treatments-xml/data/08/76/87/08768726FFC69768FF78F8ACFD44DE8A.xml

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<mods:title id="5F628771DB75E88EE8E0FCA4071AEA85">A new species of Paradella Harrison &amp; Holdich, 1982 (Crustacea: Isopoda: Sphaeromatidae) from Baja California, Mexico, with a key to East Pacific species</mods:title>
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<mods:namePart id="E91B417290E98AC794988E7E6D33FD37">Wetzer, Regina</mods:namePart>
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<taxonomicName id="47DF4DB3FFC6976AFF78F8ACFD86DB51" ID-CoL="7PPKK" authority="Harrison &amp; Holdich, 1982" authorityName="Harrison &amp; Holdich" authorityYear="1982" box="[151,560,1802,1826]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B2ABEA22FFC6976AFF78F8ACFD86DB51" bold="true" box="[151,560,1802,1826]" pageId="0" pageNumber="39">
<emphasis id="B2ABEA22FFC6976AFF78F8ACFEB4DB51" bold="true" box="[151,258,1802,1826]" italics="true" pageId="0" pageNumber="39">Paradella</emphasis>
<bibRefCitation id="E44E4BC1FFC6976AFEE7F8ACFD86DB51" author="Harrison" box="[264,560,1802,1826]" pageId="0" pageNumber="48" refString="Harrison, K. &amp; Holdich, D. M. (1982) Revision of the genera Dynamenella, Ischyromene, Dynamenopsis, and Cymodocella (Crustacea: Isopoda), including a new genus and five new species of eubranchiate sphaeromatids from Queensland waters. Journal of Crustacean Biology, 2 (1), 84 - 119." type="journal article" year="1982">Harrison &amp; Holdich, 1982</bibRefCitation>
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<taxonomicName id="47DF4DB3FFC6976AFF29F8F6FDCFDB15" ID-CoL="7PPKK" authority="Harrison &amp; Holdich, 1982: 99" authorityName="Harrison &amp; Holdich" authorityPageNumber="99" authorityYear="1982" box="[198,633,1871,1895]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B2ABEA22FFC6976AFF29F8F6FE87DB14" box="[198,305,1872,1895]" italics="true" pageId="0" pageNumber="39">Paradella</emphasis>
<treatmentCitation id="017E1021FFC6976AFED8F8E9FDCFDB15" author="Harrison" box="[311,633,1871,1894]" page="99" pageId="0" pageNumber="39" year="1982">
<bibRefCitation id="E44E4BC1FFC6976AFED8F8E9FDE5DB15" author="Harrison" box="[311,595,1871,1894]" pageId="0" pageNumber="48" refString="Harrison, K. &amp; Holdich, D. M. (1982) Revision of the genera Dynamenella, Ischyromene, Dynamenopsis, and Cymodocella (Crustacea: Isopoda), including a new genus and five new species of eubranchiate sphaeromatids from Queensland waters. Journal of Crustacean Biology, 2 (1), 84 - 119." type="journal article" year="1982">Harrison &amp; Holdich, 1982</bibRefCitation>
: 99
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.– Harrison, 1984: 386.– Harrison &amp; Ellis, 1991: 943.— Kensley &amp; Schotte, 1989: 223.— 
<treatmentCitation id="017E1021FFC6976AFDDFF8D4FD5ADBFA" author="Storey" box="[560,748,1906,1929]" page="139" pageId="0" pageNumber="39" year="2002">
<bibRefCitation id="E44E4BC1FFC6976AFDDFF8D4FD0CDBFA" author="Storey" box="[560,698,1906,1929]" pageId="0" pageNumber="49" refString="Storey, M. J. (2002) New species and a new record of sphaeromatid isopods (Crustacea) from the Andaman Sea, Thailand. In: Bruce, N. L. &amp; M. Berggren (Ed.) Proceedings of the International Workshop on the Crustacea in the Andaman Sea, Phuket Marine Biological Center, 29 November - 20 December 1998, Phuket Marine Biological Center Special Publication, 23, Vol. Phuket Marine Biology Center, Phuket, pp. 133 - 148." type="journal article" year="2002">Storey, 2002</bibRefCitation>
: 139
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.— 
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<bibRefCitation id="E44E4BC1FFC6976AFCFBF8D4FBB0DBFA" author="Bruce" box="[788,1030,1906,1929]" pageId="0" pageNumber="48" refString="Bruce, N. L. &amp; Wetzer, R. (2004) Paradella tiffany sp. nov., a distinctive sphaeromatid isopod (Crustacea: Isopoda: Sphaeromatidea) from Baja California, Mexico. Zootaxa, 623, 1 - 12." type="journal article" year="2004">Bruce &amp; Wetzer, 2004</bibRefCitation>
: 2
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.
