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<document id="0D5247EE66B733960C17AAD0503ABEBD" ID-DOI="10.5852/ejt.2016.215" ID-GBIF-Dataset="1cb872fd-ebcd-4e9d-8366-58bc5ecbd050" ID-ISSN="2118-9773" ID-Zenodo-Dep="399800" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1469458853457" checkinUser="plazi" docAuthor="Henrik Enghoff" docDate="2016" docId="03D3D100FFE4C414F028FD757B9FFC43" docLanguage="en" docName="338-1748-1-SM.pdf" docOrigin="European Journal of Taxonomy 215" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Aquattuor fasciatus Attems 1896, comb. nov." docType="treatment" docVersion="2" lastPageNumber="16" masterDocId="FFEAA978FFE9C41BF22AFFDD7A44FFBC" masterDocTitle="A mountain of millipedes IV: Species of Prionopetalum Attems, 1909, from the Udzungwa Mountains, Tanzania. With notes on “ P. ” fasciatum (Attems, 1896) and a revised species key (Diplopoda, Spirostreptida, Odontopygidae)" masterLastPageNumber="23" masterPageNumber="1" pageNumber="14" updateTime="1733842422368" updateUser="ExternalLinkService">
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<mods:title id="37B44AB272FC72A427D9552A5B94EFC3">A mountain of millipedes IV: Species of Prionopetalum Attems, 1909, from the Udzungwa Mountains, Tanzania. With notes on “ P. ” fasciatum (Attems, 1896) and a revised species key (Diplopoda, Spirostreptida, Odontopygidae)</mods:title>
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<mods:namePart id="02E86BC8E42E491DCC94C75780B7E483">Henrik Enghoff</mods:namePart>
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<mods:title id="14D261987D1A07D99A32368A201F0B97">European Journal of Taxonomy</mods:title>
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<mods:date id="342B1D58F7CB8DF74A00972F25EEA44F">2016</mods:date>
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<treatment id="03D3D100FFE4C414F028FD757B9FFC43" ID-DOI="http://doi.org/10.5281/zenodo.5695715" ID-GBIF-Taxon="156114635" ID-Zenodo-Dep="5695715" LSID="urn:lsid:plazi:treatment:03D3D100FFE4C414F028FD757B9FFC43" httpUri="http://treatment.plazi.org/id/03D3D100FFE4C414F028FD757B9FFC43" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" scope_family="Odontopygidae" scope_order="Spirostreptida">
<subSubSection id="C360339DFFE4C416F028FD757E76FD7E" box="[514,1074,679,707]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BC56016FFE4C416F028FD757E76FD7E" blockId="13.[514,1074,679,741]" box="[514,1074,679,707]" pageId="13" pageNumber="14">
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<taxonomicName id="4C7A1B95FFE4C416F028FD7579EDFD7F" ID-CoL="FZKZ" authority="Attems, 1896" authorityName="Attems" authorityYear="1896" box="[514,937,679,707]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fasciatus" status="comb. nov.">
<emphasis id="B90EBC04FFE4C416F028FD7578B5FD7D" bold="true" box="[514,753,679,706]" italics="true" pageId="13" pageNumber="14">Aquattuor fasciatus</emphasis>
(
<bibRefCitation id="EFEB1DE7FFE4C416F12AFD7579E5FD7F" author="Attems C. G." box="[768,929,680,707]" journalOrPublisher="Jahrbuch der hamburgischen wissenschaftlichen Anstalten" pageId="13" pageNumber="14" pagination="23 - 42" part="13" publicationUrl="http://biodiversitylibrary.org/page/28745426" refId="ref9638" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13 (Supplement): 23 - 42. Available from: http: // biodiversitylibrary. org / page / 28745426 [accessed 15 Jun. 2016]" title="Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden" type="book" year="1896">Attems, 1896</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23D017FFFE4C416F19BFD757E76FD7E" box="[945,1074,680,706]" pageId="13" pageNumber="14">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C360339DFFE4C416F0CDFD167909FD59" box="[743,845,715,741]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BC56016FFE4C416F0CDFD167909FD59" blockId="13.[514,1074,679,741]" box="[743,845,715,741]" pageId="13" pageNumber="14">
<figureCitation id="13417C93FFE4C416F0CDFD167909FD59" box="[743,845,715,741]" captionStart-0="Fig. 7" captionStart-1="Fig. 8" captionStartId-0="14.[189,232,1858,1884]" captionStartId-1="15.[189,232,1897,1923]" captionTargetBox-0="[224,1366,270,1826]" captionTargetBox-1="[190,1399,1261,1866]" captionTargetId-0="figure@14.[208,1379,251,1837]" captionTargetId-1="figure@15.[173,1413,1253,1873]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="Fig. 7. Aquattuor fasciatus (Attems, 1896) comb. nov., specimen from Zanzibar, Kirimkazi. A. Limbus. B C. Left gonopod. B. Anterior view. C. Posterior view. D F. Left telomere. D. Tip, basal-anterior view. E. Subdistal part, posterior view (bold arrow points to a microserrate lobe on the margin). F. Tip, apical-posterior view. Abbreviations: mbl = mesobasal lobe of coxal palette; pa = apical palette of coxa. Scale bars: A C = 0.1 mm; D F = 0.01 mm." captionText-1="Fig. 8. Aquattuor fasciatus (Attems, 1896) comb. nov., syntype of Odontopyge fasciata (NHMW 2672). A. Gonopods, anterior view. B. Left gonopod, posterior view. Scale bars = 0.5 mm. Photographs: N. Akkari." figureDoi-0="http://doi.org/10.5281/zenodo.400003" figureDoi-1="http://doi.org/10.5281/zenodo.400004" httpUri-0="https://zenodo.org/record/400003/files/figure.png" httpUri-1="https://zenodo.org/record/400004/files/figure.png" pageId="13" pageNumber="14">Figs 78</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C360339DFFE4C416F297FCD17984FC12" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8BC56016FFE4C416F297FCD178CDFC9B" blockId="13.[189,649,780,807]" box="[189,649,780,807]" pageId="13" pageNumber="14">
