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<document id="E6F7D20AE5CA7ACCEE0D344F1659379A" ID-DOI="10.11646/zootaxa.3895.1.3" ID-GBIF-Dataset="bc5d64a5-8aea-47e7-b0c6-3c84d5aa914e" ID-ISSN="1175-5326" ID-Zenodo-Dep="228001" ID-ZooBank="20E09C24-7B96-420A-9E48-FE82251D97C5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460998343819" checkinUser="plazi" docAuthor="Maier, Crystal A. &amp; Short, Andrew Edward Z." docDate="2014" docId="03BFC572F472DE6CC2E4FAFB45A682F6" docLanguage="en" docName="zt03895p072.pdf" docOrigin="Zootaxa 3895 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lutrochus Erichson" docType="key" docVersion="8" masterDocId="FF86BD0AF47EDE61C273FFA5403D802C" masterDocTitle="Review of the Lutrochidae (Coleoptera) of the Guianas and Lesser Antilles, with description of four new species" updateTime="1698563504948" updateUser="plazi">
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<mods:title id="7CE3A56C57D8AA4BF336C9C8AECBB71C">Review of the Lutrochidae (Coleoptera) of the Guianas and Lesser Antilles, with description of four new species</mods:title>
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<mods:namePart id="B02DDC39D079CE4A4230F03089A0738B">Maier, Crystal A.</mods:namePart>
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<treatment id="03BFC572F472DE6CC2E4FAFB45A682F6" ID-DOI="http://doi.org/10.5281/zenodo.6141177" ID-GBIF-Taxon="119597728" ID-Zenodo-Dep="6141177" LSID="urn:lsid:plazi:treatment:03BFC572F472DE6CC2E4FAFB45A682F6" httpUri="http://treatment.plazi.org/id/03BFC572F472DE6CC2E4FAFB45A682F6" lastPageId="13" pageId="12">
<subSubSection id="C30C27EFF472DE6DC2E4FAFB416085B7" pageId="12" type="nomenclature">
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Key to
<taxonomicName id="4C160FE7F472DE6DC283FAFB41D88554" ID-CoL="5HF6" ID-ENA="440309" authority="Erichson" authorityName="Erichson" box="[240,485,1374,1400]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="genus">
<emphasis id="B962A876F472DE6DC283FAFB41508554" bold="true" box="[240,365,1374,1400]" italics="true" pageId="12">Lutrochus</emphasis>
Erichson
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species of northern South
<collectingCountry id="F30134F4F472DE6DC141FAFB43A08554" box="[818,925,1374,1400]" name="United States of America" pageId="12">America</collectingCountry>
(after
<bibRefCitation id="EF870995F472DE6DC181FAFB44D28554" author="Maier" box="[1010,1263,1374,1400]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
, new taxa listed in BOLD)
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<keyLead id="30E7D751F472DE6DC2E4FA6245A685F1" box="[151,1435,1479,1501]" pageId="12">
1 Mesotibia with apical excavation for reception of tarsus (
<figureCitation id="132D68E1F472DE6DC177FA62437185F1" box="[772,844,1479,1501]" captionStart="FIGURES 16 17" captionStartId="9.[151,264,1396,1419]" captionText="FIGURES 16 17. Lutrochus leeanneae n. sp.: 16. Protibia and tarsus, showing patch of dense setae on protibia; 17. Mesotibia and tarsus, showing lateral expansion of mesotibia." pageId="12">Fig. 17</figureCitation>
)................................................ 2
</keyLead>
</paragraph>
<paragraph id="8BA97464F472DE6DC2E4FA4045A685D6" blockId="12.[151,1436,1479,2029]" box="[151,1435,1508,1530]" pageId="12">
<keyLead id="30E7D751F472DE6DC2E4FA4045A685D6" box="[151,1435,1508,1530]" pageId="12">
- Mesotibia lacking apical excavation for reception of tarsus (cf.
