1447,1473]" pageId="8" pageNumber="119" pagination="715 - 730" refId="ref7972" refString="Costa, W. J. E. M. &amp; Bockmann, F. A. (1994) A new genus and species of Sarcoglanidinae (Siluriformes: Trichomycteridae) from southeastern Brazil, with a re-examination of subfamilial phylogeny. Journal of Natural History, 28, 715 - 730. https: // doi. org / 10.1080 / 00222939400770331" type="journal article" year="1994">Costa 1994</bibRefCitation>
</taxonomicName>
species. Species of 
<taxonomicName id="4C3B9187ED61FFB4FEDAFA22FE50FA15" authorityName="Eigenmann" authorityYear="1917" box="[274,405,1483,1509]" class="Actinopterygii" family="Trichomycteridae" genus="Scleronema" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED61FFB4FEDAFA22FE50FA15" box="[274,405,1483,1509]" italics="true" pageId="8" pageNumber="119">Scleronema</emphasis>
</taxonomicName>
, have a yellowish background and patterns composed of rounded brown blotches (
<bibRefCitation id="EFAA97F5ED61FFB4FAFBFA22FF17F9F9" author="de Pinna, M. C. C." pageId="8" pageNumber="119" pagination="1 - 39" refId="ref9263" refString="de Pinna, M. C. C. (1989) A new sarcoglanidine catfish, phylogeny of its subfamily, and an appraisal of the phyletic status of the Trichomycterinae (Teleostei, Trichomycteridae). American Museum novitates, 2950, 1 - 39." type="journal article" year="1989">de Pinna, 1989</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FF16FA06FDE6F9F9" author="de Pinna, M. C. C. &amp; Winemiller, K. O." box="[222,547,1519,1545]" pageId="8" pageNumber="119" pagination="255 - 264" refId="ref9311" refString="de Pinna, M. C. C. &amp; Winemiller, K. O. (2000) A new species of Ammoglanis (Siluriformes: Trichomycteridae) from Venezuela. Ichthyological Exploration of Freshwaters, 11 (3), 255 - 264." type="journal article" year="2000">de Pinna &amp; Winemiller 2000</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED61FFB4FDE6FA06FC94F9F9" author="Ferrer, J. &amp; Malabarba, L. R." box="[558,849,1519,1545]" pageId="8" pageNumber="119" pagination="2" refId="ref8421" refString="Ferrer, J. &amp; Malabarba, L. R. (2020) Systematic revision of the Neotropical catfish genus Scleronema (Siluriformes: Trichomycteridae), with descriptions of six new species from Pampa grasslands. Neotropical Ichthyology, 18 (2). [published online] https: // doi. org / 10.1590 / 1982 - 0224 - 2019 - 0081" type="journal article" year="2020">Ferrer &amp; Malabarba 2020</bibRefCitation>
). A unique pattern of color of the body, composed of a yellowish background with larger dark spaced spots is observed in 
<emphasis id="B94F3616ED61FFB4FC58F9FAFBF7F9DC" box="[912,1074,1555,1581]" italics="true" pageId="8" pageNumber="119">
<taxonomicName id="4C3B9187ED61FFB4FC58F9FAFBD2F9DD" authorityName="de Pinna &amp; Starnes" authorityYear="1990" box="[912,1047,1555,1581]" class="Actinopterygii" family="Trichomycteridae" genus="Stenolicmus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">Stenolicmus</taxonomicName>
ix
</emphasis>
Wosiacki, Coutinho &amp; Montang 2011, 
<taxonomicName id="4C3B9187ED61FFB4FF15F9DEFE98F9A1" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[221,349,1591,1617]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED61FFB4FF15F9DEFE98F9A1" box="[221,349,1591,1617]" italics="true" pageId="8" pageNumber="119">M. ribeirae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FE5EF9DEFDDEF9A1" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[406,539,1591,1617]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED61FFB4FE5EF9DEFDDEF9A1" box="[406,539,1591,1617]" italics="true" pageId="8" pageNumber="119">M. bendego</emphasis>
</taxonomicName>
, however in the Amazon species, the spots are distributed irregularly along the middle of the body, while in Atlantic forest species, the spots form a series along the middle of the body. These shared characteristics suggest that 
<taxonomicName id="4C3B9187ED61FFB4FDEAF996FD6CF969" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[546,681,1663,1689]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED61FFB4FDEAF996FD6CF969" box="[546,681,1663,1689]" italics="true" pageId="8" pageNumber="119">M. bendego</emphasis>
</taxonomicName>
is more closely related to 
<taxonomicName id="4C3B9187ED61FFB4FC14F996FB98F969" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[988,1117,1663,1689]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED61FFB4FC14F996FB98F969" box="[988,1117,1663,1689]" italics="true" pageId="8" pageNumber="119">M. ribeirae</emphasis>
</taxonomicName>
from the Ribeira de Iguape ecoregion than to species in its own Fluminense ecoregion.
</paragraph>
<paragraph id="8B84EA04ED61FFB5FF0FF92EFB66FE95" blockId="8.[151,1437,150,2013]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="120" pageId="8" pageNumber="119">
The apparent sister-group relationship between 
<taxonomicName id="4C3B9187ED61FFB4FD2FF92EFCABF911" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[743,878,1735,1761]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED61FFB4FD2FF92EFD3BF910" box="[743,766,1735,1760]" italics="true" pageId="8" pageNumber="119">M</emphasis>
. 