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</paragraph>
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<subSubSection id="C8C565BBFFC6976AFF78F811FB06DBA2" box="[151,1200,1975,2001]" pageId="0" pageNumber="39" type="materials_examined">
<paragraph id="80603630FFC6976AFF78F811FB06DBA2" blockId="0.[151,1200,1975,2001]" box="[151,1200,1975,2001]" pageId="0" pageNumber="39">
<typeStatus id="5F648892FFC6976AFF78F811FF67DBA2" box="[151,209,1975,2001]" pageId="0" pageNumber="39">Type</typeStatus>
species: 
<taxonomicName id="47DF4DB3FFC6976AFED3F81EFC38DBA2" authority="Harrison &amp; Holdich, 1982" authorityName="Harrison &amp; Holdich" authorityYear="1982" box="[316,910,1975,2001]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="octaphymata">
<emphasis id="B2ABEA22FFC6976AFED3F81EFDE6DBA2" box="[316,592,1976,2001]" italics="true" pageId="0" pageNumber="39">Paradella octaphymata</emphasis>
<bibRefCitation id="E44E4BC1FFC6976AFDB7F811FC38DBA2" author="Harrison" box="[600,910,1975,2001]" pageId="0" pageNumber="48" refString="Harrison, K. &amp; Holdich, D. M. (1982) Revision of the genera Dynamenella, Ischyromene, Dynamenopsis, and Cymodocella (Crustacea: Isopoda), including a new genus and five new species of eubranchiate sphaeromatids from Queensland waters. Journal of Crustacean Biology, 2 (1), 84 - 119." type="journal article" year="1982">Harrison &amp; Holdich, 1982</bibRefCitation>
</taxonomicName>
, by original designation.
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<subSubSection id="C8C565BBFFC7976BFF78FF31FE88DDC3" pageId="1" pageNumber="40" type="diagnosis">
<paragraph id="80603630FFC7976BFF78FF31FE88DDC3" blockId="1.[151,1437,151,1783]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF78FF31FEB3DCDC" bold="true" box="[151,261,151,175]" pageId="1" pageNumber="40">Diagnosis</emphasis>
. Body vaulted, dorsal surfaces smooth or with nodular ornamentation, without setae or with sparse setae; with ability to conglobate; strongly sexually dimorphic. Pleon consisting of 4 visible segments (as determined by lateral sutures); pleonite 1 entire; sutures (except first) running to posterior margin, merging, short; dorsal surface without process, with 2 or more pairs of tubercles; posterior margin even, with 'keys'. Pleotelson anteriorly as wide as pleon, without dorsal process; posterior margin with subapical Y-shaped foramen connected to posterior by narrow slit, with ventral thickened rim; lateral margins forming ridge. Epistome anteriorly narrow, without median constriction. Penial processes basally fused, long (extending to pleopod 1 rami). Pleopod 1 rami not operculate; exopod lamellar. Pleopod 2 endopod about as long as exopod; appendix masculina inserted basally, on short proximal lobe of endopod. Uropod rami broad, lamellar.
</paragraph>
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<subSubSection id="C8C565BBFFC79768FF29FE1CFB70DD0C" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40" type="description">
<paragraph id="80603630FFC7976BFF29FE1CFF51DF7A" blockId="1.[151,1437,151,1783]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF29FE1CFEFCDDA1" bold="true" box="[198,330,442,466]" pageId="1" pageNumber="40">Description</emphasis>
. 
<emphasis id="B2ABEA22FFC7976BFEB6FE1CFE25DDA2" box="[345,403,442,465]" italics="true" pageId="1" pageNumber="40">Head</emphasis>
with rostral point present, simple, separating antennular bases; anterior margin simple, without paired incisions in front of eyes, lateral margins not laterally extended to body outline (antennules more or less ventral). 
<emphasis id="B2ABEA22FFC7976BFF78FDA6FF48DE64" box="[151,254,512,535]" italics="true" pageId="1" pageNumber="40">Pereonite</emphasis>
1 lateral margins not anteriorly produced, not laterally enclosing head; pereonites 2–7 or 5–7 with posterior margin raised, forming narrow transverse ridge (usually on pereonites 5–7); anteriorly without keys. Sternite 1 without cuticular mesial extensions. Pereonite 6 simple, dorsally without bosses, processes or marginal extensions. Pereonite 7 as wide as pereonite 6, forming part of body outline, dorsally without bosses, processes or marginal extensions or with bifid posteriorly directed process. 
<emphasis id="B2ABEA22FFC7976BFE3CFD2CFDA1DED2" box="[467,535,650,673]" italics="true" pageId="1" pageNumber="40">Coxae</emphasis>
distally wide, those of pereonites 2–7 overlapping the one behind, coxae without ventral 'lock and key' processes, without grooved articulation; those of pereonite 6 not large, not overlapping those of pereonite 7. 
<emphasis id="B2ABEA22FFC7976BFEF2FD76FEEDDE94" box="[285,347,720,743]" italics="true" pageId="1" pageNumber="40">Pleon</emphasis>
posterior margin even, as wide as pleotelson; pleonal sternite narrow; pleotelson vaulted, weakly bidomed.