<bibRefCitation id="EF870995F472DE6DC14CFA41443C85D6" author="Maier" box="[831,1025,1508,1530]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 44)......................... 11
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<keyStep id="30E26CC1F472DE6DC2E4F9A445A68686" pageId="12">
<paragraph id="8BA97464F472DE6DC2E4F9A445A5867E" blockId="12.[151,1436,1479,2029]" pageId="12">
<keyLead id="30E7D751F472DE6DC2E4F9A445A5867E" pageId="12">
2(1) Elytra explanate, epipleuron appearing broad in ventral aspect (cf.
<bibRefCitation id="EF870995F472DE6DC114F9A44416863B" author="Maier" box="[871,1067,1537,1559]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
<figureCitation id="132D68E1F472DE6DC641F9A44451863B" box="[1074,1132,1537,1559]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1887,1910]" captionTargetBox="[151,1436,337,1829]" captionTargetId="figure@5.[151,1436,337,1838]" captionTargetPageId="5" captionText="FIGURES 7 11. Lutrochus grenadensis n. sp.: 7. Habitus, dorsal view; 8. Habitus, ventral view; 9. Habitus, lateral view; 10. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 11. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228004/files/figure.png" pageId="12">Fig. 9</figureCitation>
); reddish-brown in color; prosternum strongly deflexed ventrally in anterior half (cf.
<bibRefCitation id="EF870995F472DE6DC098F9BA43968619" author="Maier" box="[747,939,1567,1589]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
<figureCitation id="132D68E1F472DE6DC1C2F9BA43CA8619" box="[945,1015,1567,1589]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1887,1910]" captionTargetBox="[151,1436,337,1829]" captionTargetId="figure@5.[151,1436,337,1838]" captionTargetPageId="5" captionText="FIGURES 7 11. Lutrochus grenadensis n. sp.: 7. Habitus, dorsal view; 8. Habitus, ventral view; 9. Habitus, lateral view; 10. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 11. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228004/files/figure.png" pageId="12">Fig. 10</figureCitation>
); apical maxillary palpomere strongly triangular (cf.
<bibRefCitation id="EF870995F472DE6DC34CF999423D867E" author="Maier" box="[319,512,1596,1618]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
<figureCitation id="132D68E1F472DE6DC075F9994274867E" box="[518,585,1596,1618]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1887,1910]" captionTargetBox="[151,1436,337,1829]" captionTargetId="figure@5.[151,1436,337,1838]" captionTargetPageId="5" captionText="FIGURES 7 11. Lutrochus grenadensis n. sp.: 7. Habitus, dorsal view; 8. Habitus, ventral view; 9. Habitus, lateral view; 10. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 11. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228004/files/figure.png" pageId="12">Fig. 11</figureCitation>
)............................................
<taxonomicName id="4C160FE7F472DE6DC62FF99945A5867E" authority="Grouvelle" authorityName="Grouvelle" box="[1116,1432,1596,1618]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="acuminatus">
<emphasis id="B962A876F472DE6DC62FF999450E867E" box="[1116,1331,1596,1618]" italics="true" pageId="12">Lutrochus acuminatus</emphasis>
Grouvelle
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA97464F472DE6DC2E4F9FF45A68686" blockId="12.[151,1436,1479,2029]" pageId="12">
<keyLead id="30E7D751F472DE6DC2E4F9FF45A68686" pageId="12">
- Elytra not explanate, epipleuron appearing narrow in ventral aspect (
<figureCitation id="132D68E1F472DE6DC10EF9FC43FB8643" box="[893,966,1625,1647]" captionStart="FIGURES 19 23" captionStartId="11.[151,264,1938,1961]" captionTargetBox="[166,1414,422,1903]" captionTargetId="figure@11.[151,1436,409,1917]" captionTargetPageId="11" captionText="FIGURES 19 23. Lutrochus wao n. sp.: 19. Habitus, dorsal view; 20. Habitus, ventral view; 21. Habitus, lateral view; 22. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 23. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228007/files/figure.png" pageId="12">Fig. 20</figureCitation>
); dark brown to black in color; prosternum very slightly or not deflexed ventrally in anterior half (
<figureCitation id="132D68E1F472DE6DC0BAF9D2432F86A1" box="[713,786,1655,1677]" captionStart="FIGURES 19 23" captionStartId="11.[151,264,1938,1961]" captionTargetBox="[166,1414,422,1903]" captionTargetId="figure@11.[151,1436,409,1917]" captionTargetPageId="11" captionText="FIGURES 19 23. Lutrochus wao n. sp.: 19. Habitus, dorsal view; 20. Habitus, ventral view; 21. Habitus, lateral view; 22. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 23. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228007/files/figure.png" pageId="12">Fig. 20</figureCitation>
); apical maxillary palpomere weakly triangular to disc-shaped (cf.