<emphasis id="B94F3616ED61FFB4FCC7F92EFCABF911" box="[783,878,1735,1761]" italics="true" pageId="8" pageNumber="119">bendego</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED61FFB4FC60F92EFBEFF911" authorityName="Costa, Lima &amp; Bizerril" authorityYear="2004" box="[936,1066,1735,1761]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED61FFB4FC60F92EFBEFF911" box="[936,1066,1735,1761]" italics="true" pageId="8" pageNumber="119">M. ribeirae</emphasis>
</taxonomicName>
is surprising given that the new species is geographically closer to 
<taxonomicName id="4C3B9187ED61FFB4FDE1F902FD6FF8F5" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[553,682,1771,1797]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="species" species="barbata">
<emphasis id="B94F3616ED61FFB4FDE1F902FD6FF8F5" box="[553,682,1771,1797]" italics="true" pageId="8" pageNumber="119">M. barbata</emphasis>
</taxonomicName>
, with the two 
<typeStatus id="548054A6ED61FFB4FC9CF902FC41F8F5" box="[852,900,1771,1797]" pageId="8" pageNumber="119">type</typeStatus>
localities separated by mere 
<quantity id="4CC347E1ED61FFB4FB1EF903FAE5F8F5" box="[1238,1312,1770,1797]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="119" unit="km" value="35.0">35 km</quantity>
in straight line. Geomorphological data, however, lend support to this biogeographic pattern. The Guapi-Macacu and S„o Jo„o drainages are separated by the Serra do Sembê, which extends from the Serra dos Órg„os Mountains to the Cabo Frio Magmatic Lineament (CFML) (
<bibRefCitation id="EFAA97F5ED61FFB4FDE6F8BEFCE0F881" author="Riccomini, C. &amp; Velazquez, V. F. &amp; Gomes, C. B." box="[558,805,1878,1905]" pageId="8" pageNumber="119" refId="ref9583" refString="Riccomini, C., Velazquez, V. F. &amp; Gomes, C. B. (2005) Tectonic controls of the Mesozoic and Cenozoic alkaline magmatism in central-southeastern Brazilian platform. In: Comin-Chiaramonti, P. &amp; Gomes, C. B. (Eds.), Mesozoic to Cenozoic Alkaline Magmatism in the Brazilian PlatformMesozoic to Cenozoic Alkaline Magmatism in the Brazilian Platform. Editora da Universidade de S &quot; o Paulo, S &quot; o Paulo, 707 pp." type="book" year="2005">
Riccomini 
<emphasis id="B94F3616ED61FFB4FD63F8BEFD27F881" box="[683,738,1879,1905]" italics="true" pageId="8" pageNumber="119">et al.</emphasis>
2005
</bibRefCitation>
). These high and ancient rocky escarpments act as a geographical barrier, which has influenced the vicariant patterns and the high diversity of freshwater fish species in the Fluminense ecoregion (
<bibRefCitation id="EFAA97F5ED61FFB4FE57F876FDA4F849" author="Lima, S. M. Q. &amp; Berbel-Filho, W. M. &amp; Araujo, T. F. &amp; Lazzarotto, H. &amp; Tatarenkov, A. &amp; Avise, J. C." box="[415,609,1951,1977]" pageId="8" pageNumber="119" pagination="1 - 8" refId="ref8792" refString="Lima, S. M. Q., Berbel-Filho, W. M., Araujo, T. F., Lazzarotto, H., Tatarenkov, A. &amp; Avise, J. C. (2017) Headwater capture evidenced by paleo-rivers reconstruction and population genetic structure of the armored catfish (Pareiorhaphis garbei) in the Serra do Mar mountains of southeastern Brazil. Frontiers in genetics, 8 (199), 1 - 8. https: // doi. org / 10.3389 / fgene. 2017.00199" type="journal article" year="2017">
Lima 
<emphasis id="B94F3616ED61FFB4FE2CF876FDD8F849" box="[484,541,1951,1977]" italics="true" pageId="8" pageNumber="119">et al.</emphasis>
2017
</bibRefCitation>
). This is reflected in some taxa with different species on each side of this divide, such as the 
<taxonomicName id="4C3B9187ED61FFB4FEB9F82AFD56F82D" authority="Costa 2008" authorityName="Costa" authorityYear="2008" box="[369,659,1987,2013]" class="Actinopterygii" family="Rivulidae" genus="Atlantirivulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED61FFB4FEB9F82AFDCEF82D" box="[369,523,1987,2013]" italics="true" pageId="8" pageNumber="119">Atlantirivulus</emphasis>
Costa 2008
</taxonomicName>
rivulids, and 
<taxonomicName id="4C3B9187ED61FFB4FCF9F82AFAFFF82D" authority="Eigenmann &amp; Eigenmann 1889" authorityName="Eigenmann &amp; Eigenmann" authorityYear="1889" box="[817,1338,1987,2013]" class="Actinopterygii" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="8" pageNumber="119" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED61FFB4FCF9F82AFC08F82D" box="[817,973,1987,2013]" italics="true" pageId="8" pageNumber="119">Parotocinclus</emphasis>
Eigenmann &amp; Eigenmann 1889
</taxonomicName>
loricari-ids, and 
<taxonomicName id="4C3B9187ED60FFB5FF3EFF7EFE97FF40" authorityName="de Pinna" authorityYear="1988" box="[246,338,151,176]" class="Actinopterygii" family="Trichomycteridae" genus="Listrura" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED60FFB5FF3EFF7EFE97FF40" box="[246,338,151,176]" italics="true" pageId="9" pageNumber="120">Listrura</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3B9187ED60FFB5FE40FF7EFC86FF41" authority="Eigenmann &amp; Ward 1907" authorityName="Eigenmann &amp; Ward" authorityYear="1907" box="[392,835,151,177]" class="Actinopterygii" family="Trichomycteridae" genus="Homodiaetus" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="genus">
<emphasis id="B94F3616ED60FFB5FE40FF7EFDD8FF41" box="[392,541,151,177]" italics="true" pageId="9" pageNumber="120">Homodiaetus</emphasis>
Eigenmann &amp; Ward 1907
</taxonomicName>
trichomycterids (
<bibRefCitation id="EFAA97F5ED60FFB5FBC1FF7EFB4FFF41" author="Koch, W. R." box="[1033,1162,151,177]" pageId="9" pageNumber="120" pagination="33 - 46" refId="ref8671" refString="Koch, W. R. (2002) Revis &quot; o taxonomica do genero Homodiaetus (Teleostei, Siluriformes, Trichomycteridae). Iheringia. Serie Zoologia, 92 (3), 33 - 46. https: // doi. org / 10.1590 / S 0073 - 47212002000300004" type="journal article" year="2002">Koch 2002</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FB5DFF7EFA5DFF41" author="Villa-Verde, L. &amp; Lazzarotto, H. &amp; Lima, S. M. Q." box="[1173,1432,151,177]" pageId="9" pageNumber="120" pagination="527 - 538" refId="ref10015" refString="Villa-Verde, L., Lazzarotto, H. &amp; Lima, S. M. Q. (2012) A new glanapterygine catfish of the genus Listrura (Siluriformes: Trichomycteridae) from southeastern Brazil, corroborated by morphological and molecular data. Neotropical Ichthyology, 10, 527 - 538. https: // doi. org / 10.1590 / S 1679 - 62252012000300005" type="journal article" year="2012">
Villa-Verde 
<emphasis id="B94F3616ED60FFB5FAD5FF7EFA93FF41" box="[1309,1366,151,177]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2012
</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FF5FFF52FE97FF25" author="Roxo, F. F. &amp; Melo, B. F. &amp; Silva, G. S. C. &amp; Oliveira, C." box="[151,338,187,213]" pageId="9" pageNumber="120" pagination="260 - 270" refId="ref9671" refString="Roxo, F. F., Melo, B. F., Silva, G. S. C. &amp; Oliveira, C. (2017) New species of Parotocinclus (Siluriformes: Loricariidae) from coastal drainages of Rio de Janeiro, southeastern Brazil. Zootaxa, 4232 (2), 260 - 270. https: // doi. org / 10.11646 / zootaxa. 4232.2.9" type="journal article" year="2017">
Roxo 
