treatments-xml/data/03/F4/0C/03F40C7EFFBE2938CCDBFB79FA86FCE5.xml
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<document id="E1338DFA9B0382246301B78A6F1F233B" ID="10.11646/zootaxa.4121.1.1" ID-CLB-Dataset="38929" ID-DOI="10.11646/zootaxa.4121.1.1" ID-GBIF-Dataset="214dc5a7-b23a-4140-b251-3a564b18bb9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="265513" ID-ZooBank="2C978846-61DD-48BD-87BE-0BC22D0CABF2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465281072039" checkinUser="plazi" docAuthor="Kelly, Michelle, Sim-Smith, Carina, Stone, Robert, Reiswig, Toufiek Samaai Henry &amp; Austin, William" docDate="2016" docId="03F40C7EFFBE2938CCDBFB79FA86FCE5" docLanguage="en" docName="zootaxa.4121.1.1.pdf" docOrigin="Zootaxa 4121 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bomba Kelly, Reiswig &amp; Samaai, gen. nov." docType="treatment" docVersion="12" lastPageNumber="29" masterDocId="FFCD7406FFA52924CC4CFF9CFFB0FFDD" masterDocTitle="New taxa and arrangements within the family Latrunculiidae (Demospongiae, Poecilosclerida)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="28" updateTime="1698662720313" updateUser="ExternalLinkService">
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<mods:title id="B70422A5BC9C088244415560EA9C7133">New taxa and arrangements within the family Latrunculiidae (Demospongiae, Poecilosclerida)</mods:title>
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<mods:namePart id="678C0EBDF99020AAADF1292034FD0420">Kelly, Michelle</mods:namePart>
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<mods:namePart id="7D010E2FBC00E20869D9811D25362FAD">Stone, Robert</mods:namePart>
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<mods:namePart id="046F9015CE216CC96EB56A16ABCFD49B">Reiswig, Toufiek Samaai Henry</mods:namePart>
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<mods:namePart id="B31C81B03C3D4C7BE7038672BAF804B8">Austin, William</mods:namePart>
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<treatment id="03F40C7EFFBE2938CCDBFB79FA86FCE5" ID-DOI="http://doi.org/10.5281/zenodo.5058025" ID-GBIF-Taxon="120817189" ID-Zenodo-Dep="5058025" LSID="urn:lsid:plazi:treatment:03F40C7EFFBE2938CCDBFB79FA86FCE5" httpUri="http://treatment.plazi.org/id/03F40C7EFFBE2938CCDBFB79FA86FCE5" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<subSubSection id="C347EEE3FFBE293FCCDBFB79FD46FB22" box="[151,758,1253,1279]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BE2BD68FFBE293FCCDBFB79FD46FB22" blockId="27.[151,758,1253,1279]" box="[151,758,1253,1279]" pageId="27" pageNumber="28">
<heading id="D0AA0A04FFBE293FCCDBFB79FD46FB22" bold="true" box="[151,758,1253,1279]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B929617AFFBE293FCCDBFB79FD46FB22" bold="true" box="[151,758,1253,1279]" pageId="27" pageNumber="28">
Genus
<taxonomicName id="4C5DC6EBFFBE293FCCA1FB79FD31FB22" ID-CoL="84MJ3" authority="Kelly, Reiswig &amp; Samaai" authorityName="Kelly, Reiswig &amp; Samaai" box="[237,641,1253,1279]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="27" pageNumber="28" phylum="Porifera" rank="genus" status="gen. nov.">
<emphasis id="B929617AFFBE293FCCA1FB79FEF3FB22" bold="true" box="[237,323,1253,1279]" italics="true" pageId="27" pageNumber="28">Bomba</emphasis>
Kelly, Reiswig &amp; Samaai
</taxonomicName>
<taxonomicNameLabel id="A21ADC01FFBE293FCEC4FB79FD46FB22" box="[648,758,1253,1279]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C347EEE3FFBE293FCCDBFAB0FC7CFA98" box="[151,972,1324,1349]" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BE2BD68FFBE293FCCDBFAB0FC7CFA98" blockId="27.[151,1437,1324,2033]" box="[151,972,1324,1349]" pageId="27" pageNumber="28">
<emphasis id="B929617AFFBE293FCCDBFAB0FE82FA98" bold="true" box="[151,306,1324,1349]" pageId="27" pageNumber="28">
<typeStatus id="54E603CAFFBE293FCCDBFAB0FF62FA98" box="[151,210,1324,1349]" pageId="27" pageNumber="28">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C5DC6EBFFBE293FCD75FAB0FD8CFA98" box="[313,572,1324,1349]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="27" pageNumber="28" phylum="Porifera" rank="species" species="endeavourensis">
<emphasis id="B929617AFFBE293FCD75FAB0FD8CFA98" box="[313,572,1324,1349]" italics="true" pageId="27" pageNumber="28">Bomba endeavourensis</emphasis>
</taxonomicName>
<emphasis id="B929617AFFBE293FCE0FFAB0FD41FA98" bold="true" box="[579,753,1324,1349]" pageId="27" pageNumber="28">gen. et sp. nov.</emphasis>
, designated herein.
