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<document id="242E090B3EF2707516F2A7DE86B288C8" ID-CLB-Dataset="21420" ID-DOI="10.5852/ejt.2020.684" ID-GBIF-Dataset="c6ceea2b-a36e-4bda-bc7b-32e66b22a899" ID-Zenodo-Dep="3959113" ID-ZooBank="B88F38A8-C3C7-47BA-8F1B-46D9F127F10B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595596762572" checkinUser="valdenar" docAuthor="Haran, Julien M., Beaudoin-Ollivier, Laurence, Benoit, Laure &amp; Kuschel, Guillermo" docDate="2020" docId="03ABCE59FFF1FF81BEB1CD756A15FE50" docLanguage="en" docName="ejt-684_haran_beaudin-olivier_benoit.pdf.imd" docOrigin="European Journal of Taxonomy 684" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Elaeidobius spatulifer Haran &amp; Beaudoin-Ollivier &amp; Benoit &amp; Kuschel 2020, comb. nov." docType="treatment" docVersion="10" lastPageNumber="9" masterDocId="FF92B621FFF6FF89BF5BCC7F6848FFCA" masterDocTitle="Revision of the palm-pollinating weevil genus Elaeidobius Kuschel, 1952 (Curculionidae, Curculioninae, Derelomini) with descriptions of two new species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="8" updateTime="1698843959693" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AE614FF7CDB599A66EE4F49FFCB37459">Revision of the palm-pollinating weevil genus Elaeidobius Kuschel, 1952 (Curculionidae, Curculioninae, Derelomini) with descriptions of two new species</mods:title>
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<mods:namePart id="87A271EA925818009626B2A2ADB4A305">Haran, Julien M.</mods:namePart>
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<mods:affiliation id="166E48F280909C5916FB86B8D61DEACF">Univ Montpellier, CIRAD, INRA, IRD, Montpellier SupAgro, Montpellier, France.</mods:affiliation>
<mods:nameIdentifier id="06E71E67079C80C193E88B59A03A1D98" type="email">julien.haran@cirad.fr</mods:nameIdentifier>
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<mods:roleTerm id="FCD00F188B8FC04E0A19B7AE7BDB7E5E">Author</mods:roleTerm>
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<mods:namePart id="D08F9782C3450BA575D2860E9C59E75B">Beaudoin-Ollivier, Laurence</mods:namePart>
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<mods:affiliation id="232EDF1E77B54A804FF13A0B9ADD25EA">Systèmes de Pérennes, CIRAD, Univ Montpellier, Montpellier, France.</mods:affiliation>
<mods:nameIdentifier id="E42068677724EAC252AD47C7AC8DC807" type="email">laurence.ollivier@cirad.fr</mods:nameIdentifier>
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<mods:namePart id="70D1BBDBC08DE204F9690A63959E0991">Benoit, Laure</mods:namePart>
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<mods:affiliation id="E78115502F3BA3F7BF89C0FF9D0A1EAA">Univ Montpellier, CIRAD, INRA, IRD, Montpellier SupAgro, Montpellier, France.</mods:affiliation>
<mods:nameIdentifier id="9BEC9BA6426EEEF7CBCDD17F77A90F1E" type="email">laure.benoit@cirad.fr</mods:nameIdentifier>
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<mods:namePart id="E86BAE04DC1CC91C888F1057EE51B4B2">Kuschel, Guillermo</mods:namePart>
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<mods:affiliation id="AEB22BB073B7333137E0A47DC627E7DD">Deceased 1 Aug. 2017.</mods:affiliation>
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<mods:title id="AC90C2499A3714D3E13BD6DF85FB9C0F">European Journal of Taxonomy</mods:title>
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<mods:date id="7B35FB524B054B3EE8635B42CE3B225D">2020</mods:date>
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<mods:number id="B404B003C726992023EEFF4958F7444B">684</mods:number>
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<treatment id="03ABCE59FFF1FF81BEB1CD756A15FE50" ID-DOI="http://doi.org/10.5281/zenodo.4331368" ID-GBIF-Taxon="165940879" ID-Zenodo-Dep="4331368" LSID="urn:lsid:plazi:treatment:03ABCE59FFF1FF81BEB1CD756A15FE50" httpUri="http://treatment.plazi.org/id/03ABCE59FFF1FF81BEB1CD756A15FE50" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection id="C3182CC4FFF1FF8EBEB1CD756C02FEEF" box="[490,1098,266,294]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BBD7F4FFFF1FF8EBEB1CD756C02FEEF" blockId="7.[490,1098,266,328]" box="[490,1098,266,294]" pageId="7" pageNumber="8">