<emphasis id="B94F3616ED60FFB5FF12FF52FED4FF25" box="[218,273,187,213]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2017
</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FE95FF52FDDCFF25" author="Lima, S. M. Q. &amp; Berbel-Filho, W. M. &amp; Araujo, T. F. &amp; Lazzarotto, H. &amp; Tatarenkov, A. &amp; Avise, J. C." box="[349,537,187,213]" pageId="9" pageNumber="120" pagination="1 - 8" refId="ref8792" refString="Lima, S. M. Q., Berbel-Filho, W. M., Araujo, T. F., Lazzarotto, H., Tatarenkov, A. &amp; Avise, J. C. (2017) Headwater capture evidenced by paleo-rivers reconstruction and population genetic structure of the armored catfish (Pareiorhaphis garbei) in the Serra do Mar mountains of southeastern Brazil. Frontiers in genetics, 8 (199), 1 - 8. https: // doi. org / 10.3389 / fgene. 2017.00199" type="journal article" year="2017">
Lima 
<emphasis id="B94F3616ED60FFB5FE57FF52FE13FF25" box="[415,470,187,213]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2017
</bibRefCitation>
). In addition, 
<bibRefCitation id="EFAA97F5ED60FFB5FD79FF52FC4FFF25" author="Santos, U., V &amp; Belei, F. A. &amp; Cioffi, M. B. &amp; Bertollo, L. A. C. &amp; Paiva, S. R. &amp; Dergam, J. A." box="[689,906,187,213]" pageId="9" pageNumber="120" pagination="2326 - 2343" refId="ref9744" refString="Santos, U., V ˆ lcker, C. M., Belei, F. A., Cioffi, M. B., Bertollo, L. A. C., Paiva, S. R. &amp; Dergam, J. A. (2009) Molecular and karyotypic phylogeography in the Neotropical Hoplias malabaricus (Erythrinidae) fish in eastern Brazil. Journal of Fish Biology, 75, 2326 - 2343. https: // doi. org / 10.1111 / j. 1095 - 8649.2009.02489. x" type="journal article" year="2009">
Santos 
<emphasis id="B94F3616ED60FFB5FCCAFF52FCFCFF25" box="[770,825,187,213]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
(2009)
</bibRefCitation>
and 
<bibRefCitation id="EFAA97F5ED60FFB5FC76FF52FB59FF25" author="Pereira, T. L. &amp; Santos, U. &amp; Schaefer, C. E. &amp; Souza, G. O. &amp; Paiva, S. R. &amp; Malabarba, L. R. &amp; Schmidt, E. E. &amp; Dergam, J. A." box="[958,1180,187,213]" pageId="9" pageNumber="120" pagination="905 - 914" refId="ref9482" refString="Pereira, T. L., Santos, U., Schaefer, C. E., Souza, G. O., Paiva, S. R., Malabarba, L. R., Schmidt, E. E. &amp; Dergam, J. A. (2013) Dispersal and vicariance of Hoplias malabaricus (Bloch, 1794) (Teleostei, Erythrinidae) populations of the Brazilian continental margin. Journal of Biogeography, 40, 905 - 914. https: // doi. org / 10.1111 / jbi. 12044" type="journal article" year="2013">
Pereira 
<emphasis id="B94F3616ED60FFB5FBDCFF52FB8EFF25" box="[1044,1099,187,213]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
(2013)
</bibRefCitation>
also demonstrated that 
<taxonomicName id="4C3B9187ED60FFB5FF5FFF36FDDAFF09" authority="(Bloch 1794)" baseAuthorityName="Bloch" baseAuthorityYear="1794" box="[151,543,223,249]" class="Actinopterygii" family="Erythrinidae" genus="Hoplias" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="malabaricus">
<emphasis id="B94F3616ED60FFB5FF5FFF36FE46FF09" box="[151,387,223,249]" italics="true" pageId="9" pageNumber="120">Hoplias malabaricus</emphasis>
(Bloch 1794)
</taxonomicName>
lineages’ from Guanabara Bay drainages are more closely related to those from the distant Paranaguá Bay, in 
<collectingRegion id="49FF24E6ED60FFB5FE16FEEAFDA9FEED" box="[478,620,259,285]" country="Brazil" name="Parana" pageId="9" pageNumber="120">Paraná State</collectingRegion>
, than to the rio S„o Jo„o basin, corroborating the effectiveness of this barrier for the freshwater aquatic biota (
<bibRefCitation id="EFAA97F5ED60FFB5FDFBFECEFCEEFEB1" author="Riccomini, C. &amp; Velazquez, V. F. &amp; Gomes, C. B." box="[563,811,294,321]" pageId="9" pageNumber="120" refId="ref9583" refString="Riccomini, C., Velazquez, V. F. &amp; Gomes, C. B. (2005) Tectonic controls of the Mesozoic and Cenozoic alkaline magmatism in central-southeastern Brazilian platform. In: Comin-Chiaramonti, P. &amp; Gomes, C. B. (Eds.), Mesozoic to Cenozoic Alkaline Magmatism in the Brazilian PlatformMesozoic to Cenozoic Alkaline Magmatism in the Brazilian Platform. Editora da Universidade de S &quot; o Paulo, S &quot; o Paulo, 707 pp." type="book" year="2005">
Riccomini 
<emphasis id="B94F3616ED60FFB5FD79FECEFD27FEB1" box="[689,738,295,321]" italics="true" pageId="9" pageNumber="120">et al</emphasis>
. 2005
</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FCF0FECEFC33FEB1" author="Lima, S. M. Q. &amp; Berbel-Filho, W. M. &amp; Araujo, T. F. &amp; Lazzarotto, H. &amp; Tatarenkov, A. &amp; Avise, J. C." box="[824,1014,295,321]" pageId="9" pageNumber="120" pagination="1 - 8" refId="ref8792" refString="Lima, S. M. Q., Berbel-Filho, W. M., Araujo, T. F., Lazzarotto, H., Tatarenkov, A. &amp; Avise, J. C. (2017) Headwater capture evidenced by paleo-rivers reconstruction and population genetic structure of the armored catfish (Pareiorhaphis garbei) in the Serra do Mar mountains of southeastern Brazil. Frontiers in genetics, 8 (199), 1 - 8. https: // doi. org / 10.3389 / fgene. 2017.00199" type="journal article" year="2017">
Lima 
<emphasis id="B94F3616ED60FFB5FCB2FECEFC77FEB1" box="[890,946,295,321]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2017
</bibRefCitation>
). Thus, the apparently geographically counter-intuitive relationships of 
<taxonomicName id="4C3B9187ED60FFB5FDC5FEA2FD54FE95" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[525,657,331,357]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED60FFB5FDC5FEA2FD54FE95" box="[525,657,331,357]" italics="true" pageId="9" pageNumber="120">M. bendego</emphasis>
</taxonomicName>
actually fits a broader biogeographical pattern.
</paragraph>
<paragraph id="8B84EA04ED60FFB5FF0FFE86FE67FD3D" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="120">
It is not ideal to describe a species based on few specimens, because obviously there is not enough material to address such basic questions as ontogenetic or intraspecific variation (
<bibRefCitation id="EFAA97F5ED60FFB5FC55FE7AFBB8FE5D" author="de Pinna, M. C. C. &amp; Reis, V. &amp; Britski, H." box="[925,1149,403,429]" pageId="9" pageNumber="120" pagination="1 - 27" refId="ref9412" refString="de Pinna, M. C. C., Reis, V. &amp; Britski, H. (2020) A new species of Trichogenes (Siluriformes, Trichomycteridae), with a discussion on the homologies of the anterior orbital bones in trichomycterids and other loricarioids. American Museum Novitates, 3951, 1 - 27. https: // doi. org / 10.1206 / 3951.1" type="journal article" year="2020">
de Pinna 
<emphasis id="B94F3616ED60FFB5FBCCFE7AFBFEFE5D" box="[1028,1083,403,429]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2020
</bibRefCitation>
). Still, some important recent discoveries on trichomycterid diversity were based on few individuals, e.g: 
<taxonomicName id="4C3B9187ED60FFB5FBD1FE5EFB52FE21" authorityName="Costa &amp; Bockmann" authorityYear="1994" box="[1049,1175,439,465]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="barbata">
<emphasis id="B94F3616ED60FFB5FBD1FE5EFBF5FE20" box="[1049,1072,439,464]" italics="true" pageId="9" pageNumber="120">M</emphasis>
. 
