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<document id="25B7B72BD77B2F943A7009425E15A833" ID-CLB-Dataset="8703" ID-DOI="10.11646/zootaxa.4825.1.1" ID-GBIF-Dataset="5ab5f5c8-481e-4d1a-8643-21e72c367278" ID-ISSN="1175-5326" ID-Zenodo-Dep="4402145" ID-ZooBank="F230F199-1C94-4E2E-9CE4-5F56212C015F" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609341789210" checkinUser="plazi" docAuthor="Schileyko, Arkady A., Vahtera, Varpu &amp; Edgecombe, Gregory D." docDate="2020" docId="03DE092DFFDFD73BFF13FA08287BDD7B" docLanguage="en" docName="zootaxa.4825.1.1.pdf" docOrigin="Zootaxa 4825 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ethmostigmus Pocock 1898" docType="treatment" docVersion="5" lastPageNumber="50" masterDocId="FFE77155FFF2D70AFF84FFE12D45DC1F" masterDocTitle="An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses" masterLastPageNumber="64" masterPageNumber="1" pageNumber="46" updateTime="1698852004925" updateUser="ExternalLinkService">
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<mods:title id="0C7DF5E4003FAE0A558B12A3351868A7">An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses</mods:title>
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<mods:namePart id="B7710BC2C6649C77C0A42B81D5BAC6CD">Schileyko, Arkady A.</mods:namePart>
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<mods:namePart id="B672F595763C96A6BA27FCBB941A5CDC">Vahtera, Varpu</mods:namePart>
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<mods:namePart id="BE86EAA4EF665CE7F87213F77907764A">Edgecombe, Gregory D.</mods:namePart>
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<treatment id="03DE092DFFDFD73BFF13FA08287BDD7B" ID-DOI="http://doi.org/10.5281/zenodo.4457019" ID-GBIF-Taxon="177020726" ID-Zenodo-Dep="4457019" LSID="urn:lsid:plazi:treatment:03DE092DFFDFD73BFF13FA08287BDD7B" httpUri="http://treatment.plazi.org/id/03DE092DFFDFD73BFF13FA08287BDD7B" lastPageId="49" lastPageNumber="50" pageId="45" pageNumber="46">
<subSubSection id="C36DEBB0FFDFD727FF13FA082C74DA37" pageId="45" pageNumber="46" type="discussion">
<subSubSection id="C36DEBB0FFDFD727FF13FA082F50DA1C" box="[151,533,1513,1540]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8BC8B83BFFDFD727FF13FA082F50DA1C" blockId="45.[151,533,1513,1576]" box="[151,533,1513,1540]" pageId="45" pageNumber="46">
<emphasis id="B9036429FFDFD727FF13FA082F50DA1C" bold="true" box="[151,533,1513,1540]" pageId="45" pageNumber="46">
(!)
<taxonomicName id="4C77C3B8FFDFD727FF38FA082F50DA1C" ID-CoL="62RHN" ID-ENA="62612" authority="Pocock, 1898" authorityName="Pocock" authorityYear="1898" box="[188,533,1513,1540]" class="Chilopoda" family="Scolopendridae" genus="Ethmostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFDFD727FF38FA082C2FDA1B" bold="true" box="[188,362,1513,1540]" italics="true" pageId="45" pageNumber="46">Ethmostigmus</emphasis>
Pocock, 1898
</taxonomicName>
</emphasis>
</paragraph>
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<subSubSection id="C36DEBB0FFDFD727FF13F9EC2C74DA37" box="[151,305,1549,1576]" pageId="45" pageNumber="46" type="description">
<paragraph id="8BC8B83BFFDFD727FF13F9EC2C74DA37" blockId="45.[151,533,1513,1576]" box="[151,305,1549,1576]" pageId="45" pageNumber="46">
<figureCitation id="134CA4BEFFDFD727FF13F9EC2C74DA37" box="[151,305,1549,1576]" captionStart-0="FIGURES 97101" captionStart-1="FIGURES 102107" captionStartId-0="48.[151,264,1693,1717]" captionStartId-1="50.[151,264,1685,1709]" captionTargetBox-0="[171,1417,200,1647]" captionTargetBox-1="[274,1314,198,1647]" captionTargetId-0="figure-19@48.[151,1436,181,1669]" captionTargetId-1="figure-19@50.[257,1331,181,1661]" captionTargetPageId-0="48" captionTargetPageId-1="50" captionText-0="FIGURES 97101. Otostigmus (Parotostigmus) scabricauda (Humbert et Saussure, 1870); Ad. male Rc 6730 97 LBS 21 and prefemora + femora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype male MNHN 460 98 posterior margin of ultimate tergite + prefemora of ultimate legs dorsally; Otostigmus (P.) diringshofeni Bücherl, 1969; Ad. male Rc 7981 99 LBS 21 + prefemora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype male MNHN 460 100 anterior margin of forcipular coxosternite + maxillae 2 ventrally; Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 101 left side of ultimate LBS + ultimate legs laterally; (as)—accessory spine, (bs)—basal suture of forcipular tooth-plate, (cp)— practically reduced spineless coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (dg)—digitiform process of ultimate prefemur, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pst)—penultimate sternite, (pts)—ultimate pretarsus, (su)—ventral spines of ultimate prefemur, (tp)—forcipular tooth-plate, (ut)—ultimate tergite." captionText-1="FIGURES 102107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi-0="http://doi.org/10.5281/zenodo.4402189" figureDoi-1="http://doi.org/10.5281/zenodo.4402191" httpUri-0="https://zenodo.org/record/4402189/files/figure.png" httpUri-1="https://zenodo.org/record/4402191/files/figure.png" pageId="45" pageNumber="46">Figs 101105</figureCitation>
</paragraph>
</subSubSection>
</subSubSection>
<subSubSection id="C36DEBB0FFDFD727FF13F9B32938DA75" box="[151,1149,1618,1642]" pageId="45" pageNumber="46" type="type_taxon">
<paragraph id="8BC8B83BFFDFD727FF13F9B32938DA75" blockId="45.[151,1149,1618,1642]" box="[151,1149,1618,1642]" pageId="45" pageNumber="46">
<emphasis id="B9036429FFDFD727FF13F9B32C66DA75" bold="true" box="[151,291,1618,1642]" pageId="45" pageNumber="46">
<typeStatus id="54CC0699FFDFD727FF13F9B32D89DA75" box="[151,204,1618,1642]" pageId="45" pageNumber="46">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C77C3B8FFDFD727FEAEF9B32FF4DA75" authority="Leach, 1817" authorityName="Leach" authorityYear="1817" box="[298,689,1618,1642]" class="Chilopoda" family="Scolopendridae" genus="Scolopendra" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="trigonopoda">