<bibRefCitation id="EF870995F472DE6DC292F931419E8686" author="Maier" box="[225,419,1684,1706]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Figs. 30, 37)......................................................................... 3
</keyLead>
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<keyStep id="30E26CC1F472DE6DC2E4F91445A686C9" pageId="12">
<paragraph id="8BA97464F472DE6DC2E4F91445A686EB" blockId="12.[151,1436,1479,2029]" box="[151,1435,1713,1735]" pageId="12">
<keyLead id="30E7D751F472DE6DC2E4F91445A686EB" box="[151,1435,1713,1735]" pageId="12">
3(2) Body size small, total length
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................................................................... 4
</keyLead>
</paragraph>
<paragraph id="8BA97464F472DE6DC2E4F96A45A686C9" blockId="12.[151,1436,1479,2029]" box="[151,1435,1743,1765]" pageId="12">
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- Body size large, total length
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................................................................... 8
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<paragraph id="8BA97464F472DE6DC2E4F94945A18711" blockId="12.[151,1436,1479,2029]" pageId="12">
<keyLead id="30E7D751F472DE6DC2E4F94945A18711" pageId="12">
4(3) Pronotum with lateral borders expanded (cf.
<bibRefCitation id="EF870995F472DE6DC0FCF949436C872E" author="Maier" box="[655,849,1772,1794]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 35); apical maxillary palpomere rounded to disc-shaped (cf.
<bibRefCitation id="EF870995F472DE6DC374F8AC41F48733" author="Maier" box="[263,457,1801,1823]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 37); apical metatarsomere setose (cf.
<bibRefCitation id="EF870995F472DE6DC112F8AC441E8733" author="Maier" box="[865,1059,1801,1823]" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 38); length
<quantity id="4CEED981F472DE6DC6B0F8AC450B8733" box="[1219,1334,1801,1823]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="12" unit="mm" value="2.85" valueMax="3.2" valueMin="2.5">2.53.2 mm</quantity>
....................................................................................
<taxonomicName id="4C160FE7F472DE6DC61DF88D45A18711" authority="Maier and Short" authorityName="Maier and Short" box="[1134,1436,1831,1853]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="maldonadoi">
<emphasis id="B962A876F472DE6DC61DF88D44C78711" box="[1134,1274,1831,1853]" italics="true" pageId="12">L. maldonadoi</emphasis>
Maier and Short
</taxonomicName>
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</paragraph>
<paragraph id="8BA97464F472DE6DC2E4F8E045A1875B" blockId="12.[151,1436,1479,2029]" pageId="12">
<keyLead id="30E7D751F472DE6DC2E4F8E045A1875B" pageId="12">
- Pronotum with lateral borders smooth with outline of body, not expanded; apical maxillary palpomere triangular (cf.