<emphasis id="B94F3616ED60FFB5FBF6FE5EFB52FE21" box="[1086,1175,439,465]" italics="true" pageId="9" pageNumber="120">barbata</emphasis>
</taxonomicName>
(n=4), 
<taxonomicName id="4C3B9187ED60FFB5FB22FE5EFA95FE21" authorityName="Mattos &amp; Lima" authorityYear="2010" box="[1258,1360,439,465]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="draco">
<emphasis id="B94F3616ED60FFB5FB22FE5EFA95FE21" box="[1258,1360,439,465]" italics="true" pageId="9" pageNumber="120">M. draco</emphasis>
</taxonomicName>
(n=2), 
<taxonomicName id="4C3B9187ED60FFB5FF5FFE32FD50FE05" authority="de Pinna &amp; Wosiacki 2002" authorityName="de Pinna &amp; Wosiacki" authorityYear="2002" box="[151,661,475,501]" class="Actinopterygii" family="Trichomycteridae" genus="Listrura" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="boticario">
<emphasis id="B94F3616ED60FFB5FF5FFE32FEA4FE05" box="[151,353,475,501]" italics="true" pageId="9" pageNumber="120">Listrura boticario</emphasis>
<bibRefCitation id="EFAA97F5ED60FFB5FEA0FE32FD50FE05" author="de Pinna, M. C. C. &amp; Wosiacki, W. B." box="[360,661,475,501]" pageId="9" pageNumber="120" pagination="720 - 726" refId="ref9357" refString="de Pinna, M. C. C. &amp; Wosiacki, W. B. (2002) A new interstitial catfish of the genus Listrura from southern Brazil (Siluriformes: Trichomycteridae: Glanapteryginae). Proceedings of the Biological Society of Washington, 115 (4), 720 - 726." type="journal article" year="2002">de Pinna &amp; Wosiacki 2002</bibRefCitation>
</taxonomicName>
(n=1), 
<taxonomicName id="4C3B9187ED60FFB5FD21FE32FAE5FE05" authority="Villa-Verde, Ferrer &amp; Malabarba 2014" authorityName="Villa-Verde, Ferrer &amp; Malabarba" authorityYear="2014" box="[745,1312,475,501]" class="Actinopterygii" family="Trichomycteridae" genus="Listrura" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="depinnai">
<emphasis id="B94F3616ED60FFB5FD21FE32FD3CFE04" box="[745,761,475,500]" italics="true" pageId="9" pageNumber="120">L</emphasis>
. 
<emphasis id="B94F3616ED60FFB5FCCFFE32FCACFE05" box="[775,873,475,501]" italics="true" pageId="9" pageNumber="120">depinnai</emphasis>
<bibRefCitation id="EFAA97F5ED60FFB5FCB8FE32FAE5FE05" author="Villa-Verde, L. &amp; Ferrer, J. &amp; Malabarba, L. R." box="[880,1312,475,501]" pageId="9" pageNumber="120" pagination="641 - 646" refId="ref10082" refString="Villa-Verde, L., Ferrer, J. &amp; Malabarba, L. R. (2014) A new species of Listrura from Laguna dos Patos System, Brazil: the southernmost record of the Glanapteryginae (Siluriformes: Trichomycteridae). Copeia, 2013 (4), 641 - 646. https: // doi. org / 10.1643 / CI- 13 - 027" type="journal article" year="2014">Villa-Verde, Ferrer &amp; Malabarba 2014</bibRefCitation>
</taxonomicName>
(n=2), and 
<taxonomicName id="4C3B9187ED60FFB5FF5FFE16FD16FDE9" authority="de Pinna, Reis &amp; Britski 2020" authorityName="de Pinna, Reis &amp; Britski" authorityYear="2020" box="[151,723,511,537]" class="Actinopterygii" family="Trichomycteridae" genus="Trichogenes" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="beagle">
<emphasis id="B94F3616ED60FFB5FF5FFE16FEB1FDE9" box="[151,372,511,537]" italics="true" pageId="9" pageNumber="120">Trichogenes beagle</emphasis>
<bibRefCitation id="EFAA97F5ED60FFB5FEB4FE16FD16FDE9" author="de Pinna, M. C. C. &amp; Reis, V. &amp; Britski, H." box="[380,723,511,537]" pageId="9" pageNumber="120" pagination="1 - 27" refId="ref9412" refString="de Pinna, M. C. C., Reis, V. &amp; Britski, H. (2020) A new species of Trichogenes (Siluriformes, Trichomycteridae), with a discussion on the homologies of the anterior orbital bones in trichomycterids and other loricarioids. American Museum Novitates, 3951, 1 - 27. https: // doi. org / 10.1206 / 3951.1" type="journal article" year="2020">de Pinna, Reis &amp; Britski 2020</bibRefCitation>
</taxonomicName>
(n=3) (
<bibRefCitation id="EFAA97F5ED60FFB5FCE2FE16FB93FDE9" author="Costa, W. J. E. M. &amp; Bockmann, F. A." box="[810,1110,511,537]" pageId="9" pageNumber="120" pagination="715 - 730" refId="ref7972" refString="Costa, W. J. E. M. &amp; Bockmann, F. A. (1994) A new genus and species of Sarcoglanidinae (Siluriformes: Trichomycteridae) from southeastern Brazil, with a re-examination of subfamilial phylogeny. Journal of Natural History, 28, 715 - 730. https: // doi. org / 10.1080 / 00222939400770331" type="journal article" year="1994">Costa &amp; Bockmann, 1994</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FBABFE16FA5DFDE9" author="de Pinna, M. C. C. &amp; Wosiacki, W. B." box="[1123,1432,511,537]" pageId="9" pageNumber="120" pagination="720 - 726" refId="ref9357" refString="de Pinna, M. C. C. &amp; Wosiacki, W. B. (2002) A new interstitial catfish of the genus Listrura from southern Brazil (Siluriformes: Trichomycteridae: Glanapteryginae). Proceedings of the Biological Society of Washington, 115 (4), 720 - 726." type="journal article" year="2002">de Pinna &amp; Wosiacki 2002</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FF5FFDCAFE40FDCD" author="Mattos, J. L. &amp; Lima, S. M. Q." box="[151,389,547,573]" pageId="9" pageNumber="120" pagination="233 - 238" refId="ref8887" refString="Mattos, J. L. &amp; Lima, S. M. Q. (2010) Microcambeva draco, a new species from north-eastern Brazil (Siluriformes: Trichomycteridae). Ichthyological Exploration of Freshwaters, 21, 233 - 238." type="journal article" year="2010">Mattos &amp; Lima 2010</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FE47FDCAFD55FDCD" author="Villa-Verde, L. &amp; Ferrer, J. &amp; Malabarba, L. R." box="[399,656,547,573]" pageId="9" pageNumber="120" pagination="641 - 646" refId="ref10082" refString="Villa-Verde, L., Ferrer, J. &amp; Malabarba, L. R. (2014) A new species of Listrura from Laguna dos Patos System, Brazil: the southernmost record of the Glanapteryginae (Siluriformes: Trichomycteridae). Copeia, 2013 (4), 641 - 646. https: // doi. org / 10.1643 / CI- 13 - 027" type="journal article" year="2014">
Villa-Verde 
<emphasis id="B94F3616ED60FFB5FDDFFDCAFD8BFDCD" box="[535,590,547,573]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2014
</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FD53FDCAFCB3FDCD" author="de Pinna, M. C. C. &amp; Reis, V. &amp; Britski, H." box="[667,886,547,573]" pageId="9" pageNumber="120" pagination="1 - 27" refId="ref9412" refString="de Pinna, M. C. C., Reis, V. &amp; Britski, H. (2020) A new species of Trichogenes (Siluriformes, Trichomycteridae), with a discussion on the homologies of the anterior orbital bones in trichomycterids and other loricarioids. American Museum Novitates, 3951, 1 - 27. https: // doi. org / 10.1206 / 3951.1" type="journal article" year="2020">de Pinna et al. 2020</bibRefCitation>
). The realities of this kind of research sometimes make it imperative to release new taxonomic knowledge on elements of biodiversity even with limited material. Species must be made available for phylogenetic, biogeographic, ecological and conservation strategies (
<bibRefCitation id="EFAA97F5ED60FFB5FA8CFD82FF33FD59" author="Silva, P. C. &amp; Malabarba, M. C. &amp; Malabarba, L. R." pageId="9" pageNumber="120" pagination="110 - 120" refId="ref9942" refString="Silva, P. C., Malabarba, M. C. &amp; Malabarba, L. R. (2019) Integrative taxonomy: Morphology and ancient DNA barcoding reveals the true identity of Astyanax taeniatus, a tetra collected by Charles Darwin during the Beagle's voyage. Zoologischer Anzeiger, 278, 110 - 120. https: // doi. org / 10.1016 / j. jcz. 2018.12.007" type="journal article" year="2019">
Silva 
<emphasis id="B94F3616ED60FFB5FA40FD82FF71FD59" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2019
</bibRefCitation>
; 
<bibRefCitation id="EFAA97F5ED60FFB5FECAFD66FE20FD59" author="de Pinna, M. C. C. &amp; Reis, V. &amp; Britski, H." box="[258,485,655,681]" pageId="9" pageNumber="120" pagination="1 - 27" refId="ref9412" refString="de Pinna, M. C. C., Reis, V. &amp; Britski, H. (2020) A new species of Trichogenes (Siluriformes, Trichomycteridae), with a discussion on the homologies of the anterior orbital bones in trichomycterids and other loricarioids. American Museum Novitates, 3951, 1 - 27. https: // doi. org / 10.1206 / 3951.1" type="journal article" year="2020">
de Pinna 
<emphasis id="B94F3616ED60FFB5FEA2FD66FE66FD59" box="[362,419,655,681]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2020
</bibRefCitation>
), especially for those occurring in highly impacted areas, such as the Guanabara Bay area in the present case.