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<taxonomicName id="4C7A1B95FFE4C416F297FCD178C1FC9B" ID-CoL="7XK7M" authority="Attems, 1896: 40" authorityName="Attems" authorityPageNumber="40" authorityYear="1896" box="[189,645,780,807]" class="Diplopoda" family="Odontopygidae" genus="Odontopyge" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B90EBC04FFE4C416F297FCD17BF5FC9B" box="[189,433,780,807]" italics="true" pageId="13" pageNumber="14">Odontopyge fasciata</emphasis>
<treatmentCitation id="0ADB4607FFE4C416F39CFCD078C1FC9B" author="Attems C. G." box="[438,645,780,807]" page="40" pageId="13" pageNumber="14" year="1896">
<bibRefCitation id="EFEB1DE7FFE4C416F39CFCD078C1FC9B" author="Attems C. G." box="[438,645,780,807]" journalOrPublisher="Jahrbuch der hamburgischen wissenschaftlichen Anstalten" pageId="13" pageNumber="14" pagination="23 - 42" part="13" publicationUrl="http://biodiversitylibrary.org/page/28745426" refId="ref9638" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13 (Supplement): 23 - 42. Available from: http: // biodiversitylibrary. org / page / 28745426 [accessed 15 Jun. 2016]" title="Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden" type="book" year="1896">Attems, 1896: 40</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="8BC56016FFE4C416F297FC9379C8FC30" blockId="13.[189,1398,846,943]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB6A4738FFE4C416F297FC9379C8FC30" pageId="13" pageNumber="14">
<taxonomicName id="4C7A1B95FFE4C416F297FC937BA9FCD4" ID-CoL="4MG36" authority="Attems, 1896" authorityName="Attems" authorityYear="1896" box="[189,493,846,872]" class="Diplopoda" family="Odontopygidae" genus="Prionopetalum" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fasciatum">
<emphasis id="B90EBC04FFE4C416F297FC937BA9FCD4" box="[189,493,846,872]" italics="true" pageId="13" pageNumber="14">Prionopetalum fasciatum</emphasis>
</taxonomicName>
<treatmentCitation id="0ADB4607FFE4C416F03CFC9378BDFCD4" author="Attems C. G." box="[534,761,846,872]" page="210" pageId="13" pageNumber="14" year="1914">
<bibRefCitation id="EFEB1DE7FFE4C416F03CFC9378BDFCD4" author="Attems C. G." box="[534,761,846,872]" pageId="13" pageNumber="14" pagination="1 - 233" refId="ref9803" refString="Attems C. G. 1914. Afrikanische Spirostreptiden nebst Uberblick uber die Spirostreptiden orbis terrarum. Zoologica Stuttgart 65 - 66: 1 - 233." year="1914">Attems 1914: 210</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ADB4607FFE4C416F11FFC937E01FCD4" author="Brolemann H. W." box="[821,1093,846,872]" page="123" pageId="13" pageNumber="14" year="1920">
<bibRefCitation id="EFEB1DE7FFE4C416F11FFC937E01FCD4" author="Brolemann H. W." box="[821,1093,846,872]" pageId="13" pageNumber="14" refId="ref9895" refString="Brolemann H. W. 1920. Myriapodes. In: Alluaud Ch. (ed.) Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale 1911 - 1912. Resultats Scientifiques. Myriapoda 3: 49 - 298. A. Schultz, Paris." year="1920">Brolemann 1920: 123</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ADB4607FFE4C416F6A8FC937F0CFCD4" author="Kraus O." box="[1154,1352,846,872]" page="86" pageId="13" pageNumber="14" year="1960">
<bibRefCitation id="EFEB1DE7FFE4C416F6A8FC937F0CFCD4" author="Kraus O." box="[1154,1352,846,872]" pageId="13" pageNumber="14" pagination="1 - 207" refId="ref10649" refString="Kraus O. 1960. Athiopische Diplopoden I. Monographie der Odontopygidae-Odontopyginae (Diplopoda, Spirostreptoidea). Annalen van het Koninklijk Museum van Belgisch-Congo 82: 1 - 207." year="1960">Kraus 1960: 86</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ADB4607FFE4C416F297FCAC7835FC30" author="VandenSpiegel D. &amp; Pierrard G." box="[189,625,881,908]" page="152" pageId="13" pageNumber="14" year="2009">
<bibRefCitation id="EFEB1DE7FFE4C416F297FCAC7835FC30" author="VandenSpiegel D. &amp; Pierrard G." box="[189,625,881,908]" pageId="13" pageNumber="14" pagination="149 - 163" refId="ref10926" refString="VandenSpiegel D. &amp; Pierrard G. 2009. Revision du genre Prionopetalum (Odontopygidae, Diplopoda) et descriptions de nouvelles especes d'Afrique de l'Est. Journal of Afrotropical Zoology 5: 149 - 163." year="2009">VandenSpiegel &amp; Pierrard 2009: 152</bibRefCitation>
</treatmentCitation>
. —
<bibRefCitation id="EFEB1DE7FFE4C416F088FCAC79C2FC37" author="Enghoff H. &amp; Hoffman R. L. &amp; Howell K. M." box="[674,902,881,907]" pageId="13" pageNumber="14" pagination="51 - 113" refId="ref10405" refString="Enghoff H., Hoffman R. L. &amp; Howell K. M. 2016. Checklist of the millipedes (Diplopoda) of Tanzania. Journal of East African Natural History 105: 51 - 113. http: // dx. doi. org / 10.2982 / 028.105.0103" year="2016">
Enghoff
<emphasis id="B90EBC04FFE4C416F123FCAF7907FC37" box="[777,835,881,907]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
2016
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.
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<paragraph id="8BC56016FFE4C416F297FC487984FC12" blockId="13.[189,1398,846,943]" box="[189,960,916,943]" pageId="13" pageNumber="14">
<emphasis id="B90EBC04FFE4C416F297FC487B70FC13" box="[189,308,917,943]" italics="true" pageId="13" pageNumber="14">Aquattuor</emphasis>
aff.