</paragraph>
<paragraph id="80603630FFC7976BFF29FCB3FC93DFE7" blockId="1.[151,1437,151,1783]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF29FCB3FE84DF5F" box="[198,306,789,812]" italics="true" pageId="1" pageNumber="40">Antennule</emphasis>
peduncle with basal articles mesially not in contact, peduncle articles 1 and 2 robust, article 3 slender; article 1 not produced; article 2 approximately 0.5 as long as article 1; with articles 2 and 3 colinear, article 3 longer than article 2; flagellum shorter than peduncle, longer than peduncular article 3. 
<emphasis id="B2ABEA22FFC7976BFC5AFCFCFBB8DF02" box="[949,1038,858,881]" italics="true" pageId="1" pageNumber="40">Antenna</emphasis>
peduncle articles all colinear, of similar thickness, less robust than peduncle articles of antennule.
</paragraph>
<paragraph id="80603630FFC7976BFF29FC06FBE9DFC4" blockId="1.[151,1437,151,1783]" box="[198,1119,928,951]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF29FC06FE9FDFC4" box="[198,297,928,951]" italics="true" pageId="1" pageNumber="40">Epistome</emphasis>
anteriorly not prominently extended, elongate, posteriorly enclosing labrum.
</paragraph>
<paragraph id="80603630FFC7976BFF29FC64FEBAD817" blockId="1.[151,1437,151,1783]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF29FC64FE9BDFAA" box="[198,301,962,985]" italics="true" pageId="1" pageNumber="40">Mandible</emphasis>
incisor wide, multicuspid; lacinia mobilis present; spine row normal, molar process gnathal surface with transverse ridges, rounded. 
<emphasis id="B2ABEA22FFC7976BFE54FC43FD97DF8F" box="[443,545,997,1020]" italics="true" pageId="1" pageNumber="40">Maxillule</emphasis>
lateral lobe with RS, some or all serrate, mesial lobe with 4 major heavily RS. 
<emphasis id="B2ABEA22FFC7976BFA8BFC43FF0DD86C" italics="true" pageId="1" pageNumber="40">Maxilla</emphasis>
with nodular setae on middle and lateral lobes nodular. 
<emphasis id="B2ABEA22FFC7976BFCF8FBAEFC3CD86C" box="[791,906,1032,1055]" italics="true" pageId="1" pageNumber="40">Maxilliped</emphasis>
palp articles 2–4 mesial margins lobate, article 2 not expanded; endite distal margin truncate, with clubbed RS, anteromesial (upper) marginal ridge without long curved serrate RS.
</paragraph>
<paragraph id="80603630FFC7976BFF29FBD6FBE8D962" blockId="1.[151,1437,151,1783]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF29FBD6FE9BD8F4" box="[198,301,1136,1159]" italics="true" pageId="1" pageNumber="40">Pereopod</emphasis>
1 ambulatory; dactylus secondary unguis simple; setae on merus and propodus short. Pereopod 1 (or 1–3), inferior margins of merus, carpus and propodus without conspicuous RS. Pereopod 2 similar in proportion to pereopod 3, dactylus with secondary unguis simple, short and stout. Pereopods 3–7 dactylus with secondary unguis simple. Pereopods with inferior margins of ischium to carpus bearing dense setulose fringe; ischium superior margin with sinuate acute RS, pereopods 1–3 (or 1–4) ischium superior margin without long stiff slender setae.
</paragraph>
<paragraph id="80603630FFC7976BFF29FABBFCB7D947" blockId="1.[151,1437,151,1783]" box="[198,769,1309,1332]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF29FABBFEB0D947" box="[198,262,1309,1332]" italics="true" pageId="1" pageNumber="40">Penes</emphasis>
tapering smoothly from midlength, apex acute.
</paragraph>
<paragraph id="80603630FFC7976BFF29FAE6FEF4DAFC" blockId="1.[151,1437,151,1783]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF29FAE6FE82D924" box="[198,308,1344,1367]" italics="true" pageId="1" pageNumber="40">Pleopod 1</emphasis>
exopod with longitudinal axis weakly oblique, distally subtruncate or truncate; endopod of similar proportions to exopod, mesial margin lamellar, distally angular, endopod proximomesial heel present. 
<emphasis id="B2ABEA22FFC7976BFB41FAC5FAD0D909" box="[1198,1382,1379,1402]" italics="true" pageId="1" pageNumber="40">Pleopods 1 and 2</emphasis>
exopod distal margins not deeply serrate. 
<emphasis id="B2ABEA22FFC7976BFDADFA23FCA9D9EF" box="[578,799,1413,1436]" italics="true" pageId="1" pageNumber="40">Appendix masculina</emphasis>
with straight margins, longer than and extending beyond endopod, distally acute. 
<emphasis id="B2ABEA22FFC7976BFE74FA0EFDB1D9CC" box="[411,519,1448,1471]" italics="true" pageId="1" pageNumber="40">Pleopod 3</emphasis>
exopod transverse suture present, endopod transverse ridges absent, endopod of similar proportions to exopod. 
