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<document id="22A2C0669C8A2B4CFB3FF74AA2A9FD58" ID-CLB-Dataset="21955" ID-DOI="10.5852/ejt.2017.368" ID-GBIF-Dataset="7d707e31-1104-4ed3-8493-ef9302b98796" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838412" ID-ZooBank="E4AEAA16-A546-48B8-99BA-2E0152C8C6F0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590087673171" checkinUser="carolina" docAuthor="Zlatkov, Boyan &amp; Huemer, Peter" docDate="2017" docId="03E9996C144D4E7FB5356E588128DA1A" docLanguage="en" docName="ejt-2017-368.pdf.imf" docOrigin="European Journal of Taxonomy 368" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Phtheochroa Stephens 1829" docType="key" docVersion="7" lastPageNumber="3" masterDocId="FFD0E114144F4E7DB7706C11845EDC35" masterDocTitle="Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="3" updateTime="1698840706466" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="360036A1511D5A07590E828F1485F585">Allopatric cryptic diversity in the alpine species complex Phtheochroa frigidana s. lat. (Lepidoptera: Tortricidae)</mods:title>
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<mods:name id="2FFDA698C1121511A9806CB3BE3A2DA7" type="personal">
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<mods:namePart id="146D6B1301E1F53BB2B33FF5B9195E15">Zlatkov, Boyan</mods:namePart>
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<mods:affiliation id="0E14BCC00BFB19631FA78C380026366B">Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. &amp; National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria. &amp; urn: lsid: zoobank. org: author: 3538 D 3 F 5 - 6822 - 403 F-B 9 E 6 - E 8 A 95 E 884404 &amp; Corresponding author: bzlatkov @ gmail. com</mods:affiliation>
<mods:nameIdentifier id="F1EFAB2E966ABF52CEC69D92CD2F206A" type="email">bzlatkov@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C60B907AA28AF58C1B5303683A12E847">Huemer, Peter</mods:namePart>
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<mods:affiliation id="5E7E23DC9FF1D9DE88AC19EB901B606E">Tiroler Landesmuseen Betriebsgesellschaft m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria. &amp; Email: p. huemer @ tiroler-landesmuseen. at &amp; urn: lsid: zoobank. org: author: 4 E 6 F 001 E-DB 9 D- 460 C-AB 39 - 0 CE 7 CC 773 B 59</mods:affiliation>
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<mods:title id="A5DE974D9204126783A998C818039AFF">European Journal of Taxonomy</mods:title>
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<mods:date id="DDF9420736C35FBAD9E19E45A43143BF">2017</mods:date>
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<mods:number id="3E8A64F6BE1322EF37D8D6C32F789CBC">2017-11-21</mods:number>
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<treatment id="03E9996C144D4E7FB5356E588128DA1A" ID-DOI="http://doi.org/10.5281/zenodo.3852274" ID-GBIF-Taxon="164246396" ID-Zenodo-Dep="3852274" LSID="urn:lsid:plazi:treatment:03E9996C144D4E7FB5356E588128DA1A" httpUri="http://treatment.plazi.org/id/03E9996C144D4E7FB5356E588128DA1A" lastPageNumber="3" pageId="2" pageNumber="3">
<subSubSection id="C35A7BF1144D4E7FB5356E5887B1DE56" box="[581,1007,584,611]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BFF287A144D4E7FB5356E5887B1DE56" blockId="2.[581,1007,584,611]" box="[581,1007,584,611]" pageId="2" pageNumber="3">
<heading id="D0B79F16144D4E7FB5356E5887B1DE56" box="[581,1007,584,611]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
Genus
<taxonomicName id="4C4053F9144D4E7FB5E76E5987B1DE56" ID-CoL="92JXC" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" box="[663,1007,584,611]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B934F468144D4E7FB5E76E59876DDE56" bold="true" box="[663,819,584,611]" italics="true" pageId="2" pageNumber="3">Phtheochroa</emphasis>
Stephens, 1829
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C35A7BF1144D4E7FB7CD6E9B8690DEDE" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8BFF287A144D4E7FB7CD6E9B8690DEDE" blockId="2.[189,1399,650,747]" pageId="2" pageNumber="3">
The analysis of genitalia structures of everted vesicae in male genitalia and DNA barcodes revealed the existence of five European species in the
<taxonomicName id="4C4053F9144D4E7FB5AF6EBF87B2DEFD" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[735,1004,686,712]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F468144D4E7FB5AF6EBF87B2DEFD" box="[735,1004,686,712]" italics="true" pageId="2" pageNumber="3">Phtheochroa frigidana</emphasis>
</taxonomicName>
group. The key to the species is followed by detailed taxonomic descriptions.