<emphasis id="B90EBC04FFE4C416F340FC497859FC12" box="[362,541,916,942]" italics="true" pageId="13" pageNumber="14">claudiahempae</emphasis>
<bibRefCitation id="EFEB1DE7FFE4C416F011FC4979F8FC12" author="Enghoff H. &amp; Frederiksen S. B." box="[571,956,916,943]" pageId="13" pageNumber="14" pagination="1 - 25" refId="ref10334" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" year="2015">Enghoff &amp; Frederiksen 2015: 19</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C360339DFFE4C416F297FC0878C8FA92" pageId="13" pageNumber="14" type="materials_examined">
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<emphasis id="B90EBC04FFE4C416F297FC087BC8FC53" bold="true" box="[189,396,981,1007]" pageId="13" pageNumber="14">Material studied</emphasis>
</heading>
</paragraph>
<paragraph id="8BC56016FFE4C416F2C0FBD97B1EFBA2" blockId="13.[189,1235,1028,1090]" box="[234,346,1028,1054]" pageId="13" pageNumber="14">
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<typeStatus id="54C1DEB4FFE4C416F2C0FBD97B1EFBA2" box="[234,346,1028,1054]" pageId="13" pageNumber="14" type="syntype">
<emphasis id="B90EBC04FFE4C416F2C0FBD97B1EFBA2" bold="true" box="[234,346,1028,1054]" pageId="13" pageNumber="14">Syntypes</emphasis>
</typeStatus>
</heading>
</paragraph>
<paragraph id="8BC56016FFE4C416F297FBF57E97FBFE" blockId="13.[189,1235,1028,1090]" box="[189,1235,1063,1090]" pageId="13" pageNumber="14">
<materialsCitation id="3B126A4BFFE4C416F297FBF57E8BFBFE" box="[189,1231,1063,1090]" collectionCode="ZMUH" country="Tanzania" location="Kibueni" pageId="13" pageNumber="14" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Zanzibar">
<collectingCountry id="F36D2086FFE4C416F297FBF57B13FBFE" box="[189,343,1064,1090]" name="Tanzania" pageId="13" pageNumber="14">TANZANIA</collectingCountry>
:
<specimenCount id="9D7CAB9FFFE4C416F348FBF57BCAFBFE" box="[354,398,1064,1090]" count="1" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
(+ one extra set of gonopods),
<specimenCount id="9D7CAB9FFFE4C416F0D6FBFA7977FBFD" box="[764,819,1063,1089]" count="6" pageId="13" pageNumber="14" type="female">6 ♀♀</specimenCount>
/juvs,
<collectingRegion id="49BEAEF4FFE4C416F1ACFBFA79B4FBFD" box="[902,1008,1063,1089]" country="Tanzania" name="Kaskazini Unguja" pageId="13" pageNumber="14">Zanzibar</collectingRegion>
,
<location id="8EA536CDFFE4C416F1D6FBFA7E1FFBFD" LSID="urn:lsid:plazi:treatment:03D3D100FFE4C414F028FD757B9FFC43:8EA536CDFFE4C416F1D6FBFA7E1FFBFD" box="[1020,1115,1063,1089]" country="Tanzania" name="Kibueni" pageId="13" pageNumber="14" stateProvince="Zanzibar">Kibueni</location>
(
<collectionCode id="ED6BF8D3FFE4C416F640FBF57E83FBFE" box="[1130,1223,1064,1090]" collectionName="Germany, Hamburg, Universitaet von Hamburg, Zoologisches Institut und Zoologisches Museum" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="13" pageNumber="14">ZMUH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC56016FFE4C416F2C0FBB47BE2FB3F" blockId="13.[189,1398,1129,1226]" box="[234,422,1129,1155]" pageId="13" pageNumber="14">
<heading id="D08DD77AFFE4C416F2C0FBB47BE2FB3F" bold="true" box="[234,422,1129,1155]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<emphasis id="B90EBC04FFE4C416F2C0FBB47BE2FB3F" bold="true" box="[234,422,1129,1155]" pageId="13" pageNumber="14">Other material</emphasis>
</heading>
</paragraph>
<paragraph id="8BC56016FFE4C416F297FB517E59FB75" blockId="13.[189,1398,1129,1226]" pageId="13" pageNumber="14">
<materialsCitation id="3B126A4BFFE4C416F297FB517F36FB1A" box="[189,1394,1160,1191]" collectingDate="1979-06-17" collectionCode="ZMUC" collectorName="M. Stoltze" country="Tanzania" location="Kizimkazi" pageId="13" pageNumber="14" specimenCount="3" specimenCount-male="3" stateProvince="Zanzibar">
<collectingCountry id="F36D2086FFE4C416F297FB517B13FB1A" box="[189,343,1164,1190]" name="Tanzania" pageId="13" pageNumber="14">TANZANIA</collectingCountry>
:
<specimenCount id="9D7CAB9FFFE4C416F34BFB517BE3FB1A" box="[353,423,1160,1190]" count="3" pageId="13" pageNumber="14" type="male">3 ♂♂</specimenCount>
,
<collectingRegion id="49BEAEF4FFE4C416F39BFB51785FFB1A" box="[433,539,1164,1190]" country="Tanzania" name="Kaskazini Unguja" pageId="13" pageNumber="14">Zanzibar</collectingRegion>
,
<location id="8EA536CDFFE4C416F00CFB5178E7FB1A" LSID="urn:lsid:plazi:treatment:03D3D100FFE4C414F028FD757B9FFC43:8EA536CDFFE4C416F00CFB5178E7FB1A" box="[550,675,1164,1190]" country="Tanzania" name="Kizimkazi" pageId="13" pageNumber="14" stateProvince="Zanzibar">Kizimkazi</location>
, at foot of mango tree,
<date id="FFC446D6FFE4C416F19AFB517E0CFB1A" box="[944,1096,1164,1191]" pageId="13" pageNumber="14" value="1979-06-17">
<collectingDate id="EF80BF3EFFE4C416F19AFB517E0CFB1A" box="[944,1096,1164,1191]" pageId="13" pageNumber="14" value="1979-06-17">17 Jun. 1979</collectingDate>
</date>
,
<collectorName id="268F05C0FFE4C416F67EFB507E89FB1A" box="[1108,1229,1164,1191]" pageId="13" pageNumber="14">M. Stoltze</collectorName>
leg. (
<collectionCode id="ED6BF8D3FFE4C416F724FB517F2DFB1A" box="[1294,1385,1164,1190]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="13" pageNumber="14">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B126A4BFFE4C416F297FB727E5DFB75" box="[189,1049,1195,1226]" collectingDate="1979-06-16" collectionCode="ZMUC" collectorName="M. Stoltze" location="Zanzibar" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Zanzibar">
<specimenCount id="9D7CAB9FFFE4C416F297FB727AA9FB75" box="[189,237,1195,1225]" count="1" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
,