</paragraph>
<paragraph id="8B84EA04ED60FFB5FF0FFD3FFAA4FC1D" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="120">
<emphasis id="B94F3616ED60FFB5FF0FFD3FFE7EFD01" bold="true" box="[199,443,726,753]" pageId="9" pageNumber="120">Conservation status.</emphasis>
The Guanabara Bay is composed of approximately 40 rivers and streams, and the region has been sampled by many naturalists since the 19 
<superScript id="7C4E474CED60FFB5FD0FFD13FD11FCF8" attach="left" box="[711,724,762,776]" fontSize="6" pageId="9" pageNumber="120">th</superScript>
century. The 
<typeStatus id="548054A6ED60FFB5FCA3FD12FC5EFCE5" box="[875,923,763,789]" pageId="9" pageNumber="120">type</typeStatus>
locality of the new species is near large urban conglomerates and rural properties concentrating approximately 70% of the metropolitan region of 
<collectingRegion id="49FF24E6ED60FFB5FB31FCF6FF17FCAD" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="120">Rio de Janeiro State</collectingRegion>
, including a major industrial center (
<bibRefCitation id="EFAA97F5ED60FFB5FD49FCAAFC48FCAD" author="Bizerril, C. R. S. F. &amp; Primo, P. B. S." box="[641,909,835,861]" pageId="9" pageNumber="120" refId="ref7831" refString="Bizerril, C. R. S. F. &amp; Primo, P. B. S. (2001) Peixes de Aguas Interiores do Estado do Rio de Janeiro. FEMAR-SEMADS, Rio de Janeiro, 417 pp." type="book" year="2001">Bizerril &amp; Primo 2001</bibRefCitation>
). The rio Guapi-Macacu basin has an area of ca. 
<quantity id="4CC347E1ED60FFB5FF08FC8EFEE8FC73" box="[192,301,871,899]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="120" unit="km" value="1.6">1.600 km</quantity>
², making it the largest drainage flowing into the Guanabara bay. Inserted in the inner portion of the basin, with its headwaters in the Serra dos Órg„os mountain range, it is considered as a biodiversity pocket of the Guanabara Bay region. It has been heavily impacted by unrestrained urban occupation, intensified in the mid-1930, deforestation of riparian forests, fires, diversion of river courses and water pollution (
<bibRefCitation id="EFAA97F5ED60FFB5FB9AFC3AFA91FC1D" author="Bizerril, C. R. S. F. &amp; Primo, P. B. S." box="[1106,1364,979,1005]" pageId="9" pageNumber="120" refId="ref7831" refString="Bizerril, C. R. S. F. &amp; Primo, P. B. S. (2001) Peixes de Aguas Interiores do Estado do Rio de Janeiro. FEMAR-SEMADS, Rio de Janeiro, 417 pp." type="book" year="2001">Bizerril &amp; Primo 2001</bibRefCitation>
).
</paragraph>
<paragraph id="8B84EA04ED60FFB5FF0FFC1EFC9EFA89" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="120">
The intense degradation of the Guanabara’s Bay aquatic environments is evidenced by its numerous threatened freshwater fish species, such as the ‘endangered’ 
<taxonomicName id="4C3B9187ED60FFB5FD18FBF2FBA4FBC5" authority="Steindachner 1877" authorityName="Steindachner" authorityYear="1877" box="[720,1121,1051,1077]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="insignis">
<emphasis id="B94F3616ED60FFB5FD18FBF2FC44FBC4" box="[720,897,1051,1076]" italics="true" pageId="9" pageNumber="120">Brycon insignis</emphasis>
Steindachner 1877
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED60FFB5FBA6FBF2FEE6FBA9" authority="Myers 1924" authorityName="Myers" authorityYear="1924" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="flammeus">
<emphasis id="B94F3616ED60FFB5FBA6FBF2FA59FBC5" box="[1134,1436,1051,1077]" italics="true" pageId="9" pageNumber="120">Hyphessobrycon flammeus</emphasis>
Myers 1924
</taxonomicName>
, and 
<taxonomicName id="4C3B9187ED60FFB5FEAAFBD6FCC7FBA9" authority="Myers 1925" authorityName="Myers" authorityYear="1925" box="[354,770,1086,1113]" class="Actinopterygii" family="Characidae" genus="Spintherobolus" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="broccae">
<emphasis id="B94F3616ED60FFB5FEAAFBD6FDA8FBA9" box="[354,621,1087,1113]" italics="true" pageId="9" pageNumber="120">Spintherobolus broccae</emphasis>
Myers 1925
</taxonomicName>
, and the ‘critically endangered’ 
<taxonomicName id="4C3B9187ED60FFB5FBBDFBD6FE8DFB8D" authority="Travassos 1944" authorityName="Travassos" authorityYear="1944" class="Actinopterygii" family="Crenuchidae" genus="Characidium" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="grajahuense">
<emphasis id="B94F3616ED60FFB5FBBDFBD6FA59FBA9" box="[1141,1436,1087,1113]" italics="true" pageId="9" pageNumber="120">Characidium grajahuense</emphasis>
Travassos 1944
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED60FFB5FE9CFB8AFD3FFB8D" authority="de Pinna 1988" authorityName="de Pinna" authorityYear="1988" box="[340,762,1123,1149]" class="Actinopterygii" family="Trichomycteridae" genus="Listrura" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="nematopteryx">