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF1144D4E7FB7CD6F038062DF19" box="[189,1084,786,813]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BFF287A144D4E7FB7CD6F038062DF19" blockId="2.[189,1084,786,813]" box="[189,1084,786,813]" pageId="2" pageNumber="3">
<heading id="D0B79F16144D4E7FB7CD6F038062DF19" bold="true" box="[189,1084,786,813]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="B934F468144D4E7FB7CD6F038062DF19" bold="true" box="[189,1084,786,813]" pageId="2" pageNumber="3">
Key to the
<taxonomicName id="4C4053F9144D4E7FB6356F038607DF18" ID-CoL="4GX97" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[325,601,786,813]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F468144D4E7FB6356F038607DF18" bold="true" box="[325,601,786,813]" italics="true" pageId="2" pageNumber="3">Phtheochroa frigidana</emphasis>
</taxonomicName>
species group, based on male genitalia
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35A7BF1144D4E7FB7CD6F508128DA1A" pageId="2" pageNumber="3" type="key">
<key id="FFD17F4B144D4E7FB7CD6F508128DA1A" pageId="2" pageNumber="3">
<keyStep id="30B430DF144D4E7FB7CD6F508128DFF1" pageId="2" pageNumber="3">
<paragraph id="8BFF287A144D4E7FB7CD6F508128DF4A" blockId="2.[189,1399,833,964]" pageId="2" pageNumber="3">
<keyLead id="30B18B4F144D4E7FB7CD6F508128DF4A" pageId="2" pageNumber="3">
1. Vesica with a single long dorsal diverticulum bearing cornutus (
<figureCitation id="137B34FF144D4E7FB4A36F508049DF6E" box="[979,1047,833,859]" captionStart="Fig" captionStartId="14.[189,232,1893,1919]" captionTargetBox="[250,1346,269,1849]" captionTargetId="figure@14.[238,1349,266,1851]" captionTargetPageId="14" captionText="Fig. 7. Phalli with vesica everted of Phtheochroa spp. A–B. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – C–D. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. A, C: left. B, D: dorsal. Abbreviations as in Figs 5–6. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838426" httpUri="https://zenodo.org/record/3838426/files/figure.png" pageId="2" pageNumber="3">Fig. 7</figureCitation>
A–B). Phallic process straight (
<figureCitation id="137B34FF144D4E7FB7836F758511DF4B" box="[243,335,868,894]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="2" pageNumber="3">Fig. 4H</figureCitation>
), phallus almost straight .............................................................