<collectingRegion id="49BEAEF4FFE4C416F2D2FB727B26FB75" box="[248,354,1199,1225]" country="Tanzania" name="Kaskazini Unguja" pageId="13" pageNumber="14">Zanzibar</collectingRegion>
, the sultans palace,
<date id="FFC446D6FFE4C416F07EFB7278A9FB75" box="[596,749,1199,1226]" pageId="13" pageNumber="14" value="1979-06-16">
<collectingDate id="EF80BF3EFFE4C416F07EFB7278A9FB75" box="[596,749,1199,1226]" pageId="13" pageNumber="14" value="1979-06-16">16 Jun. 1979</collectingDate>
</date>
,
<collectorName id="268F05C0FFE4C416F0D3FB6D7937FB75" box="[761,883,1199,1226]" pageId="13" pageNumber="14">M. Stoltze</collectorName>
leg. (
<collectionCode id="ED6BF8D3FFE4C416F19CFB727E56FB75" box="[950,1042,1199,1225]" collectionName="Denmark, Kobenhavn [= Copenhagen], University of Copenhagen, Zoological Museum" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="13" pageNumber="14">ZMUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC56016FFE4C416F297FB2C78C8FA92" blockId="13.[189,1399,1265,1326]" pageId="13" pageNumber="14">
In addition, Dr. Nesrine Akkari has kindly examined
<typeStatus id="54C1DEB4FFE4C416F107FB2C79D0FAB7" box="[813,916,1265,1291]" pageId="13" pageNumber="14" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C7A1B95FFE4C416F191FB2C7E78FAB7" box="[955,1084,1265,1291]" class="Diplopoda" family="Odontopygidae" genus="Odontopyge" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B90EBC04FFE4C416F191FB2C7E78FAB7" box="[955,1084,1265,1291]" italics="true" pageId="13" pageNumber="14">O. fasciata</emphasis>
</taxonomicName>
in
<collectionCode id="ED6BF8D3FFE4C416F648FB2C7E86FAB7" box="[1122,1218,1265,1291]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="14">NHMW</collectionCode>
and has placed information and images at my disposal.
</paragraph>
</subSubSection>
<subSubSection id="C360339DFFE4C414F297FA8B7B9FFC43" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BC56016FFE4C416F297FA8B7B6BFACC" blockId="13.[189,1399,1366,1580]" box="[189,303,1366,1392]" pageId="13" pageNumber="14">
<heading id="D08DD77AFFE4C416F297FA8B7B6BFACC" bold="true" box="[189,303,1366,1392]" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
<emphasis id="B90EBC04FFE4C416F297FA8B7B6BFACC" bold="true" box="[189,303,1366,1392]" pageId="13" pageNumber="14">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BC56016FFE4C416F297FA5978A7F997" blockId="13.[189,1399,1366,1580]" pageId="13" pageNumber="14">
The single intact male among the examined
<typeStatus id="54C1DEB4FFE4C416F0F9FA59797EFA22" box="[723,826,1412,1438]" pageId="13" pageNumber="14" type="syntype">syntypes</typeStatus>
has 53 podous rings (no apodous rings in front of the telson) and a diameter of
<quantity id="4C82CDF3FFE4C416F069FA7A78F5FA7E" box="[579,689,1447,1474]" metricMagnitude="-3" metricUnit="m" metricValue="1.91" pageId="13" pageNumber="14" unit="mm" value="1.91">1.91 mm</quantity>
.
<collectingRegion id="49BEAEF4FFE4C416F097FA7A7895FA7D" box="[701,721,1447,1473]" country="Gambia" name="Upper River" pageId="13" pageNumber="14">A</collectingRegion>
<typeStatus id="54C1DEB4FFE4C416F0F1FA757972FA7E" box="[731,822,1448,1474]" pageId="13" pageNumber="14" type="syntype">syntype</typeStatus>
at
<collectionCode id="ED6BF8D3FFE4C416F174FA7579FAFA7E" box="[862,958,1448,1474]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="14">NHMW</collectionCode>
has the same number of rings and a diameter of
<quantity id="4C82CDF3FFE4C416F365FA177BF8FA59" box="[335,444,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.86" pageId="13" pageNumber="14" unit="mm" value="1.86">1.86 mm</quantity>
. The specimens from Kizimkazi are all broken, the males have maximum body diameters of
<quantity id="4C82CDF3FFE4C416F37FFA307845F9B4" box="[341,513,1517,1544]" metricMagnitude="-3" metricUnit="m" metricValue="1.585" metricValueMax="1.62" metricValueMin="1.55" pageId="13" pageNumber="14" unit="mm" value="1.585" valueMax="1.62" valueMin="1.55">1.551.62 mm</quantity>
. The male from the sultans palace has 51 podous rings (no apodous rings in front of the telson) and a diameter of
<quantity id="4C82CDF3FFE4C416F059F9CC789BF997" box="[627,735,1553,1579]" metricMagnitude="-3" metricUnit="m" metricValue="1.86" pageId="13" pageNumber="14" unit="mm" value="1.86">1.86 mm</quantity>
.
</paragraph>
<paragraph id="8BC56016FFE4C416F297F98F7B7FF92C" blockId="13.[189,1398,1618,1680]" pageId="13" pageNumber="14">
The limbus (
<figureCitation id="13417C93FFE4C416F37DF98F7BDBF9D0" box="[343,415,1618,1644]" captionStart="Fig. 7" captionStartId="14.[189,232,1858,1884]" captionTargetBox="[224,1366,270,1826]" captionTargetId="figure@14.[208,1379,251,1837]" captionTargetPageId="14" captionText="Fig. 7. Aquattuor fasciatus (Attems, 1896) comb. nov., specimen from Zanzibar, Kirimkazi. A. Limbus. B C. Left gonopod. B. Anterior view. C. Posterior view. D F. Left telomere. D. Tip, basal-anterior view. E. Subdistal part, posterior view (bold arrow points to a microserrate lobe on the margin). F. Tip, apical-posterior view. Abbreviations: mbl = mesobasal lobe of coxal palette; pa = apical palette of coxa. Scale bars: A C = 0.1 mm; D F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.400003" httpUri="https://zenodo.org/record/400003/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
<collectingRegion id="49BEAEF4FFE4C416F3B5F98F7BF2F9D0" box="[415,438,1618,1644]" country="Gambia" name="Upper River" pageId="13" pageNumber="14">A</collectingRegion>
) consists of large, rectangular, easily detached flaps, characteristic of the genus
<emphasis id="B90EBC04FFE4C416F297F9AB7B70F92C" box="[189,308,1654,1680]" italics="true" pageId="13" pageNumber="14">Aquattuor</emphasis>
.