<emphasis id="B9036429FFDFD727FEAEF9B32F68DA75" box="[298,557,1618,1642]" italics="true" pageId="45" pageNumber="46">Scolopendra trigonopoda</emphasis>
Leach, 1817
</taxonomicName>
(by subsequent designation of
<bibRefCitation id="EFE6C5CAFFDFD727FC6FF9B32937DA75" author="Attems, C." box="[1003,1138,1618,1642]" pageId="45" pageNumber="46" refId="ref45522" refString="Attems, C. (1930) Myriapoda 2. Scolopendromorpha. Das Tierreich. Vol. 54. Walter de Gruyter et Co. Publ., Berlin, 308 pp." type="book" year="1930">Attems 1930</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFDFD73BFF13F979287BDD7B" lastPageId="49" lastPageNumber="50" pageId="45" pageNumber="46" type="diagnosis">
<subSubSection id="C36DEBB0FFDFD727FF13F9792F09DBE9" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="8BC8B83BFFDFD727FF13F9792F09DBE9" blockId="45.[151,1437,1688,2039]" pageId="45" pageNumber="46">
<emphasis id="B9036429FFDFD727FF13F9792C57DAAD" bold="true" box="[151,274,1688,1714]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
Median tooth of labrum well developed. Forcipular tooth-plates present, trochantero-prefemoral processes virtually absent (
<figureCitation id="134CA4BEFFDFD727FE19F95D2CBBDAC9" box="[413,510,1724,1750]" captionStart="FIGURES 102107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="45" pageNumber="46">Fig. 102</figureCitation>
). Tergites consistently lacking longitudinal keels. Sternites with paramedian sutures and/or sulci both developed to varying degrees (figs
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<bibRefCitation id="EFE6C5CAFFDFD727FCEAF90129CBDAE4" author="Schileyko, A. &amp; Stoev, P." box="[878,1166,1760,1787]" pageId="45" pageNumber="46" pagination="247 - 280" refId="ref50391" refString="Schileyko, A. &amp; Stoev, P. (2016) Scolopendromorpha of New Guinea and adjacent islands (Myriapoda, Chilopoda). Zootaxa, 4147 (3), 247 - 280. https: // doi. org / 10.11646 / zootaxa. 4147.3.3" type="journal article" year="2016">Schileyko &amp; Stoev 2016</bibRefCitation>
). LBS 7 with spiracles, spiracles virtually without atrium (
<figureCitation id="134CA4BEFFDFD727FD92F8E52F32DB01" box="[534,631,1796,1822]" captionStart="FIGURES 102107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="45" pageNumber="46">Fig. 103</figureCitation>
). Legs with tarsal spur(s). Coxopleural process (
<figureCitation id="134CA4BEFFDFD727FB08F8E529AFDB00" box="[1164,1258,1796,1823]" captionStart="FIGURES 102107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="45" pageNumber="46">Fig. 105</figureCitation>
) from moderate (fig.
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Schileyko &amp; Stagl) to very long and large (
<figureCitation id="134CA4BEFFDFD727FD74F8C92E17DB5D" box="[752,850,1832,1858]" captionStart="FIGURES 102107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="45" pageNumber="46">Fig. 104</figureCitation>
), only short in some Australian species (e.g.,
<taxonomicName id="4C77C3B8FFDFD727FACBF8C82C12DB79" authority="Koch, 1983" authorityName="Koch" authorityYear="1983" class="Chilopoda" family="Scolopendridae" genus="Ethmostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="curtipes">
<emphasis id="B9036429FFDFD727FACBF8C82D89DB79" italics="true" pageId="45" pageNumber="46">E. curtipes</emphasis>
<bibRefCitation id="EFE6C5CAFFDFD727FF56F8AD2C12DB79" author="Koch, L. E." box="[210,343,1868,1894]" pageId="45" pageNumber="46" pagination="799 - 833" refId="ref48255" refString="Koch, L. E. (1983) Revision of the Australian centipedes of the genus Cormocephalus Newport (Chilopoda: Scolopendridae: Scolopendrinae). Australian Journal of Zoology, 31, 799 - 833." type="journal article" year="1983">Koch, 1983</bibRefCitation>
</taxonomicName>
). Ultimate legs of “common” shape (
<figureCitation id="134CA4BEFFDFD727FD74F8AD2E15DB79" box="[752,848,1868,1894]" captionStart="FIGURES 97101" captionStartId="48.[151,264,1693,1717]" captionTargetBox="[171,1417,200,1647]" captionTargetId="figure-19@48.[151,1436,181,1669]" captionTargetPageId="48" captionText="FIGURES 97101. Otostigmus (Parotostigmus) scabricauda (Humbert et Saussure, 1870); Ad. male Rc 6730 97 LBS 21 and prefemora + femora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype male MNHN 460 98 posterior margin of ultimate tergite + prefemora of ultimate legs dorsally; Otostigmus (P.) diringshofeni Bücherl, 1969; Ad. male Rc 7981 99 LBS 21 + prefemora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype male MNHN 460 100 anterior margin of forcipular coxosternite + maxillae 2 ventrally; Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 101 left side of ultimate LBS + ultimate legs laterally; (as)—accessory spine, (bs)—basal suture of forcipular tooth-plate, (cp)— practically reduced spineless coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (dg)—digitiform process of ultimate prefemur, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pst)—penultimate sternite, (pts)—ultimate pretarsus, (su)—ventral spines of ultimate prefemur, (tp)—forcipular tooth-plate, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402189" httpUri="https://zenodo.org/record/4402189/files/figure.png" pageId="45" pageNumber="46">Fig. 101</figureCitation>
), but in a few species—for example in
<taxonomicName id="4C77C3B8FFDFD727FB7AF8AD2F44DB94" authority="rubripes (Brandt, 1840)" authorityName="rubripes (Brandt" authorityYear="1840" baseAuthorityName="Brandt" baseAuthorityYear="1840" class="Chilopoda" family="Scolopendridae" genus="Ethmostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="45" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="rubripes" subSpecies="rubripes">
<emphasis id="B9036429FFDFD727FB7AF8AD2C1EDB95" italics="true" pageId="45" pageNumber="46">Ethmostigmus rubripes rubripes</emphasis>
(Brandt, 1840)
</taxonomicName>