<bibRefCitation id="EF870995F472DE6DC73AF8E14176875B" author="Maier" pageId="12" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 30); apical metatarsomere almost entirely glabrous (
<figureCitation id="132D68E1F472DE6DC103F8C44385875B" box="[880,952,1889,1911]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1887,1910]" captionTargetBox="[151,1436,337,1829]" captionTargetId="figure@5.[151,1436,337,1838]" captionTargetPageId="5" captionText="FIGURES 7 11. Lutrochus grenadensis n. sp.: 7. Habitus, dorsal view; 8. Habitus, ventral view; 9. Habitus, lateral view; 10. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 11. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228004/files/figure.png" pageId="12">Fig. 10</figureCitation>
); length 3.0
<quantity id="4CEED981F472DE6DC645F8C44443875B" box="[1078,1150,1889,1911]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="12" unit="mm" value="4.5">4.5 mm</quantity>
..................... 5
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<paragraph id="8BA97464F472DE6DC2E4F8DA45A187B9" blockId="12.[151,1436,1479,2029]" box="[151,1436,1919,1941]" pageId="12">
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5(4) Body strongly wedge-shaped...............................................................
<taxonomicName id="4C160FE7F472DE6DC68FF82545A187B9" authority="Sharp" authorityName="Sharp" box="[1276,1436,1919,1941]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="vestitus">
<emphasis id="B962A876F472DE6DC68FF825456387B9" box="[1276,1374,1920,1941]" italics="true" pageId="12">L. vestitus</emphasis>
Sharp
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA97464F472DE6DC2E4F83845A6879E" blockId="12.[151,1436,1479,2029]" box="[151,1435,1948,1970]" pageId="12">
<keyLead id="30E7D751F472DE6DC2E4F83845A6879E" box="[151,1435,1948,1970]" pageId="12">- Body ovate or weakly wedge-shaped..................................................................... 6</keyLead>
</paragraph>
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<paragraph id="8BA97464F472DE6DC2E4F81F45A187C1" blockId="12.[151,1436,1479,2029]" pageId="12">
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6(5) Apical metatarsomere pubescent for at least half of length when viewed dorsally (
<figureCitation id="132D68E1F472DE6DC189F81F447F87FC" box="[1018,1090,1978,2000]" captionStart="FIGURES 19 23" captionStartId="11.[151,264,1938,1961]" captionTargetBox="[166,1414,422,1903]" captionTargetId="figure@11.[151,1436,409,1917]" captionTargetPageId="11" captionText="FIGURES 19 23. Lutrochus wao n. sp.: 19. Habitus, dorsal view; 20. Habitus, ventral view; 21. Habitus, lateral view; 22. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 23. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228007/files/figure.png" pageId="12">Fig. 22</figureCitation>
); parameres of aedaegus with weak crenulations at border (
<figureCitation id="132D68E1F472DE6DC3CDF87241C287C1" box="[446,511,2007,2029]" captionStart="FIGURES 19 23" captionStartId="11.[151,264,1938,1961]" captionTargetBox="[166,1414,422,1903]" captionTargetId="figure@11.[151,1436,409,1917]" captionTargetPageId="11" captionText="FIGURES 19 23. Lutrochus wao n. sp.: 19. Habitus, dorsal view; 20. Habitus, ventral view; 21. Habitus, lateral view; 22. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 23. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228007/files/figure.png" pageId="12">Fig 23</figureCitation>
).................................................................