</paragraph>
<paragraph id="8BC56016FFE4C416F297F96A78D8F887" blockId="13.[189,1399,1719,1851]" pageId="13" pageNumber="14">
The gonopods (
<figureCitation id="13417C93FFE4C416F35EF96A7BFAF96D" box="[372,446,1719,1745]" captionStart="Fig. 7" captionStartId="14.[189,232,1858,1884]" captionTargetBox="[224,1366,270,1826]" captionTargetId="figure@14.[208,1379,251,1837]" captionTargetPageId="14" captionText="Fig. 7. Aquattuor fasciatus (Attems, 1896) comb. nov., specimen from Zanzibar, Kirimkazi. A. Limbus. B C. Left gonopod. B. Anterior view. C. Posterior view. D F. Left telomere. D. Tip, basal-anterior view. E. Subdistal part, posterior view (bold arrow points to a microserrate lobe on the margin). F. Tip, apical-posterior view. Abbreviations: mbl = mesobasal lobe of coxal palette; pa = apical palette of coxa. Scale bars: A C = 0.1 mm; D F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.400003" httpUri="https://zenodo.org/record/400003/files/figure.png" pageId="13" pageNumber="14">Figs 7</figureCitation>
BF, 8) are indistinguishable from those of
<emphasis id="B90EBC04FFE4C416F195F96A7EB5F96D" box="[959,1265,1719,1745]" italics="true" pageId="13" pageNumber="14">Aquattuor claudiahempae</emphasis>
<bibRefCitation id="EFEB1DE7FFE4C416F6D2F96A7BD1F949" author="Enghoff H. &amp; Frederiksen S. B." pageId="13" pageNumber="14" pagination="1 - 25" refId="ref10334" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" year="2015">Enghoff &amp; Frederiksen, 2015</bibRefCitation>
(cf. their fig. 15). Noticeable special similarities include the profile of the apical palette (
<emphasis id="B90EBC04FFE4C416F337F9237B7EF8AB" box="[285,314,1790,1815]" italics="true" pageId="13" pageNumber="14">pa</emphasis>
), the well-developed mesobasal lobe (
<emphasis id="B90EBC04FFE4C416F128F920796AF8AB" box="[770,814,1789,1815]" italics="true" pageId="13" pageNumber="14">mbl</emphasis>
) of the coxal palette, and the partly micro-serrate margin of the telomere (
<figureCitation id="13417C93FFE4C416F3F3F8FD785AF887" box="[473,542,1824,1851]" captionStart="Fig. 7" captionStartId="14.[189,232,1858,1884]" captionTargetBox="[224,1366,270,1826]" captionTargetId="figure@14.[208,1379,251,1837]" captionTargetPageId="14" captionText="Fig. 7. Aquattuor fasciatus (Attems, 1896) comb. nov., specimen from Zanzibar, Kirimkazi. A. Limbus. B C. Left gonopod. B. Anterior view. C. Posterior view. D F. Left telomere. D. Tip, basal-anterior view. E. Subdistal part, posterior view (bold arrow points to a microserrate lobe on the margin). F. Tip, apical-posterior view. Abbreviations: mbl = mesobasal lobe of coxal palette; pa = apical palette of coxa. Scale bars: A C = 0.1 mm; D F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.400003" httpUri="https://zenodo.org/record/400003/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
E, arrow).
</paragraph>
<paragraph id="8BC56016FFE4C414F297F8BF799DFE44" blockId="13.[189,1399,1890,2022]" lastBlockId="15.[189,1399,267,505]" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
The characteristic limbus was not noted by previous authors.
<bibRefCitation id="EFEB1DE7FFE4C416F1A7F8BF7E73F8C1" author="Attems C. G." box="[909,1079,1890,1917]" journalOrPublisher="Jahrbuch der hamburgischen wissenschaftlichen Anstalten" pageId="13" pageNumber="14" pagination="23 - 42" part="13" publicationUrl="http://biodiversitylibrary.org/page/28745426" refId="ref9638" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13 (Supplement): 23 - 42. Available from: http: // biodiversitylibrary. org / page / 28745426 [accessed 15 Jun. 2016]" title="Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden" type="book" year="1896">Attems (1896)</bibRefCitation>
and
<bibRefCitation id="EFEB1DE7FFE4C416F65BF8BF7F0DF8C0" author="Brolemann H. W." box="[1137,1353,1890,1916]" pageId="13" pageNumber="14" refId="ref9895" refString="Brolemann H. W. 1920. Myriapodes. In: Alluaud Ch. (ed.) Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale 1911 - 1912. Resultats Scientifiques. Myriapoda 3: 49 - 298. A. Schultz, Paris." year="1920">Brolemann (1920)</bibRefCitation>
did not mention the limbus at all.