—these legs may be much shortened and broadened (becoming pracrically “pincershaped”) as a result of geographic variability (see
<bibRefCitation id="EFE6C5CAFFDFD727FD4EF8752956DBB1" author="Schileyko, A. &amp; Stagl, V." box="[714,1043,1940,1966]" pageId="45" pageNumber="46" pagination="67 - 137" refId="ref50342" refString="Schileyko, A. &amp; Stagl, V. (2004) The collection of scolopendromorph centipedes (Chilopoda) in the Natural History Museum in Vienna: a critical re-evaluation of former taxonomic identifications. Annalen des Naturhistorischen Museum Wien, 105 Β, 67 - 137." type="journal article" year="2004">Schileyko &amp; Stagl 2004: 120</bibRefCitation>
). Prefemur of the ultimate leg with spines plus well-developed corner spine (
<figureCitation id="134CA4BEFFDFD727FDE8F8592F97DBCD" box="[620,722,1976,2002]" captionStart="FIGURES 97101" captionStartId="48.[151,264,1693,1717]" captionTargetBox="[171,1417,200,1647]" captionTargetId="figure-19@48.[151,1436,181,1669]" captionTargetPageId="48" captionText="FIGURES 97101. Otostigmus (Parotostigmus) scabricauda (Humbert et Saussure, 1870); Ad. male Rc 6730 97 LBS 21 and prefemora + femora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype male MNHN 460 98 posterior margin of ultimate tergite + prefemora of ultimate legs dorsally; Otostigmus (P.) diringshofeni Bücherl, 1969; Ad. male Rc 7981 99 LBS 21 + prefemora of ultimate legs ventrally; Otostigmus (P.) salticus Schileyko, Iorio &amp; Coulis, 2018; Ad. holotype male MNHN 460 100 anterior margin of forcipular coxosternite + maxillae 2 ventrally; Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 101 left side of ultimate LBS + ultimate legs laterally; (as)—accessory spine, (bs)—basal suture of forcipular tooth-plate, (cp)— practically reduced spineless coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (dg)—digitiform process of ultimate prefemur, (p2)—pretarsus of maxilla 2, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pst)—penultimate sternite, (pts)—ultimate pretarsus, (su)—ventral spines of ultimate prefemur, (tp)—forcipular tooth-plate, (ut)—ultimate tergite." figureDoi="http://doi.org/10.5281/zenodo.4402189" httpUri="https://zenodo.org/record/4402189/files/figure.png" pageId="45" pageNumber="46">Figs 101</figureCitation>
,
<figureCitation id="134CA4BEFFDFD727FD5BF8592E48DBCC" box="[735,781,1976,2003]" captionStart="FIGURES 102107" captionStartId="50.[151,264,1685,1709]" captionTargetBox="[274,1314,198,1647]" captionTargetId="figure-19@50.[257,1331,181,1661]" captionTargetPageId="50" captionText="FIGURES 102107. Ethmostigmus rubripes spinosus (Newport, 1845); Juv. Rc 7174 102 forcipular segment ventrally; Ethmostigmus rubripes platycephalus (Newport, 1845); Ad. Rc 7514 103 left pleuron of LBS 3 laterally; Ad. Rc 7513 104 LBS 21 ventrally; Sad. СDT 105 coxopleural processes + prefemora of ultimate legs ventrally; Rhysida longipes (Newport, 1845); Ad. Rc 6683 106 anterior margin of forcipular coxosternite ventrally 107 coxopleural processes, legs 20 + ultimate legs laterally; (ap)—apical spine, (bs)—basal suture of forcipular tooth-plate, (cp)—coxopleural process, (csp)—corner spine of ultimate prefemur, (cx)—coxopleuron, (im)—intersclerite membrane, (lc)—lateral spines of coxopleural process, (pf)—ultimate prefemur, (pr)—trochantero-prefemoral process, (pts)—ultimate pretarsus, (rpf)—abnormally small (regenerated) right ultimate prefemur, (rpr)—rudiment of trochantero-prefemoral process, (s)—open spiracle without atrium, (scs)— spine at place of corner spine of ultimate prefemur, (su)—spines of ultimate prefemur, (t1)—ultimate tarsus 1, (tp)—forcipular tooth-plates, (ust)—ultimate sternite." figureDoi="http://doi.org/10.5281/zenodo.4402191" httpUri="https://zenodo.org/record/4402191/files/figure.png" pageId="45" pageNumber="46">105</figureCitation>
); claw-shaped pretarsus in most species neither elongated nor enlarged, rarely as long as tarsus 2.
</paragraph>
</subSubSection>
<caption id="DF08E8B3FFDCD724FF13F9192CC8DBF6" ID-DOI="http://doi.org/10.5281/zenodo.4402183" ID-Zenodo-Dep="4402183" httpUri="https://zenodo.org/record/4402183/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,264,1784,1808]" targetBox="[186,1400,200,1740]" targetPageId="46">
<paragraph id="8BC8B83BFFDCD724FF13F9192CC8DBF6" blockId="46.[151,1437,1784,2025]" pageId="46" pageNumber="47">
<emphasis id="B9036429FFDCD724FF13F9192C13DB0F" bold="true" box="[151,342,1784,1808]" pageId="46" pageNumber="47">FIGURES 8488.</emphasis>
<taxonomicName id="4C77C3B8FFDCD724FED9F9182FE6DB0F" authority="Imhoff, 1854" authorityName="Imhoff" authorityYear="1854" box="[349,675,1784,1809]" class="Chilopoda" family="Scolopendridae" genus="Alipes" kingdom="Animalia" order="Scolopendromorpha" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="multicostis">
<emphasis id="B9036429FFDCD724FED9F9182F57DB0E" box="[349,530,1785,1809]" italics="true" pageId="46" pageNumber="47">Alipes multicostis</emphasis>
Imhoff, 1854
</taxonomicName>
; Live specimen CDT (photo by Dmitry Telnov)
<emphasis id="B9036429FFDCD724FB13F91929F5DB0F" bold="true" box="[1175,1200,1784,1808]" pageId="46" pageNumber="47">84</emphasis>
general view dorsally;
<taxonomicName id="4C77C3B8FFDCD724FF13F8FC2C9DDB2A" authority="Lucas, 1864" authorityName="Lucas" authorityYear="1864" box="[151,472,1821,1845]" class="Chilopoda" family="Scolopendridae" genus="Alipes" kingdom="Animalia" order="Scolopendromorpha" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="grandidieri">
<emphasis id="B9036429FFDCD724FF13F8FC2C14DB2A" box="[151,337,1821,1845]" italics="true" pageId="46" pageNumber="47">Alipes grandidieri</emphasis>
Lucas, 1864
</taxonomicName>
; BMNH 1959.3.8
<emphasis id="B9036429FFDCD724FD12F8FD2FEADB2B" bold="true" box="[662,687,1820,1844]" pageId="46" pageNumber="47">85</emphasis>
distal part of ultimate leg laterally;