<emphasis id="B2ABEA22FFC7976BFE52FA6DFD9AD991" box="[445,556,1483,1506]" italics="true" pageId="1" pageNumber="40">Pleopod 4</emphasis>
rami without PMS; exopod with thickened transverse ridges present, transverse suture absent, lateral margin not thickened, with short simple marginal setae; endopod with thickened transverse ridges present; mesial margin with deep distal notch, proximomesial lobe present. 
<emphasis id="B2ABEA22FFC7976BFC2EF9B6FB98DA54" box="[961,1070,1552,1575]" italics="true" pageId="1" pageNumber="40">Pleopod 5</emphasis>
exopod with thickened transverse ridges present, transverse suture present, incomplete, lateral margin with short simple setae, not thickened, distally usually with 3 discrete scale patches forming protruding lobes; endopod with thickened transverse ridges present, with proximomesial lobe.
</paragraph>
<paragraph id="80603630FFC7976BFF29F93DFCC9DA84" blockId="1.[151,1437,151,1783]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF29F93DFEAEDAC1" box="[198,280,1691,1714]" italics="true" pageId="1" pageNumber="40">Uropod</emphasis>
rami not strongly flattened, not forming part of continuous body outline; exopod (of adult male) similar in length to endopod, lamellar, lateral margin simple, finely serrate or smooth, distally broadly rounded or narrowly rounded or distally acute; endopod lamellar, distally broadly rounded.
</paragraph>
<paragraph id="80603630FFC7976BFF78F883FAA5DB0D" blockId="1.[151,1436,1829,1918]" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF78F883FF51DB4F" box="[151,231,1829,1852]" italics="true" pageId="1" pageNumber="40">Female</emphasis>
. Mouthparts of female not metamorphosed. Marsupium formed from 4 pairs of oostegites, anterior pocket absent, posterior pocket absent, oostegites not overlapping at mid-line. Ornamentation of pereonites, pleon and pleotelson less developed than in male; uropod rami of similar proportions, lateral margin of exopod not upturned.
</paragraph>
<paragraph id="80603630FFC79768FF78F80CFCCBDD2F" blockId="1.[151,1436,1962,2020]" lastBlockId="2.[151,1436,151,761]" lastPageId="2" lastPageNumber="41" pageId="1" pageNumber="40">
<emphasis id="B2ABEA22FFC7976BFF78F80CFEE2DBB1" bold="true" box="[151,340,1962,1986]" pageId="1" pageNumber="40">Species included</emphasis>
: 
<emphasis id="B2ABEA22FFC7976BFE8BF80DFE44DBB1" box="[356,498,1963,1986]" italics="true" pageId="1" pageNumber="40">P. a c u t i t e l s o n</emphasis>
(
<bibRefCitation id="E44E4BC1FFC7976BFDEAF80CFCA7DBB2" author="Menzies" box="[517,785,1962,1985]" pageId="1" pageNumber="49" refString="Menzies, R. J. &amp; Glynn, P. W. (1968) The common marine isopod Crustacea of Puerto Rico. Studies on the fauna of Curacao and other Caribbean Islands, 27 (104), 1 - 133." type="journal article" year="1968">Menzies &amp; Glynn, 1968</bibRefCitation>
), 
<collectingCountry id="F8C876A0FFC7976BFCC9F80CFC1CDBB2" box="[806,938,1962,1985]" name="Puerto Rico" pageId="1" pageNumber="40">Puerto Rico</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC7976BFC58F80DFB57DBB2" authority="(Menzies, 1962b)" baseAuthorityName="Menzies" baseAuthorityYear="1962" box="[951,1249,1962,1986]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B2ABEA22FFC7976BFC58F80DFBA1DBB1" box="[951,1047,1963,1986]" italics="true" pageId="1" pageNumber="40">P. bakeri</emphasis>
(
<bibRefCitation id="E44E4BC1FFC7976BFBC8F80CFB6FDBB2" author="Menzies" box="[1063,1241,1962,1985]" pageId="1" pageNumber="48" refString="Menzies, R. J. (1962 b) The zoogeography, ecology, and systematics of the Chilean marine isopods. Reports of the Lund University Chile Expedition 1948 - 49. 42. Lunds Universitets Arsskrifter, N. F. Avd. 2, 57 (11), 1 - 162." type="journal article" year="1962" yearSuffix="b">Menzies, 1962b</bibRefCitation>
)
</taxonomicName>
, 
<collectingCountry id="F8C876A0FFC7976BFB02F80CFA9CDBB2" box="[1261,1322,1962,1985]" name="Chile" pageId="1" pageNumber="40">Chile</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC7976BFAD8F80DFEE1DB97" authority="(Menzies, 1962a)" baseAuthorityName="Menzies" baseAuthorityYear="1962" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="40" phylum="Arthropoda" rank="species" species="dianae">
<emphasis id="B2ABEA22FFC7976BFAD8F80DFA2ADBB1" box="[1335,1436,1963,1986]" italics="true" pageId="1" pageNumber="40">P. dianae</emphasis>
(
<bibRefCitation id="E44E4BC1FFC7976BFF71F86BFEF9DB97" author="Menzies" box="[158,335,1997,2020]" pageId="1" pageNumber="48" refString="Menzies, R. J. (1962 a) The marine isopod fauna of Bahia de San Quintin Baja California, Mexico. Pacific Naturalist, 3 (11), 339 - 348." type="journal article" year="1962" yearSuffix="a">Menzies, 1962a</bibRefCitation>
)
</taxonomicName>
, Baja California, 
<collectingCountry id="F8C876A0FFC7976BFDF4F86BFDDBDB97" box="[539,621,1997,2020]" name="Mexico" pageId="1" pageNumber="40">Mexico</collectingCountry>
(now widely translocated by international shipping, e.g. 