</paragraph>
</subSubSection>
<subSubSection id="C347EEE3FFBE293FCC8BFACCFE66F858" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8BE2BD68FFBE293FCC8BFACCFE66F858" blockId="27.[151,1437,1324,2033]" pageId="27" pageNumber="28">
<emphasis id="B929617AFFBE293FCC8BFACCFEF1FAB4" bold="true" box="[199,321,1360,1385]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
Small, low-lying hemispherical to pedunculate
<taxonomicName id="4C5DC6EBFFBE293FCF2AFACCFBB9FAB4" box="[870,1033,1360,1385]" class="Demospongiae" family="Latrunculiidae" kingdom="Animalia" order="Poecilosclerida" pageId="27" pageNumber="28" phylum="Porifera" rank="family">Latrunculiidae</taxonomicName>
with a smooth or wrinkled surface which may be raised in places into tough, horn-like bundles of megascleres. Megascleres are large sinuous styles that are lightly terminally spined. Microscleres are unusual anisodiscorhabds with only three whorls of projections: a basal whorl, a median whorl, and an apical whorl. The basal whorl is composed of bifurcate projections that may be terminally rounded or sharp. The median whorl, located slightly above centre on the shaft, is similar to the basal whorl in all aspects of morphology except that the bifurcate projections may be splayed horizontally. The apical whorl consists of doublet cylindrical projections, the tips of which may be sharp or bluntly terminated in the form of a chiaster, the edges ringed with minute spines. The doublet projections of the basal and median whorls are horizontally arrayed perpendicular to the shaft, while those of the apical whorl are vertically orientated, parallel with the spicule shaft. Projections are generally smooth but may be sparsely acanthose with spines in patches and linear arrays. The basal whorl is orientated obliquely and slightly away from the median and apical whorls which are orientated obliquely and slightly forward. The anisodiscorhabd progenitor consists of differentiated basal, median, and apical whorls from the earliest stages. Ectosome forms a leathery casing formed by a tangential crosshatching of megascleres topped with a patchy palisade of single, well-spaced microscleres. Choanosome is soft, detaching easily from the ectosome, permeated with thick, strong fibres. Megascleres and microscleres are scattered between the fibres.
</paragraph>
</subSubSection>
<subSubSection id="C347EEE3FFBE293FCC8BF80CFE1EF810" pageId="27" pageNumber="28" type="etymology">
<paragraph id="8BE2BD68FFBE293FCC8BF80CFE1EF810" blockId="27.[151,1437,1324,2033]" pageId="27" pageNumber="28">
<emphasis id="B929617AFFBE293FCC8BF80CFEFEF874" bold="true" box="[199,334,1936,1961]" pageId="27" pageNumber="28">Etymology.</emphasis>
Named for the
<typeStatus id="54E603CAFFBE293FCDB3F80CFDD1F874" box="[511,609,1936,1961]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
which has the morphology of a floating sea mine as seen from above the water (
<emphasis id="B929617AFFBE293FCCA9F828FE81F810" box="[229,305,1972,1997]" italics="true" pageId="27" pageNumber="28">bomba</emphasis>
, mine; L.).