<taxonomicName id="4C77C3B8FFDCD724FBA6F8FC28DCDB2A" authority="Pocock, 1896" authorityName="Pocock" authorityYear="1896" box="[1058,1433,1821,1845]" class="Chilopoda" family="Scolopendridae" genus="Alipes" kingdom="Animalia" order="Scolopendromorpha" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="appendiculatus">
<emphasis id="B9036429FFDCD724FBA6F8FC2846DB2A" box="[1058,1283,1821,1845]" italics="true" pageId="46" pageNumber="47">Alipes appendiculatus</emphasis>
Pocock, 1896
</taxonomicName>
; BMNH1950.3.9.2
<emphasis id="B9036429FFDCD724FEDCF8A12C34DB47" bold="true" box="[344,369,1856,1880]" pageId="46" pageNumber="47">86</emphasis>
forcipular segment ventrally
<emphasis id="B9036429FFDCD724FD2EF8A12F86DB47" bold="true" box="[682,707,1856,1880]" pageId="46" pageNumber="47">87</emphasis>
LBS 410 dorsally;
<taxonomicName id="4C77C3B8FFDCD724FC26F8A029AFDB46" authority="Lucas, 1864" authorityName="Lucas" authorityYear="1864" box="[930,1258,1857,1881]" class="Chilopoda" family="Scolopendridae" genus="Alipes" kingdom="Animalia" order="Scolopendromorpha" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="grandidieri">
<emphasis id="B9036429FFDCD724FC26F8A0291BDB46" box="[930,1118,1857,1881]" italics="true" pageId="46" pageNumber="47">Alipes grandidieri</emphasis>
Lucas, 1864
</taxonomicName>
; BMNH1959.3.8
<emphasis id="B9036429FFDCD724FF13F8852DF5DB63" bold="true" box="[151,176,1892,1916]" pageId="46" pageNumber="47">88</emphasis>
LBS 21 + prefemora of ultimate legs ventrally; (
<emphasis id="B9036429FFDCD724FD45F8842F9CDB62" bold="true" box="[705,729,1893,1917]" pageId="46" pageNumber="47">bs</emphasis>
)—basal suture of forcipular tooth-plate, (
<emphasis id="B9036429FFDCD724FB17F88429E9DB62" bold="true" box="[1171,1196,1893,1917]" pageId="46" pageNumber="47">cp</emphasis>
)—very short spineless coxopleural process, (
<emphasis id="B9036429FFDCD724FEF0F8682CC9DBBE" bold="true" box="[372,396,1929,1953]" pageId="46" pageNumber="47">cx</emphasis>
)—coxopleuron, (
<emphasis id="B9036429FFDCD724FDC4F8692F12DBBF" bold="true" box="[576,599,1928,1952]" pageId="46" pageNumber="47">pf</emphasis>
)—ultimate prefemur, (
<emphasis id="B9036429FFDCD724FCC5F8682E1FDBBE" bold="true" box="[833,858,1929,1953]" pageId="46" pageNumber="47">pr</emphasis>
)—trochantero-prefemoral process, (
<emphasis id="B9036429FFDCD724FB48F86929A7DBBF" bold="true" box="[1228,1250,1928,1952]" pageId="46" pageNumber="47">t1</emphasis>
)—flattened tarsus 1, (
<emphasis id="B9036429FFDCD724FF38F84D2D94DBDB" bold="true" box="[188,209,1964,1988]" pageId="46" pageNumber="47">t2</emphasis>
)—flattened tarsus 2, (
<emphasis id="B9036429FFDCD724FE3AF84C2C99DBDA" bold="true" box="[446,476,1965,1989]" pageId="46" pageNumber="47">T5</emphasis>
)—terite 5, (
<emphasis id="B9036429FFDCD724FDE4F84D2F2ADBDB" bold="true" box="[608,623,1964,1988]" pageId="46" pageNumber="47">ti</emphasis>
)—flattened tibia, (
<emphasis id="B9036429FFDCD724FCBCF84C2E0ADBDA" bold="true" box="[824,847,1965,1989]" pageId="46" pageNumber="47">tk</emphasis>
)—tergal longitudinal keels, (
<emphasis id="B9036429FFDCD724FB07F84C29DFDBDA" bold="true" box="[1155,1178,1965,1989]" pageId="46" pageNumber="47">tp</emphasis>
)—forcipular tooth-plate, (
<emphasis id="B9036429FFDCD724FF24F8302D85DBF6" bold="true" box="[160,192,2001,2025]" pageId="46" pageNumber="47">ust</emphasis>
)—ultimate sternite.
</paragraph>
</caption>
<caption id="DF08E8B3FFDDD725FF13F95A2EBBDBEC" ID-DOI="http://doi.org/10.5281/zenodo.4402185" ID-Zenodo-Dep="4402185" httpUri="https://zenodo.org/record/4402185/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,264,1723,1747]" targetBox="[181,1399,207,1674]" targetPageId="47">
<paragraph id="8BC8B83BFFDDD725FF13F95A2EBBDBEC" blockId="47.[151,1437,1723,2036]" pageId="47" pageNumber="48">
<emphasis id="B9036429FFDDD725FF13F95A2C11DACC" bold="true" box="[151,340,1723,1747]" pageId="47" pageNumber="48">FIGURES 8996.</emphasis>
<taxonomicName id="4C77C3B8FFDDD725FEDDF95D2E55DACB" authority="Jangi &amp; Dass, 1984" authorityName="Jangi &amp; Dass" authorityYear="1984" box="[345,784,1723,1748]" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="coonoorensis">
<emphasis id="B9036429FFDDD725FEDDF95D2F07DACC" box="[345,578,1724,1747]" italics="true" pageId="47" pageNumber="48">Digitipes coonoorensis</emphasis>
Jangi &amp; Dass, 1984
</taxonomicName>
; Male CES 091088
<emphasis id="B9036429FFDDD725FC5DF95A2EB7DACC" bold="true" box="[985,1010,1723,1747]" pageId="47" pageNumber="48">89</emphasis>
femur of right ultimate leg medially;
<taxonomicName id="4C77C3B8FFDDD725FAEAF95D2D8BDAE8" class="Chilopoda" family="Scolopendridae" genus="Digitipes" kingdom="Animalia" order="Scolopendromorpha" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFDDD725FAEAF95D2D8BDAE8" italics="true" pageId="47" pageNumber="48">Digitipes</emphasis>
</taxonomicName>
sp.; Ad. Rc 7565
<emphasis id="B9036429FFDDD725FE0DF93E2CE7DAE8" bold="true" box="[393,418,1759,1783]" pageId="47" pageNumber="48">90</emphasis>
clypeal part of head ventrally
<emphasis id="B9036429FFDDD725FD58F93E2FB0DAE8" bold="true" box="[732,757,1759,1783]" pageId="47" pageNumber="48">91</emphasis>
forcipular segment ventrally
<emphasis id="B9036429FFDDD725FBA0F93E2978DAE8" bold="true" box="[1060,1085,1759,1783]" pageId="47" pageNumber="48">92</emphasis>
LBS 1315 ventrally;
<taxonomicName id="4C77C3B8FFDDD725FAADF90128D9DAE8" box="[1321,1436,1760,1783]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFDDD725FAADF90128D9DAE8" box="[1321,1436,1760,1783]" italics="true" pageId="47" pageNumber="48">Otostigmus</emphasis>
</taxonomicName>
<taxonomicName id="4C77C3B8FFDDD725FF13F8E52C69DB04" authorityName="Haase" authorityYear="1887" box="[151,300,1795,1820]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="multidens" subGenus="Otostigmus">
(
<emphasis id="B9036429FFDDD725FF24F8E52DF7DB04" box="[160,178,1796,1819]" italics="true" pageId="47" pageNumber="48">O</emphasis>
.)