<bibRefCitation id="E44E4BC1FFC7976BFB0CF86BFA2EDB97" author="Bey" box="[1251,1432,1997,2020]" pageId="1" pageNumber="48" refString="Bey, A., Rezig, M., Ben Souissi, J. &amp; Dridi, M. S. (2001) First record of Paradella diana e (Isopoda Sphaeromatidae) in the southern part of Tunis Lake. Bulletin de la Societe Zoologique de France, 126 (1 - 2), 220 - 223." type="journal article" year="2001">
Bey 
<emphasis id="B2ABEA22FFC7976BFAF7F86BFAE6DB97" box="[1304,1360,1997,2020]" italics="true" pageId="1" pageNumber="40">et al.</emphasis>
, 2001
</bibRefCitation>
, 
<bibRefCitation id="E44E4BC1FFC49768FF78FF3EFEC4DCDC" author="Hass" box="[151,370,152,175]" pageId="2" pageNumber="48" refString="Hass, C. G. &amp; Knott, B. (1998) Sphaeromatid isopods from the Swan River, Western Australia: diversity, distribution, and geographic sources. Crustaceana, 71, 36 - 46." type="journal article" year="1998">Hass &amp; Knott, 1998</bibRefCitation>
); 
<taxonomicName id="47DF4DB3FFC49768FE69FF3EFD95DCDC" box="[390,547,152,175]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="garsonorum" status="sp. nov.">
<emphasis id="B2ABEA22FFC49768FE69FF3EFD95DCDC" box="[390,547,152,175]" italics="true" pageId="2" pageNumber="41">P. garsonorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9985759FFC49768FDC4FF31FD37DCDC" box="[555,641,151,175]" pageId="2" pageNumber="41" rank="species">
<emphasis id="B2ABEA22FFC49768FDC4FF31FD37DCDC" bold="true" box="[555,641,151,175]" pageId="2" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
, Baja California, 
<collectingCountry id="F8C876A0FFC49768FCADFF3EFC21DCDC" box="[834,919,152,175]" name="Mexico" pageId="2" pageNumber="41">Mexico</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC49768FC4DFF3EFB03DCDC" authority="Muller, 1995" authorityName="Muller" authorityYear="1995" box="[930,1205,152,175]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="harrisoni">
<emphasis id="B2ABEA22FFC49768FC4DFF3EFB96DCDC" box="[930,1056,152,175]" italics="true" pageId="2" pageNumber="41">P. harrisoni</emphasis>
<bibRefCitation id="E44E4BC1FFC49768FBC7FF3EFB03DCDC" author="Muller" box="[1064,1205,152,175]" pageId="2" pageNumber="49" refString="Muller, H. - G. (1995) Sphaeromatidae from the Watamu area, Kenya description of a new genus and four new species (Isopoda, Flabellifera). Crustaceana, 68, 350 - 381." type="journal article" year="1995">Müller, 1995</bibRefCitation>
</taxonomicName>
; 
<collectingCountry id="F8C876A0FFC49768FB2DFF3EFABDDCDC" box="[1218,1291,152,175]" name="Kenya" pageId="2" pageNumber="41">Kenya</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC49768FAF9FF3EFDBADCA2" authority="Shoukr, Mona &amp; Rizk, 1987" authorityName="Shoukr, Mona &amp; Rizk" authorityYear="1987" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="heptaphymata">
<emphasis id="B2ABEA22FFC49768FAF9FF3EFF7ADCA2" italics="true" pageId="2" pageNumber="41">P. heptaphymata</emphasis>
<bibRefCitation id="E44E4BC1FFC49768FF3BFF1CFDBADCA2" author="Shoukr" box="[212,524,186,209]" pageId="2" pageNumber="49" refString="Shoukr, F. A., Mona, M. H. &amp; Rizk, E. T. (1987) Paradella heptaphymata: a new isopod species (Crustacea, Isopoda) from marine Egyptian water. Delta Journal of Science, 11 (1), 508 - 522." type="journal article" year="1987">Shoukr, Mona &amp; Rizk, 1987</bibRefCitation>
</taxonomicName>
, Suez Canal, 
<collectingCountry id="F8C876A0FFC49768FD4DFF1CFD53DCA2" box="[674,741,186,209]" name="Egypt" pageId="2" pageNumber="41">Egypt</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC49768FD1EFF1CFB0BDCA2" authority="Harrison &amp; Holdich, 1982" authorityName="Harrison &amp; Holdich" authorityYear="1982" box="[753,1213,186,209]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="octaphymata">
<emphasis id="B2ABEA22FFC49768FD1EFF1CFC20DCA2" box="[753,918,186,209]" italics="true" pageId="2" pageNumber="41">P. octaphymata</emphasis>