</paragraph>
</subSubSection>
<subSubSection id="C347EEE3FFBE2938CC8BF844FA86FCE5" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BE2BD68FFBE2938CC8BF844FD5FFEB8" blockId="27.[151,1437,1324,2033]" lastBlockId="28.[151,1436,151,824]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B929617AFFBE293FCC8BF844FE8BF82C" bold="true" box="[199,315,2008,2033]" pageId="27" pageNumber="28">Remarks.</emphasis>
The anisodiscorhabds of
<taxonomicName id="4C5DC6EBFFBE293FCE20F844FD0BF82C" box="[620,699,2008,2033]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="27" pageNumber="28" phylum="Porifera" rank="genus" status="gen. nov.">
<emphasis id="B929617AFFBE293FCE20F844FD0BF82C" box="[620,699,2008,2033]" italics="true" pageId="27" pageNumber="28">Bomba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21ADC01FFBE293FCE8BF844FC85F82C" box="[711,821,2008,2033]" pageId="27" pageNumber="28" rank="genus">
<emphasis id="B929617AFFBE293FCE8BF844FC85F82C" bold="true" box="[711,821,2008,2033]" pageId="27" pageNumber="28">gen. nov.</emphasis>
</taxonomicNameLabel>
are very unusual in that they appear to be a highly reduced form with only three whorls of projections; there is no evidence of a manubrium or subsidiary whorl or apex. Although the microsclere appears superficially to be the same at each end, the basal and apical whorls are at slightly different angles to the shaft (the apical whorl projecting forwards) and the doublet projections on the apical whorl are on a different plane compared to those of the median and basal whorls. The megascleres in this genus are the longest known to date in the
<taxonomicName id="4C5DC6EBFFB92938CE48FEBBFD1AFE9D" box="[516,682,295,320]" class="Demospongiae" family="Latrunculiidae" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="family">Latrunculiidae</taxonomicName>
, averaging over 600 µm and reaching well over 700 µm in the case of the
<typeStatus id="54E603CAFFB92938CC92FED1FEBEFEB8" box="[222,270,333,357]" pageId="28" pageNumber="29">type</typeStatus>
species
<taxonomicName id="4C5DC6EBFFB92938CD21FED1FD89FEB8" box="[365,569,332,357]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="species" species="endeavourensis">
<emphasis id="B929617AFFB92938CD21FED1FECEFEB9" box="[365,382,333,356]" italics="true" pageId="28" pageNumber="29">B</emphasis>
.
<emphasis id="B929617AFFB92938CDC0FED0FD89FEB8" box="[396,569,332,357]" italics="true" pageId="28" pageNumber="29">endeavourensis</emphasis>
</taxonomicName>
<emphasis id="B929617AFFB92938CE0DFED0FD5FFEB8" bold="true" box="[577,751,332,357]" pageId="28" pageNumber="29">gen. et sp. nov.</emphasis>
</paragraph>
<paragraph id="8BE2BD68FFB92938CC8BFEECFD93FD7A" blockId="28.[151,1436,151,824]" pageId="28" pageNumber="29">
<typeStatus id="54E603CAFFB92938CC8BFEECFF4FFE55" box="[199,255,368,392]" pageId="28" pageNumber="29">Type</typeStatus>
species
<taxonomicName id="4C5DC6EBFFB92938CD29FEF3FDDDFE55" box="[357,621,367,392]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="species" species="endeavourensis">
<emphasis id="B929617AFFB92938CD29FEF3FDDDFE55" box="[357,621,367,392]" italics="true" pageId="28" pageNumber="29">Bomba endeavourensis</emphasis>
</taxonomicName>
<emphasis id="B929617AFFB92938CE34FEF3FC82FE55" bold="true" box="[632,818,367,392]" pageId="28" pageNumber="29">gen. et sp. nov.</emphasis>
can be compared to species of
<taxonomicName id="4C5DC6EBFFB92938C8E1FEF3FA88FE55" box="[1197,1336,367,392]" class="Demospongiae" family="Latrunculiidae" genus="Tsitsikamma" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="genus">
<emphasis id="B929617AFFB92938C8E1FEF3FA88FE55" box="[1197,1336,367,392]" italics="true" pageId="28" pageNumber="29">Tsitsikamma</emphasis>
</taxonomicName>
(Family
<taxonomicName id="4C5DC6EBFFB92938CCDBFE08FE8CFE70" box="[151,316,404,429]" class="Demospongiae" family="Latrunculiidae" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="family">Latrunculiidae</taxonomicName>
) from South African waters, which also have an ectosomal casing that is extraordinarily tough and leathery and composed of a thick, paratangential cross-hatch of megascleres. However,
<taxonomicName id="4C5DC6EBFFB92938C835FE2BFAB4FE0D" box="[1145,1284,439,464]" class="Demospongiae" family="Latrunculiidae" genus="Tsitsikamma" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="genus">
<emphasis id="B929617AFFB92938C835FE2BFAB4FE0D" box="[1145,1284,439,464]" italics="true" pageId="28" pageNumber="29">Tsitsikamma</emphasis>
</taxonomicName>
<typeStatus id="54E603CAFFB92938C940FE24FA8CFE0D" box="[1292,1340,440,464]" pageId="28" pageNumber="29">type</typeStatus>
species,
<taxonomicName id="4C5DC6EBFFB92938CCDBFE41FE41FE29" authority="Samaai &amp; Kelly, 2002" authorityName="Samaai &amp; Kelly" authorityYear="2002" box="[151,497,476,501]" class="Demospongiae" family="Latrunculiidae" genus="Tsitsikamma" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="species" species="favus">
<emphasis id="B929617AFFB92938CCDBFE41FF14FE29" box="[151,164,477,500]" italics="true" pageId="28" pageNumber="29">T</emphasis>
.