<taxonomicName id="4C4053F9144D4E7FB3406F758128DF4A" ID-CoL="4GX97" authority="(Guenee, 1845)" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Guenee" baseAuthorityYear="1845" box="[1072,1398,867,895]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="frigidana">
<emphasis id="B934F468144D4E7FB3406F7580E5DF4B" bold="true" box="[1072,1211,867,894]" italics="true" pageId="2" pageNumber="3">P. frigidana</emphasis>
(Guenée, 1845)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFF287A144D4E7FB7B16F998128DFF1" blockId="2.[189,1399,833,964]" pageId="2" pageNumber="3">
<keyLead id="30B18B4F144D4E7FB7B16F998128DFF1" pageId="2" pageNumber="3">– Vesica with two distinct diverticula, each with cornutus. Phallic process apically curved to the right, phallus curved....................................................................................................................................2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B430DF144D4E7FB7CD6FFD8129D879" pageId="2" pageNumber="3">
<paragraph id="8BFF287A144D4E7FB7CD6FFD8129D81C" blockId="2.[189,1399,1004,1101]" pageId="2" pageNumber="3">
<keyLead id="30B18B4F144D4E7FB7CD6FFD8129D81C" pageId="2" pageNumber="3">
2. Vesica with dorsal and ventral diverticulum protruding from its small right part (
<figureCitation id="137B34FF144D4E7FB3C16FFD80A6D832" box="[1201,1272,1004,1031]" captionStart="Fig" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[262,1281,269,1845]" captionTargetId="figure@12.[253,1334,265,1851]" captionTargetPageId="12" captionText="Fig. 6. Phalli with vesica everted of Phtheochroa spp. A–B. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – C–D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype.A, C: left. B, D: dorsal. Abbreviations: dd = dorsal diverticulum; vd = ventral diverticulum; other abbreviations as in Fig. 5. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838424" httpUri="https://zenodo.org/record/3838424/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
A–B) ....... ..............................................................................................................................
<taxonomicName id="4C4053F9144D4E7FB3F6681E814FD81F" ID-CoL="B3ZCD" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[1158,1297,1039,1066]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alpinana" status="sp. nov.">
<emphasis id="B934F468144D4E7FB3F6681E814FD81F" bold="true" box="[1158,1297,1039,1066]" italics="true" pageId="2" pageNumber="3">P. alpinana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074913144D4E7FB26A681E8129D81C" box="[1306,1399,1039,1065]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BFF287A144D4E7FB7B168228129D879" blockId="2.[189,1399,1004,1101]" box="[193,1399,1074,1101]" pageId="2" pageNumber="3">
<keyLead id="30B18B4F144D4E7FB7B168228129D879" box="[193,1399,1074,1101]" pageId="2" pageNumber="3">– Vesica with right and left diverticulum protruding from its large right part..................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B430DF144D4E7FB7CD68658128D90B" pageId="2" pageNumber="3">
<paragraph id="8BFF287A144D4E7FB7CD68658129D8E0" blockId="2.[189,1399,1139,1342]" pageId="2" pageNumber="3">
<keyLead id="30B18B4F144D4E7FB7CD68658129D8E0" pageId="2" pageNumber="3">
3. Forewing upperside pale yellow, hindwing pale grey. Main part of vesica in lateral left view angular (
<figureCitation id="137B34FF144D4E7FB7836886850BD884" box="[243,341,1175,1201]" captionStart="Fig" captionStartId="14.[189,232,1893,1919]" captionTargetBox="[250,1346,269,1849]" captionTargetId="figure@14.[238,1349,266,1851]" captionTargetPageId="14" captionText="Fig. 7. Phalli with vesica everted of Phtheochroa spp. A–B. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – C–D. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. A, C: left. B, D: dorsal. Abbreviations as in Figs 5–6. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838426" httpUri="https://zenodo.org/record/3838426/files/figure.png" pageId="2" pageNumber="3">Fig. 7C</figureCitation>
). The angle between the axes of phallus and right diverticulum 120–140° (
<figureCitation id="137B34FF144D4E7FB27568868137D884" box="[1285,1385,1175,1201]" captionStart="Fig" captionStartId="14.[189,232,1893,1919]" captionTargetBox="[250,1346,269,1849]" captionTargetId="figure@14.[238,1349,266,1851]" captionTargetPageId="14" captionText="Fig. 7. Phalli with vesica everted of Phtheochroa spp. A–B. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – C–D. P. cantabriana sp. nov., Spain, Picos de Europa National Park, holotype. A, C: left. B, D: dorsal. Abbreviations as in Figs 5–6. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838426" httpUri="https://zenodo.org/record/3838426/files/figure.png" pageId="2" pageNumber="3">Fig. 7D</figureCitation>
). Phallic process slender, not strongly curved (
<figureCitation id="137B34FF144D4E7FB47568AB8713D8E1" box="[773,845,1210,1236]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
I–J) ..............................
