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<document id="52827F1EAE83DA05904335D53A395BD2" ID-CLB-Dataset="39861" ID-DOI="10.11646/zootaxa.3986.2.5" ID-GBIF-Dataset="0d76d10a-3a7e-495d-9489-869cef93683d" ID-ISSN="1175-5326" ID-Zenodo-Dep="243017" ID-ZooBank="3DA5305F-4D0B-4031-8240-9574842B336F" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1461262227298" checkinUser="plazi" docAuthor="Kamiński, Marcin Jan" docDate="2015" docId="03FC87F6D5402F38FF6AC7F9FA70FB5E" docLanguage="en" docName="zt03986p226.pdf" docOrigin="Zootaxa 3986 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quadrideres simplicipes" docType="key" docVersion="12" lastPageNumber="224" masterDocId="FFC5FF8ED5472F3FFFFDC521FFECFF8D" masterDocTitle="A taxonomic revision of the Quadrideres simplicipes species-group (Coleoptera: Tenebrionidae: Pedinini), including description of a new species from Tanzania" masterLastPageNumber="226" masterPageNumber="217" pageNumber="224" updateTime="1739538280245" updateUser="ExternalLinkService">
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<mods:title id="3BBF7DEFD20DF8082BA132CE5548B1F0">A taxonomic revision of the Quadrideres simplicipes species-group (Coleoptera: Tenebrionidae: Pedinini), including description of a new species from Tanzania</mods:title>
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<mods:namePart id="73F192B88743B92ADFC575E45C7F7442">Kamiński, Marcin Jan</mods:namePart>
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<treatment id="03FC87F6D5402F38FF6AC7F9FA70FB5E" ID-DOI="http://doi.org/10.5281/zenodo.6114064" ID-GBIF-Taxon="119637310" ID-Zenodo-Dep="6114064" LSID="urn:lsid:plazi:treatment:03FC87F6D5402F38FF6AC7F9FA70FB5E" httpUri="http://treatment.plazi.org/id/03FC87F6D5402F38FF6AC7F9FA70FB5E" lastPageNumber="224" pageId="7" pageNumber="224" scope_family="Tenebrionidae" scope_genus="Quadrideres" scope_order="Coleoptera">
<subSubSection id="C34F656BD5402F38FF6AC7F9FCCAFD7F" box="[151,806,727,754]" pageId="7" pageNumber="224" type="nomenclature">
<paragraph id="8BEA36E0D5402F38FF6AC7F9FCCAFD7F" blockId="7.[151,806,727,754]" box="[151,806,727,754]" pageId="7" pageNumber="224">
<heading id="D0A2818CD5402F38FF6AC7F9FCCAFD7F" bold="true" box="[151,806,727,754]" fontSize="11" level="1" pageId="7" pageNumber="224" reason="1">
<emphasis id="B921EAF2D5402F38FF6AC7F9FCCAFD7F" bold="true" box="[151,806,727,754]" pageId="7" pageNumber="224">
Key to the species of the
<taxonomicName id="4C554D63D5402F38FE37C7F6FD9FFD7C" baseAuthorityName="Gebien" baseAuthorityYear="1910" box="[458,627,727,753]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5402F38FE37C7F6FD9FFD7C" bold="true" box="[458,627,727,753]" italics="true" pageId="7" pageNumber="224">Q. simplicipes</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F656BD5402F38FF6AC63CFA70FB5E" pageId="7" pageNumber="224" type="key">
<keyStep id="30A12E45D5402F38FF6AC63CFA70FCE3" pageId="7" pageNumber="224">
<paragraph id="8BEA36E0D5402F38FF6AC63CFA77FCDC" blockId="7.[151,1436,797,1235]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC63CFA77FCDC" pageId="7" pageNumber="224">
1 Inner contours of male protibae with conspicuous postmedian dilatation....................................................................................................