<emphasis id="B962A876F472DE6DC76EF87245A187C1" bold="true" box="[1309,1436,2007,2029]" pageId="12">
<taxonomicName id="4C160FE7F472DE6DC76EF872455D87C1" box="[1309,1376,2007,2029]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="wao" status="sp. nov.">
<emphasis id="B962A876F472DE6DC76EF872455D87C1" bold="true" box="[1309,1376,2007,2029]" italics="true" pageId="12">L. wao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251150DF472DE6DC715F87245A187C1" box="[1382,1436,2007,2029]" pageId="12" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA97464F473DE6CC2E4FF3245A680E6" blockId="13.[151,1436,151,730]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FF3245A680E6" pageId="13">
- Apical metatarsomere pubescent for significantly less than half of length when viewed dorsally (
<figureCitation id="132D68E1F473DE6CC6F2FF3244F58081" box="[1153,1224,151,173]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1887,1910]" captionTargetBox="[151,1436,337,1829]" captionTargetId="figure@5.[151,1436,337,1838]" captionTargetPageId="5" captionText="FIGURES 7 11. Lutrochus grenadensis n. sp.: 7. Habitus, dorsal view; 8. Habitus, ventral view; 9. Habitus, lateral view; 10. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 11. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228004/files/figure.png" pageId="13">Fig. 10</figureCitation>
); parameres of aedaegus smooth, lacking weak crenulations at border (
<figureCitation id="132D68E1F473DE6CC0CEFF1142C080E6" box="[701,765,180,202]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1887,1910]" captionTargetBox="[151,1436,337,1829]" captionTargetId="figure@5.[151,1436,337,1838]" captionTargetPageId="5" captionText="FIGURES 7 11. Lutrochus grenadensis n. sp.: 7. Habitus, dorsal view; 8. Habitus, ventral view; 9. Habitus, lateral view; 10. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 11. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228004/files/figure.png" pageId="13">Fig 11</figureCitation>
)..................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E26CC1F473DE6CC2E4FF7445AB816C" pageId="13">
<paragraph id="8BA97464F473DE6CC2E4FF7445A18129" blockId="13.[151,1436,151,730]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FF7445A18129" pageId="13">
7(6) Apical protarsomere two-toned, dark brown to nearly apex, testaceous at apex; femora light brown dorsally and dark brown ventrally; tibiae dark brown (
<figureCitation id="132D68E1F473DE6CC38AFF4A420E8129" box="[505,563,239,261]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1741,1764]" captionTargetBox="[151,1434,193,1713]" captionTargetId="figure@3.[151,1436,193,1720]" captionTargetPageId="3" captionText="FIGURES 3 6. Lutrochus funkae n. sp.: 3. Habitus, dorsal view; 4. Habitus, ventral view; 5. Habitus, lateral view; 6. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228003/files/figure.png" pageId="13">Fig. 5</figureCitation>
); apex of parameres not extended apically (
<figureCitation id="132D68E1F473DE6CC1A6FF4A44368129" box="[981,1035,239,261]" captionStart="FIGURES 3 6" captionStartId="3.[151,264,1741,1764]" captionTargetBox="[151,1434,193,1713]" captionTargetId="figure@3.[151,1436,193,1720]" captionTargetPageId="3" captionText="FIGURES 3 6. Lutrochus funkae n. sp.: 3. Habitus, dorsal view; 4. Habitus, ventral view; 5. Habitus, lateral view; 6. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228003/files/figure.png" pageId="13">Fig 6</figureCitation>
)....................
<emphasis id="B962A876F473DE6CC775FF4A45A18129" bold="true" box="[1286,1436,239,261]" pageId="13">
<taxonomicName id="4C160FE7F473DE6CC775FF4A45588129" box="[1286,1381,239,261]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="funkae" status="sp. nov.">
<emphasis id="B962A876F473DE6CC775FF4A45588129" bold="true" box="[1286,1381,239,261]" italics="true" pageId="13">L. funkae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251150DF473DE6CC71FFF4A45A18129" box="[1388,1436,239,261]" pageId="13" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BA97464F473DE6CC2E4FEA845AB816C" blockId="13.[151,1436,151,730]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FEA845AB816C" pageId="13">
- Apical protarsomere, femora, and tibae entirely reddish-brown to testaceous (
<figureCitation id="132D68E1F473DE6CC1B4FEA943C6810E" box="[967,1019,268,290]" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1887,1910]" captionTargetBox="[151,1436,337,1829]" captionTargetId="figure@5.[151,1436,337,1838]" captionTargetPageId="5" captionText="FIGURES 7 11. Lutrochus grenadensis n. sp.: 7. Habitus, dorsal view; 8. Habitus, ventral view; 9. Habitus, lateral view; 10. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 11. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228004/files/figure.png" pageId="13">Fig 8</figureCitation>
); apex of parameres extended apically (
<figureCitation id="132D68E1F473DE6CC709FEA940C78113" captionStart="FIGURES 7 11" captionStartId="5.[151,264,1887,1910]" captionTargetBox="[151,1436,337,1829]" captionTargetId="figure@5.[151,1436,337,1838]" captionTargetPageId="5" captionText="FIGURES 7 11. Lutrochus grenadensis n. sp.: 7. Habitus, dorsal view; 8. Habitus, ventral view; 9. Habitus, lateral view; 10. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 11. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228004/files/figure.png" pageId="13">Fig 11</figureCitation>
)...............................................................................