<emphasis id="B94F3616ED60FFB5FE9CFB8AFD8AFB8C" box="[340,591,1123,1148]" italics="true" pageId="9" pageNumber="120">Listrura nematopteryx</emphasis>
de Pinna 1988
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED60FFB5FCCDFB8AFAC8FB8D" authority="(Valenciennes 1821)" baseAuthorityName="Valenciennes" baseAuthorityYear="1821" box="[773,1293,1123,1149]" class="Actinopterygii" family="Rivulidae" genus="Kryptolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="B94F3616ED60FFB5FCCDFB8AFBDBFB8D" box="[773,1054,1123,1149]" italics="true" pageId="9" pageNumber="120">Kryptolebias brasiliensis</emphasis>
(Valenciennes 1821)
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED60FFB5FAD2FB8AFE12FB51" authority="(Ladiges 1934)" baseAuthorityName="Ladiges" baseAuthorityYear="1934" class="Actinopterygii" family="Rivulidae" genus="Leptolebias" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="marmoratus">
<emphasis id="B94F3616ED60FFB5FAD2FB8AFEE5FB50" italics="true" pageId="9" pageNumber="120">Leptolebias marmoratus</emphasis>
(Ladiges 1934)
</taxonomicName>
, 
<taxonomicName id="4C3B9187ED60FFB5FE2DFB6EFC64FB51" authority="(Myers 1942)" baseAuthorityName="Myers" baseAuthorityYear="1942" box="[485,929,1159,1185]" class="Actinopterygii" family="Cynolebiidae" genus="Leptopanchax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="splendens">
<emphasis id="B94F3616ED60FFB5FE2DFB6EFD3EFB51" box="[485,763,1159,1185]" italics="true" pageId="9" pageNumber="120">Leptopanchax splendens</emphasis>
(Myers 1942)
</taxonomicName>
, and 
<taxonomicName id="4C3B9187ED60FFB5FC28FB6EFAE3FB51" authority="(Myers, 1942)" baseAuthorityName="Myers" baseAuthorityYear="1942" box="[992,1318,1159,1185]" class="Actinopterygii" family="Cynolebiidae" genus="Leptopanchax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="opalescens">
<emphasis id="B94F3616ED60FFB5FC28FB6EFBBEFB51" box="[992,1147,1159,1185]" italics="true" pageId="9" pageNumber="120">L. opalescens</emphasis>
(Myers, 1942)
</taxonomicName>
(ICMBio, 2018). While species previously considered extinct such as 
<taxonomicName id="4C3B9187ED60FFB5FCE5FB42FC7DFB35" baseAuthorityName="Myers" baseAuthorityYear="1942" box="[813,952,1195,1221]" class="Actinopterygii" family="Cynolebiidae" genus="Leptopanchax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="splendens">
<emphasis id="B94F3616ED60FFB5FCE5FB42FC7DFB35" box="[813,952,1195,1221]" italics="true" pageId="9" pageNumber="120">L. splendens</emphasis>
</taxonomicName>
, were later rediscovered in remnant natural areas of the Guanabara Bay region (
<bibRefCitation id="EFAA97F5ED60FFB5FDF1FB26FCD7FB1A" author="Costa, W. J. E. M. &amp; Mattos, J. L. O. &amp; Amorim, P. F." box="[569,786,1231,1258]" pageId="9" pageNumber="120" pagination="345 - 347" refId="ref8203" refString="Costa, W. J. E. M., Mattos, J. L. O. &amp; Amorim, P. F. (2019 b) Rediscovery of Leptopanchax splendens (Cyprinodontiformes: Aplocheilidae): a seasonal killifish from the Atlantic Forest of south-eastern Brazil that was recently considered extinct. Journal of Fish Biology, 94, 345 - 347. https: // doi. org / 10.1111 / jfb. 13898" type="journal article" year="2019">
Costa 
<emphasis id="B94F3616ED60FFB5FD4CFB26FD7DFB19" box="[644,696,1231,1257]" italics="true" pageId="9" pageNumber="120">et al</emphasis>
. 2019b
</bibRefCitation>
), others like 
<taxonomicName id="4C3B9187ED60FFB5FC62FB26FA9BFB19" authority="Costa 2019" authorityName="Costa" authorityYear="2019" box="[938,1374,1231,1257]" class="Actinopterygii" family="Cynolebiidae" genus="Leptopanchax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyprinodontiformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="sanguineus">
<emphasis id="B94F3616ED60FFB5FC62FB26FB15FB18" box="[938,1232,1231,1257]" italics="true" pageId="9" pageNumber="120">Leptopanchax sanguineus</emphasis>
Costa 2019
</taxonomicName>
, may already be extinct (Costa 2019). Considering that 
<taxonomicName id="4C3B9187ED60FFB5FD1BFB1AFC27FAFD" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[723,994,1267,1293]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED60FFB5FD1BFB1AFC27FAFD" box="[723,994,1267,1293]" italics="true" pageId="9" pageNumber="120">Microcambeva bendego</emphasis>
</taxonomicName>
was only collected in a single locality consisting in a highly degraded habitat of well-sampled drainage, that recent efforts failed in the obtaining additional specimens, and its area is occupation of less than 
<quantity id="4CC347E1ED60FFB5FCF9FAD2FCBFFAA5" box="[817,890,1339,1365]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="120" unit="km" value="20.0">20 km</quantity>
<superScript id="7C4E474CED60FFB5FCB1FAD3FC44FAB8" attach="left" box="[889,897,1338,1352]" fontSize="6" pageId="9" pageNumber="120">2</superScript>
, the B2ab(iii) critically endangered category is recommended as an obvious preventive measure (IUCN 2012).