<bibRefCitation id="E44E4BC1FFC49768FC71FF1CFB0BDCA2" author="Harrison" box="[926,1213,186,209]" pageId="2" pageNumber="48" refString="Harrison, K. &amp; Holdich, D. M. (1982) Revision of the genera Dynamenella, Ischyromene, Dynamenopsis, and Cymodocella (Crustacea: Isopoda), including a new genus and five new species of eubranchiate sphaeromatids from Queensland waters. Journal of Crustacean Biology, 2 (1), 84 - 119." type="journal article" year="1982">Harrison &amp; Holdich, 1982</bibRefCitation>
</taxonomicName>
, 
<emphasis id="B2ABEA22FFC49768FB25FF1CFAE7DCA1" bold="true" box="[1226,1361,186,210]" pageId="2" pageNumber="41">
<typeStatus id="5F648892FFC49768FB25FF1CFB4DDCA1" box="[1226,1275,186,210]" pageId="2" pageNumber="41">type</typeStatus>
species
</emphasis>
, Great Barrier Reef, Queensland and 
<collectingCountry id="F8C876A0FFC49768FE09FF7BFDF5DC87" box="[486,579,221,244]" name="Vietnam" pageId="2" pageNumber="41">Vietnam</collectingCountry>
, South 
<collectingCountry id="F8C876A0FFC49768FD77FF7BFD6EDC87" box="[664,728,221,244]" name="China" pageId="2" pageNumber="41">China</collectingCountry>
Sea (
<bibRefCitation id="E44E4BC1FFC49768FCF6FF7BFBEBDC87" author="Kussakin" box="[793,1117,221,244]" pageId="2" pageNumber="48" refString="Kussakin, O. G. &amp; Malyutina, M. V. (1993) Sphaeromatidae (Crustacea: Isopoda: Flabellifera) from the South China Sea. Invertebrate Taxonomy, 7, 1167 - 203." type="journal article" year="1993">Kussakin &amp; Malyutina, 1993</bibRefCitation>
); 
<taxonomicName id="47DF4DB3FFC49768FB9CFF7BFA2EDC87" authority="Glynn, 1970" authorityName="Glynn" authorityYear="1970" box="[1139,1432,221,244]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="plicatura">
<emphasis id="B2ABEA22FFC49768FB9CFF7BFB47DC87" box="[1139,1265,221,244]" italics="true" pageId="2" pageNumber="41">P. plicatura</emphasis>
(
<bibRefCitation id="E44E4BC1FFC49768FAECFF7BFA26DC87" author="Glynn" box="[1283,1424,221,244]" pageId="2" pageNumber="48" refString="Glynn, P. W. (1970) A systematic study of the Sphaeromatidae (Crustacea: Isopoda) of Isla Margarita, Venezuela, with description of three new species. Memoria Sociedad de Ciencias Naturales La Salle, 30 (85), 5 - 48." type="journal article" year="1970">Glynn, 1970</bibRefCitation>
)
</taxonomicName>
, 
<collectingCountry id="F8C876A0FFC49768FF78FEA6FEBEDD64" box="[151,264,256,279]" name="Venezuela" pageId="2" pageNumber="41">Venezuela</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC49768FEFDFEA6FD5ADD64" authority="Menzies &amp; Glynn, 1968" authorityName="Menzies &amp; Glynn" authorityYear="1968" box="[274,748,256,279]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="quadripunctata">
<emphasis id="B2ABEA22FFC49768FEFDFEA6FE66DD64" box="[274,464,256,279]" italics="true" pageId="2" pageNumber="41">P. quadripunctata</emphasis>
(
<bibRefCitation id="E44E4BC1FFC49768FE31FEA6FD52DD64" author="Menzies" box="[478,740,256,279]" pageId="2" pageNumber="49" refString="Menzies, R. J. &amp; Glynn, P. W. (1968) The common marine isopod Crustacea of Puerto Rico. Studies on the fauna of Curacao and other Caribbean Islands, 27 (104), 1 - 133." type="journal article" year="1968">Menzies &amp; Glynn, 1968</bibRefCitation>
)
</taxonomicName>
, 
<collectingCountry id="F8C876A0FFC49768FD18FEA6FCCFDD64" box="[759,889,256,279]" name="Puerto Rico" pageId="2" pageNumber="41">Puerto Rico</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC49768FC6BFEA6FB36DD64" authority="Glynn, 1968" authorityName="Glynn" authorityYear="1968" box="[900,1152,256,279]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="setosa">
<emphasis id="B2ABEA22FFC49768FC6BFEA6FC56DD64" box="[900,992,256,279]" italics="true" pageId="2" pageNumber="41">P. setosa</emphasis>
(Glynn, 1968)
</taxonomicName>
, Pacific 
<collectingCountry id="F8C876A0FFC49768FB33FEA6FA82DD64" box="[1244,1332,256,279]" name="Panama" pageId="2" pageNumber="41">Panama</collectingCountry>
; 