<emphasis id="B962A876F473DE6CC6BAFE8F45AB816C" bold="true" box="[1225,1430,298,320]" pageId="13">
<taxonomicName id="4C160FE7F473DE6CC6BAFE8F4564816C" box="[1225,1369,298,320]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="grenadensis" status="sp. nov.">
<emphasis id="B962A876F473DE6CC6BAFE8F4564816C" bold="true" box="[1225,1369,298,320]" italics="true" pageId="13">L. grenadensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251150DF473DE6CC713FE8F45AB816C" box="[1376,1430,298,320]" pageId="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E26CC1F473DE6CC2E4FEE245A18156" pageId="13">
<paragraph id="8BA97464F473DE6CC2E4FEE245A18171" blockId="13.[151,1436,151,730]" box="[151,1436,327,349]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FEE245A18171" box="[151,1436,327,349]" pageId="13">
8(3) Apical protarsomere entirely glabrous (cf.
<bibRefCitation id="EF870995F473DE6CC00AFEE243068171" author="Maier" box="[633,827,327,349]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
<figureCitation id="132D68E1F473DE6CC132FEE243B48171" box="[833,905,327,349]" captionStart="FIGURES 19 23" captionStartId="11.[151,264,1938,1961]" captionTargetBox="[166,1414,422,1903]" captionTargetId="figure@11.[151,1436,409,1917]" captionTargetPageId="11" captionText="FIGURES 19 23. Lutrochus wao n. sp.: 19. Habitus, dorsal view; 20. Habitus, ventral view; 21. Habitus, lateral view; 22. Diagrammatic view of the protibia and tarsus, showing areas of dense pubescence (pubescent areas shaded in gray); 23. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228007/files/figure.png" pageId="13">Fig. 23</figureCitation>
)....................
<taxonomicName id="4C160FE7F473DE6CC60DFEED45A18171" authority="Maier and Short" authorityName="Maier and Short" box="[1150,1436,327,349]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="gustafsoni">
<emphasis id="B962A876F473DE6CC60DFEED44C78171" box="[1150,1274,327,349]" italics="true" pageId="13">L. gustafsoni</emphasis>
Maier and Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA97464F473DE6CC2E4FEC045A18156" blockId="13.[151,1436,151,730]" box="[151,1436,356,378]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FEC045A18156" box="[151,1436,356,378]" pageId="13">
- Apical protarsomere at least partially pubescent (cf.
<bibRefCitation id="EF870995F473DE6CC0A5FEC143A58156" author="Maier" box="[726,920,356,378]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
<figureCitation id="132D68E1F473DE6CC1EDFEC143DF8156" box="[926,994,356,378]" captionStart="FIGURE 18" captionStartId="9.[151,250,1458,1480]" captionTargetBox="[151,1401,193,1375]" captionTargetId="figure@9.[151,1435,193,1376]" captionTargetPageId="9" captionText="FIGURE 18. Lutrochus grenadensis n. sp. (St. George population): Aedeagus, dorsal and lateral views, showing probable injury (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228006/files/figure.png" pageId="13">Fig. 18</figureCitation>
).................................. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E26CC1F473DE6CC2E4FE2445A18199" pageId="13">
<paragraph id="8BA97464F473DE6CC2E4FE2445A181BB" blockId="13.[151,1436,151,730]" box="[151,1436,385,408]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FE2445A181BB" box="[151,1436,385,408]" pageId="13">
9(8) Apex of protibia of male with distinct patch of long golden setae apically (
<figureCitation id="132D68E1F473DE6CC1D8FE2443CC81BB" box="[939,1009,385,407]" captionStart="FIGURES 16 17" captionStartId="9.[151,264,1396,1419]" captionText="FIGURES 16 17. Lutrochus leeanneae n. sp.: 16. Protibia and tarsus, showing patch of dense setae on protibia; 17. Mesotibia and tarsus, showing lateral expansion of mesotibia." pageId="13">Fig. 16</figureCitation>
)...................