<emphasis id="B929617AFFB92938CCFFFE40FF5EFE29" box="[179,238,476,500]" italics="true" pageId="28" pageNumber="29">favus</emphasis>
Samaai &amp; Kelly, 2002
</taxonomicName>
, and
<taxonomicName id="4C5DC6EBFFB92938CE60FE41FB7DFE29" authority="Samaai, Gibbons, Kelly &amp; Davies-Coleman, 2003" authorityName="Samaai, Gibbons, Kelly &amp; Davies-Coleman" authorityYear="2003" box="[556,1229,476,501]" class="Demospongiae" family="Latrunculiidae" genus="Tsitsikamma" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="species" species="scurra">
<emphasis id="B929617AFFB92938CE60FE41FD89FE29" box="[556,569,477,500]" italics="true" pageId="28" pageNumber="29">T</emphasis>
.
<emphasis id="B929617AFFB92938CE0BFE41FD20FE29" box="[583,656,477,500]" italics="true" pageId="28" pageNumber="29">scurra</emphasis>
<bibRefCitation id="EFCCC099FFB92938CED4FE40FB7DFE29" author="Samaai" box="[664,1229,476,501]" pageId="28" pageNumber="29" refString="Samaai, T., Gibbons, M. J., Kelly, M. &amp; Davies-Coleman, M. (2003) South African Latrunculiidae (Porifera: Demospongiae: Poecilosclerida): descriptions of new species of Latrunculia du Bocage, Strongylodesma Levi, and Tsitsikamma Samaai &amp; Kelly. Zootaxa, 371, 1 - 26." type="journal article" year="2003">Samaai, Gibbons, Kelly &amp; Davies-Coleman, 2003</bibRefCitation>
</taxonomicName>
, differ in that they are constructed of large, rigid chambers, not a single sack as
<taxonomicName id="4C5DC6EBFFB92938CF01FD9DFBABFDC5" box="[845,1051,511,536]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="species" species="endeavourensis">
<emphasis id="B929617AFFB92938CF01FD9DFCEEFDC5" box="[845,862,513,536]" italics="true" pageId="28" pageNumber="29">B</emphasis>
.
<emphasis id="B929617AFFB92938CF22FE63FBABFDC5" box="[878,1051,511,536]" italics="true" pageId="28" pageNumber="29">endeavourensis</emphasis>
</taxonomicName>
<emphasis id="B929617AFFB92938C86FFE63FB65FDC5" bold="true" box="[1059,1237,511,536]" pageId="28" pageNumber="29">gen. et sp. nov.</emphasis>
Both genera also have species with unusual anisodiscorhabds featuring three rings of uniform doublet projections (or triplets in the case of
<taxonomicName id="4C5DC6EBFFB92938CCB2FDDBFE37FDBD" box="[254,391,583,608]" class="Demospongiae" family="Latrunculiidae" genus="Tsitsikamma" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="genus">
<emphasis id="B929617AFFB92938CCB2FDDBFE37FDBD" box="[254,391,583,608]" italics="true" pageId="28" pageNumber="29">Tsitsikamma</emphasis>
</taxonomicName>
species) with chiaster-like terminations. The specific form and ornamentation of the microscleres in
<taxonomicName id="4C5DC6EBFFB92938CD05FDF0FE28FD58" box="[329,408,620,645]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="genus" status="gen. nov.">
<emphasis id="B929617AFFB92938CD05FDF0FE28FD58" box="[329,408,620,645]" italics="true" pageId="28" pageNumber="29">Bomba</emphasis>
</taxonomicName>
<emphasis id="B929617AFFB92938CDECFDF0FDBAFD58" bold="true" box="[416,522,620,645]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="A21ADC01FFB92938CDECFDF0FDBAFD58" box="[416,522,620,645]" pageId="28" pageNumber="29" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
are quite different, however, and no other
<taxonomicName id="4C5DC6EBFFB92938CFA0FDF0FB3FFD58" box="[1004,1167,620,645]" class="Demospongiae" family="Latrunculiidae" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="family">Latrunculiidae</taxonomicName>
has been recorded with protruding, horn-like spicule tracts.