<bibRefCitation id="EFEB1DE7FFE4C416F034F85879E7F823" author="VandenSpiegel D. &amp; Pierrard G." box="[542,931,1925,1951]" pageId="13" pageNumber="14" pagination="149 - 163" refId="ref10926" refString="VandenSpiegel D. &amp; Pierrard G. 2009. Revision du genre Prionopetalum (Odontopygidae, Diplopoda) et descriptions de nouvelles especes d'Afrique de l'Est. Journal of Afrotropical Zoology 5: 149 - 163." year="2009">VandenSpiegel &amp; Pierrard (2009</bibRefCitation>
: fig. 1b) illustrated the limbus, but their SEM micrograph only showed an irregularly wavy margin. The large limbus lobes were probably all broken off in the specimens examined by these authors (compare the left part of
<figureCitation id="13417C93FFE4C416F6A4F8167E90F85A" box="[1166,1236,1995,2022]" captionStart="Fig. 7" captionStartId="14.[189,232,1858,1884]" captionTargetBox="[224,1366,270,1826]" captionTargetId="figure@14.[208,1379,251,1837]" captionTargetPageId="14" captionText="Fig. 7. Aquattuor fasciatus (Attems, 1896) comb. nov., specimen from Zanzibar, Kirimkazi. A. Limbus. B C. Left gonopod. B. Anterior view. C. Posterior view. D F. Left telomere. D. Tip, basal-anterior view. E. Subdistal part, posterior view (bold arrow points to a microserrate lobe on the margin). F. Tip, apical-posterior view. Abbreviations: mbl = mesobasal lobe of coxal palette; pa = apical palette of coxa. Scale bars: A C = 0.1 mm; D F = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.400003" httpUri="https://zenodo.org/record/400003/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
<collectingRegion id="49BEAEF4FFE4C416F6FEF8117EADF85A" box="[1236,1257,1996,2022]" country="Gambia" name="Upper River" pageId="13" pageNumber="14">A</collectingRegion>
with fig 1b in
<bibRefCitation id="EFEB1DE7FFE6C414F2F7FED6781BFE99" author="VandenSpiegel D. &amp; Pierrard G." box="[221,607,267,293]" pageId="15" pageNumber="16" pagination="149 - 163" refId="ref10926" refString="VandenSpiegel D. &amp; Pierrard G. 2009. Revision du genre Prionopetalum (Odontopygidae, Diplopoda) et descriptions de nouvelles especes d'Afrique de l'Est. Journal of Afrotropical Zoology 5: 149 - 163." year="2009">VandenSpiegel &amp; Pierrard 2009</bibRefCitation>
). In the
<typeStatus id="54C1DEB4FFE6C414F0EEFED1796FFE9A" box="[708,811,268,294]" pageId="15" pageNumber="16" type="syntype">syntypes</typeStatus>
from
<collectionCode id="ED6BF8D3FFE6C414F15CFED17988FE9A" box="[886,972,268,294]" country="Germany" httpUri="http://grbio.org/cool/82c5-17ij" name="Zoologisches Institut und Zoologisches Museum, Universitat Hamburg" pageId="15" pageNumber="16">ZMUH</collectionCode>
the limbus seems to be completely worn off. However, Dr. Nesrine Akkari has kindly provided images based on the
<typeStatus id="54C1DEB4FFE6C414F640FEF27E95FEF5" box="[1130,1233,303,329]" pageId="15" pageNumber="16" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C7A1B95FFE6C414F6DCFEF27F33FEF5" box="[1270,1399,303,329]" class="Diplopoda" family="Odontopygidae" genus="Odontopyge" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B90EBC04FFE6C414F6DCFEF27F33FEF5" box="[1270,1399,303,329]" italics="true" pageId="15" pageNumber="16">O. fasciata</emphasis>
</taxonomicName>
in
<collectionCode id="ED6BF8D3FFE6C414F2F4FE8F7B7AFED0" box="[222,318,338,364]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="15" pageNumber="16">NHMW</collectionCode>
which clearly show the
<emphasis id="B90EBC04FFE6C414F040FE8F78A5FED0" box="[618,737,338,364]" italics="true" pageId="15" pageNumber="16">Aquattuor</emphasis>
<typeStatus id="54C1DEB4FFE6C414F0C1FE8F795AFED0" box="[747,798,338,364]" pageId="15" pageNumber="16">type</typeStatus>
of limbus, as well as confirm the great similarity in gonopod structure between this species and
<emphasis id="B90EBC04FFE6C414F0C6FEA87986FE33" box="[748,962,373,399]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F0C6FEA87945FE33" box="[748,769,373,399]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
. claudiahempae
</emphasis>
(
<figureCitation id="13417C93FFE6C414F1FEFEA87E58FE33" box="[980,1052,373,399]" captionStart="Fig. 8" captionStartId="15.[189,232,1897,1923]" captionTargetBox="[190,1399,1261,1866]" captionTargetId="figure@15.[173,1413,1253,1873]" captionTargetPageId="15" captionText="Fig. 8. Aquattuor fasciatus (Attems, 1896) comb. nov., syntype of Odontopyge fasciata (NHMW 2672). A. Gonopods, anterior view. B. Left gonopod, posterior view. Scale bars = 0.5 mm. Photographs: N. Akkari." figureDoi="http://doi.org/10.5281/zenodo.400004" httpUri="https://zenodo.org/record/400004/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
). The gonopods of the intact male
<typeStatus id="54C1DEB4FFE6C414F2D5FE447B1EFE0F" box="[255,346,409,435]" pageId="15" pageNumber="16" type="syntype">syntype</typeStatus>
(now dissected by me) closely agree with the illustrations in
<figureCitation id="13417C93FFE6C414F66BFE457EE8FE0E" box="[1089,1196,408,434]" captionStart-0="Fig. 7" captionStart-1="Fig. 8" captionStartId-0="14.[189,232,1858,1884]" captionStartId-1="15.[189,232,1897,1923]" captionTargetBox-0="[224,1366,270,1826]" captionTargetBox-1="[190,1399,1261,1866]" captionTargetId-0="figure@14.[208,1379,251,1837]" captionTargetId-1="figure@15.[173,1413,1253,1873]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="Fig. 7. Aquattuor fasciatus (Attems, 1896) comb. nov., specimen from Zanzibar, Kirimkazi. A. Limbus. B C. Left gonopod. B. Anterior view. C. Posterior view. D F. Left telomere. D. Tip, basal-anterior view. E. Subdistal part, posterior view (bold arrow points to a microserrate lobe on the margin). F. Tip, apical-posterior view. Abbreviations: mbl = mesobasal lobe of coxal palette; pa = apical palette of coxa. Scale bars: A C = 0.1 mm; D F = 0.01 mm." captionText-1="Fig. 8. Aquattuor fasciatus (Attems, 1896) comb. nov., syntype of Odontopyge fasciata (NHMW 2672). A. Gonopods, anterior view. B. Left gonopod, posterior view. Scale bars = 0.5 mm. Photographs: N. Akkari." figureDoi-0="http://doi.org/10.5281/zenodo.400003" figureDoi-1="http://doi.org/10.5281/zenodo.400004" httpUri-0="https://zenodo.org/record/400003/files/figure.png" httpUri-1="https://zenodo.org/record/400004/files/figure.png" pageId="15" pageNumber="16">Figs 78</figureCitation>
. The extra set of gonopods among the
<typeStatus id="54C1DEB4FFE6C414F39EFE61785FFE6A" box="[436,539,444,470]" pageId="15" pageNumber="16" type="syntype">syntypes</typeStatus>
very much resembles fig.