<emphasis id="B962A876F473DE6CC692FE2745A881B4" bold="true" box="[1249,1429,386,408]" pageId="13">
<taxonomicName id="4C160FE7F473DE6CC692FE27455D81B4" box="[1249,1376,386,408]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="leeanneae" status="sp. nov.">
<emphasis id="B962A876F473DE6CC692FE27455D81B4" bold="true" box="[1249,1376,386,408]" italics="true" pageId="13">L. leeanneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A251150DF473DE6CC715FE2745A881B4" box="[1382,1429,386,408]" pageId="13" rank="species">n. sp</taxonomicNameLabel>
</emphasis>
.
</keyLead>
</paragraph>
<paragraph id="8BA97464F473DE6CC2E4FE3A45A18199" blockId="13.[151,1436,151,730]" box="[151,1436,415,437]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FE3A45A18199" box="[151,1436,415,437]" pageId="13">
- Apex of protibia of male with setae of similar length on entire surface (cf.
<bibRefCitation id="EF870995F473DE6CC1D8FE3A44518199" author="Maier" box="[939,1132,415,437]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
<figureCitation id="132D68E1F473DE6CC601FE3A448B8199" box="[1138,1206,415,437]" captionStart="FIGURES 12 15" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[173,1428,421,1875]" captionTargetId="figure@8.[151,1436,409,1880]" captionTargetPageId="8" captionText="FIGURES 12 15. Lutrochus leeanneae n. sp.: 12. Habitus, dorsal view; 13. Habitus, ventral view; 14. Habitus, lateral view; 15. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228005/files/figure.png" pageId="13">Fig. 15</figureCitation>
)............... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E26CC1F473DE6CC2E4FE1945A68206" pageId="13">
<paragraph id="8BA97464F473DE6CC2E4FE1945A681C3" blockId="13.[151,1436,151,730]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FE1945A681C3" pageId="13">
10(9) Apical protarsomere with discrete pubescent patch in basal third, apical two-thirds glabrous (cf.
<bibRefCitation id="EF870995F473DE6CC60CFE19457F81FE" author="Maier" box="[1151,1346,444,466]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
<figureCitation id="132D68E1F473DE6CC73BFE1945B281FE" box="[1352,1423,444,466]" captionStart="FIGURE 18" captionStartId="9.[151,250,1458,1480]" captionTargetBox="[151,1401,193,1375]" captionTargetId="figure@9.[151,1435,193,1376]" captionTargetPageId="9" captionText="FIGURE 18. Lutrochus grenadensis n. sp. (St. George population): Aedeagus, dorsal and lateral views, showing probable injury (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228006/files/figure.png" pageId="13">Fig. 18</figureCitation>
); body without metallic sheen dorsally (cf.
<bibRefCitation id="EF870995F473DE6CC003FE7C430C81C3" author="Maier" box="[624,817,473,495]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
<figureCitation id="132D68E1F473DE6CC144FE7C434181C3" box="[823,892,473,495]" captionStart="FIGURES 12 15" captionStartId="8.[151,264,1901,1924]" captionTargetBox="[173,1428,421,1875]" captionTargetId="figure@8.[151,1436,409,1880]" captionTargetPageId="8" captionText="FIGURES 12 15. Lutrochus leeanneae n. sp.: 12. Habitus, dorsal view; 13. Habitus, ventral view; 14. Habitus, lateral view; 15. Aedeagus, dorsal and lateral views (scale bar = 1.0 mm)." httpUri="https://zenodo.org/record/228005/files/figure.png" pageId="13">Fig. 14</figureCitation>
).....................