<emphasis id="B2ABEA22FFC49768FAD1FEA6FF74DD4A" italics="true" pageId="2" pageNumber="41">P. t o m l e - klek</emphasis>
<bibRefCitation id="E44E4BC1FFC49768FF27FE84FEE7DD4A" author="Storey" box="[200,337,290,313]" pageId="2" pageNumber="49" refString="Storey, M. J. (2002) New species and a new record of sphaeromatid isopods (Crustacea) from the Andaman Sea, Thailand. In: Bruce, N. L. &amp; M. Berggren (Ed.) Proceedings of the International Workshop on the Crustacea in the Andaman Sea, Phuket Marine Biological Center, 29 November - 20 December 1998, Phuket Marine Biological Center Special Publication, 23, Vol. Phuket Marine Biology Center, Phuket, pp. 133 - 148." type="journal article" year="2002">Storey, 2002</bibRefCitation>
, Phuket, 
<collectingCountry id="F8C876A0FFC49768FE5CFE84FDA3DD4A" box="[435,533,290,313]" name="Thailand" pageId="2" pageNumber="41">Thailand</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC49768FDCFFE84FCF0DD4A" authority="Muller, 1991" authorityName="Muller" authorityYear="1991" box="[544,838,290,313]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="B2ABEA22FFC49768FDCFFE84FD02DD4A" box="[544,692,290,313]" italics="true" pageId="2" pageNumber="41">P. tuberculata</emphasis>
<bibRefCitation id="E44E4BC1FFC49768FD54FE84FCF0DD4A" author="Muller" box="[699,838,290,313]" pageId="2" pageNumber="49" refString="Muller, H. - G. (1991) Sphaeromatidae from coral reefs on the Society Islands, French Polynesia (Crustacea: Isopoda). Cahiers de Biologie Marine, 32, 83 - 104." type="journal article" year="1991">Müller, 1991</bibRefCitation>
</taxonomicName>
, Society Islands (Tahiti); 
<emphasis id="B2ABEA22FFC49768FBB7FE84FB42DD4A" box="[1112,1268,290,313]" italics="true" pageId="2" pageNumber="41">P. t u m i d i c a u d a</emphasis>
(
<bibRefCitation id="E44E4BC1FFC49768FAE9FE84FA26DD4A" author="Glynn" box="[1286,1424,290,313]" pageId="2" pageNumber="48" refString="Glynn, P. W. (1970) A systematic study of the Sphaeromatidae (Crustacea: Isopoda) of Isla Margarita, Venezuela, with description of three new species. Memoria Sociedad de Ciencias Naturales La Salle, 30 (85), 5 - 48." type="journal article" year="1970">Glynn, 1970</bibRefCitation>
), 
<collectingCountry id="F8C876A0FFC49768FF78FEE3FEBEDD2F" box="[151,264,325,348]" name="Venezuela" pageId="2" pageNumber="41">Venezuela</collectingCountry>
; 
<taxonomicName id="47DF4DB3FFC49768FEFDFEE3FDDEDD2F" authority="Bruce &amp; Wetzer, 2004" authorityName="Bruce &amp; Wetzer" authorityYear="2004" box="[274,616,325,348]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="species" species="tiffany">
<emphasis id="B2ABEA22FFC49768FEFDFEE3FEC6DD2F" box="[274,368,325,348]" italics="true" pageId="2" pageNumber="41">P. tiffany</emphasis>
<bibRefCitation id="E44E4BC1FFC49768FE98FEE3FDDEDD2F" author="Bruce" box="[375,616,325,348]" pageId="2" pageNumber="48" refString="Bruce, N. L. &amp; Wetzer, R. (2004) Paradella tiffany sp. nov., a distinctive sphaeromatid isopod (Crustacea: Isopoda: Sphaeromatidea) from Baja California, Mexico. Zootaxa, 623, 1 - 12." type="journal article" year="2004">Bruce &amp; Wetzer, 2004</bibRefCitation>
</taxonomicName>
, Baja California, 
<collectingCountry id="F8C876A0FFC49768FCCBFEE3FCCFDD2F" box="[804,889,325,348]" name="Mexico" pageId="2" pageNumber="41">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="80603630FFC49768FF29FECEFB70DD0C" blockId="2.[151,1436,151,761]" box="[198,1222,360,383]" pageId="2" pageNumber="41">
Known undescribed species: 
<taxonomicName id="47DF4DB3FFC49768FE11FECEFDDFDD0C" box="[510,617,360,383]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B2ABEA22FFC49768FE11FECEFDDFDD0C" box="[510,617,360,383]" italics="true" pageId="2" pageNumber="41">Paradella</emphasis>
</taxonomicName>
sp. Baja California, 
<collectingCountry id="F8C876A0FFC49768FCAAFECEFC2FDD0C" box="[837,921,360,383]" name="Mexico" pageId="2" pageNumber="41">Mexico</collectingCountry>
, from mangrove prop-roots.