<emphasis id="B9036429FFDDD725FF4DF8E22C69DB04" box="[201,300,1795,1819]" italics="true" pageId="47" pageNumber="48">multidens</emphasis>
</taxonomicName>
<emphasis id="B9036429FFDDD725FEB7F8E22CCEDB04" box="[307,395,1795,1819]" italics="true" pageId="47" pageNumber="48">nordicus</emphasis>
<bibRefCitation id="EFE6C5CAFFDDD725FE17F8E22F79DB04" author="Schileyko" box="[403,572,1795,1819]" pageId="47" pageNumber="48" pagination="73 - 87" refId="ref49933" refString="Schileyko, Ark. A. (1995) Scolopendromorph centipedes of Viet-Nam (Chilopoda) Part 2. Arthropoda Selecta, 4 (2), 73 - 87." type="journal article" year="1995">Schileyko, 1995</bibRefCitation>
; Ad. Rc 6545
<emphasis id="B9036429FFDDD725FD57F8E22FA9DB04" bold="true" box="[723,748,1795,1819]" pageId="47" pageNumber="48">93</emphasis>
LBS 21 ventrally; Ad. Rc 6747
<emphasis id="B9036429FFDDD725FBBBF8E2291DDB04" bold="true" box="[1087,1112,1795,1819]" pageId="47" pageNumber="48">94</emphasis>
anterior part of head ventrally;
<taxonomicName id="4C77C3B8FFDDD725FF13F8C92F27DB5F" authority="(Kohlrausch, 1881)" baseAuthorityName="Kohlrausch" baseAuthorityYear="1881" box="[151,610,1831,1856]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="astenus" subGenus="Otostigmus">
<emphasis id="B9036429FFDDD725FF13F8C92C4FDB20" box="[151,266,1832,1855]" italics="true" pageId="47" pageNumber="48">Otostigmus</emphasis>
(
<emphasis id="B9036429FFDDD725FE9FF8C92C71DB20" box="[283,308,1832,1855]" italics="true" pageId="47" pageNumber="48">O.</emphasis>
)
<emphasis id="B9036429FFDDD725FEC0F8C92CD5DB20" box="[324,400,1832,1855]" italics="true" pageId="47" pageNumber="48">astenus</emphasis>
(Kohlrausch, 1881)
</taxonomicName>
; MCZ DNA 102463
<emphasis id="B9036429FFDDD725FCBBF8C62E1DDB20" bold="true" box="[831,856,1831,1855]" pageId="47" pageNumber="48">95</emphasis>
pretarsus of maxilla 2; Sad. Rc 7507
<emphasis id="B9036429FFDDD725FB61F8C629BBDB20" bold="true" box="[1253,1278,1831,1855]" pageId="47" pageNumber="48">96</emphasis>
forcipular segment ventrally; (
<emphasis id="B9036429FFDDD725FEC3F8AD2C27DB7B" bold="true" box="[327,354,1868,1892]" pageId="47" pageNumber="48">ap</emphasis>
)—apical spines of coxopleural process, (
<emphasis id="B9036429FFDDD725FC9CF8AD2E6ADB7B" bold="true" box="[792,815,1868,1892]" pageId="47" pageNumber="48">as</emphasis>
)—accessory spines, (
<emphasis id="B9036429FFDDD725FB90F8AD2969DB7B" bold="true" box="[1044,1068,1868,1892]" pageId="47" pageNumber="48">bs</emphasis>
)—basal suture of forcipular toothplate, (
<emphasis id="B9036429FFDDD725FF58F8912DABDB97" bold="true" box="[220,238,1904,1928]" pageId="47" pageNumber="48">cl</emphasis>
)—clypeus, (
<emphasis id="B9036429FFDDD725FEF6F8912CCFDB97" bold="true" box="[370,394,1904,1928]" pageId="47" pageNumber="48">cx</emphasis>
)—coxopleuron, (
<emphasis id="B9036429FFDDD725FDBAF88E2F10DB98" bold="true" box="[574,597,1903,1927]" pageId="47" pageNumber="48">df</emphasis>
)—conical distomedial process of ultimate femur, (
<emphasis id="B9036429FFDDD725FBD7F88E291EDB98" bold="true" box="[1107,1115,1903,1927]" pageId="47" pageNumber="48">f</emphasis>
)—ultimate femur, (
<emphasis id="B9036429FFDDD725FAA1F8912869DB97" bold="true" box="[1317,1324,1904,1928]" pageId="47" pageNumber="48">l</emphasis>
)—labrum, (
<emphasis id="B9036429FFDDD725FF24F8752DF7DBB3" bold="true" box="[160,178,1940,1964]" pageId="47" pageNumber="48">lc</emphasis>
)—lateral spine of coxopleural process, (
<emphasis id="B9036429FFDDD725FDDBF8752F3EDBB3" bold="true" box="[607,635,1940,1964]" pageId="47" pageNumber="48">lsc</emphasis>
)—sternal median longitudinal sulcus, (
<emphasis id="B9036429FFDDD725FB9DF8752979DBB3" bold="true" box="[1049,1084,1940,1964]" pageId="47" pageNumber="48">md</emphasis>
)—medial longitudinal depression of ultimate femur, (
<emphasis id="B9036429FFDDD725FEECF8562CC6DBD0" bold="true" box="[360,387,1975,1999]" pageId="47" pageNumber="48">p2</emphasis>
)—pretarsus of maxilla 2, (
<emphasis id="B9036429FFDDD725FD21F8592F84DBCF" bold="true" box="[677,705,1976,2000]" pageId="47" pageNumber="48">pd</emphasis>
)—sternal posterior depression, (
<emphasis id="B9036429FFDDD725FB9EF8592976DBCF" bold="true" box="[1050,1075,1976,2000]" pageId="47" pageNumber="48">pr</emphasis>
)—trochantero-prefemoral process, (
<emphasis id="B9036429FFDDD725FF24F83D2DFADBEB" bold="true" box="[160,191,2012,2036]" pageId="47" pageNumber="48">psl</emphasis>
)—sternal paramedian sulci, (
<emphasis id="B9036429FFDDD725FE68F83D2F46DBEB" bold="true" box="[492,515,2012,2036]" pageId="47" pageNumber="48">tp</emphasis>
)—forcipular tooth-plate, (
<emphasis id="B9036429FFDDD725FC95F83D2E74DBEB" bold="true" box="[785,817,2012,2036]" pageId="47" pageNumber="48">ust</emphasis>
)—ultimate sternite.