<taxonomicName id="4C4053F9144D4E7FB32F68AB814FD8E0" ID-CoL="B3ZCG" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[1119,1297,1210,1237]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cantabriana" status="sp. nov.">
<emphasis id="B934F468144D4E7FB32F68AB814FD8E0" bold="true" box="[1119,1297,1210,1237]" italics="true" pageId="2" pageNumber="3">P. cantabriana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074913144D4E7FB26A68AA8129D8E0" box="[1306,1399,1211,1237]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BFF287A144D4E7FB7B168CF8128D90B" blockId="2.[189,1399,1139,1342]" pageId="2" pageNumber="3">
<keyLead id="30B18B4F144D4E7FB7B168CF8128D90B" pageId="2" pageNumber="3">
– Forewing upperside ground colour deep yellow, hindwing dark grey (
<figureCitation id="137B34FF144D4E7FB34A68CC8021D8C2" box="[1082,1151,1245,1271]" captionStart="Fig" captionStartId="4.[189,232,1824,1850]" captionTargetBox="[230,1357,265,1782]" captionTargetId="figure@4.[230,1357,265,1782]" captionTargetPageId="4" captionText="Fig. 1. Adults of Phtheochroa spp. A–D. P. schawerdae (Rebel, 1908) comb. nov. A. ♂, Bulgaria, Pirin Mts. B. ♂, Bulgaria, Rila Mts. C. ♀, Bulgaria, Rila Mts. D. ♂, Republic of Macedonia, Korab Mts. – E–F. P. alpinana sp. nov., France, Alpes Maritimes. E. Holotype, ♂. F. Paratype, ♀. – G. P. apenninana sp. nov., holotype, ♂, Italy, Gran Sasso National Park. – H. P. frigidana (Guenée, 1845) stat. rev., ♂, neotype of Eupoecilia frigidana, Andorra, Pyrenees. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype, ♂. J. Paratype, ♂. Scale bar = 5 mm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838414" httpUri="https://zenodo.org/record/3838414/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
A–D, G). Main part of vesica in lateral left view ovoid or subspherical. The angle between the axes of phallus and right diverticulum usually less than 120°. Phallic process robust..................................................4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B430DF144D4E7FB7CD69748128DA1A" pageId="2" pageNumber="3">
<paragraph id="8BFF287A144D4E7FB7CD69748129D9F3" blockId="2.[189,1399,1381,1584]" pageId="2" pageNumber="3">
<keyLead id="30B18B4F144D4E7FB7CD69748129D9F3" pageId="2" pageNumber="3">
4. The angle between the axes of phallus and right diverticulum a maximum of 90°. The right part of vesica large, heart-shaped in dorsal view (
<figureCitation id="137B34FF144D4E7FB47A69998735D996" box="[778,875,1416,1443]" captionStart="Fig" captionStartId="12.[189,232,1894,1920]" captionTargetBox="[262,1281,269,1845]" captionTargetId="figure@12.[253,1334,265,1851]" captionTargetPageId="12" captionText="Fig. 6. Phalli with vesica everted of Phtheochroa spp. A–B. P. alpinana sp. nov., France, Alpes Maritimes, holotype. – C–D. P. apenninana sp. nov., Italy, Gran Sasso National Park, holotype.A, C: left. B, D: dorsal. Abbreviations: dd = dorsal diverticulum; vd = ventral diverticulum; other abbreviations as in Fig. 5. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838424" httpUri="https://zenodo.org/record/3838424/files/figure.png" pageId="2" pageNumber="3">Fig. 6D</figureCitation>
). Phallic process blunt, curved (
<figureCitation id="137B34FF144D4E7FB38069998166D997" box="[1264,1336,1416,1442]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
F–G) ......................................................................................................................