<emphasis id="B921EAF2D5402F38FC68C61AFB72FCDC" bold="true" box="[917,1182,827,849]" pageId="7" pageNumber="224">
<taxonomicName id="4C554D63D5402F38FC68C61AFBE2FCDC" baseAuthorityName="Gebien" baseAuthorityYear="1911" box="[917,1038,827,849]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="interioris">
<emphasis id="B921EAF2D5402F38FC68C61AFBE2FCDC" bold="true" box="[917,1038,827,849]" italics="true" pageId="7" pageNumber="224">Q. interioris</emphasis>
</taxonomicName>
species-group
</emphasis>
(see Kamiński 2013a: 93)
</keyLead>
</paragraph>
<paragraph id="8BEA36E0D5402F38FF6AC678FA70FCE3" blockId="7.[151,1436,797,1235]" box="[151,1436,856,878]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC678FA70FCE3" box="[151,1436,856,878]" pageId="7" pageNumber="224">- Inner contours of male protibae without postmedian dilatation................................................ (2)</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E45D5402F38FF6AC654FA70FC24" pageId="7" pageNumber="224">
<paragraph id="8BEA36E0D5402F38FF6AC654FA70FC06" blockId="7.[151,1436,797,1235]" box="[151,1436,885,908]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC654FA70FC06" box="[151,1436,885,908]" pageId="7" pageNumber="224">
2 Secondary intervals and epipleurae of elytra with fine punctures.........
<emphasis id="B921EAF2D5402F38FC3AC657FB2AFC01" bold="true" box="[967,1222,886,908]" pageId="7" pageNumber="224">
<taxonomicName id="4C554D63D5402F38FC3AC657FBD9FC01" baseAuthorityName="Gerstaecker" baseAuthorityYear="1871" box="[967,1077,886,908]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="B921EAF2D5402F38FC3AC657FBD9FC01" bold="true" box="[967,1077,886,908]" italics="true" pageId="7" pageNumber="224">Q. scutatus</emphasis>
</taxonomicName>
species-group
</emphasis>
(see
<bibRefCitation id="EFC44B11D5402F38FB04C654FA8AFC06" author="Koch" box="[1273,1382,885,907]" pageId="7" pageNumber="224" refString="Koch, C. (1956) Exploration du Parc National de l'Upemba. II. Tenebrionidae (Coleoptera, Polyphaga), Opatrinae, First part: Platynotini, Litoborini and Loensini. M. Hayez, Bruxelles, 472 pp." type="book" year="1956">Koch 1956</bibRefCitation>
: 192)
</keyLead>
</paragraph>
<paragraph id="8BEA36E0D5402F38FF6AC6B5FA70FC24" blockId="7.[151,1436,797,1235]" box="[151,1436,915,937]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC6B5FA70FC24" box="[151,1436,915,937]" pageId="7" pageNumber="224">
- Secondary intervals and epipleurae of elytra with coarse punctures..................
<emphasis id="B921EAF2D5402F38FBB1C6B2FA96FC24" bold="true" box="[1100,1402,915,937]" pageId="7" pageNumber="224">
<taxonomicName id="4C554D63D5402F38FBB1C6B2FB3AFC24" baseAuthorityName="Gebien" baseAuthorityYear="1910" box="[1100,1238,915,937]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5402F38FBB1C6B2FB3AFC24" bold="true" box="[1100,1238,915,937]" italics="true" pageId="7" pageNumber="224">Q. simplicipes</emphasis>
</taxonomicName>
species-group...
</emphasis>
(3)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E45D5402F38FF6AC692FA77FC65" pageId="7" pageNumber="224">
<paragraph id="8BEA36E0D5402F38FF6AC692FA77FC44" blockId="7.[151,1436,797,1235]" box="[151,1435,945,969]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC692FA77FC44" box="[151,1435,945,969]" pageId="7" pageNumber="224">
3 5th abdominal ventrite without margin (
<figureCitation id="136E2A65D5402F38FDAAC692FD71FC44" box="[599,669,947,969]" captionStart="FIGURES 511" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[171,1416,199,1894]" captionTargetId="figure@3.[165,1421,195,1900]" captionText="FIGURES 511. Diagnostic characters proposed for the representatives of the Quadrideres simplicipes species-group. Pronotum (57), 5th ventrite (9, 10) and male protibia (11). Q. kazimierzi sp. nov. (57, 10), Q. parallelus (8, 11), Q. simplicipes (9)." figureDoi="http://doi.org/10.5281/zenodo.14871226" httpUri="https://zenodo.org/record/14871226/files/figure.png" pageId="7" pageNumber="224">Fig. 10</figureCitation>
)............................................................ (4)
</keyLead>
</paragraph>
<paragraph id="8BEA36E0D5402F38FF6AC6F2FA77FC65" blockId="7.[151,1436,797,1235]" box="[151,1435,976,1000]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC6F2FA77FC65" box="[151,1435,976,1000]" pageId="7" pageNumber="224">
- 5th abdominal ventrite with complete margin (
<figureCitation id="136E2A65D5402F38FD6BC6F3FD3CFC65" box="[662,720,978,1000]" captionStart="FIGURES 511" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[171,1416,199,1894]" captionTargetId="figure@3.[165,1421,195,1900]" captionText="FIGURES 511. Diagnostic characters proposed for the representatives of the Quadrideres simplicipes species-group. Pronotum (57), 5th ventrite (9, 10) and male protibia (11). Q. kazimierzi sp. nov. (57, 10), Q. parallelus (8, 11), Q. simplicipes (9)." figureDoi="http://doi.org/10.5281/zenodo.14871226" httpUri="https://zenodo.org/record/14871226/files/figure.png" pageId="7" pageNumber="224">Fig. 9</figureCitation>
)..............................................