<heading id="D0F5C823FFF1FF8EBEB1CD756C02FEEF" box="[490,1098,266,294]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="4C0204CCFFF1FF8EBEB1CD756B89FEEC" authority="(Marshall, 1950)" authorityName="Haran &amp; Beaudoin-Ollivier &amp; Benoit &amp; Kuschel" authorityYear="2020" baseAuthorityName="Marshall" baseAuthorityYear="1950" box="[490,961,266,294]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer" status="comb. nov.">
<emphasis id="B976A35DFFF1FF8EBEB1CD756ABDFEEF" bold="true" box="[490,757,266,293]" italics="true" pageId="7" pageNumber="8">Elaeidobius spatulifer</emphasis>
(
<bibRefCitation id="EF9302BEFFF1FF8EBC5FCD746BF0FEEC" author="Marshall" box="[772,952,267,294]" firstAuthor="Marshall" pageId="7" pageNumber="8" pagination="725 - 750" refId="ref16691" refString="Marshall G. A. K. 1950. LXV. - New Curculionidae (Col.) from tropical Africa. Journal of Natural History Series 12, 3 (33): 725 - 750. https: // doi. org / 10.1080 / 00222935008654101" type="journal article" year="1950">Marshall, 1950</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2451E26FFF1FF8EBC93CD746C02FEEF" box="[968,1098,267,293]" pageId="7" pageNumber="8" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3182CC4FFF1FF8EBDE0CD516B31FE82" box="[699,889,302,328]" pageId="7" pageNumber="8" type="description">
<paragraph id="8BBD7F4FFFF1FF8EBDE0CD516B31FE82" blockId="7.[490,1098,266,328]" box="[699,889,302,328]" pageId="7" pageNumber="8">
<figureCitation id="133963CAFFF1FF8EBDE0CD516B50FE82" box="[699,792,302,328]" captionStart="Fig" captionStartId="8.[189,231,1805,1831]" captionTargetBox="[270,1357,796,1756]" captionTargetId="figure@8.[241,1390,776,1768]" captionTargetPageId="8" captionText="Fig. 1. Habitus of males and females of species of Elaeidobius Kuschel, 1952. A. E. bilineatus (Hustache, 1924) comb. nov., Dakar, Senegal. B. E. spatulifer (Marshall, 1950) comb. nov., holotype, Thysville, Democratic Republic of Congo (MRAC). C. E. subvittatus (Faust, 1898), Dakar, Senegal. D. E. kamerunicus (Faust, 1898) comb. nov. (JHAR00209). E. E. pilimargo Haran &amp; Kuschel sp. nov. (JHAR00409). F. E. piliventris Haran &amp; Kuschel sp. nov. (JHAR02182). G. E. plagiatus (Fåhraeus, 1844) comb. nov. (JHAR00272). H. E. singularis (Faust, 1898) comb. nov. (JHAR00283). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3959115" httpUri="https://zenodo.org/record/3959115/files/figure.png" pageId="7" pageNumber="8">Figs 1B</figureCitation>
,
<figureCitation id="133963CAFFF1FF8EBC7ECD516B02FE82" box="[805,842,302,328]" captionStart="Fig" captionStartId="11.[189,232,1885,1911]" captionTargetBox="[225,1361,291,1818]" captionTargetId="figure@11.[189,1399,262,1835]" captionTargetPageId="11" captionText="Fig. 2. Head and prothorax in lateral view of species of Elaeidobius Kuschel, 1952 (Part 1). A. E. bilineatus (Hustache, 1924) comb. nov., Dakar, Senegal. B. E. spatulifer (Marshall, 1950) comb. nov., holotype, Thysville, Democratic Republic of Congo (MRAC). C. E. subvittatus (Faust, 1898), Dakar, Senegal. D. E. kamerunicus (Faust, 1898) comb. nov. (JHAR00209). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.3959117" httpUri="https://zenodo.org/record/3959117/files/figure.png" pageId="7" pageNumber="8">2B</figureCitation>
,