<taxonomicName id="4C4053F9144D4E7FB33C69BD815DD9F0" ID-CoL="B3ZCF" authority="Zlatkov &amp; Huemer, 2017" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" box="[1100,1283,1451,1478]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="apenninana" status="sp. nov.">
<emphasis id="B934F468144D4E7FB33C69BD815DD9F0" bold="true" box="[1100,1283,1451,1478]" italics="true" pageId="2" pageNumber="3">P. apenninana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2074913144D4E7FB26369BD8129D9F3" box="[1299,1399,1452,1478]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="8BFF287A144D4E7FB7B169DE8128DA1A" blockId="2.[189,1399,1381,1584]" pageId="2" pageNumber="3">
<keyLead id="30B18B4F144D4E7FB7B169DE8128DA1A" pageId="2" pageNumber="3">
– The angle between the axes of phallus and right diverticulum 100–120°. The right part of vesica small, not heart-shaped (
<figureCitation id="137B34FF144D4E7FB55069E386F0DA39" box="[544,686,1522,1549]" captionStart="Fig" captionStartId="9.[189,232,1856,1882]" captionTargetBox="[199,1390,280,1807]" captionTargetId="figure@9.[189,1398,265,1814]" captionTargetPageId="9" captionText="Fig. 5. Phalli with vesica everted, Phtheochroa schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. –C–D. Republic of Macedonia, Korab Mts.A, C: left.B, D: dorsal.Abbreviations:gs = gonopore sclerotization; ld = left diverticulum; rd = right diverticulum; svd = small ventral diverticulum; vpp = ventral phallic process. Scale bar = 250 µm." figureDoi="http://doi.org/10.5281/zenodo.3838422" httpUri="https://zenodo.org/record/3838422/files/figure.png" pageId="2" pageNumber="3">Fig. 5B, D</figureCitation>
). Phallic process large, sharp-tipped and strongly curved (
<figureCitation id="137B34FF144D4E7FB7836A048562DA1A" box="[243,316,1557,1583]" captionStart="Fig" captionStartId="8.[189,232,1824,1850]" captionTargetBox="[189,1398,1117,1787]" captionTargetId="figure@8.[189,1398,1117,1787]" captionTargetPageId="8" captionText="Fig. 4. Ventral phallic process of Phtheochroa spp., ventral view. A–C. P. schawerdae (Rebel, 1908) comb. nov. A–B. Bulgaria, Rila Mts. C. Republic of Macedonia, Korab Mts. – D–E. P. alpinana sp. nov., France, Alpes Maritimes. D. Holotype. E. Paratype. – F–G. P. apenninana sp. nov., Italy, Gran Sasso National Park. F. Paratype. G. Holotype. – H. P. frigidana (Guenée, 1845) stat. rev., Andorra, Pyrenees, neotype of Eupoecilia frigidana. – I–J. P. cantabriana sp. nov., Spain, Picos de Europa National Park. I. Holotype. J. Paratype. Scale bar = 100 µm, all to scale." figureDoi="http://doi.org/10.5281/zenodo.3838420" httpUri="https://zenodo.org/record/3838420/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
A–C) .........................................................................................
<taxonomicName id="4C4053F9144D4E7FB3666A048128DA1A" ID-CoL="92YKZ" authority="(Rebel, 1908)" authorityName="Zlatkov &amp; Huemer" authorityYear="2017" baseAuthorityName="Rebel" baseAuthorityYear="1908" box="[1046,1398,1557,1584]" class="Insecta" family="Tortricidae" genus="Phtheochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="schawerdae">
<emphasis id="B934F468144D4E7FB3666A04809BDA05" bold="true" box="[1046,1221,1557,1584]" italics="true" pageId="2" pageNumber="3">P. schawerdae</emphasis>
(Rebel, 1908)
</taxonomicName>
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</paragraph>
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</key>
</subSubSection>
</treatment>
</document>
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