<taxonomicName id="4C554D63D5402F38FAECC6F3FA77FC65" baseAuthorityName="Gebien" baseAuthorityYear="1910" box="[1297,1435,978,1000]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5402F38FAECC6F3FA77FC65" bold="true" box="[1297,1435,978,1000]" italics="true" pageId="7" pageNumber="224">Q. simplicipes</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E45D5402F38FF6AC6D1FE27FBD3" pageId="7" pageNumber="224">
<paragraph id="8BEA36E0D5402F38FF6AC6D1FA70FBAE" blockId="7.[151,1436,797,1235]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC6D1FA70FBAE" pageId="7" pageNumber="224">
4 Pronotal base with strong lateral indentations (
<figureCitation id="136E2A65D5402F38FD5BC6D1FD0DFB88" box="[678,737,1008,1030]" captionStart="FIGURES 511" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[171,1416,199,1894]" captionTargetId="figure@3.[165,1421,195,1900]" captionText="FIGURES 511. Diagnostic characters proposed for the representatives of the Quadrideres simplicipes species-group. Pronotum (57), 5th ventrite (9, 10) and male protibia (11). Q. kazimierzi sp. nov. (57, 10), Q. parallelus (8, 11), Q. simplicipes (9)." figureDoi="http://doi.org/10.5281/zenodo.14871226" httpUri="https://zenodo.org/record/14871226/files/figure.png" pageId="7" pageNumber="224">Fig. 7</figureCitation>
). Emargination of pronotal base interrupted laterally (
<figureCitation id="136E2A65D5402F38FB13C6D1FAC6FB88" box="[1262,1322,1008,1030]" captionStart="FIGURES 511" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[171,1416,199,1894]" captionTargetId="figure@3.[165,1421,195,1900]" captionText="FIGURES 511. Diagnostic characters proposed for the representatives of the Quadrideres simplicipes species-group. Pronotum (57), 5th ventrite (9, 10) and male protibia (11). Q. kazimierzi sp. nov. (57, 10), Q. parallelus (8, 11), Q. simplicipes (9)." figureDoi="http://doi.org/10.5281/zenodo.14871226" httpUri="https://zenodo.org/record/14871226/files/figure.png" pageId="7" pageNumber="224">Fig. 7</figureCitation>
). Pronotum widest at 0.75 of its length (
<figureCitation id="136E2A65D5402F38FE11C12CFDCAFBAE" box="[492,550,1037,1059]" captionStart="FIGURES 511" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[171,1416,199,1894]" captionTargetId="figure@3.[165,1421,195,1900]" captionText="FIGURES 511. Diagnostic characters proposed for the representatives of the Quadrideres simplicipes species-group. Pronotum (57), 5th ventrite (9, 10) and male protibia (11). Q. kazimierzi sp. nov. (57, 10), Q. parallelus (8, 11), Q. simplicipes (9)." figureDoi="http://doi.org/10.5281/zenodo.14871226" httpUri="https://zenodo.org/record/14871226/files/figure.png" pageId="7" pageNumber="224">Fig. 7</figureCitation>
)......................................................