</paragraph>
</subSubSection>
<subSubSection id="C8C565BBFFC49768FF78FE2CFC6DDEE7" pageId="2" pageNumber="41" type="discussion">
<paragraph id="80603630FFC49768FF78FE2CFD57DE7A" blockId="2.[151,1436,151,761]" pageId="2" pageNumber="41">
<emphasis id="B2ABEA22FFC49768FF78FE2CFF42DDD2" box="[151,244,394,417]" italics="true" pageId="2" pageNumber="41">Remarks</emphasis>
. Current generic diagnoses in the 
<taxonomicName id="47DF4DB3FFC49768FD80FE2CFCADDDD2" box="[623,795,394,417]" class="Malacostraca" family="Sphaeromatidae" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="family">Sphaeromatidae</taxonomicName>
are more restrictive than previously. A critical revision of those species placed in 
<taxonomicName id="47DF4DB3FFC49768FE7DFE0BFD0CDDB7" authority="Hansen, 1905" authorityName="Hansen" authorityYear="1905" box="[402,698,429,452]" class="Malacostraca" family="Sphaeromatidae" genus="Dynamenella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B2ABEA22FFC49768FE7DFE0BFD96DDB7" box="[402,544,429,452]" italics="true" pageId="2" pageNumber="41">Dynamenella</emphasis>
<bibRefCitation id="E44E4BC1FFC49768FDC8FE0BFD0CDDB7" author="Hansen" box="[551,698,429,452]" pageId="2" pageNumber="48" refString="Hansen, H. J. (1905) On the propagation, structure, and classification of the family Sphaeromidae. Quarterly Journal of Microscopical Science, 49, 69 - 135." type="journal article" year="1905">Hansen, 1905</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="E44E4BC1FFC49768FD1CFE0BFBBEDDB7" author="Harrison" box="[755,1032,429,452]" pageId="2" pageNumber="48" refString="Harrison, K. &amp; Holdich, D. M. (1982) Revision of the genera Dynamenella, Ischyromene, Dynamenopsis, and Cymodocella (Crustacea: Isopoda), including a new genus and five new species of eubranchiate sphaeromatids from Queensland waters. Journal of Crustacean Biology, 2 (1), 84 - 119." type="journal article" year="1982">Harrison &amp; Holdich 1982</bibRefCitation>
), would likely see transfer of some of those species and some of the numerous 
<emphasis id="B2ABEA22FFC49768FDB6FE76FD58DD94" box="[601,750,464,487]" italics="true" pageId="2" pageNumber="41">incertae sedis</emphasis>
species into 
<taxonomicName id="47DF4DB3FFC49768FC90FE76FC5FDD94" box="[895,1001,464,487]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B2ABEA22FFC49768FC90FE76FC5FDD94" box="[895,1001,464,487]" italics="true" pageId="2" pageNumber="41">Paradella</emphasis>
</taxonomicName>
, notably those species with merging Yshaped pleonal sutures and a long appendix masculina.
</paragraph>
<paragraph id="80603630FFC49768FF29FDB3FC6DDEE7" blockId="2.[151,1436,151,761]" pageId="2" pageNumber="41">
<taxonomicName id="47DF4DB3FFC49768FF29FDB3FE87DE5F" box="[198,305,533,556]" class="Malacostraca" family="Sphaeromatidae" genus="Paradella" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B2ABEA22FFC49768FF29FDB3FE87DE5F" box="[198,305,533,556]" italics="true" pageId="2" pageNumber="41">Paradella</emphasis>
</taxonomicName>
can best be identified by males having a distinct dorsally-directed, Y-shaped and posteriorly closed pleotelson foramen, long, tapering and basally fused penial processes, a long and basally narrow appendix masculina that usually extends beyond the distal margin of the endopod, both sexes with two fused Y-shaped pleonal sutures, entirely lamellar pleopod 1 rami and pereopod dactyli with a simple secondary unguis.
</paragraph>
</subSubSection>
<subSubSection id="C8C565BBFFC49768FF29FD04FD44DE8A" pageId="2" pageNumber="41" type="distribution">
<paragraph id="80603630FFC49768FF29FD04FD44DE8A" blockId="2.[151,1436,151,761]" pageId="2" pageNumber="41">
<emphasis id="B2ABEA22FFC49768FF29FD04FEFEDECA" box="[198,328,674,697]" italics="true" pageId="2" pageNumber="41">Distribution</emphasis>
. Widespread in shallow-water habitats in temperate and tropical regions of the Pacific Ocean, Indian Ocean and tropical western Atlantic; probably world wide, but not recorded from the eastern Atlantic or Mediterranean, nor from temperate northwestern Pacific (
<collectingCountry id="F8C876A0FFC49768FDBAFD44FD20DE8A" box="[597,662,738,761]" name="Japan" pageId="2" pageNumber="41">Japan</collectingCountry>
, 
<collectingCountry id="F8C876A0FFC49768FD4EFD44FD51DE8A" box="[673,743,738,761]" name="South Korea" pageId="2" pageNumber="41">Korea</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>