<figureCitation id="133963CAFFF1FF8EBC0DCD516B31FE82" box="[854,889,302,328]" captionStart="Fig" captionStartId="18.[189,232,1781,1807]" captionTargetBox="[203,1381,343,1664]" captionTargetId="figure@18.[189,1398,326,1679]" captionTargetPageId="18" captionText="Fig. 4. Penis of males of species of the genus Elaeidobius Kuschel, 1952 in dorsal (left) and lateral (right) view. A. E. bilineatus (Hustache, 1924) comb. nov., Dakar, Senegal. B. E. spatulifer (Marshall, 1950) comb. nov., holotype, Thysville, Democratic Republic of Congo (MRAC). C. E. subvittatus (Faust, 1898), Dakar, Senegal. D. E. kamerunicus (Faust, 1898) comb. nov. (JHAR00209). E. E. pilimargo Haran &amp; Kuschel sp. nov. (JHAR00409). F. E. piliventris Haran &amp; Kuschel sp. nov. (JHAR02182). G. E. plagiatus (Fåhraeus, 1844) comb. nov. (JHAR00272). H. E. singularis (Faust, 1898) comb. nov. (JHAR00283). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3959121" httpUri="https://zenodo.org/record/3959121/files/figure.png" pageId="7" pageNumber="8">4B</figureCitation>
</paragraph>
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<subSubSection id="C3182CC4FFF1FF8EBFE6CD0F6AF8FE43" box="[189,688,367,394]" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="8BBD7F4FFFF1FF8EBFE6CD0F6AF8FE43" blockId="7.[189,688,367,394]" box="[189,688,367,394]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="AB125861FFF1FF8EBFE6CD0F6AF8FE43" box="[189,688,367,394]" pageId="7" pageNumber="8">
<treatmentCitation id="0AA3595EFFF1FF8EBFE6CD0F6AE5FE43" author="Marshall G. A. K." box="[189,685,367,394]" page="264" pageId="7" pageNumber="8" year="1950">
<taxonomicName id="4C0204CCFFF1FF8EBFE6CD0F6AE5FE43" authority="Marshall, 1950: 264" authorityName="Marshall" authorityPageNumber="264" authorityYear="1950" box="[189,685,367,394]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer">
<emphasis id="B976A35DFFF1FF8EBFE6CD0F69FCFE43" box="[189,436,367,393]" italics="true" pageId="7" pageNumber="8">Prosoestus spatulifer</emphasis>
<bibRefCitation id="EF9302BEFFF1FF8EBEE7CD106AE5FE43" author="Marshall" box="[444,685,367,394]" firstAuthor="Marshall" pageId="7" pageNumber="8" pagination="725 - 750" refId="ref16691" refString="Marshall G. A. K. 1950. LXV. - New Curculionidae (Col.) from tropical Africa. Journal of Natural History Series 12, 3 (33): 725 - 750. https: // doi. org / 10.1080 / 00222935008654101" type="journal article" year="1950">Marshall, 1950: 264</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3182CC4FFF1FF8EBFE6CDCE6BBAF894" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BBD7F4FFFF1FF8EBFE6CDCE69E2FE01" blockId="7.[189,426,433,459]" box="[189,426,433,459]" pageId="7" pageNumber="8">
<heading id="D0F5C823FFF1FF8EBFE6CDCE69E2FE01" bold="true" box="[189,426,433,459]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B976A35DFFF1FF8EBFE6CDCE69E2FE01" bold="true" box="[189,426,433,459]" pageId="7" pageNumber="8">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFB1CD9F6914FE30" blockId="7.[189,1399,480,647]" box="[234,348,480,506]" pageId="7" pageNumber="8">
<emphasis id="B976A35DFFF1FF8EBFB1CD9F6914FE30" bold="true" box="[234,348,480,506]" pageId="7" pageNumber="8">
<typeStatus id="54B9C1EDFFF1FF8EBFB1CD9F6914FE30" box="[234,348,480,506]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CE7C6A65FD4C" blockId="7.[189,1399,480,647]" pageId="7" pageNumber="8">
<materialsCitation id="3B6A7512FFF1FF8EBFE6CE7C6A65FD4C" ID-GBIF-Occurrence="2817897303" collectingDate="1935-11" collectionCode="MRAC" country="Democratic Republic of the Congo" location="DEMOCRATIC REPUBLIC OF CONGO" pageId="7" pageNumber="8" specimenCount="3" specimenCount-male="3" typeStatus="holotype">
<collectingCountry id="F3153FDFFFF1FF8EBFE6CE7C6AEDFDD7" box="[189,677,515,541]" name="Democratic Republic of the Congo" pageId="7" pageNumber="8">DEMOCRATIC REPUBLIC OF CONGO</collectingCountry>
• “Musée du Congo.Thysville. -
<date id="FFBC598FFFF1FF8EBB4CCE7D6CCDFDD7" box="[1047,1157,514,541]" pageId="7" pageNumber="8" value="1935-11">
<collectingDate id="EFF8A067FFF1FF8EBB4CCE7D6CCDFDD7" box="[1047,1157,514,541]" pageId="7" pageNumber="8" value="1935-11">XI–1935</collectingDate>
</date>
. 2445. J Ghesquière” “
<typeStatus id="54B9C1EDFFF1FF8EBF9CCE596934FD8A" box="[199,380,550,576]" pageId="7" pageNumber="8" type="holotype">HOLOTYPUS</typeStatus>
.