<quantity id="4C82CDF3FFE6C414F16DFE667937FE69" box="[839,883,443,469]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="15" pageNumber="16" unit="in" value="6.0">6 in</quantity>
<bibRefCitation id="EFEB1DE7FFE6C414F15CFE617E66FE6A" author="Attems C. G." box="[886,1058,443,470]" journalOrPublisher="Jahrbuch der hamburgischen wissenschaftlichen Anstalten" pageId="15" pageNumber="16" pagination="23 - 42" part="13" publicationUrl="http://biodiversitylibrary.org/page/28745426" refId="ref9638" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13 (Supplement): 23 - 42. Available from: http: // biodiversitylibrary. org / page / 28745426 [accessed 15 Jun. 2016]" title="Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden" type="book" year="1896">Attems (1896)</bibRefCitation>
, which is based on macerated gonopods, and they were most probably the model for this drawing.
</paragraph>
<caption id="DF05309EFFE7C415F297F89F7882F855" ID-DOI="http://doi.org/10.5281/zenodo.400003" httpUri="https://zenodo.org/record/400003/files/figure.png" pageId="14" pageNumber="15" targetBox="[224,1366,270,1826]" targetPageId="14">
<paragraph id="8BC56016FFE7C415F297F89F7882F855" blockId="14.[189,1399,1858,2025]" pageId="14" pageNumber="15">
<emphasis id="B90EBC04FFE7C415F297F89F7B4EF8E0" bold="true" box="[189,266,1858,1884]" pageId="14" pageNumber="15">Fig. 7.</emphasis>
<taxonomicName id="4C7A1B95FFE7C415F33AF89F78EAF8E1" authority="Attems, 1896" authorityName="Attems" authorityYear="1896" box="[272,686,1858,1885]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="fasciatus" status="comb. nov.">
<emphasis id="B90EBC04FFE7C415F33AF89F7BB2F8E0" box="[272,502,1858,1884]" italics="true" pageId="14" pageNumber="15">Aquattuor fasciatus</emphasis>
(
<bibRefCitation id="EFEB1DE7FFE7C415F02FF89F78E1F8E1" author="Attems C. G." box="[517,677,1858,1885]" journalOrPublisher="Jahrbuch der hamburgischen wissenschaftlichen Anstalten" pageId="14" pageNumber="15" pagination="23 - 42" part="13" publicationUrl="http://biodiversitylibrary.org/page/28745426" refId="ref9638" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13 (Supplement): 23 - 42. Available from: http: // biodiversitylibrary. org / page / 28745426 [accessed 15 Jun. 2016]" title="Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden" type="book" year="1896">Attems, 1896</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23D017FFFE7C415F09FF89F797CF8E0" box="[693,824,1858,1884]" pageId="14" pageNumber="15">comb. nov.</taxonomicNameLabel>
, specimen from Zanzibar, Kirimkazi.
<emphasis id="B90EBC04FFE7C415F6D8F89F7F43F8E0" bold="true" box="[1266,1287,1858,1884]" pageId="14" pageNumber="15">A</emphasis>
. Limbus.
<emphasis id="B90EBC04FFE7C415F297F8B87AB0F8C3" bold="true" box="[189,244,1893,1920]" pageId="14" pageNumber="15">BC</emphasis>
. Left gonopod. B. Anterior view. C. Posterior view.
<emphasis id="B90EBC04FFE7C415F153F8B879EBF8C3" bold="true" box="[889,943,1893,1920]" pageId="14" pageNumber="15">DF</emphasis>
. Left telomere. D. Tip, basal-anterior view. E. Subdistal part, posterior view (bold arrow points to a microserrate lobe on the margin). F. Tip, apical-posterior view. Abbreviations:
<emphasis id="B90EBC04FFE7C415F05FF87678E5F879" box="[629,673,1963,1989]" italics="true" pageId="14" pageNumber="15">mbl</emphasis>
= mesobasal lobe of coxal palette;
<emphasis id="B90EBC04FFE7C415F611F8717E1CF879" box="[1083,1112,1964,1989]" italics="true" pageId="14" pageNumber="15">pa</emphasis>
= apical palette of coxa. Scale bars: AC = 0.1 mm; DF = 0.01 mm.
</paragraph>
</caption>
<paragraph id="8BC56016FFE6C414F297FDFD781BFC26" blockId="15.[189,1399,544,922]" pageId="15" pageNumber="16">
The
<typeStatus id="54C1DEB4FFE6C414F2D8FDFD7B1DFD86" box="[242,345,544,570]" pageId="15" pageNumber="16" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C7A1B95FFE6C414F3AAFDFD7BACFD86" box="[384,488,544,570]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fasciatus">
<emphasis id="B90EBC04FFE6C414F3AAFDFD7BACFD86" box="[384,488,544,570]" italics="true" pageId="15" pageNumber="16">fasciatus</emphasis>
</taxonomicName>
as well as the
<collectingRegion id="49BEAEF4FFE6C414F0BCFDFD78BAFD86" box="[662,766,544,570]" country="Tanzania" name="Kaskazini Unguja" pageId="15" pageNumber="16">
<collectingCountry id="F36D2086FFE6C414F0BCFDFD78BAFD86" box="[662,766,544,570]" name="Tanzania" pageId="15" pageNumber="16">Zanzibar</collectingCountry>
</collectingRegion>
specimens collected by M. Stoltze are very similar to
<emphasis id="B90EBC04FFE6C414F297FD997BCAFDE1" box="[189,398,579,606]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F297FD997A96FDE2" box="[189,210,580,606]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
. claudiahempae
</emphasis>
<bibRefCitation id="EFEB1DE7FFE6C414F3B9FD9E78B7FDE2" author="Enghoff H. &amp; Frederiksen S. B." box="[403,755,579,606]" pageId="15" pageNumber="16" pagination="1 - 25" refId="ref10334" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" year="2015">Enghoff &amp; Frederiksen (2015)</bibRefCitation>
, and it is possible that the latter name should be regarded as a junior synonym of
<emphasis id="B90EBC04FFE6C414F3C8FDBA782AFD3D" box="[482,622,615,641]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F3C8FDBA7BB3FD3D" box="[482,503,615,641]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
.
<taxonomicName id="4C7A1B95FFE6C414F02CFDBA782AFD3D" box="[518,622,615,641]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fasciatus">fasciatus</taxonomicName>
</emphasis>
. However, the body diameter of the specimens from Kizimkazi is at the high end, and partly larger, compared to
<emphasis id="B90EBC04FFE6C414F131FD5779B5FD18" box="[795,1009,650,676]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F131FD577974FD18" box="[795,816,650,676]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
. claudiahempae
</emphasis>
from Mt.