</paragraph>
</caption>
<caption id="DF08E8B3FFC2D73AFF13F97C2E84DBE6" ID-DOI="http://doi.org/10.5281/zenodo.4402189" ID-Zenodo-Dep="4402189" httpUri="https://zenodo.org/record/4402189/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,264,1693,1717]" targetBox="[171,1417,200,1647]" targetPageId="48">
<paragraph id="8BC8B83BFFC2D73AFF13F97C2E84DBE6" blockId="48.[151,1437,1693,2041]" pageId="48" pageNumber="49">
<emphasis id="B9036429FFC2D73AFF13F97C2C24DAAA" bold="true" box="[151,353,1693,1717]" pageId="48" pageNumber="49">FIGURES 97101.</emphasis>
<taxonomicName id="4C77C3B8FFC2D73AFEECF97C297DDAAA" authority="(Humbert et Saussure, 1870)" baseAuthorityName="Humbert et Saussure" baseAuthorityYear="1870" box="[360,1080,1693,1717]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="scabricauda" subGenus="Parotostigmus">
<emphasis id="B9036429FFC2D73AFEECF97C2C9EDAAB" box="[360,475,1693,1716]" italics="true" pageId="48" pageNumber="49">Otostigmus</emphasis>
(
<emphasis id="B9036429FFC2D73AFE6EF97F2F38DAAA" box="[490,637,1694,1717]" italics="true" pageId="48" pageNumber="49">Parotostigmus</emphasis>
)
<emphasis id="B9036429FFC2D73AFD08F97C2E4CDAAA" box="[652,777,1693,1717]" italics="true" pageId="48" pageNumber="49">scabricauda</emphasis>
(Humbert et Saussure, 1870)
</taxonomicName>
; Ad. male Rc 6730
<emphasis id="B9036429FFC2D73AFA86F97C285EDAAA" bold="true" box="[1282,1307,1693,1717]" pageId="48" pageNumber="49">97</emphasis>
LBS 21 and prefemora + femora of ultimate legs ventrally;
<taxonomicName id="4C77C3B8FFC2D73AFD07F9202994DAC6" authority="Schileyko, Iorio &amp; Coulis, 2018" authorityName="Schileyko, Iorio &amp; Coulis" authorityYear="2018" box="[643,1233,1729,1753]" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="salticus" subGenus="Parotostigmus">
<emphasis id="B9036429FFC2D73AFD07F9202FB3DAC7" box="[643,758,1729,1752]" italics="true" pageId="48" pageNumber="49">Otostigmus</emphasis>
(
<emphasis id="B9036429FFC2D73AFC83F9232E5EDAC6" box="[775,795,1730,1753]" italics="true" pageId="48" pageNumber="49">P.</emphasis>
)
<emphasis id="B9036429FFC2D73AFCAEF9202E32DAC6" box="[810,887,1729,1753]" italics="true" pageId="48" pageNumber="49">salticus</emphasis>
<bibRefCitation id="EFE6C5CAFFC2D73AFC04F9202994DAC6" author="Schileyko, A. &amp; Iorio, E. &amp; Coulis, M." box="[896,1233,1729,1753]" pageId="48" pageNumber="49" pagination="559 - 574" refId="ref50439" refString="Schileyko, A., Iorio, E. &amp; Coulis, M. (2018) A contribution to the knowledge of scolopendromorph centipedes of Martinique Island, with descriptions of two new species (Chilopoda: Scolopendromorpha). Zootaxa, 4486 (4), 559 - 574. https: // doi. org / 10.11646 / zootaxa. 4486.4.9" type="journal article" year="2018">Schileyko, Iorio &amp; Coulis, 2018</bibRefCitation>
</taxonomicName>
; Ad. holotype male MNHN 460
<emphasis id="B9036429FFC2D73AFE91F9042C6BDAE2" bold="true" box="[277,302,1765,1789]" pageId="48" pageNumber="49">98</emphasis>
posterior margin of ultimate tergite + prefemora of ultimate legs dorsally;
<taxonomicName id="4C77C3B8FFC2D73AFB9CF9042D88DB3E" authority="Bucherl, 1969" authorityName="Bucherl" authorityYear="1969" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="diringshofeni" subGenus="Parotostigmus">
<emphasis id="B9036429FFC2D73AFB9CF90429CEDAE3" box="[1048,1163,1765,1788]" italics="true" pageId="48" pageNumber="49">Otostigmus</emphasis>
(
<emphasis id="B9036429FFC2D73AFB1DF90729E8DAE2" box="[1177,1197,1766,1789]" italics="true" pageId="48" pageNumber="49">P.</emphasis>
)
<emphasis id="B9036429FFC2D73AFB3DF9042805DAE2" box="[1209,1344,1765,1789]" italics="true" pageId="48" pageNumber="49">diringshofeni</emphasis>
Bücherl, 1969
</taxonomicName>
; Ad. male Rc 7981
<emphasis id="B9036429FFC2D73AFE11F8E82CEBDB3E" bold="true" box="[405,430,1801,1825]" pageId="48" pageNumber="49">99</emphasis>
LBS 21 + prefemora of ultimate legs ventrally;
<taxonomicName id="4C77C3B8FFC2D73AFC1CF8E82D88DB5A" authority="Schileyko, Iorio &amp; Coulis, 2018" authorityName="Schileyko, Iorio &amp; Coulis" authorityYear="2018" class="Chilopoda" family="Scolopendridae" genus="Otostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="salticus" subGenus="Parotostigmus">
<emphasis id="B9036429FFC2D73AFC1CF8E8294EDB3F" box="[920,1035,1801,1824]" italics="true" pageId="48" pageNumber="49">Otostigmus</emphasis>
(
<emphasis id="B9036429FFC2D73AFB9FF8EB296BDB3E" box="[1051,1070,1802,1825]" italics="true" pageId="48" pageNumber="49">P.</emphasis>
)
<emphasis id="B9036429FFC2D73AFBB8F8E829CCDB3E" box="[1084,1161,1801,1825]" italics="true" pageId="48" pageNumber="49">salticus</emphasis>
<bibRefCitation id="EFE6C5CAFFC2D73AFB14F8E82D88DB5A" author="Schileyko, A. &amp; Iorio, E. &amp; Coulis, M." pageId="48" pageNumber="49" pagination="559 - 574" refId="ref50439" refString="Schileyko, A., Iorio, E. &amp; Coulis, M. (2018) A contribution to the knowledge of scolopendromorph centipedes of Martinique Island, with descriptions of two new species (Chilopoda: Scolopendromorpha). Zootaxa, 4486 (4), 559 - 574. https: // doi. org / 10.11646 / zootaxa. 4486.4.9" type="journal article" year="2018">Schileyko, Iorio &amp; Coulis, 2018</bibRefCitation>
</taxonomicName>
; Ad. holotype male MNHN 460
<emphasis id="B9036429FFC2D73AFD98F8CC2F07DB5A" bold="true" box="[540,578,1837,1861]" pageId="48" pageNumber="49">100</emphasis>
anterior margin of forcipular coxosternite + maxillae 2 ventrally;