</paragraph>
<paragraph id="8B84EA04ED60FFB5FF0FFA6AFEDAF881" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="120">
The coastal basins of Atlantic Forest harbor many endemic and restricted-range freshwater fish species. However, the habitat loss along five centuries of colonization has left its mark, with several of those species currently threatened (
<bibRefCitation id="EFAA97F5ED60FFB5FED3FA22FE3FFA15" author="Abilhoa, V. &amp; Braga, R. R. &amp; Bornatowski, H. &amp; Vitule, J. R. S." box="[283,506,1483,1509]" pageId="9" pageNumber="120" pagination="259 - 282" refId="ref7464" refString="Abilhoa, V., Braga, R. R., Bornatowski, H. &amp; Vitule, J. R. S. (2011). Fishes of the Atlantic Rain Forest Streams: Ecological Patterns and Conservation. In: Grillo, O. &amp; Venora, G. (Eds.), Changing Diversity in Changing Environment. Intech, Rijeka, pp. 259 - 282. https: // doi. org / 10.5772 / 24540" type="book chapter" year="2011">
Abilhoa 
<emphasis id="B94F3616ED60FFB5FEB6FA22FE75FA15" box="[382,432,1483,1509]" italics="true" pageId="9" pageNumber="120">et al</emphasis>
. 2011
</bibRefCitation>
). To some of these ‘endangered’ species, the creation or expansion of conservation units encompassing small hydrographic basins and associated riparian zones in southeastern 
<collectingCountry id="F32CAA94ED60FFB5FB7FFA06FB3EF9F9" box="[1207,1275,1519,1545]" name="Brazil" pageId="9" pageNumber="120">Brazil</collectingCountry>
are important measures (
<bibRefCitation id="EFAA97F5ED60FFB5FED8F9FAFE3AF9DD" author="Menezes, N. A. &amp; Weitzman, S. H. &amp; Oyakawa, O. T. &amp; Lima, F. C. T. &amp; Castro, R. M. &amp; Weitzman, M. J." box="[272,511,1555,1581]" pageId="9" pageNumber="120" refId="ref8931" refString="Menezes, N. A., Weitzman, S. H., Oyakawa, O. T., Lima, F. C. T., Correa e Castro, R. M. &amp; Weitzman, M. J. (2007) Peixes de agua doce da Mata Atlantica: lista preliminar das especies e comentarios sobre conservacao de peixes de agua doce neotropicais. Museu de Zoologia da Universidade de S &quot; o Paulo, S &quot; o Paulo, 407 pp." type="book" year="2007">
Menezes 
<emphasis id="B94F3616ED60FFB5FEB7F9FAFE77F9DD" box="[383,434,1555,1581]" italics="true" pageId="9" pageNumber="120">et al</emphasis>
. 2007
</bibRefCitation>
). In the estuarine portion of rio Guapi-Macacu basin, there are two conservation units, the Guanabara Ecological Station (GES) between the municipalities of Itaboraí, Guapimirim, S„o Gonçalo and Magé, covering an area of 
<quantity id="4CC347E1ED60FFB5FE32F9B5FD9AF987" box="[506,607,1628,1655]" metricMagnitude="6" metricUnit="m" metricValue="1.42" pageId="9" pageNumber="120" unit="km" value="1420.0">1,420km</quantity>
², and the Guapi-Mirim Environmental Protection Area (GMEPA), covering an area of 
<quantity id="4CC347E1ED60FFB5FEF3F997FE73F96B" box="[315,438,1662,1691]" metricMagnitude="5" metricUnit="m" metricValue="1.3824999999999998" pageId="9" pageNumber="120" unit="km" value="138.25">138.25 km</quantity>
². Outside the protected areas this region is severely impacted, with the riparian zone deforested, diversion of river courses to agriculture, and the introduction of exotic species (
<bibRefCitation id="EFAA97F5ED60FFB5FB42F94AFA55F94D" author="Bizerril, C. R. S. F. &amp; Primo, P. B. S." box="[1162,1424,1699,1725]" pageId="9" pageNumber="120" refId="ref7831" refString="Bizerril, C. R. S. F. &amp; Primo, P. B. S. (2001) Peixes de Aguas Interiores do Estado do Rio de Janeiro. FEMAR-SEMADS, Rio de Janeiro, 417 pp." type="book" year="2001">Bizerril &amp; Primo 2001</bibRefCitation>
). Although the 
<typeStatus id="548054A6ED60FFB5FEFBF92EFEA6F911" box="[307,355,1735,1761]" pageId="9" pageNumber="120">type</typeStatus>
locality of 
<taxonomicName id="4C3B9187ED60FFB5FE2EF92EFDA9F911" authorityName="Medeiros &amp; Moreira &amp; Pinna &amp; Lima" authorityYear="2020" box="[486,620,1735,1761]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="bendego">
<emphasis id="B94F3616ED60FFB5FE2EF92EFDA9F911" box="[486,620,1735,1761]" italics="true" pageId="9" pageNumber="120">M. bendego</emphasis>
</taxonomicName>
is close to the buffer zone of the GES inside in GMEPA, its geographic distribution is limited to the middle section of the rio Guapiaçu. A measure that will contribute to the conservation, not only of the new species, but also of other aquatic species, is the expansion of the GES area to the middle portions of the rio Guapi-Macacu river basin to protect most freshwater habitats and their associated biota (
<bibRefCitation id="EFAA97F5ED60FFB5FB2BF8DAFED6F881" author="Azevedo-Santos, V. M. &amp; Frederico, R. &amp; Fagundes, C. K. &amp; Pompeu, P. S. &amp; Pelicice, F. M. &amp; Padial, A. A. &amp; Nogueira, M. G. &amp; Fearnside, P. M. &amp; Lima, L. B. &amp; Daga, V. S. &amp; Oliveira, F. J. &amp; Vitule, J. R. S. &amp; Callisto, M. &amp; Agostinho, A. A. &amp; Esteves, F. A. &amp; Lima-Junior, D. P. &amp; Sabino, J. &amp; Mormul, R. P. &amp; Grasel, D. &amp; Zuanon, J. &amp; Vilella, F. S. &amp; Henry, R." pageId="9" pageNumber="120" pagination="442 - 448" refId="ref7641" refString="Azevedo-Santos, V. M., Frederico, R. G, Fagundes, C. K., Pompeu, P. S., Pelicice, F. M., Padial, A. A., Nogueira, M. G., Fearnside, P. M., Lima, L. B., Daga, V. S., Oliveira, F. J., Vitule, J. R. S., Callisto, M., Agostinho, A. A., Esteves, F. A., Lima-Junior, D. P., Magalh &quot; es, A. L. B., Sabino, J., Mormul, R. P., Grasel, D., Zuanon, J., Vilella, F. S. &amp; Henry, R. (2019) Pretected areas: A focus on Brazilian freshwater biodiversity. Diversity and Distributions, 25, 442 - 448. https: // doi. org / 10.1111 / ddi. 12871" type="journal article" year="2019">
Azevedo-Santos 
<emphasis id="B94F3616ED60FFB5FF5FF8BEFF0AF881" box="[151,207,1879,1905]" italics="true" pageId="9" pageNumber="120">et al.</emphasis>
2019
</bibRefCitation>
).
</paragraph>
<paragraph id="8B84EA04ED60FFB6FF0FF893FDBDFE95" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,150,357]" lastPageId="10" lastPageNumber="121" pageId="9" pageNumber="120">
<emphasis id="B94F3616ED60FFB5FF0FF893FD5CF864" bold="true" box="[199,665,1914,1941]" pageId="9" pageNumber="120">
Comparative material. 
<collectingRegion id="49FF24E6ED60FFB5FE28F893FD51F864" box="[480,660,1914,1941]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="120">Rio de Janeiro</collectingRegion>
:
</emphasis>
<taxonomicName id="4C3B9187ED60FFB5FD69F892FBD1F865" authority=": MZUSP" authorityName="MZUSP" box="[673,1044,1915,1941]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="9" pageNumber="120" phylum="Chordata" rank="species" species="barbata">
<emphasis id="B94F3616ED60FFB5FD69F892FC6DF865" box="[673,936,1915,1941]" italics="true" pageId="9" pageNumber="120">Microcambeva barbata</emphasis>
: MZUSP
</taxonomicName>
43678, 
<quantity id="4CC347E1ED60FFB5FBB9F892FB10F865" box="[1137,1237,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="2.4699999999999998" pageId="9" pageNumber="120" unit="mm" value="24.7">24.7 mm</quantity>
SL, 
<typeStatus id="548054A6ED60FFB5FAC4F892FAB4F865" box="[1292,1393,1915,1941]" pageId="9" pageNumber="120" type="holotype">holotype</typeStatus>
, rio S„o Jo„o. MZUSP 43679, 1, 
<quantity id="4CC347E1ED60FFB5FE2FF876FD88F849" box="[487,589,1951,1977]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="9" pageNumber="120" unit="mm" value="18.7">18.7 mm</quantity>
SL, 
<typeStatus id="548054A6ED60FFB5FD4FF876FD2EF849" box="[647,747,1951,1977]" pageId="9" pageNumber="120" type="paratype">paratype</typeStatus>
. DZSJRP-Pisces 13861, 
<quantity id="4CC347E1ED60FFB5FBC0F877FBABF849" box="[1032,1134,1950,1977]" metricMagnitude="-2" metricUnit="m" metricValue="2.51" pageId="9" pageNumber="120" unit="mm" value="25.1">25.1 mm</quantity>
SL, affluent rio S„o Jo„o. MNRJ 49371, 
<quantity id="4CC347E1ED60FFB5FEF4F82AFE5AF82D" box="[316,415,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="2.27" pageId="9" pageNumber="120" unit="mm" value="22.7">22.7 mm</quantity>
SL, rio Aldeia Velha. MNRJ 47108, 
<quantity id="4CC347E1ED60FFB5FCF3F82AFC5BF82D" box="[827,926,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="9" pageNumber="120" unit="mm" value="19.6">19.6 mm</quantity>
SL, rio Aldeia Velha. MNRJ 37572, 
<quantity id="4CC347E1ED60FFB5FAF1F82AFA58F82D" box="[1337,1437,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="9" pageNumber="120" unit="mm" value="26.3">26.3 mm</quantity>
SL, rio S„o Jo„o. 