<collectingRegion id="49BEAEF4FFE6C414F659FD547F41FD1F" box="[1139,1285,649,675]" country="Tanzania" name="Kilimanjaro" pageId="15" pageNumber="16">Kilimanjaro</collectingRegion>
. The two intact specimens from
<collectingRegion id="49BEAEF4FFE6C414F3E1FD707877FD7B" box="[459,563,685,711]" country="Tanzania" name="Kaskazini Unguja" pageId="15" pageNumber="16">
<collectingCountry id="F36D2086FFE6C414F3E1FD707877FD7B" box="[459,563,685,711]" name="Tanzania" pageId="15" pageNumber="16">Zanzibar</collectingCountry>
</collectingRegion>
examined here, viz., the intact
<typeStatus id="54C1DEB4FFE6C414F180FD707E41FD7B" box="[938,1029,685,711]" pageId="15" pageNumber="16" type="syntype">syntype</typeStatus>
and the male from the sultans palace, are definitely much larger. The latter specimens are more similar in size to
<emphasis id="B90EBC04FFE6C414F652FD0D7ED5FD56" box="[1144,1169,720,746]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F652FD0D7EC9FD56" box="[1144,1165,720,746]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
.
</emphasis>
cf.
<emphasis id="B90EBC04FFE6C414F696FD0D7F2BFD56" box="[1212,1391,720,746]" italics="true" pageId="15" pageNumber="16">claudiahempae</emphasis>
, recorded from the
<collectingRegion id="49BEAEF4FFE6C414F3B7FD297831FCB1" box="[413,629,755,782]" country="Tanzania" name="Morogoro" pageId="15" pageNumber="16">Morogoro Region</collectingRegion>
, Kilosa District, Rubeho Mts by
<bibRefCitation id="EFEB1DE7FFE6C414F62EFD2E7F37FCB2" author="Enghoff H. &amp; Frederiksen S. B." box="[1028,1395,755,782]" pageId="15" pageNumber="16" pagination="1 - 25" refId="ref10334" refString="Enghoff H. &amp; Frederiksen S. B. 2015. A mountain of millipedes II: The genus Aquattuor Frederiksen, 2013 - five new species from the Udzungwa Mountains and one from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 150: 1 - 25. http: // dx. doi. org / 10.5852 / ejt. 2015.150" year="2015">Enghoff &amp; Frederiksen (2015)</bibRefCitation>
.
<bibRefCitation id="EFEB1DE7FFE6C414F297FCCA7B2FFC8D" author="Attems C. G." box="[189,363,790,817]" journalOrPublisher="Jahrbuch der hamburgischen wissenschaftlichen Anstalten" pageId="15" pageNumber="16" pagination="23 - 42" part="13" publicationUrl="http://biodiversitylibrary.org/page/28745426" refId="ref9638" refString="Attems C. G. 1896. Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden. Jahrbuch der hamburgischen wissenschaftlichen Anstalten 13 (Supplement): 23 - 42. Available from: http: // biodiversitylibrary. org / page / 28745426 [accessed 15 Jun. 2016]" title="Beschreibung der von Dr. Stuhlmann in Ost-Afrika gesammelten Myriopoden" type="book" year="1896">Attems (1896)</bibRefCitation>
gave the (horizontal) diameter of
<emphasis id="B90EBC04FFE6C414F13CFCCA79E7FC8D" box="[790,931,791,817]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F13CFCCA796FFC8D" box="[790,811,791,817]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
.
<taxonomicName id="4C7A1B95FFE6C414F111FCCA79E7FC8D" box="[827,931,791,817]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fasciatus">fasciatus</taxonomicName>
</emphasis>
as
<quantity id="4C82CDF3FFE6C414F1F9FCCB7E5BFC8D" box="[979,1055,790,817]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="mm" value="2.0">2 mm</quantity>
.
<bibRefCitation id="EFEB1DE7FFE6C414F604FCCB7F4EFC8C" author="Brolemann H. W." box="[1070,1290,790,816]" pageId="15" pageNumber="16" refId="ref9895" refString="Brolemann H. W. 1920. Myriapodes. In: Alluaud Ch. (ed.) Voyage de Ch. Alluaud et R. Jeannel en Afrique Orientale 1911 - 1912. Resultats Scientifiques. Myriapoda 3: 49 - 298. A. Schultz, Paris." year="1920">Brolemann (1920)</bibRefCitation>
gave 55 podous rings (= “56 segments … 1 segment apode”) and a diameter of
<quantity id="4C82CDF3FFE6C414F602FCE77ED0FCE8" box="[1064,1172,826,852]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="15" pageNumber="16" unit="mm" value="1.7">1.70 mm</quantity>
for an adult male. Whereas “
<quantity id="4C82CDF3FFE6C414F31CFC807B3BFCCB" box="[310,383,861,887]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="mm" value="2.0">2 mm</quantity>
” is much more than any other specimen in this complex, Brolemanns values would fit
<emphasis id="B90EBC04FFE6C414F297FC5D7BD5FC26" box="[189,401,896,922]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F297FC5D7A96FC26" box="[189,210,896,922]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
. claudiahempae
</emphasis>
by extrapolation.
</paragraph>
<paragraph id="8BC56016FFE6C414F297FC1C7B9FFC43" blockId="15.[189,1399,961,1023]" pageId="15" pageNumber="16">
The situation is obviously complicated, and for the time being,
<emphasis id="B90EBC04FFE6C414F19AFC1F7E7DFC60" box="[944,1081,962,988]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F19AFC1F7981FC60" box="[944,965,962,988]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
.
<taxonomicName id="4C7A1B95FFE6C414F1FBFC1F7E7DFC60" box="[977,1081,962,988]" class="Diplopoda" family="Odontopygidae" genus="Aquattuor" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="fasciatus">fasciatus</taxonomicName>
</emphasis>
and
<emphasis id="B90EBC04FFE6C414F65FFC1F7F0EFC67" box="[1141,1354,961,988]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="49BEAEF4FFE6C414F65FFC1F7ECEFC60" box="[1141,1162,962,988]" country="Gambia" name="Upper River" pageId="15" pageNumber="16">A</collectingRegion>
. claudiahempae
</emphasis>
are kept as separate species.
</paragraph>
</subSubSection>
</treatment>
</document>