<taxonomicName id="4C77C3B8FFC2D73AFB63F8CC2CAFDB76" authority="(Newport, 1845)" baseAuthorityName="Newport" baseAuthorityYear="1845" class="Chilopoda" family="Scolopendridae" genus="Ethmostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subSpecies" species="rubripes" subSpecies="spinosus">
<emphasis id="B9036429FFC2D73AFB63F8CC2C70DB76" italics="true" pageId="48" pageNumber="49">Ethmostigmus rubripes spinosus</emphasis>
(Newport, 1845)
</taxonomicName>
; Juv. Rc 7174
<emphasis id="B9036429FFC2D73AFD05F8B02FE2DB76" bold="true" box="[641,679,1873,1897]" pageId="48" pageNumber="49">101</emphasis>
left side of ultimate LBS + ultimate legs laterally; (
<emphasis id="B9036429FFC2D73AFB3AF8B02990DB76" bold="true" box="[1214,1237,1873,1897]" pageId="48" pageNumber="49">as</emphasis>
)—accessory spine, (
<emphasis id="B9036429FFC2D73AFF24F8942DFDDB92" bold="true" box="[160,184,1909,1933]" pageId="48" pageNumber="49">bs</emphasis>
)—basal suture of forcipular tooth-plate, (
<emphasis id="B9036429FFC2D73AFDECF8942FC4DB92" bold="true" box="[616,641,1909,1933]" pageId="48" pageNumber="49">cp</emphasis>
)— practically reduced spineless coxopleural process, (
<emphasis id="B9036429FFC2D73AFB38F894299ADB92" bold="true" box="[1212,1247,1909,1933]" pageId="48" pageNumber="49">csp</emphasis>
)—corner spine of ultimate prefemur, (
<emphasis id="B9036429FFC2D73AFEE7F8782C3EDBAE" bold="true" box="[355,379,1945,1969]" pageId="48" pageNumber="49">cx</emphasis>
)—coxopleuron, (
<emphasis id="B9036429FFC2D73AFDB5F8782F09DBAE" bold="true" box="[561,588,1945,1969]" pageId="48" pageNumber="49">dg</emphasis>
)—digitiform process of ultimate prefemur, (
<emphasis id="B9036429FFC2D73AFB93F8782977DBAE" bold="true" box="[1047,1074,1945,1969]" pageId="48" pageNumber="49">p2</emphasis>
)—pretarsus of maxilla 2, (
<emphasis id="B9036429FFC2D73AFACCF878281ADBAE" bold="true" box="[1352,1375,1945,1969]" pageId="48" pageNumber="49">pf</emphasis>
)—ultimate prefemur, (
<emphasis id="B9036429FFC2D73AFED7F85C2C29DBCA" bold="true" box="[339,364,1981,2005]" pageId="48" pageNumber="49">pr</emphasis>
)—trochantero-prefemoral process, (
<emphasis id="B9036429FFC2D73AFD62F85C2E43DBCA" bold="true" box="[742,774,1981,2005]" pageId="48" pageNumber="49">pst</emphasis>
)—penultimate sternite, (
<emphasis id="B9036429FFC2D73AFB8FF85C296EDBCA" bold="true" box="[1035,1067,1981,2005]" pageId="48" pageNumber="49">pts</emphasis>
)—ultimate pretarsus, (
<emphasis id="B9036429FFC2D73AFA98F85C2871DBCA" bold="true" box="[1308,1332,1981,2005]" pageId="48" pageNumber="49">su</emphasis>
)—ventral spines of ultimate prefemur, (
<emphasis id="B9036429FFC2D73AFE47F8002C9FDBE6" bold="true" box="[451,474,2017,2041]" pageId="48" pageNumber="49">tp</emphasis>
)—forcipular tooth-plate, (
<emphasis id="B9036429FFC2D73AFD6CF8002FBADBE6" bold="true" box="[744,767,2017,2041]" pageId="48" pageNumber="49">ut</emphasis>
)—ultimate tergite.
</paragraph>
</caption>
<subSubSection id="C36DEBB0FFC3D73BFF43FF772F71DCE7" pageId="49" pageNumber="50" type="description">
<paragraph id="8BC8B83BFFC3D73BFF43FF772CB0DCCB" blockId="49.[151,1437,150,357]" pageId="49" pageNumber="50">
<emphasis id="B9036429FFC3D73BFF43FF772CECDCAF" bold="true" box="[199,425,150,176]" pageId="49" pageNumber="50">Number of species.</emphasis>
17 (
<bibRefCitation id="EFE6C5CAFFC3D73BFE5FFF772E5DDCAF" author="Edgecombe, G. D. &amp; Bonato, L." box="[475,792,150,177]" pageId="49" pageNumber="50" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe &amp; Bonato 2011</bibRefCitation>
), 22 (
<bibRefCitation id="EFE6C5CAFFC3D73BFCD2FF77296EDCAE" author="Bonato, L. &amp; Chagas-Junior, A. &amp; Edgecombe, G. D. &amp; Lewis, J. G. E. &amp; Minelli, A. &amp; Pereira, L. A. &amp; Shelley, R. M. &amp; Stoev, P. &amp; Zapparoli, M." box="[854,1067,150,177]" pageId="49" pageNumber="50" refId="ref45701" refString="Bonato, L., Chagas-Junior, A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. &amp; Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: http: // chilobase. biologia. unipd. it (accessed 2 August 2016)" type="url" year="2016">
Bonato
<emphasis id="B9036429FFC3D73BFC34FF762EA4DCAF" box="[944,993,150,176]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. 2016
</bibRefCitation>
), 15 (
<bibRefCitation id="EFE6C5CAFFC3D73BFBEDFF7728D5DCAF" author="Joshi, J. &amp; Edgecombe, G. D." box="[1129,1424,150,176]" pageId="49" pageNumber="50" pagination="1316 - 1335" refId="ref47976" refString="Joshi, J. &amp; Edgecombe, G. D. (2018) Molecular phylogeny and systematics of the centipede genus Ethmostigmus Pocock (Chilopoda: Scolopendromorpha) from peninsular India. Invertebrate Systematics, 32, 1316 - 1335. https: // doi. org / 10.1071 / IS 18030" type="journal article" year="2018">Joshi &amp; Edgecombe 2018</bibRefCitation>
), 19 (
<bibRefCitation id="EFE6C5CAFFC3D73BFF46FF5B2CACDCCB" author="Joshi, J. &amp; Edgecombe, G. D." box="[194,489,186,212]" pageId="49" pageNumber="50" pagination="1 - 10" refId="ref48029" refString="Joshi, J. &amp; Edgecombe, G. D. (2019) Evolutionary biogeography of the centipede genus Ethmostigmus from Peninsular India: testing an ancient vicariance hypothesis for Old World tropical diversity. BMC Evolutionary Biology, 19 (41), 1 - 10. https: // doi. org / 10.1186 / s 12862 - 019 - 1367 - 6" type="journal article" year="2019">Joshi &amp; Edgecombe 2019</bibRefCitation>
).