</paragraph>
<paragraph id="8BE2BD68FFB92938CC8BFD28FA86FCE5" blockId="28.[151,1436,151,824]" pageId="28" pageNumber="29">
The microscleres of
<taxonomicName id="4C5DC6EBFFB92938CDE0FD29FDC9FD10" box="[428,633,692,717]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="species" species="endeavourensis">
<emphasis id="B929617AFFB92938CDE0FD29FE0DFD11" box="[428,445,693,716]" italics="true" pageId="28" pageNumber="29">B</emphasis>
.
<emphasis id="B929617AFFB92938CD80FD28FDC9FD10" box="[460,633,692,717]" italics="true" pageId="28" pageNumber="29">endeavourensis</emphasis>
</taxonomicName>
<emphasis id="B929617AFFB92938CECCFD28FC9FFD10" bold="true" box="[640,815,692,717]" pageId="28" pageNumber="29">gen. et sp. nov.</emphasis>
are grossly comparable to species of the South African genus
<taxonomicName id="4C5DC6EBFFB92938CCA0FD4BFCF1FD2D" authority="Samaai, Govender &amp; Kelly, 2004" authorityName="Samaai, Govender &amp; Kelly" authorityYear="2004" box="[236,833,727,752]" class="Demospongiae" family="Latrunculiidae" genus="Cyclacanthia" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="genus">
<emphasis id="B929617AFFB92938CCA0FD4BFE31FD2D" box="[236,385,727,752]" italics="true" pageId="28" pageNumber="29">Cyclacanthia</emphasis>
<bibRefCitation id="EFCCC099FFB92938CDDAFD4BFCF1FD2D" author="Samaai" box="[406,833,727,752]" pageId="28" pageNumber="29" refString="Samaai, T., Govender, V. &amp; Kelly, M. (2004) Cyclacanthia n. g. (Demospongiae: Poecilosclerida: Latrunculiidae incertae sedis), a new genus of marine sponges from South African waters, and description of two new species. Zootaxa, 725, 1 - 18." type="journal article" year="2004">Samaai, Govender &amp; Kelly, 2004</bibRefCitation>
</taxonomicName>
(Family
<taxonomicName id="4C5DC6EBFFB92938CF8FFD4BFBDBFD2D" box="[963,1131,727,752]" class="Demospongiae" family="Latrunculiidae" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="family">Latrunculiidae</taxonomicName>
), which have acanthose isospinodiscorhabds with three whorls of spines positioned equidistantly along the shaft, but which also bear a manubrium and apical spike. Figure H illustrates the diagnostic microscleres that define
<taxonomicName id="4C5DC6EBFFB92938C83AFC83FB75FCE5" box="[1142,1221,799,824]" class="Demospongiae" family="Latrunculiidae" genus="Bomba" kingdom="Animalia" order="Poecilosclerida" pageId="28" pageNumber="29" phylum="Porifera" rank="genus" status="gen. nov.">
<emphasis id="B929617AFFB92938C83AFC83FB75FCE5" box="[1142,1221,799,824]" italics="true" pageId="28" pageNumber="29">Bomba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21ADC01FFB92938C880FC83FA86FCE5" box="[1228,1334,799,824]" pageId="28" pageNumber="29" rank="genus">
<emphasis id="B929617AFFB92938C880FC83FA86FCE5" bold="true" box="[1228,1334,799,824]" pageId="28" pageNumber="29">gen. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>