<emphasis id="B921EAF2D5402F38FB3BC12CFA70FBAE" bold="true" box="[1222,1436,1037,1059]" pageId="7" pageNumber="224">
<taxonomicName id="4C554D63D5402F38FB3BC12CFAA4FBAE" authorityName="Kamiński" authorityYear="2015" box="[1222,1352,1037,1059]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="kazimierzi" status="sp. nov.">
<emphasis id="B921EAF2D5402F38FB3BC12CFAA4FBAE" bold="true" box="[1222,1352,1037,1059]" italics="true" pageId="7" pageNumber="224">Q. kazimierzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125789D5402F38FAB2C12CFA70FBAE" box="[1359,1436,1037,1059]" pageId="7" pageNumber="224" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEA36E0D5402F38FF6AC10AFE27FBD3" blockId="7.[151,1436,797,1235]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC10AFE27FBD3" pageId="7" pageNumber="224">
- Pronotal base with fine lateral indentations (
<figureCitation id="136E2A65D5402F38FD78C10BFD52FBCD" box="[645,702,1066,1088]" captionStart="FIGURES 511" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[171,1416,199,1894]" captionTargetId="figure@3.[165,1421,195,1900]" captionText="FIGURES 511. Diagnostic characters proposed for the representatives of the Quadrideres simplicipes species-group. Pronotum (57), 5th ventrite (9, 10) and male protibia (11). Q. kazimierzi sp. nov. (57, 10), Q. parallelus (8, 11), Q. simplicipes (9)." figureDoi="http://doi.org/10.5281/zenodo.14871226" httpUri="https://zenodo.org/record/14871226/files/figure.png" pageId="7" pageNumber="224">Fig. 8</figureCitation>
). Emargination of pronotal base complete (
<figureCitation id="136E2A65D5402F38FBA6C10BFB78FBCD" box="[1115,1172,1066,1088]" captionStart="FIGURES 511" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[171,1416,199,1894]" captionTargetId="figure@3.[165,1421,195,1900]" captionText="FIGURES 511. Diagnostic characters proposed for the representatives of the Quadrideres simplicipes species-group. Pronotum (57), 5th ventrite (9, 10) and male protibia (11). Q. kazimierzi sp. nov. (57, 10), Q. parallelus (8, 11), Q. simplicipes (9)." figureDoi="http://doi.org/10.5281/zenodo.14871226" httpUri="https://zenodo.org/record/14871226/files/figure.png" pageId="7" pageNumber="224">Fig. 8</figureCitation>
). Lateral sides of pronotum widest at the base … (5)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E45D5402F38FF6AC144FA70FB5E" pageId="7" pageNumber="224">
<paragraph id="8BEA36E0D5402F38FF6AC144FA70FB15" blockId="7.[151,1436,797,1235]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC144FA70FB15" pageId="7" pageNumber="224">
5 Protibae with shallow distal cavity on the inner side. Punctuation on the head, pronotum and elytra coarse. Body size: 10.0 12.0 mm.....................................................................................
<taxonomicName id="4C554D63D5402F38FAC8C1A3FA70FB15" authorityName="Koch" authorityYear="1956" box="[1333,1436,1154,1176]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="robynsi">
<emphasis id="B921EAF2D5402F38FAC8C1A3FA70FB15" bold="true" box="[1333,1436,1154,1176]" italics="true" pageId="7" pageNumber="224">Q. robynsi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEA36E0D5402F38FF6AC181FA70FB5E" blockId="7.[151,1436,797,1235]" pageId="7" pageNumber="224">
<keyLead id="30A495D5D5402F38FF6AC181FA70FB5E" pageId="7" pageNumber="224">
- Protibae with relatively deep distal cavity on the inner side (
<figureCitation id="136E2A65D5402F38FCD5C181FC82FB3B" box="[808,878,1184,1206]" captionStart="FIGURES 511" captionStartId="3.[151,264,1924,1947]" captionTargetBox="[171,1416,199,1894]" captionTargetId="figure@3.[165,1421,195,1900]" captionText="FIGURES 511. Diagnostic characters proposed for the representatives of the Quadrideres simplicipes species-group. Pronotum (57), 5th ventrite (9, 10) and male protibia (11). Q. kazimierzi sp. nov. (57, 10), Q. parallelus (8, 11), Q. simplicipes (9)." figureDoi="http://doi.org/10.5281/zenodo.14871226" httpUri="https://zenodo.org/record/14871226/files/figure.png" pageId="7" pageNumber="224">Fig. 11</figureCitation>
). Punctuation on the head, pronotum and elytra relatively fine. Body size:
<quantity id="4CAD9B05D5402F38FE83C19CFE11FB5E" box="[382,509,1213,1235]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="7" pageNumber="224" unit="mm" value="9.5" valueMax="10.5" valueMin="8.5">8.510.5 mm</quantity>
.................................................................
<taxonomicName id="4C554D63D5402F38FAE1C19CFA70FB5E" authorityName="Ancey" authorityYear="1877" box="[1308,1436,1213,1235]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="224" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B921EAF2D5402F38FAE1C19CFA70FB5E" bold="true" box="[1308,1436,1213,1235]" italics="true" pageId="7" pageNumber="224">Q. parallelus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>