<taxonomicName id="4C0204CCFFF1FF8EBEDCCE5969BFFD8A" authorityName="Marshall" authorityYear="1950" box="[391,503,550,576]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer">
<emphasis id="B976A35DFFF1FF8EBEDCCE5969BFFD8A" box="[391,503,550,576]" italics="true" pageId="7" pageNumber="8">spatulifer</emphasis>
</taxonomicName>
<specimenCount id="9D04B4C6FFF1FF8EBEA5CE596A5EFD8A" box="[510,534,550,576]" pageId="7" pageNumber="8" type="male">♂</specimenCount>
. Marsh [red label]” “
<taxonomicName id="4C0204CCFFF1FF8EBC55CE586C4DFD8A" authorityName="Marshall" authorityYear="1950" box="[782,1029,550,576]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer">
<emphasis id="B976A35DFFF1FF8EBC55CE586C4DFD8A" box="[782,1029,550,576]" italics="true" pageId="7" pageNumber="8">Prosoestus spatulifer</emphasis>
</taxonomicName>
. Mshl.
<typeStatus id="54B9C1EDFFF1FF8EBB05CE596CEFFD8A" box="[1118,1191,550,576]" pageId="7" pageNumber="8">TYPE</typeStatus>
<specimenCount id="9D04B4C6FFF1FF8EBBF5CE596C8CFD8A" box="[1198,1220,550,576]" pageId="7" pageNumber="8" type="male">♂</specimenCount>
” “
<typeStatus id="54B9C1EDFFF1FF8EBBB9CE596D1AFD8A" box="[1250,1362,550,576]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
<specimenCount id="9D04B4C6FFF1FF8EBA02CE596D39FD8A" box="[1369,1393,550,576]" pageId="7" pageNumber="8" type="male">♂</specimenCount>
.
<taxonomicName id="4C0204CCFFF1FF8EBFE6CE356975FDA9" authorityName="Faust" authorityYear="1899" box="[189,317,586,611]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B976A35DFFF1FF8EBFE6CE356975FDA9" box="[189,317,586,611]" italics="true" pageId="7" pageNumber="8">Prosoestus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C0204CCFFF1FF8EBE14CE3669F7FDA9" authorityName="Marshall" authorityYear="1950" box="[335,447,585,611]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer">
<emphasis id="B976A35DFFF1FF8EBE14CE3669F7FDA9" box="[335,447,585,611]" italics="true" pageId="7" pageNumber="8">spatulifer</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF9302BEFFF1FF8EBE89CE366ACBFDAE" author="Marshall" box="[466,643,585,612]" firstAuthor="Marshall" pageId="7" pageNumber="8" pagination="725 - 750" refId="ref16691" refString="Marshall G. A. K. 1950. LXV. - New Curculionidae (Col.) from tropical Africa. Journal of Natural History Series 12, 3 (33): 725 - 750. https: // doi. org / 10.1080 / 00222935008654101" type="journal article" year="1950">Marshall 1950</bibRefCitation>
. Kuschel 2009 [red label]” “
<emphasis id="B976A35DFFF1FF8EBCBDCE366CB2FDA9" box="[998,1274,585,611]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C0204CCFFF1FF8EBCBDCE366C3EFDA9" box="[998,1142,585,611]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Elaeidobius</taxonomicName>
.