</paragraph>
<paragraph id="8BC8B83BFFC3D73BFF43FF3F2F71DCE7" blockId="49.[151,1437,150,357]" box="[199,564,222,248]" pageId="49" pageNumber="50">
<emphasis id="B9036429FFC3D73BFF43FF3F2CF2DCE7" bold="true" box="[199,439,222,248]" pageId="49" pageNumber="50">Sexual dimorphism.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36DEBB0FFC3D73BFF43FEE3287BDD7B" pageId="49" pageNumber="50" type="discussion">
<paragraph id="8BC8B83BFFC3D73BFF43FEE3287BDD7B" blockId="49.[151,1437,150,357]" pageId="49" pageNumber="50">
<emphasis id="B9036429FFC3D73BFF43FEE32C7EDD03" bold="true" box="[199,315,258,284]" pageId="49" pageNumber="50">Remarks.</emphasis>
Treated as a genus in
<bibRefCitation id="EFE6C5CAFFC3D73BFDB5FEE32EF1DD03" author="Edgecombe, G. D. &amp; Bonato, L." box="[561,948,258,285]" pageId="49" pageNumber="50" refId="ref47498" refString="Edgecombe, G. D. &amp; Bonato, L. (2011) Chilopoda-taxonomic overview. Order Scolopendromorpha. In: Minelli, A. (Ed.), Treatise on Zoology-Anatomy, Taxonomy, Biology. The Myriapoda. Vol. 1. Brill, Leiden, pp. 392 - 407." type="book" year="2011">Edgecombe &amp; Bonato (2011: 402)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFC3D73BFC3BFEE32980DD02" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[959,1221,258,285]" pageId="49" pageNumber="50" pagination="4 - 20" refId="ref51271" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2012 a) Evolution of blindness in scolopendromorph centipedes (Chilopoda: Scolopendromorpha): insight from an expanded sampling of molecular data. Cladistics, 28, 4 - 20. https: // doi. org / 10.1111 / j. 1096 - 0031.2011.00361. x" type="journal article" year="2012">
Vahtera
<emphasis id="B9036429FFC3D73BFB9FFEE22916DD03" box="[1051,1107,258,284]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(2012a: 7
</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFC3D73BFB55FEE32817DD02" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." box="[1233,1362,258,285]" pageId="49" pageNumber="50" pagination="225 - 248" refId="ref51336" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2012 b) Spiracle structure in scolopendromorph centipedes (Chilopoda: Scolopendromorpha) and its contribution to phylogenetics. Zoomorphology, 131, 225 - 248. https: // doi. org / 10.1007 / s 00435 - 012 - 0157 - 0" type="journal article" year="2012">2012b: 235</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFC3D73BFAD8FEE32D80DD5E" author="Vahtera, V. &amp; Edgecombe, G. D. &amp; Giribet, G." pageId="49" pageNumber="50" pagination="578 - 602" refId="ref51397" refString="Vahtera, V., Edgecombe, G. D. &amp; Giribet, G. (2013). Phylogenetics of scolopendromorph centipedes: can denser taxon sampling improve an artificial classification? Invertebrate Systematics, 27 (5), 578 - 602. https: // doi. org / 10.1071 / IS 13035" type="journal article" year="2013">2013: 584</bibRefCitation>
),
<bibRefCitation id="EFE6C5CAFFC3D73BFF5CFEC72F73DD5E" author="Schileyko, A. &amp; Stoev, P." box="[216,566,294,321]" pageId="49" pageNumber="50" pagination="247 - 280" refId="ref50391" refString="Schileyko, A. &amp; Stoev, P. (2016) Scolopendromorpha of New Guinea and adjacent islands (Myriapoda, Chilopoda). Zootaxa, 4147 (3), 247 - 280. https: // doi. org / 10.11646 / zootaxa. 4147.3.3" type="journal article" year="2016">Schileyko &amp; Stoev (2016: 258)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFC3D73BFDC6FEC72EF9DD5E" author="Joshi, J. &amp; Edgecombe, G. D." box="[578,956,294,321]" pageId="49" pageNumber="50" pagination="1316 - 1335" refId="ref47976" refString="Joshi, J. &amp; Edgecombe, G. D. (2018) Molecular phylogeny and systematics of the centipede genus Ethmostigmus Pocock (Chilopoda: Scolopendromorpha) from peninsular India. Invertebrate Systematics, 32, 1316 - 1335. https: // doi. org / 10.1071 / IS 18030" type="journal article" year="2018">Joshi &amp; Edgecombe (2018: 1316)</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFC3D73BFC43FEC729B7DD5E" author="Siriwut, W. &amp; Edgecombe, G. D. &amp; Sutcharit, C. &amp; Tongkerd, P. &amp; Panha, S." box="[967,1266,294,321]" pageId="49" pageNumber="50" pagination="1005 - 1049" refId="ref51057" refString="Siriwut, W., Edgecombe, G. D., Sutcharit, C., Tongkerd, P. &amp; Panha, S. (2018) Systematic revision and phylogenetic reassessment of the centipede genera Rhysida Wood, 1862 and Alluropus Silvestri, 1912 (Chilopoda: Scolopendromorpha) in Southeast Asia, with further discussion of the subfamily Otostigminae. Invertebrate Systematics, 32, 1005 - 1049. https: // doi. org / 10.1071 / IS 17081" type="journal article" year="2018">
Siriwut
<emphasis id="B9036429FFC3D73BFBA4FEC6291DDD5F" box="[1056,1112,294,320]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(2018: 1005)
</bibRefCitation>
,
<bibRefCitation id="EFE6C5CAFFC3D73BFB79FEC72C17DD7B" author="Joshi, J. &amp; Edgecombe, G. D." pageId="49" pageNumber="50" pagination="1 - 10" refId="ref48029" refString="Joshi, J. &amp; Edgecombe, G. D. (2019) Evolutionary biogeography of the centipede genus Ethmostigmus from Peninsular India: testing an ancient vicariance hypothesis for Old World tropical diversity. BMC Evolutionary Biology, 19 (41), 1 - 10. https: // doi. org / 10.1186 / s 12862 - 019 - 1367 - 6" type="journal article" year="2019">Joshi &amp; Edgecombe (2019: 1)</bibRefCitation>
. The most recent morphological account on
<taxonomicName id="4C77C3B8FFC3D73BFCC0FEAB2EA6DD7B" authorityName="Pocock" authorityYear="1898" box="[836,995,330,356]" class="Chilopoda" family="Scolopendridae" genus="Ethmostigmus" kingdom="Animalia" order="Scolopendromorpha" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B9036429FFC3D73BFCC0FEAB2EA6DD7B" box="[836,995,330,356]" italics="true" pageId="49" pageNumber="50">Ethmostigmus</emphasis>
</taxonomicName>
is
<bibRefCitation id="EFE6C5CAFFC3D73BFB80FEAB287FDD7B" author="Joshi, J. &amp; Edgecombe, G. D." box="[1028,1338,330,356]" pageId="49" pageNumber="50" pagination="1316 - 1335" refId="ref47976" refString="Joshi, J. &amp; Edgecombe, G. D. (2018) Molecular phylogeny and systematics of the centipede genus Ethmostigmus Pocock (Chilopoda: Scolopendromorpha) from peninsular India. Invertebrate Systematics, 32, 1316 - 1335. https: // doi. org / 10.1071 / IS 18030" type="journal article" year="2018">Joshi &amp; Edgecombe (2018)</bibRefCitation>
.
</paragraph>
</subSubSection>
</subSubSection>
</treatment>
</document>