<emphasis id="B94F3616ED63FFB6FE93FF7FFE1FFF41" bold="true" box="[347,474,150,177]" pageId="10" pageNumber="121">
<collectingRegion id="49FF24E6ED63FFB6FE93FF7FFE10FF41" box="[347,469,150,177]" country="Brazil" name="Sao Paulo" pageId="10" pageNumber="121">São Paulo</collectingRegion>
:
</emphasis>
<taxonomicName id="4C3B9187ED63FFB6FE28FF7EFC97FF41" authority=": MZUSP" authorityName="MZUSP" box="[480,850,151,177]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="121" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED63FFB6FE28FF7EFD22FF41" box="[480,743,151,177]" italics="true" pageId="10" pageNumber="121">Microcambeva ribeirae</emphasis>
: MZUSP
</taxonomicName>
84301, 
<quantity id="4CC347E1ED63FFB6FC63FF7EFBCBFF41" box="[939,1038,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="4.779999999999999" pageId="10" pageNumber="121" unit="mm" value="47.8">47.8 mm</quantity>
SL, 
<typeStatus id="548054A6ED63FFB6FB8AFF7EFB62FF41" box="[1090,1191,151,177]" pageId="10" pageNumber="121" type="holotype">holotype</typeStatus>
, rio S„o Lourencinho. MZUSP 79953, 7, 
<quantity id="4CC347E1ED63FFB6FEAEFF53FDCDFF25" box="[358,520,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="3.125" metricValueMax="3.66" metricValueMin="2.59" pageId="10" pageNumber="121" unit="mm" value="31.25" valueMax="36.6" valueMin="25.9">25.9–36.6 mm</quantity>
SL, 
<typeStatus id="548054A6ED63FFB6FDF3FF52FD6CFF25" box="[571,681,187,213]" pageId="10" pageNumber="121" type="paratype">paratypes</typeStatus>
, rio Espraiado. MZUSP 74699, 10, 35.0– 
<quantity id="4CC347E1ED63FFB6FBB8FF53FB16FF25" box="[1136,1235,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" pageId="10" pageNumber="121" unit="mm" value="47.5">47.5 mm</quantity>
SL, 
<typeStatus id="548054A6ED63FFB6FACDFF52FAB6FF25" box="[1285,1395,187,213]" pageId="10" pageNumber="121" type="paratype">paratypes</typeStatus>
, rio Faú. MNRJ14304, 3, 
<quantity id="4CC347E1ED63FFB6FE4EFF37FDE2FF09" box="[390,551,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.0949999999999998" metricValueMax="3.2399999999999998" metricValueMin="2.95" pageId="10" pageNumber="121" unit="mm" value="30.95" valueMax="32.4" valueMin="29.5">29.5–32.4 mm</quantity>
SL, 
<typeStatus id="548054A6ED63FFB6FD92FF36FD0DFF09" box="[602,712,223,249]" pageId="10" pageNumber="121" type="paratype">paratypes</typeStatus>
, tributary of rio S„o Lourencinho. MNRJ 37165, 
<quantity id="4CC347E1ED63FFB6FB20FF37FA8FFF09" box="[1256,1354,222,249]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" pageId="10" pageNumber="121" unit="mm" value="35.1">35.1 mm</quantity>
SL, rio Jacupiranga. 
<emphasis id="B94F3616ED63FFB6FEE3FEEBFE4EFEED" bold="true" box="[299,395,258,285]" pageId="10" pageNumber="121">
<collectingRegion id="49FF24E6ED63FFB6FEE3FEEBFE43FEED" box="[299,390,258,285]" country="Brazil" name="Parana" pageId="10" pageNumber="121">Paraná</collectingRegion>
:
</emphasis>
<taxonomicName id="4C3B9187ED63FFB6FE5AFEEAFCC7FEED" authority=": UFRGS" authorityName="UFRGS" box="[402,770,259,285]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="121" phylum="Chordata" rank="species" species="ribeirae">
<emphasis id="B94F3616ED63FFB6FE5AFEEAFD5CFEED" box="[402,665,259,285]" italics="true" pageId="10" pageNumber="121">Microcambeva ribeirae</emphasis>
: UFRGS
</taxonomicName>
24759, 
<quantity id="4CC347E1ED63FFB6FC94FEEAFC05FEED" box="[860,960,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="10" pageNumber="121" unit="mm" value="37.7">37.7 mm</quantity>
SL, tributary of rio Guaraqueçaba. 
<emphasis id="B94F3616ED63FFB6FA85FEEBFA59FEEC" bold="true" box="[1357,1436,258,284]" pageId="10" pageNumber="121">
<collectingRegion id="49FF24E6ED63FFB6FA85FEEBFA52FEEC" box="[1357,1431,258,284]" country="Brazil" name="Bahia" pageId="10" pageNumber="121">Bahia</collectingRegion>
:
</emphasis>
<taxonomicName id="4C3B9187ED63FFB6FF5FFECEFE11FEB1" authority=": MCP" authorityName="MCP" box="[151,468,295,321]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="121" phylum="Chordata" rank="species" species="draco">
<emphasis id="B94F3616ED63FFB6FF5FFECEFE4DFEB1" box="[151,392,295,321]" italics="true" pageId="10" pageNumber="121">Microcambeva draco</emphasis>
: MCP
</taxonomicName>
17796, 2, 24.6– 
<quantity id="4CC347E1ED63FFB6FD59FECEFD3CFEB1" box="[657,761,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="10" pageNumber="121" unit="mm" value="22.6">22.6 mm</quantity>
, 
<typeStatus id="548054A6ED63FFB6FCCFFECEFCA9FEB1" box="[775,876,295,321]" pageId="10" pageNumber="121" type="holotype">holotype</typeStatus>
, rio Jucuruçu. 
<taxonomicName id="4C3B9187ED63FFB6FBDFFECEFA59FEB1" authority=": MZUSP" authorityName="MZUSP" box="[1047,1436,295,321]" class="Actinopterygii" family="Trichomycteridae" genus="Microcambeva" kingdom="Animalia" order="Siluriformes" pageId="10" pageNumber="121" phylum="Chordata" rank="species" species="jucuensis">
<emphasis id="B94F3616ED63FFB6FBDFFECEFAEAFEB0" box="[1047,1327,295,321]" italics="true" pageId="10" pageNumber="121">Microcambeva jucuensis</emphasis>
: MZUSP
</taxonomicName>
91641, 11, 
<quantity id="4CC347E1ED63FFB6FEDCFEA3FE72FE95" box="[276,439,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.205" metricValueMax="2.46" metricValueMin="1.95" pageId="10" pageNumber="121" unit="mm" value="22.05" valueMax="24.6" valueMin="19.5">19.5–24.6 mm</quantity>
SL, rio Formate.
</paragraph>
</subSubSection>
</treatment>
</document>