<taxonomicName id="4C0204CCFFF1FF8EBBDECE366CBEFDA9" authorityName="Marshall" authorityYear="1950" box="[1157,1270,585,611]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer">spatulifer</taxonomicName>
.
</emphasis>
(
<bibRefCitation id="EF9302BEFFF1FF8EBA57CE3668B4FD4D" author="Marshall" firstAuthor="Marshall" pageId="7" pageNumber="8" pagination="725 - 750" refId="ref16691" refString="Marshall G. A. K. 1950. LXV. - New Curculionidae (Col.) from tropical Africa. Journal of Natural History Series 12, 3 (33): 725 - 750. https: // doi. org / 10.1080 / 00222935008654101" type="journal article" year="1950">Marshall 1950</bibRefCitation>
). Kuschel 2009”;
<collectionCode id="ED13E78AFFF1FF8EBE94CE136A61FD4C" box="[463,553,620,646]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="7" pageNumber="8" type="Museum">MRAC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFB1CED1692FFD02" blockId="7.[189,1399,686,853]" box="[234,359,686,712]" pageId="7" pageNumber="8">
<emphasis id="B976A35DFFF1FF8EBFB1CED1692FFD02" bold="true" box="[234,359,686,712]" pageId="7" pageNumber="8">
<typeStatus id="54B9C1EDFFF1FF8EBFB1CED1692FFD02" box="[234,359,686,712]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CEAE6A63FC9F" blockId="7.[189,1399,686,853]" pageId="7" pageNumber="8">
<materialsCitation id="3B6A7512FFF1FF8EBFE6CEAE6C38FCFB" ID-GBIF-Occurrence="2817897324" collectionCode="MRAC" country="Democratic Republic of the Congo" location="DEMOCRATIC REPUBLIC OF CONGO" pageId="7" pageNumber="8" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectingCountry id="F3153FDFFFF1FF8EBFE6CEAE6AFEFD21" box="[189,694,721,747]" name="Democratic Republic of the Congo" pageId="7" pageNumber="8">DEMOCRATIC REPUBLIC OF CONGO</collectingCountry>
• same collection data as for holotype; “
<typeStatus id="54B9C1EDFFF1FF8EBBFDCEAE6D1BFD21" box="[1190,1363,721,747]" pageId="7" pageNumber="8" type="paratype">PARATYPUS</typeStatus>
.
<specimenCount id="9D04B4C6FFF1FF8EBA3BCEAE6D3EFD21" box="[1376,1398,721,747]" pageId="7" pageNumber="8" type="female">♀</specimenCount>
[red label]” “
<taxonomicName id="4C0204CCFFF1FF8EBE02CE8A6A18FCC4" authorityName="Marshall" authorityYear="1950" box="[345,592,756,782]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer">
<emphasis id="B976A35DFFF1FF8EBE02CE8A6A18FCC4" box="[345,592,756,782]" italics="true" pageId="7" pageNumber="8">Prosoestus spatulifer</emphasis>
</taxonomicName>
. Mshl. Cotype
<specimenCount id="9D04B4C6FFF1FF8EBC53CE8B6B54FCC4" box="[776,796,756,782]" pageId="7" pageNumber="8" type="female">♀</specimenCount>
” “
<typeStatus id="54B9C1EDFFF1FF8EBC66CE8B6BE2FCC4" box="[829,938,756,782]" pageId="7" pageNumber="8" type="paratype">Paratype</typeStatus>
.
<taxonomicName id="4C0204CCFFF1FF8EBCEDCE8A6C7EFCC4" authorityName="Faust" authorityYear="1899" box="[950,1078,757,782]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B976A35DFFF1FF8EBCEDCE8A6C7EFCC4" box="[950,1078,757,782]" italics="true" pageId="7" pageNumber="8">Prosoestus</emphasis>
</taxonomicName>
.