<taxonomicName id="4C160FE7F473DE6CC6FBFE7F45A681C3" authority="Maier &amp; Short" authorityName="Maier &amp; Short" box="[1160,1435,473,495]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="cauraensis">
<emphasis id="B962A876F473DE6CC6FBFE7F453781C3" box="[1160,1290,474,495]" italics="true" pageId="13">L. cauraensis</emphasis>
Maier &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA97464F473DE6CC2E4FE5245A68206" blockId="13.[151,1436,151,730]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FE5245A68206" pageId="13">
- Apical protarsomere pubescent, although more sparse in apical half (cf.
<bibRefCitation id="EF870995F473DE6CC1EBFE5244668221" author="Maier" box="[920,1115,503,525]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 58); body with bluish-violet metallic sheen dorsally (cf.
<bibRefCitation id="EF870995F473DE6CC399FDB142918206" author="Maier" box="[490,684,532,554]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 55).................................
<taxonomicName id="4C160FE7F473DE6CC6FAFDB045A68206" authority="Maier &amp; Short" authorityName="Maier &amp; Short" box="[1161,1435,532,554]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="violaceous">
<emphasis id="B962A876F473DE6CC6FAFDB045348206" box="[1161,1289,532,554]" italics="true" pageId="13">L. violaceous</emphasis>
Maier &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E26CC1F473DE6CC2E4FD9445A682F6" pageId="13">
<paragraph id="8BA97464F473DE6CC2E4FD9445A682AE" blockId="13.[151,1436,151,730]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FD9445A682AE" pageId="13">
11(1) Metasternum with medial circular glabrous patch (cf.
<bibRefCitation id="EF870995F473DE6CC09DFD944389826B" author="Maier" box="[750,948,561,583]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 45); scutellum diamond-shaped (cf.
<bibRefCitation id="EF870995F473DE6CC73BFD9441768249" author="Maier" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 46); elytron with seven rows of shallow punctures (cf.
<bibRefCitation id="EF870995F473DE6CC1F9FDEA44718249" author="Maier" box="[906,1100,591,613]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 40); apical maxillary palpomere narrow (cf.
<bibRefCitation id="EF870995F473DE6CC307FDC9420B82AE" author="Maier" box="[372,566,620,642]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 43).........................................
<taxonomicName id="4C160FE7F473DE6CC60FFDC845A682AE" authority="Maier &amp; Short" authorityName="Maier &amp; Short" box="[1148,1435,620,642]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="meridaensis">
<emphasis id="B962A876F473DE6CC60FFDC8453482AE" box="[1148,1289,620,642]" italics="true" pageId="13">L. meridaensis</emphasis>
Maier &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BA97464F473DE6CC2E4FD2F45A682F6" blockId="13.[151,1436,151,730]" pageId="13">
<keyLead id="30E7D751F473DE6CC2E4FD2F45A682F6" pageId="13">
- Metasternum without medial circular glabrous patch (cf.
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Fig. 49); scutellum triangular (cf.
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Fig. 48); elytral punctation dense, confused, not in rows (cf.
<bibRefCitation id="EF870995F473DE6CC119FD02440D8291" author="Maier" box="[874,1072,679,701]" pageId="13" refString="Maier, C. A. &amp; Short, A. E. Z. (2013) A revision of the Lutrochidae (Coleoptera) of Venezuela, with description of six new species and key. Zootaxa, 3637 (3), 285 - 307. http: // dx. doi. org / 10.11646 / zootaxa. 3637.3.3" type="journal article" year="2013">Maier &amp; Short 2013</bibRefCitation>
Fig. 52); apical maxillary palpomere wide (cf.
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Fig. 53).................................................
<taxonomicName id="4C160FE7F473DE6CC6D7FD6045A682F6" authority="Maier &amp; Short" authorityName="Maier &amp; Short" box="[1188,1435,708,730]" class="Insecta" family="Lutrochidae" genus="Lutrochus" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="minutus">
<emphasis id="B962A876F473DE6CC6D7FD60453782F6" box="[1188,1290,709,730]" italics="true" pageId="13">L. minutus</emphasis>
Maier &amp; Short
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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