<taxonomicName id="4C0204CCFFF1FF8EBB1ECE8B6CFDFCC4" authorityName="Marshall" authorityYear="1950" box="[1093,1205,756,782]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer">
<emphasis id="B976A35DFFF1FF8EBB1ECE8B6CFDFCC4" box="[1093,1205,756,782]" italics="true" pageId="7" pageNumber="8">spatulifer</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF9302BEFFF1FF8EBB9FCE8B6D3AFCC5" author="Marshall" box="[1220,1394,756,783]" firstAuthor="Marshall" pageId="7" pageNumber="8" pagination="725 - 750" refId="ref16691" refString="Marshall G. A. K. 1950. LXV. - New Curculionidae (Col.) from tropical Africa. Journal of Natural History Series 12, 3 (33): 725 - 750. https: // doi. org / 10.1080 / 00222935008654101" type="journal article" year="1950">Marshall 1950</bibRefCitation>
. Kuschel 2009” “
<emphasis id="B976A35DFFF1FF8EBE25CF686AC5FCFB" box="[382,653,791,817]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C0204CCFFF1FF8EBE25CF686A46FCFB" box="[382,526,791,817]" class="Insecta" family="Curculionidae" genus="Elaeidobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Elaeidobius</taxonomicName>
.
<taxonomicName id="4C0204CCFFF1FF8EBD43CF686AC1FCFB" authorityName="Marshall" authorityYear="1950" box="[536,649,791,817]" class="Insecta" family="Curculionidae" genus="Prosoestus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="spatulifer">spatulifer</taxonomicName>
.
</emphasis>
(
<bibRefCitation id="EF9302BEFFF1FF8EBDC0CF686B02FCF8" author="Marshall" box="[667,842,791,818]" firstAuthor="Marshall" pageId="7" pageNumber="8" pagination="725 - 750" refId="ref16691" refString="Marshall G. A. K. 1950. LXV. - New Curculionidae (Col.) from tropical Africa. Journal of Natural History Series 12, 3 (33): 725 - 750. https: // doi. org / 10.1080 / 00222935008654101" type="journal article" year="1950">Marshall 1950</bibRefCitation>
). Kuschel 2009”;
<collectionCode id="ED13E78AFFF1FF8EBB41CF686C38FCFB" box="[1050,1136,791,817]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="7" pageNumber="8" type="Museum">MRAC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B6A7512FFF1FF8EBBDCCF686A63FC9F" ID-GBIF-Occurrence="2817897350" collectionCode="NHMUK" country="Democratic Republic of the Congo" location="DEMOCRATIC REPUBLIC OF CONGO" pageId="7" pageNumber="8" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D04B4C6FFF1FF8EBBDCCF686CFBFCF8" box="[1159,1203,791,818]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED13E78AFFF1FF8EBEEECF446A60FC9F" box="[437,552,827,853]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="8" type="Museum">NHMUK</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFB1CF0369EEFC5C" blockId="7.[189,1121,892,954]" box="[234,422,892,918]" pageId="7" pageNumber="8">
<emphasis id="B976A35DFFF1FF8EBFB1CF0369EEFC5C" bold="true" box="[234,422,892,918]" pageId="7" pageNumber="8">Other material</emphasis>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CFE06C29FC73" blockId="7.[189,1121,892,954]" box="[189,1121,927,954]" pageId="7" pageNumber="8">
<materialsCitation id="3B6A7512FFF1FF8EBFE6CFE06C29FC73" ID-GBIF-Occurrence="2817897323" box="[189,1121,927,954]" collectionCode="NHMUK, NZAC" collectorName="G. Kuschel" pageId="7" pageNumber="8" specimenCount="1">
Other specimens identified and labelled by
<collectorName id="26F71A99FFF1FF8EBDE2CFE06B75FC73" box="[697,829,927,953]" pageId="7" pageNumber="8">G. Kuschel</collectorName>
at
<collectionCode id="ED13E78AFFF1FF8EBC3ACFDF6B98FC70" box="[865,976,928,954]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="8" type="Museum">NHMUK</collectionCode>
and
<collectionCode id="ED13E78AFFF1FF8EBB52CFE06C15FC73" box="[1033,1117,927,953]" collectionName="New Zealand, Auckland, Landcare Research, New Zealand Arthropod Collection" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="7" pageNumber="8" type="Other institutional type">NZAC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CF9F6934FC31" blockId="7.[189,380,992,1019]" box="[189,380,992,1019]" pageId="7" pageNumber="8">
<emphasis id="B976A35DFFF1FF8EBFE6CF9F697EFC30" bold="true" box="[189,310,992,1018]" pageId="7" pageNumber="8">Diagnosis</emphasis>
(♂
<specimenCount id="9D04B4C6FFF1FF8EBE04CF9E693AFC31" box="[351,370,993,1019]" pageId="7" pageNumber="8" type="female">♀</specimenCount>
)
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6C86F69B2FBE0" blockId="7.[189,506,1039,1066]" box="[189,506,1039,1066]" pageId="7" pageNumber="8">
BODY LENGTH.
<quantity id="4CFAD2AAFFF1FF8EBE3EC87069BDFBE0" box="[357,501,1039,1066]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="7" pageNumber="8" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
.
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6C82E6BB3FB44" blockId="7.[189,1398,1104,1166]" pageId="7" pageNumber="8">COLOUR. Derm dorsally reddish brown, ventrally dark brown, elytra vaguely darker on interstria 2 and laterally on interstriae 6 and 7; antennae and legs reddish or yellowish.</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6C8C96CF7FB38" blockId="7.[189,1399,1205,1267]" pageId="7" pageNumber="8">HEAD. Forehead not impressed in either sex. Rostrum in male about as long as prothorax, in female 1.2 × as long, moderately robust and downcurved, with a slight postmental elevation in male.</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6C9656A8DFAB0" blockId="7.[189,1399,1306,1402]" pageId="7" pageNumber="8">PROTHORAX. Uniformly coloured, lacking stripes and foveae; lateral carina in dorsal view uninterrupted at apex, with conspicuous cuticular expansion in male, carina in lateral view moderately upcurved in male, gently in female towards apical collar.</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6C9DD6AC7F9C8" blockId="7.[189,1399,1442,1539]" pageId="7" pageNumber="8">ELYTRA. A low costa on interstria 9 gradually fading towards end; pubescence inconspicuous, very sparse, with a row of more conspicuous hairs on interstriae. Prosternal process of male variable in size, the apex rounded without a lateral lobe.</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CA556C80F9AD" blockId="7.[189,1399,1577,1639]" pageId="7" pageNumber="8">
ABDOMEN. Tergites 1–6 divided on midline; pruinose patches on tergites 5–7; tergite
<quantity id="4CFAD2AAFFF1FF8EBBD6CA556CF1F989" box="[1165,1209,1577,1604]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="7" pageNumber="8" unit="in" value="7.0">7 in</quantity>
male impinging slightly into 6, subtruncate at apex; plectral rows on basal two thirds, each with 12 pegs.
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CAF16BE1F924" blockId="7.[189,1399,1678,1775]" pageId="7" pageNumber="8">
GENITALIA. Penis as long as last four ventrites; body deeply pigmented, 2.9 × as long as apodemes (ratio w/l: 0.19), widest near basal third, then tapering to less than one third of its width, widening moderately to an asymmetrical apex (
<figureCitation id="133963CAFFF1FF8EBEB0CAAB6A00F925" box="[491,584,1748,1775]" captionStart="Fig" captionStartId="18.[189,232,1781,1807]" captionTargetBox="[203,1381,343,1664]" captionTargetId="figure@18.[189,1398,326,1679]" captionTargetPageId="18" captionText="Fig. 4. Penis of males of species of the genus Elaeidobius Kuschel, 1952 in dorsal (left) and lateral (right) view. A. E. bilineatus (Hustache, 1924) comb. nov., Dakar, Senegal. B. E. spatulifer (Marshall, 1950) comb. nov., holotype, Thysville, Democratic Republic of Congo (MRAC). C. E. subvittatus (Faust, 1898), Dakar, Senegal. D. E. kamerunicus (Faust, 1898) comb. nov. (JHAR00209). E. E. pilimargo Haran &amp; Kuschel sp. nov. (JHAR00409). F. E. piliventris Haran &amp; Kuschel sp. nov. (JHAR02182). G. E. plagiatus (Fåhraeus, 1844) comb. nov. (JHAR00272). H. E. singularis (Faust, 1898) comb. nov. (JHAR00283). Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3959121" httpUri="https://zenodo.org/record/3959121/files/figure.png" pageId="7" pageNumber="8">Fig. 4B</figureCitation>
); internal sac without sclerite.
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CB696907F8FA" blockId="7.[189,335,1814,1840]" box="[189,335,1814,1840]" pageId="7" pageNumber="8">
<heading id="D0F5C823FFF1FF8EBFE6CB696907F8FA" bold="true" box="[189,335,1814,1840]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B976A35DFFF1FF8EBFE6CB696907F8FA" bold="true" box="[189,335,1814,1840]" pageId="7" pageNumber="8">Life history</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CB3B6BBAF894" blockId="7.[189,1010,1860,1887]" box="[189,1010,1860,1887]" pageId="7" pageNumber="8">
Unknown,
<taxonomicName id="4C0204CCFFF1FF8EBE64CB3A6AECF895" box="[319,676,1861,1887]" class="Liliopsida" family="Arecaceae" genus="Elaeis" kingdom="Plantae" order="Arecales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="guineensis">
<emphasis id="B976A35DFFF1FF8EBE64CB3A6A44F894" box="[319,524,1861,1887]" italics="true" pageId="7" pageNumber="8">Elaeis guineensis</emphasis>
(Arecaceae)
</taxonomicName>
, oil palm, on inflorescences.
</paragraph>
</subSubSection>
<subSubSection id="C3182CC4FFF1FF8EBFE6CBF96D44F805" box="[189,1292,1926,1999]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8BBD7F4FFFF1FF8EBFE6CBF9691EF86A" blockId="7.[189,342,1926,1952]" box="[189,342,1926,1952]" pageId="7" pageNumber="8">
<heading id="D0F5C823FFF1FF8EBFE6CBF9691EF86A" bold="true" box="[189,342,1926,1952]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B976A35DFFF1FF8EBFE6CBF9691EF86A" bold="true" box="[189,342,1926,1952]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD7F4FFFF1FF8EBFE6CBCB6D44F805" blockId="7.[189,1292,1972,1999]" box="[189,1292,1972,1999]" pageId="7" pageNumber="8">
<emphasis id="B976A35DFFF1FF8EBFE6CBCB6908F804" bold="true" box="[189,320,1972,1998]" pageId="7" pageNumber="8">
<collectingCountry id="F3153FDFFFF1FF8EBFE6CBCB6908F804" box="[189,320,1972,1998]" name="Cameroon" pageId="7" pageNumber="8">Cameroon</collectingCountry>
</emphasis>
(Yaoundé);
<emphasis id="B976A35DFFF1FF8EBE89CBCB6B1BF804" bold="true" box="[466,851,1972,1998]" pageId="7" pageNumber="8">
<collectingCountry id="F3153FDFFFF1FF8EBE89CBCB6B1BF804" box="[466,851,1972,1998]" name="Democratic Republic of the Congo" pageId="7" pageNumber="8">Democratic Republic of Congo</collectingCountry>
</emphasis>
(Thysville);
<emphasis id="B976A35DFFF1FF8EBCB7CBCA6CC8F804" bold="true" box="[1004,1152,1972,1999]" pageId="7" pageNumber="8">
<collectingCountry id="F3153FDFFFF1FF8EBCB7CBCA6CC8F804" box="[1004,1152,1972,1999]" name="Ivory Coast" pageId="7" pageNumber="8">Ivory Coast</collectingCountry>
</emphasis>
(Toumodi).
</paragraph>
</subSubSection>
<subSubSection id="C3182CC4FFFEFF81BFE6CD746A15FE50" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BBD7F4FFFFEFF81BFE6CD746967FEEF" blockId="8.[189,303,267,293]" box="[189,303,267,293]" pageId="8" pageNumber="9">
<heading id="D0F5C823FFFEFF81BFE6CD746967FEEF" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B976A35DFFFEFF81BFE6CD746967FEEF" bold="true" box="[189,303,267,293]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD7F4FFFFEFF81BFE6CD466A15FE50" blockId="8.[189,1399,313,410]" pageId="8" pageNumber="9">The species is distinguished by a uniform dark colour, and in the male by a prominent spade-like prosternal process and lack of a postmental tubercle. No DNA-grade material of this species could be obtained in the course of this study.</paragraph>
</subSubSection>
</treatment>
</document>
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